Genus ( / ˈ dʒ iː n ə s / ; pl. : genera / ˈ dʒ ɛ n ər ə / ) is a taxonomic rank above species and below family as used in the biological classification of living and fossil organisms as well as viruses . In binomial nomenclature , the genus name forms the first part of the binomial species name for each species within the genus.
102-475: Around 80, see text . Musa is one of three genera in the family Musaceae . The genus includes 83 species of flowering plants producing edible bananas and plantains , and fiber ( abacá ), used to make paper and cloth. Though they grow as high as trees , banana and plantain plants are not woody and their apparent " stem " is made up of the bases of the huge leaf stalks . Thus, they are technically gigantic herbaceous plants . Banana plants are among
204-475: A synthetic population . In horticulture , the term stable hybrid is used to describe an annual plant that, if grown and bred in a small monoculture free of external pollen (e.g., an air-filtered greenhouse) produces offspring that are "true to type" with respect to phenotype; i.e., a true-breeding organism. Hybridization can occur in the hybrid zones where the geographical ranges of species, subspecies, or distinct genetic lineages overlap. For example,
306-460: A bactrian camel and a dromedary . There are many examples of felid hybrids , including the liger . The oldest-known animal hybrid bred by humans is the kunga equid hybrid produced as a draft animal and status symbol 4,500 years ago in Umm el-Marra , present-day Syria . The first known instance of hybrid speciation in marine mammals was discovered in 2014. The clymene dolphin ( Stenella clymene )
408-419: A petiole , and a terminal leaf blade. The false trunk is an aggregation of leaf sheaths; only when the plant is ready to flower does a true stem grow up through the sheath and droop back down. At the end of this stem, a peduncle forms (with M. ingens having the second-longest peduncle known, exceeded only by Agave salmiana ), bearing many female flowers protected by large purple-red bracts. The extension of
510-555: A bear shot by a hunter in the Northwest Territories confirmed the existence of naturally occurring and fertile grizzly–polar bear hybrids . Hybridization between reproductively isolated species often results in hybrid offspring with lower fitness than either parental. However, hybrids are not, as might be expected, always intermediate between their parents (as if there were blending inheritance), but are sometimes stronger or perform better than either parental lineage or variety,
612-508: A considerable seed yield advantage over open pollinated varieties. Hybrid seed dominates the commercial maize seed market in the United States, Canada and many other major maize-producing countries. In a hybrid, any trait that falls outside the range of parental variation (and is thus not simply intermediate between its parents) is considered heterotic. Positive heterosis produces more robust hybrids, they might be stronger or bigger; while
714-504: A cross between a female horse and a male donkey, and the hinny, a cross between a female donkey and a male horse. Pairs of complementary types like the mule and hinny are called reciprocal hybrids. Polar bears and brown bears are another case of a hybridizing species pairs, and introgression among non-sister species of bears appears to have shaped the Ursidae family tree. Among many other mammal crosses are hybrid camels , crosses between
816-501: A degree that none of the originally genetically distinct population remains. In agriculture and animal husbandry , the Green Revolution 's use of conventional hybridization increased yields by breeding high-yielding varieties . The replacement of locally indigenous breeds, compounded with unintentional cross-pollination and crossbreeding (genetic mixing), has reduced the gene pools of various wild and indigenous breeds resulting in
918-407: A different organism is called a chimera . Hybrids are not always intermediates between their parents such as in blending inheritance (a now discredited theory in modern genetics by particulate inheritance ), but can show hybrid vigor , sometimes growing larger or taller than either parent. The concept of a hybrid is interpreted differently in animal and plant breeding, where there is interest in
1020-602: A donkey as the father. A variety of mechanisms limit the success of hybridization, including the large genetic difference between most species. Barriers include morphological differences, differing times of fertility, mating behaviors and cues, and physiological rejection of sperm cells or the developing embryo. Some act before fertilization; others after it. In plants, some barriers to hybridization include blooming period differences, different pollinator vectors, inhibition of pollen tube growth, somatoplastic sterility, cytoplasmic-genic male sterility and structural differences of
