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97-724: It underlies the Pliocene epoch Etchegoin Formation . It preserves fossils dating back to the Cretaceous period . This article about a specific stratigraphic formation in California is a stub . You can help Misplaced Pages by expanding it . This article related to the Cretaceous period is a stub . You can help Misplaced Pages by expanding it . Pliocene The Pliocene ( / ˈ p l aɪ . ə s iː n , ˈ p l aɪ . oʊ -/ PLY -ə-seen, PLY -oh- ; also Pleiocene )

194-469: A monotreme , appeared. The predatory South American phorusrhacids were rare in this time; among the last was Titanis , a large phorusrhacid that migrated to North America and rivaled mammals as top predator. Other birds probably evolved at this time, some modern (such as the genera Cygnus , Bubo , Struthio and Corvus ), some now extinct. Alligators and crocodiles died out in Europe as

291-526: A broader aridification trend. The EMCI ended 18 million years ago, giving way to the Middle Miocene Warm Interval (MMWI), the warmest part of which was the MMCO that began 16 million years ago. As the world transitioned into the MMCO, carbon dioxide concentrations varied between 300 and 500 ppm. Global annual mean surface temperature during the MMCO was about 18.4 °C. MMCO warmth was driven by

388-519: A clade of large terrestrial predatory crocodyliformes distantly related to modern crocodilians, from which they likely diverged over 180 million years ago, are known from the Miocene of South America. The last Desmostylians thrived during this period before becoming the only extinct marine mammal order. The pinnipeds , which appeared near the end of the Oligocene, became more aquatic. A prominent genus

485-561: A cooler, drier climate. C 4 grasses, which are able to assimilate carbon dioxide and water more efficiently than C 3 grasses, expanded to become ecologically significant near the end of the Miocene between 6 and 7 million years ago, although they did not expand northward during the Late Miocene. The expansion of grasslands and radiations among terrestrial herbivores correlates to fluctuations in CO 2 . One study, however, has attributed

582-473: A good analogue for future warmer climates caused by anthropogenic global warming , with this being especially true of the global climate during the Middle Miocene Climatic Optimum (MMCO), because the last time carbon dioxide levels were comparable to projected future atmospheric carbon dioxide levels resulting from anthropogenic climate change was during the MMCO. The Ross Sea margin of

679-435: A good model for a "living fossil". Eucalyptus fossil leaves occur in the Miocene of New Zealand , where the genus is not native today, but have been introduced from Australia . Both marine and continental fauna were fairly modern, although marine mammals were less numerous. Only in isolated South America and Australia did widely divergent fauna exist. In Eurasia, genus richness shifted southward to lower latitudes from

776-406: A large neocortex relative to body mass and stone tool manufacture. Improvements in dating methods and in the use of climate proxies have provided scientists with the means to test hypotheses of the evolution of human ancestors. Early hypotheses of the evolution of human traits emphasized the selective pressures produced by particular habitats. For example, many scientists have long favored

873-579: A major expansion of Antarctic glaciers. This severed the connection between the Indian Ocean and the Mediterranean Sea and formed the present land connection between Afro-Arabia and Eurasia. The subsequent uplift of mountains in the western Mediterranean region and a global fall in sea levels combined to cause a temporary drying up of the Mediterranean Sea (known as the Messinian salinity crisis ) near

970-480: A major expansion of grass-grazer ecosystems . Herds of large, swift grazers were hunted by predators across broad sweeps of open grasslands , displacing desert, woodland, and browsers . The higher organic content and water retention of the deeper and richer grassland soils , with long-term burial of carbon in sediments, produced a carbon and water vapor sink. This, combined with higher surface albedo and lower evapotranspiration of grassland, contributed to

1067-429: A particular habitat. The change to a cooler, drier, more seasonal climate had considerable impacts on Pliocene vegetation, reducing tropical species worldwide. Deciduous forests proliferated, coniferous forests and tundra covered much of the north, and grasslands spread on all continents (except Antarctica). Eastern Africa in particular saw a huge expansion of C 4 grasslands. Tropical forests were limited to

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1164-578: A significant drop in atmospheric carbon dioxide levels. Both continental and oceanic thermal gradients in mid-latitudes during the Early Miocene were very similar to those in the present. Global cooling caused the East Asian Summer Monsoon (EASM) to begin to take on its modern form during the Early Miocene. From 22.1 to 19.7 Ma, the Xining Basin experienced relative warmth and humidity amidst

1261-695: A significant local decline along the northeastern coast of Australia during the Tortonian, most likely due to warming seawater. Cetaceans attained their greatest diversity during the Miocene, with over 20 recognized genera of baleen whales in comparison to only six living genera. This diversification correlates with emergence of gigantic macro-predators such as megatoothed sharks and raptorial sperm whales . Prominent examples are O. megalodon and L. melvillei . Other notable large sharks were O. chubutensis , Isurus hastalis , and Hemipristis serra . Crocodilians also showed signs of diversification during

1358-953: A tight band around the equator, and in addition to dry savannahs , deserts appeared in Asia and Africa. Both marine and continental faunas were essentially modern, although continental faunas were a bit more primitive than today. The land mass collisions meant great migration and mixing of previously isolated species, such as in the Great American Interchange . Herbivores got bigger, as did specialized predators. In North America, rodents , large mastodons and gomphotheres , and opossums continued successfully, while hoofed animals ( ungulates ) declined, with camel , deer , and horse all seeing populations recede. Three-toed horses ( Nannippus ), oreodonts , protoceratids , and chalicotheres became extinct. Borophagine dogs and Agriotherium became extinct, but other carnivores including

1455-627: A zone of low rainfall in the Late Miocene. The Indian Plate continued to collide with the Eurasian Plate , creating new mountain ranges and uplifting the Tibetan Plateau , resulting in the rain shadowing and aridification of the Asian interior. The Tian Shan experienced significant uplift in the Late Miocene, blocking westerlies from coming into the Tarim Basin and drying it as a result. At

1552-540: Is a key finding of research into a lake-sediment core obtained in Eastern Siberia, which is of exceptional importance because it has provided the longest continuous late Cenozoic land-based sedimentary record thus far. During the late Zanclean, Italy remained relatively warm and humid. Central Asia became more seasonal during the Pliocene, with colder, drier winters and wetter summers, which contributed to an increase in

1649-688: Is the epoch in the geologic time scale that extends from 5.33 to 2.58 million years ago (Ma). It is the second and most recent epoch of the Neogene Period in the Cenozoic Era . The Pliocene follows the Miocene Epoch and is followed by the Pleistocene Epoch. Prior to the 2009 revision of the geologic time scale, which placed the four most recent major glaciations entirely within the Pleistocene,

1746-567: The Hardangervidda plateau to 1200 m in the Early Pliocene. In Southern Sweden similar movements elevated the South Swedish highlands leading to a deflection of the ancient Eridanos river from its original path across south-central Sweden into a course south of Sweden. The Pliocene is bookended by two significant events in the evolution of human ancestors. The first is the appearance of

1843-676: The Isthmus of Panama during the Pliocene, making possible the Great American Interchange and bringing a nearly complete end to South America's distinctive native ungulate fauna, though other South American lineages like its predatory mammals were already extinct by this point and others like xenarthrans continued to do well afterwards. The formation of the Isthmus had major consequences on global temperatures, since warm equatorial ocean currents were cut off and an Atlantic cooling cycle began, with cold Arctic and Antarctic waters decreasing temperatures in

1940-539: The Middle Miocene Climate Transition (MMCT). Abrupt increases in opal deposition indicate this cooling was driven by enhanced drawdown of carbon dioxide via silicate weathering. The MMCT caused a sea surface temperature (SST) drop of approximately 6 °C in the North Atlantic. The drop in benthic foraminiferal δ O values was most noticeable in the waters around Antarctica, suggesting cooling

2037-555: The Netherlands the Pliocene is divided into these stages (old to young): Brunssumian C, Reuverian A, Reuverian B, Reuverian C, Praetiglian , Tiglian A, Tiglian B, Tiglian C1-4b, Tiglian C4c, Tiglian C5, Tiglian C6 and Eburonian . The exact correlations between these local stages and the International Commission on Stratigraphy (ICS) stages is not established. During the Pliocene epoch (5.3 to 2.6 million years ago (Ma)),

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2134-471: The North Atlantic and North Pacific Ocean beds. Mid-latitude glaciation was probably underway before the end of the epoch. The global cooling that occurred during the Pliocene may have accelerated on the disappearance of forests and the spread of grasslands and savannas. During the Pliocene the earth climate system response shifted from a period of high frequency-low amplitude oscillation dominated by

2231-550: The carbon cycle occurred approximately 6 Ma, causing continental carbon reservoirs to no longer expand during cold spells, as they had done during cold periods in the Oligocene and most of the Miocene. At the end of the Miocene, global temperatures rose again as the amplitude of Earth's obliquity increased, which caused increased aridity in Central Asia. Around 5.5 Ma, the EAWM underwent a period of rapid intensification. Life during

2328-461: The geology of the Himalaya occurred during that epoch, affecting monsoonal patterns in Asia, which were interlinked with glacial periods in the northern hemisphere. The Miocene faunal stages from youngest to oldest are typically named according to the International Commission on Stratigraphy : Regionally, other systems are used, based on characteristic land mammals; some of them overlap with

2425-489: The hominin Australopithecus anamensis in the early Pliocene, around 4.2 million years ago. The second is the appearance of Homo , the genus that includes modern humans and their closest extinct relatives, near the end of the Pliocene at 2.6 million years ago. Key traits that evolved among hominins during the Pliocene include terrestrial bipedality and, by the end of the Pliocene, encephalized brains (brains with