1122-1065: A form * muku . From there, the term was borrowed into the Austronesian languages of the area, and migrated across Asia, via the Dravidian languages of India, into Persian , Greek, and Arabic as a Wanderwort : The late Latin term musa was later chosen by Carl Linnaeus in 1753, as the name for the genus. From the time of Linnaeus until the 1940s, different types of edible bananas and plantains were given Linnaean binomial names, such as Musa cavendishii , as if they were species. In fact, edible bananas have an extremely complicated origin involving hybridization , mutation , and finally selection by humans. Most edible bananas are seedless ( parthenocarpic ), hence sterile, so they are propagated vegetatively. The giving of species names to what are actually very complex, largely asexual, hybrids (mostly of two species of wild bananas, Musa acuminata and Musa balbisiana ) led to endless confusion in banana botany . In
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#17330923368911224-465: A group from section Musa to which M. schizocarpa has also contributed, and a group of hybrids between section Musa and section Callimusa . Banana and plantains are the fourth most produced food globally surpassed only by the staple crops of rice , wheat and maize . Plants of the Musa spp. including roots, flowers and fruits have been used in the folk medicine cultures of Africa, Asia, India and
1326-548: A group of about 50 natural hybrids between Australian blacktip shark and the larger common blacktip shark was found by Australia's eastern coast in 2012. Russian sturgeon and American paddlefish were hybridized in captivity when sperm from the paddlefish and eggs from the sturgeon were combined, unexpectedly resulting in viable offspring. This hybrid is called a sturddlefish . The two genera Asymmetron and Branchiostoma are able to produce viable hybrid offspring, even if none have lived into adulthood so far, despite
1428-530: A history dating back to 1887, when M.P. Sagot published "Sur le genre Bananier", where the genus Musa was first formally classified. In this article, Sagot arranged the Musa species into three groups, although no section names were assigned to them. The grouping was based on morphological traits, establishing the trio as bananas with fleshy fruit, ornamental bananas with upright inflorescences and bracts that were vibrantly colored, and bananas that were giant in size. Five years after Sagot's article, J.G. Baker made
1530-643: A later homonym of a validly published name is a nomen illegitimum or nom. illeg. ; for a full list refer to the International Code of Nomenclature for algae, fungi, and plants and the work cited above by Hawksworth, 2010. In place of the "valid taxon" in zoology, the nearest equivalent in botany is " correct name " or "current name" which can, again, differ or change with alternative taxonomic treatments or new information that results in previously accepted genera being combined or split. Prokaryote and virus codes of nomenclature also exist which serve as
1632-621: A long time and redescribed as new by a range of subsequent workers, or if a range of genera previously considered separate taxa have subsequently been consolidated into one. For example, the World Register of Marine Species presently lists 8 genus-level synonyms for the sperm whale genus Physeter Linnaeus, 1758, and 13 for the bivalve genus Pecten O.F. Müller, 1776. Within the same kingdom, one generic name can apply to one genus only. However, many names have been assigned (usually unintentionally) to two or more different genera. For example,
1734-439: A long time, both intentionally for purposes such as biological control , and unintentionally, as with accidental escapes of individuals. Introductions can drastically affect populations, including through hybridization. There is a kind of continuum with three semi-distinct categories dealing with anthropogenic hybridization: hybridization without introgression, hybridization with widespread introgression (backcrossing with one of
1836-494: A mate among the species that raised it, instead of the species of its biological parents. Cagebird breeders sometimes breed bird hybrids known as mules between species of finch , such as goldfinch × canary . Among amphibians, Japanese giant salamanders and Chinese giant salamanders have created hybrids that threaten the survival of Japanese giant salamanders because of competition for similar resources in Japan. Among fish,
1938-432: A phenomenon called heterosis, hybrid vigour, or heterozygote advantage . This is most common with plant hybrids. A transgressive phenotype is a phenotype that displays more extreme characteristics than either of the parent lines. Plant breeders use several techniques to produce hybrids, including line breeding and the formation of complex hybrids. An economically important example is hybrid maize (corn), which provides
2040-409: A reference for designating currently accepted genus names as opposed to others which may be either reduced to synonymy, or, in the case of prokaryotes, relegated to a status of "names without standing in prokaryotic nomenclature". An available (zoological) or validly published (botanical) name that has been historically applied to a genus but is not regarded as the accepted (current/valid) name for