2522-634: The savannah hypothesis . This proposes that the evolution of terrestrial bipedality and other traits was an adaptive response to Pliocene climate change that transformed forests into more open savannah . This was championed by Grafton Elliot Smith in his 1924 book, The Evolution of Man , as "the unknown world beyond the trees", and was further elaborated by Raymond Dart as the killer ape theory . Other scientists, such as Sherwood L. Washburn , emphasized an intrinsic model of hominin evolution. According to this model, early evolutionary developments triggered later developments. The model placed little emphasis on

2619-493: The weasel family diversified, and dogs and short-faced bears did well. Ground sloths , huge glyptodonts , and armadillos came north with the formation of the Isthmus of Panama. The latitudinal diversity gradient among terrestrial North American mammals became established during this epoch some time after 4 Ma. In Eurasia rodents did well, while primate distribution declined. Elephants , gomphotheres and stegodonts were successful in Asia (the largest land mammals of

2716-523: The " Zanclean flood ". Also during the early Miocene (specifically the Aquitanian and Burdigalian Stages), the apes first evolved, began diversifying, and became widespread throughout the Old World . Around the end of this epoch, the ancestors of humans had split away from the ancestors of the chimpanzees and had begun following their own evolutionary path during the final Messinian Stage (7.5–5.3 Ma) of

2813-465: The 41,000-year period of Earth's obliquity to one of low-frequency, high-amplitude oscillation dominated by the 100,000-year period of the orbital eccentricity characteristic of the Pleistocene glacial-interglacial cycles. During the late Pliocene and early Pleistocene, 3.6 to 2.6 Ma, the Arctic was much warmer than it is at the present day (with summer temperatures some 8 °C warmer than today). That

2910-673: The Central Paratethys, cut off from sources of freshwater input by its separation from the Eastern Paratethys. From 13.36 to 12.65 Ma, the Central Paratethys was characterised by open marine conditions, before the reopening of the Bârlad Strait resulted in a shift to brackish-marine conditions in the Central Paratethys, causing the Badenian-Sarmatian Extinction Event. As a result of the Bârlad Strait's reopening,

3007-511: The Cretaceous and the early Cenozoic . This may have contributed to further cooling of the oceans worldwide. The Pliocene seas were alive with sea cows , seals , sea lions , sharks and whales . Miocene The Miocene ( / ˈ m aɪ . ə s iː n , - oʊ -/ MY -ə-seen, -⁠oh- ) is the first geological epoch of the Neogene Period and extends from about 23.03 to 5.333 million years ago (Ma). The Miocene

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3104-826: The Early to the Middle Miocene. Europe's large mammal diversity significantly declined during the Late Miocene. In the Early Miocene, several Oligocene groups were still diverse, including nimravids , entelodonts , and three-toed equids. As in the previous Oligocene Epoch, oreodonts were still diverse, only to disappear in the earliest Pliocene. During the later Miocene mammals were more modern, with easily recognizable canids , bears , red pandas , procyonids , equids , beavers , deer , camelids , and whales , along with now-extinct groups like borophagine canids , certain gomphotheres , three-toed horses , and hornless rhinos like Teleoceras and Aphelos . The late Miocene also marks

3201-580: The Earth's climate became cooler and drier, as well as more seasonal, marking a transition between the relatively warm Miocene to the cooler Pleistocene . However, the beginning of the Pliocene was marked by an increase in global temperatures relative to the cooler Messinian . This increase was related to the 1.2 million year obliquity amplitude modulation cycle . By 3.3-3.0 Ma, during the Mid-Piacenzian Warm Period (mPWP), global average temperature

3298-677: The East Antarctic Ice Sheet (EAIS) was highly dynamic during the Early Miocene. The Miocene began with the Early Miocene Cool Event (Mi-1) around 23 million years ago, which marked the start of the Early Miocene Cool Interval (EMCI). This cool event occurred immediately after the Oligocene-Miocene Transition (OMT) during a major expansion of Antarctica's ice sheets, but was not associated with

3395-526: The East Asian Winter Monsoon (EAWM) became stronger synchronously with a southward shift of the subarctic front. Greenland may have begun to have large glaciers as early as 8 to 7 Ma, although the climate for the most part remained warm enough to support forests there well into the Pliocene. Zhejiang, China was noticeably more humid than today. In the Great Rift Valley of Kenya , there

3492-457: The LMC; extratropical sea surface temperatures dropped substantially by approximately 7–9 °C. 41 kyr obliquity cycles became the dominant orbital climatic control 7.7 Ma and this dominance strengthened 6.4 Ma. Benthic δ O values show significant glaciation occurred from 6.26 to 5.50 Ma, during which glacial-interglacial cycles were governed by the 41 kyr obliquity cycle. A major reorganisation of

3589-514: The Late Cretaceous, are known from the Miocene of Patagonia, represented by the mole-like Necrolestes . The youngest known representatives of metatherians (the broader grouping to which marsupials belong) in Europe, Asia and Africa are known from the Miocene, including the European herpetotheriid Amphiperatherium , the peradectids Siamoperadectes and Sinoperadectes from Asia, and

3686-646: The Late Miocene, the Earth's climate began to display a high degree of similarity to that of the present day . The 173 kyr obliquity modulation cycle governed by Earth's interactions with Saturn became detectable in the Late Miocene. By 12 Ma, Oregon was a savanna akin to that of the western margins of the Sierra Nevada of northern California . Central Australia became progressively drier, although southwestern Australia experienced significant wettening from around 12 to 8 Ma. The South Asian Winter Monsoon (SAWM) underwent strengthening ~9.2–8.5 Ma. From 7.9 to 5.8 Ma,

3783-567: The Miocene Epoch was mostly supported by the two newly formed biomes , kelp forests and grasslands . Grasslands allow for more grazers, such as horses , rhinoceroses , and hippos . Ninety-five percent of modern plants existed by the end of this epoch . Modern bony fish genera were established. A modern-style latitudinal biodiversity gradient appeared ~15 Ma. The coevolution of gritty , fibrous, fire-tolerant grasses and long-legged gregarious ungulates with high-crowned teeth , led to

3880-484: The Miocene. As in the Oligocene before it, grasslands continued to expand, and forests to dwindle. In the seas of the Miocene, kelp forests made their first appearance and soon became one of Earth's most productive ecosystems. The plants and animals of the Miocene were recognizably modern. Mammals and birds were well established. Whales , pinnipeds , and kelp spread. The Miocene is of particular interest to geologists and palaeoclimatologists because major phases of

3977-449: The Miocene. The largest form among them was a gigantic caiman Purussaurus which inhabited South America. Another gigantic form was a false gharial Rhamphosuchus , which inhabited modern age India . A strange form, Mourasuchus also thrived alongside Purussaurus . This species developed a specialized filter-feeding mechanism, and it likely preyed upon small fauna despite its gigantic size. The youngest members of Sebecidae ,

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4074-534: The North American Great Plains and in Argentina . The global trend was towards increasing aridity caused primarily by global cooling reducing the ability of the atmosphere to absorb moisture, particularly after 7 to 8 million years ago. Uplift of East Africa in the late Miocene was partly responsible for the shrinking of tropical rain forests in that region, and Australia got drier as it entered

4171-648: The Oligocene–Miocene transgression. As the southern Andes rose in the Middle Miocene (14–12 million years ago) the resulting rain shadow originated the Patagonian Desert to the east. Far northern Australia was monsoonal during the Miocene. Although northern Australia is often believed to have been much wetter during the Miocene, this interpretation may be an artefact of preservation bias of riparian and lacustrine plants; this finding has itself been challenged by other papers. Western Australia, like today,

4268-503: The Pliocene also included the Gelasian Stage, which lasted from 2.59 to 1.81 Ma, and is now included in the Pleistocene. As with other older geologic periods, the geological strata that define the start and end are well-identified but the exact dates of the start and end of the epoch are slightly uncertain. The boundaries defining the Pliocene are not set at an easily identified worldwide event but rather at regional boundaries between

4365-613: The Pliocene contains the Dacian (roughly equal to the Zanclean) and Romanian (roughly equal to the Piacenzian and Gelasian together) stages. As usual in stratigraphy, there are many other regional and local subdivisions in use. In Britain , the Pliocene is divided into the following stages (old to young): Gedgravian, Waltonian , Pre-Ludhamian, Ludhamian, Thurnian, Bramertonian or Antian, Pre-Pastonian or Baventian, Pastonian and Beestonian . In

4462-559: The Pliocene were such proboscideans as Deinotherium , Anancus , and Mammut borsoni , ) though proboscidean diversity declined significantly during the Late Pliocene. Hyraxes migrated north from Africa. Horse diversity declined, while tapirs and rhinos did fairly well. Bovines and antelopes were successful; some camel species crossed into Asia from North America. Hyenas and early saber-toothed cats appeared, joining other predators including dogs, bears, and weasels. Africa

4559-598: The Qiongdongnan Basin in the northern South China Sea indicates the Pearl River was a major source of sediment flux into the sea during the Early Miocene and was a major fluvial system as in the present. During the Oligocene and Early Miocene, the coast of northern Brazil, Colombia, south-central Peru , central Chile and large swathes of inland Patagonia were subject to a marine transgression . The transgressions in

4656-533: The Western Atlantic, assemblages of bivalves exhibited remarkable stasis with regards to their basal metabolic rates throughout the various climatic changes of the Pliocene. The Pliocene was a high water mark for species diversity among Caribbean corals. From 5 to 2 Ma, coral species origination rates were relatively high in the Caribbean, although a noticeable extinction event and drop in diversity occurred at