2142-405: A species, such as between different breeds . Single cross hybrids result from the cross between two true-breeding organisms which produces an F1 hybrid (first filial generation). The cross between two different homozygous lines produces an F1 hybrid that is heterozygous ; having two alleles , one contributed by each parent and typically one is dominant and the other recessive . Typically,
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#17330923368912244-427: A taxon; however, the names published in suppressed works are made unavailable via the relevant Opinion dealing with the work in question. In botany, similar concepts exist but with different labels. The botanical equivalent of zoology's "available name" is a validly published name . An invalidly published name is a nomen invalidum or nom. inval. ; a rejected name is a nomen rejiciendum or nom. rej. ;
2346-455: A total of c. 520,000 published names (including synonyms) as at end 2019, increasing at some 2,500 published generic names per year. "Official" registers of taxon names at all ranks, including genera, exist for a few groups only such as viruses and prokaryotes, while for others there are compendia with no "official" standing such as Index Fungorum for fungi, Index Nominum Algarum and AlgaeBase for algae, Index Nominum Genericorum and
2448-577: Is a hybrid of two Atlantic species, the spinner and striped dolphins . In 2019, scientists confirmed that a skull found 30 years earlier was a hybrid between the beluga whale and narwhal , dubbed the narluga . Hybridization between species is common in birds. Hybrid birds are purposefully bred by humans, but hybridization is also common in the wild. Waterfowl have a particularly high incidence of hybridization, with at least 60% of species known to produce hybrids with another species. Among ducks , mallards widely hybridize with many other species, and
2550-407: Is a hybridization test widely used in genetics to determine whether two separately isolated mutants that have the same (or similar) phenotype are defective in the same gene or in different genes (see Complementation (genetics) article). If a hybrid organism containing the genomes of two different mutant parental organisms displays a wild type phenotype, it is ordinarily considered that
2652-571: Is a particularly common mechanism for speciation in plants, and is now known to be fundamental to the evolutionary history of plants. Plants frequently form polyploids , individuals with more than two copies of each chromosome. Whole genome doubling has occurred repeatedly in plant evolution. When two plant species hybridize, the hybrid may double its chromosome count by incorporating the entire nuclear genome of both parents, resulting in offspring that are reproductively incompatible with either parent because of different chromosome counts. Human impact on
2754-405: Is a triploid, with three sets of chromosomes, all derived from Musa acuminata , which is designated by the letter "A". When Musa balbisiana is involved, the letter "B" is used to denote its genome. Thus, the cultivar 'Rajapuri' may be called Musa (AAB Group) 'Rajapuri'. 'Rajapuri' is also a triploid, expected to have two sets of chromosomes from Musa acuminata and one from Musa balbisiana . In
2856-659: Is also more occasionally done in the livestock and pet trades; some well-known wild × domestic hybrids are beefalo and wolfdogs . Human selective breeding of domesticated animals and plants has also resulted in the development of distinct breeds (usually called cultivars in reference to plants); crossbreeds between them (without any wild stock ) are sometimes also imprecisely referred to as "hybrids". Hybrid humans existed in prehistory. For example, Neanderthals and anatomically modern humans are thought to have interbred as recently as 40,000 years ago. Mythological hybrids appear in human culture in forms as diverse as
2958-451: Is always sterile worker ants (and because ants are haplodiploid , unfertilized eggs become males). Without mating with males of the other species, the queens are unable to produce workers, and will fail to establish a colony of their own. Plant species hybridize more readily than animal species, and the resulting hybrids are fertile more often. Many plant species are the result of hybridization, combined with polyploidy , which duplicates
3060-514: Is based on GRIN (where this gives the species), regrouped according to Wong et al. [A] and [C] indicate known placement in the former sections Australimusa and Callimusa , respectively. The native distribution of the genus Musa includes most of the Indomalayan realm and parts of north-eastern Australasia . It has been introduced to many other parts of the world with tropical or subtropical climates. Musa species are used as food plants by
3162-596: Is bright orange, with a high level of beta carotene . Fe'i bananas are no longer very important for food, as imported foods have grown in popularity, although some have ritual significance. Investigations are under way to use the Fe'i karat bananas (the name derives from " carrot " due to the intense orange-yellow color of the fruit) in prevention of childhood blindness in Pohnpei . Fe'i bananas probably derive mainly from Musa maclayi , although their origins are not as well understood as
Musa (genus) - Misplaced Pages Continue