4753-476: The abundance of C 4 plants across the region. In the Loess Plateau , δ13C values of occluded organic matter increased by 2.5% while those of pedogenic carbonate increased by 5% over the course of the Late Miocene and Pliocene, indicating increased aridification. Further aridification of Central Asia was caused by the development of Northern Hemisphere glaciation during the Late Pliocene. A sediment core from

4850-487: The activity of the Columbia River Basalts and enhanced by decreased albedo from the reduction of deserts and expansion of forests. Climate modelling suggests additional, currently unknown, factors also worked to create the warm conditions of the MMCO. The MMCO saw the expansion of the tropical climatic zone to much larger than its current size. The July ITCZ, the zone of maximal monsoonal rainfall, moved to

4947-472: The aridity of the former. Unequivocally-recognizable dabbling ducks , plovers , typical owls , cockatoos and crows appear during the Miocene. By the epoch's end, all or almost all modern bird groups are believed to have been present; the few post-Miocene bird fossils which cannot be placed in the evolutionary tree with full confidence are simply too badly preserved, rather than too equivocal in character. Marine birds reached their highest diversity ever in

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5044-619: The beginning of the Miocene, the northern margin of the Arabian plate, then part of the African landmass, collided with Eurasia; as a result, the Tethys seaway continued to shrink and then disappeared as Africa collided with Eurasia in the Turkish – Arabian region. The first step of this closure occurred 20 Ma, reducing water mass exchange by 90%, while the second step occurred around 13.8 Ma, coincident with

5141-668: The climate cooled. Venomous snake genera continued to increase as more rodents and birds evolved. Rattlesnakes first appeared in the Pliocene. The modern species Alligator mississippiensis , having evolved in the Miocene, continued into the Pliocene, except with a more northern range; specimens have been found in very late Miocene deposits of Tennessee . Giant tortoises still thrived in North America, with genera like Hesperotestudo . Madtsoid snakes were still present in Australia. The amphibian order Allocaudata became extinct. In

5238-481: The climate slowly cooled towards a series of ice ages . The Miocene boundaries are not marked by distinct global events but by regionally defined transitions from the warmer Oligocene to the cooler Pliocene Epoch. During the Early Miocene, Afro-Arabia collided with Eurasia, severing the connection between the Mediterranean and Indian Oceans, and allowing the interchange of fauna between Eurasia and Africa, including

5335-566: The course of this epoch . The youngest representatives of Choristodera , an extinct order of aquatic reptiles that first appeared in the Middle Jurassic , are known from the Miocene of Europe, belonging to the genus Lazarussuchus , which had been the only known surviving genus of the group since the beginning of the Eocene. The last known representatives of the archaic primitive mammal order Meridiolestida , which dominated South America during

5432-570: The dispersal of proboscideans and hominoids into Eurasia. During the late Miocene, the connections between the Atlantic and Mediterranean closed, causing the Mediterranean Sea to almost completely evaporate. This event is referred to as the " Messinian salinity crisis ". Then, at the Miocene–Pliocene boundary, the Strait of Gibraltar opened, and the Mediterranean refilled. That event is referred to as

5529-475: The early-mid Pliocene (5 Ma – 3 Ma), after three-million-year intervals with modern or glacial ice volume became longer and collapse occurs only at times when warmer global temperature coincide with strong austral summer insolation anomalies. Continents continued to drift , moving from positions possibly as far as 250 km from their present locations to positions only 70 km from their current locations. South America became linked to North America through

5626-508: The end of the Miocene due to increased habitat uniformity. The expansion of grasslands in North America also led to an explosive radiation among snakes. Previously, snakes were a minor component of the North American fauna, but during the Miocene, the number of species and their prevalence increased dramatically with the first appearances of vipers and elapids in North America and the significant diversification of Colubridae (including

5723-422: The end of the Miocene. The Paratethys underwent a significant transgression during the early Middle Miocene. Around 13.8 Ma, during a global sea level drop, the Eastern Paratethys was cut off from the global ocean by the closure of the Bârlad Strait, effectively turning it into a saltwater lake. From 13.8 to 13.36 Ma, an evaporite period similar to the later Messinian salinity crisis in the Mediterranean ensued in

5820-501: The end of this interval. Oceans continued to be relatively warm during the Pliocene, though they continued cooling. The Arctic ice cap formed , drying the climate and increasing cool shallow currents in the North Atlantic. Deep cold currents flowed from the Antarctic. The formation of the Isthmus of Panama about 3.5 million years ago cut off the final remnant of what was once essentially a circum-equatorial current that had existed since

5917-612: The enigmatic Saint Bathans Mammal . Microbial life in the igneous crust of the Fennoscandian Shield shifted from being dominated by methanogens to being primarily composed of sulphate-reducing prokaryotes . The change resulted from fracture reactivation during the Pyrenean-Alpine orogeny, enabling sulphate-reducing microbes to permeate into the Fennoscandian Shield via descending surficial waters. Diatom diversity

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6014-461: The evolution of both groups into modern representatives. The early Miocene Saint Bathans Fauna is the only Cenozoic terrestrial fossil record of the landmass, showcasing a wide variety of not only bird species, including early representatives of clades such as moa , kiwi and adzebills , but also a diverse herpetofauna of sphenodontians , crocodiles and turtles as well as a rich terrestrial mammal fauna composed of various species of bats and

6111-499: The expansion of grasslands not to a CO 2 drop but to the increasing seasonality and aridity, coupled with a monsoon climate, which made wildfires highly prevalent compared to before. The Late Miocene expansion of grasslands had cascading effects on the global carbon cycle, evidenced by the imprint it left in carbon isotope records. Cycads between 11.5 and 5 million years ago began to rediversify after previous declines in variety due to climatic changes, and thus modern cycads are not

6208-653: The extinction of the last-surviving members of the hyaenodonts . Islands began to form between South and North America in the Late Miocene, allowing ground sloths like Thinobadistes to island-hop to North America. The expansion of silica-rich C 4 grasses led to worldwide extinctions of herbivorous species without high-crowned teeth . Mustelids diversified into their largest forms as terrestrial predators like Ekorus , Eomellivora , and Megalictis and bunodont otters like Enhydriodon and Sivaonyx appeared. Eulipotyphlans were widespread in Europe, being less diverse in Southern Europe than farther north due to

6305-421: The findings of marine invertebrate fossils of both Atlantic and Pacific affinity in La Cascada Formation . Connection would have occurred through narrow epicontinental seaways that formed channels in a dissected topography . The Antarctic Plate started to subduct beneath South America 14 million years ago in the Miocene, forming the Chile Triple Junction . At first the Antarctic Plate subducted only in

6402-491: The first time since the Cretaceous , with North American rodents and primates mixing with southern forms. Litopterns and the notoungulates , South American natives, were mostly wiped out, except for the macrauchenids and toxodonts , which managed to survive. Small weasel-like carnivorous mustelids , coatis and short-faced bears migrated from the north. Grazing glyptodonts , browsing giant ground sloths and smaller caviomorph rodents , pampatheres , and armadillos did

6499-411: The human lineage) appeared in Africa at the very end of the Miocene, including Sahelanthropus , Orrorin , and an early form of Ardipithecus ( A. kadabba ). The chimpanzee–human divergence is thought to have occurred at this time. The evolution of bipedalism in apes at the end of the Miocene instigated an increased rate of faunal turnover in Africa. In contrast, European apes met their end at

6596-413: The influence of seasonal variation. More recent research has emphasized the variability selection hypothesis, which proposes that variability in climate fostered development of hominin traits. Improved climate proxies show that the Pliocene climate of east Africa was highly variable, suggesting that adaptability to varying conditions was more important in driving hominin evolution than the steady pressure of

6693-422: The lake levels of the Eastern Paratethys dropped as it once again became a sea. The Fram Strait opened during the Miocene and acted as the only throughflow for Atlantic Water into the Arctic Ocean until the Quaternary period. Due to regional uplift of the continental shelf, this water could not move through the Barents Seaway in the Miocene. The modern day Mekong Delta took shape after 8 Ma. Geochemistry of

6790-496: The north, increasing precipitation over southern China whilst simultaneously decreasing it over Indochina during the EASM. Western Australia was at this time characterised by exceptional aridity. In Antarctica, average summer temperatures on land reached 10 °C. In the oceans, the lysocline shoaled by approximately half of a kilometre during warm phases that corresponded to orbital eccentricity maxima. The MMCO ended around 14 million years ago, when global temperatures fell in

6887-480: The northern South China Sea shows an increase in dust storm activity during the middle Pliocene. The South Asian Summer Monsoon (SASM) increased in intensity after 2.95 Ma, likely because of enhanced cross-equatorial pressure caused by the reorganisation of the Indonesian Throughflow. In the south-central Andes , an arid period occurred from 6.1 to 5.2 Ma, with another occurring from 3.6 to 3.3 Ma. These arid periods are coincident with global cold periods, during which

6984-673: The now-separated Atlantic Ocean. Africa 's collision with Europe formed the Mediterranean Sea , cutting off the remnants of the Tethys Ocean . The border between the Miocene and the Pliocene is also the time of the Messinian salinity crisis . During the Late Pliocene, the Himalayas became less active in their uplift, as evidenced by sedimentation changes in the Bengal Fan . The land bridge between Alaska and Siberia ( Beringia )

7081-504: The onset of continental glaciation in the Arctic, suggesting that vegetation changes in Australia during the Pliocene behaved similarly to during the Late Pleistocene and were likely characterised by comparable cycles of aridity and humidity. The equatorial Pacific Ocean sea surface temperature gradient was considerably lower than it is today. Mean sea surface temperatures in the east were substantially warmer than today but similar in

7178-400: The opposite, migrating to the north and thriving there. The marsupials remained the dominant Australian mammals, with herbivore forms including wombats and kangaroos , and the huge Diprotodon . Carnivorous marsupials continued hunting in the Pliocene, including dasyurids , the dog-like thylacine and cat-like Thylacoleo . The first rodents arrived in Australia. The modern platypus ,

7275-523: The origin of many modern genera such as Nerodia , Lampropeltis , Pituophis and Pantherophis ). Arthropods were abundant, including in areas such as Tibet where they have traditionally been thought to be undiverse. Neoisopterans diversified and expanded into areas they previously were absent from, such as Madagascar and Australia. In the oceans, brown algae , called kelp , proliferated, supporting new species of sea life, including otters , fish and various invertebrates . Corals suffered

7372-602: The position of the Southern Hemisphere westerlies shifted northward and disrupted the South American Low Level Jet, which brings moisture to southeastern South America. From around 3.8 Ma to about 3.3 Ma, North Africa experienced an extended humid period. In northwestern Africa, tropical forests extended up to Cape Blanc during the Zanclean until around 3.5 Ma. During the Piacenzian, from about 3.5 to 2.6 Ma,

7469-463: The possible herpetotheriid Morotodon from the late Early Miocene of Uganda. Approximately 100 species of apes lived during this time , ranging throughout Africa, Asia and Europe and varying widely in size, diet, and anatomy. Due to scanty fossil evidence it is unclear which ape or apes contributed to the modern hominid clade, but molecular evidence indicates this ape lived between 18 and 13 million years ago. The first hominins ( bipedal apes of

7566-828: The preceding Oligocene and following Pliocene Epochs: Continents continued to drift toward their present positions. Of the modern geologic features, only the land bridge between South America and North America was absent, although South America was approaching the western subduction zone in the Pacific Ocean , causing both the rise of the Andes and a southward extension of the Meso-American peninsula. Mountain building took place in western North America , Europe , and East Asia . Both continental and marine Miocene deposits are common worldwide with marine outcrops common near modern shorelines. Well studied continental exposures occur in

7663-505: The recent", referring to the essentially modern marine mollusc fauna. In the official timescale of the ICS , the Pliocene is subdivided into two stages . From youngest to oldest they are: The Piacenzian is sometimes referred to as the Late Pliocene, whereas the Zanclean is referred to as the Early Pliocene. In the system of In the Paratethys area (central Europe and parts of western Asia)

7760-499: The region was forested at irregular intervals and contained a significant Saharan palaeoriver until 3.35 Ma, when trade winds began to dominate over fluvial transport of pollen. Around 3.26 Ma, a strong aridification event that was followed by a return to more humid conditions, which was itself followed by another aridification around 2.7 Ma. From 2.6 to 2.4 Ma, vegetation zones began repeatedly shifting latitudinally in response to glacial-interglacial cycles. The climate of eastern Africa

7857-468: The scene. Bears, dogs and weasels (originally from North America) joined cats, hyenas and civets as the African predators, forcing hyenas to adapt as specialized scavengers. Most mustelids in Africa declined as a result of increased competition from the new predators, although Enhydriodon omoensis remained an unusually successful terrestrial predator. South America was invaded by North American species for

7954-753: The southernmost tip of Patagonia, meaning that the Chile Triple Junction lay near the Strait of Magellan . As the southern part of Nazca Plate and the Chile Rise became consumed by subduction the more northerly regions of the Antarctic Plate begun to subduct beneath Patagonia so that the Chile Triple Junction advanced to the north over time. The asthenospheric window associated to the triple junction disturbed previous patterns of mantle convection beneath Patagonia inducing an uplift of ca. 1 km that reversed

8051-400: The surrounding environment. Anthropologists tended to focus on intrinsic models while geologists and vertebrate paleontologists tended to put greater emphasis on habitats. Alternatives to the savanna hypothesis include the woodland/forest hypothesis, which emphasizes the evolution of hominins in closed habitats, or hypotheses emphasizing the influence of colder habitats at higher latitudes or

8148-504: The warmer Miocene and the relatively cooler Pleistocene. The upper boundary was set at the start of the Pleistocene glaciations. Charles Lyell (later Sir Charles) gave the Pliocene its name in Principles of Geology (volume 3, 1833). The word pliocene comes from the Greek words πλεῖον ( pleion , "more") and καινός ( kainos , "new" or "recent") and means roughly "continuation of

8245-590: The west coast of South America are thought to be caused by a regional phenomenon while the steadily rising central segment of the Andes represents an exception. While there are numerous registers of Oligocene–Miocene transgressions around the world it is doubtful that these correlate. It is thought that the Oligocene–Miocene transgression in Patagonia could have temporarily linked the Pacific and Atlantic Oceans, as inferred from

8342-495: The west. This condition has been described as a permanent El Niño state, or “El Padre.” Several mechanisms have been proposed for this pattern, including increased tropical cyclone activity . The extent of the West Antarctic Ice Sheet oscillated at the 40 kyr period of Earth's obliquity. Ice sheet collapse occurred when the global average temperature was 3 °C warmer than today and carbon dioxide concentration

8439-423: Was Allodesmus . A ferocious walrus , Pelagiarctos may have preyed upon other species of pinnipeds including Allodesmus . Furthermore, South American waters witnessed the arrival of Megapiranha paranensis , which were considerably larger than modern age piranhas . New Zealand 's Miocene fossil record is particularly rich. Marine deposits showcase a variety of cetaceans and penguins , illustrating

8536-470: Was 2–3 °C higher than today, while carbon dioxide levels were the same as today (400 ppm). Global sea level was about 25 m higher, though its exact value is uncertain. The northern hemisphere ice sheet was ephemeral before the onset of extensive glaciation over Greenland that occurred in the late Pliocene around 3 Ma. The formation of an Arctic ice cap is signaled by an abrupt shift in oxygen isotope ratios and ice-rafted cobbles in

8633-610: Was a gradual and progressive trend of increasing aridification, though it was not unidirectional, and wet humid episodes continued to occur. Between 7 and 5.3 Ma, temperatures dropped sharply again in the Late Miocene Cooling (LMC), most likely as a result of a decline in atmospheric carbon dioxide and a drop in the amplitude of Earth's obliquity, and the Antarctic ice sheet was approaching its present-day size and thickness. Ocean temperatures plummeted to near-modern values during

8730-458: Was arid, particularly so during the Middle Miocene. Climates remained moderately warm, although the slow global cooling that eventually led to the Pleistocene glaciations continued. Although a long-term cooling trend was well underway, there is evidence of a warm period during the Miocene when the global climate rivalled that of the Oligocene . The climate of the Miocene has been suggested as

8827-506: Was at 400 ppmv. This resulted in open waters in the Ross Sea . Global sea-level fluctuation associated with ice-sheet collapse was probably up to 7 meters for the west Antarctic and 3 meters for the east Antarctic. Model simulations are consistent with reconstructed ice-sheet oscillations and suggest a progression from a smaller to a larger West Antarctic ice sheet in the last 5 million years. Intervals of ice sheet collapse were much more common in

8924-480: Was dominated by hoofed animals, and primates continued their evolution, with australopithecines (some of the first hominins ) and baboon-like monkeys such as the Dinopithecus appearing in the late Pliocene. Rodents were successful, and elephant populations increased. Cows and antelopes continued diversification and overtook pigs in numbers of species. Early giraffes appeared. Horses and modern rhinos came onto

9021-452: Was first flooded near the start of the Pliocene, allowing marine organisms to spread between the Arctic and Pacific Oceans. The bridge would continue to be periodically flooded and restored thereafter. Pliocene marine formations are exposed in northeast Spain , southern California , New Zealand , and Italy . During the Pliocene parts of southern Norway and southern Sweden that had been near sea level rose. In Norway this rise elevated

9118-671: Was inversely correlated with carbon dioxide levels and global temperatures during the Miocene. Most modern lineages of diatoms appeared by the Late Miocene. There is evidence from oxygen isotopes at Deep Sea Drilling Program sites that ice began to build up in Antarctica about 36 Ma during the Eocene . Further marked decreases in temperature during the Middle Miocene at 15 Ma probably reflect increased ice growth in Antarctica. It can therefore be assumed that East Antarctica had some glaciers during

9215-627: Was most intense there. Around this time the Mi3b glacial event (a massive expansion of Antarctic glaciers) occurred. The East Antarctic Ice Sheet (EAIS) markedly stabilised following the MMCT. The intensification of glaciation caused a decoherence of sediment deposition from the 405 kyr eccentricity cycle. The MMWI ended about 11 Ma, when the Late Miocene Cool Interval (LMCI) started. A major but transient warming occurred around 10.8-10.7 Ma. During

9312-402: Was named by Scottish geologist Charles Lyell ; the name comes from the Greek words μείων ( meíōn , "less") and καινός ( kainós , "new") and means "less recent" because it has 18% fewer modern marine invertebrates than the Pliocene has. The Miocene followed the Oligocene and preceded the Pliocene. As Earth went from the Oligocene through the Miocene and into the Pliocene,

9409-541: Was very similar to what it is today. Unexpectedly, the expansion of grasslands in eastern Africa during this epoch appears to have been decoupled from aridification and not caused by it, as evidenced by their asynchrony. Southwestern Australia hosted heathlands , shrublands , and woodlands with a greater species diversity compared to today during the Middle and Late Pliocene. Three different aridification events occurred around 2.90, 2.59, and 2.56 Ma, and may have been linked to

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