3264-547: Is derived from Latin hybrida , used for crosses such as of a tame sow and a wild boar. The term came into popular use in English in the 19th century, though examples of its use have been found from the early 17th century. Conspicuous hybrids are popularly named with portmanteau words , starting in the 1920s with the breeding of tiger–lion hybrids ( liger and tigon ). From the point of view of animal and plant breeders, there are several kinds of hybrid formed from crosses within
3366-596: Is discouraged by both the International Code of Zoological Nomenclature and the International Code of Nomenclature for algae, fungi, and plants , there are some five thousand such names in use in more than one kingdom. For instance, A list of generic homonyms (with their authorities), including both available (validly published) and selected unavailable names, has been compiled by the Interim Register of Marine and Nonmarine Genera (IRMNG). The type genus forms
3468-464: Is grown for fiber. Genus The composition of a genus is determined by taxonomists . The standards for genus classification are not strictly codified, so different authorities often produce different classifications for genera. There are some general practices used, however, including the idea that a newly defined genus should fulfill these three criteria to be descriptively useful: Moreover, genera should be composed of phylogenetic units of
3570-460: Is somewhat arbitrary. Although all species within a genus are supposed to be "similar", there are no objective criteria for grouping species into genera. There is much debate among zoologists about whether enormous, species-rich genera should be maintained, as it is extremely difficult to come up with identification keys or even character sets that distinguish all species. Hence, many taxonomists argue in favor of breaking down large genera. For instance,
3672-427: Is that hybrid individuals can form a "bridge" transmitting potentially helpful genes from one species to another when the hybrid backcrosses with one of its parent species, a process called introgression . Hybrids can also cause speciation , either because the hybrids are genetically incompatible with their parents and not each other, or because the hybrids occupy a different niche than either parent. Hybridization
3774-474: Is the type species , and the generic name is permanently associated with the type specimen of its type species. Should the specimen turn out to be assignable to another genus, the generic name linked to it becomes a junior synonym and the remaining taxa in the former genus need to be reassessed. In zoological usage, taxonomic names, including those of genera, are classified as "available" or "unavailable". Available names are those published in accordance with
3876-515: Is the proper time to give up on a population that is becoming a hybrid swarm, or to try and save the still existing pure individuals. Once a population becomes a complete mixture, the goal becomes to conserve those hybrids to avoid their loss. Conservationists treat each case on its merits, depending on detecting hybrids within the population. It is nearly impossible to formulate a uniform hybridization policy, because hybridization can occur beneficially when it occurs "naturally", and when hybrid swarms are
3978-621: The International Code of Zoological Nomenclature ; the earliest such name for any taxon (for example, a genus) should then be selected as the " valid " (i.e., current or accepted) name for the taxon in question. Consequently, there will be more available names than valid names at any point in time; which names are currently in use depending on the judgement of taxonomists in either combining taxa described under multiple names, or splitting taxa which may bring available names previously treated as synonyms back into use. "Unavailable" names in zoology comprise names that either were not published according to
4080-799: The International Plant Names Index for plants in general, and ferns through angiosperms, respectively, and Nomenclator Zoologicus and the Index to Organism Names for zoological names. Totals for both "all names" and estimates for "accepted names" as held in the Interim Register of Marine and Nonmarine Genera (IRMNG) are broken down further in the publication by Rees et al., 2020 cited above. The accepted names estimates are as follows, broken down by kingdom: The cited ranges of uncertainty arise because IRMNG lists "uncertain" names (not researched therein) in addition to known "accepted" names;
4182-638: The Minotaur , blends of animals, humans and mythical beasts such as centaurs and sphinxes , and the Nephilim of the Biblical apocrypha described as the wicked sons of fallen angels and attractive women. Hybridization between species plays an important role in evolution, though there is much debate about its significance. Roughly 25% of plants and 10% of animals are known to form hybrids with at least one other species. One example of an adaptive benefit to hybridization
Musa (genus) - Misplaced Pages Continue
4284-540: The larvae of some Lepidoptera species, including the giant leopard moth and other Hypercompe species, including H. albescens (only recorded on Musa ), H. eridanus , and H. icasia . A number of distinct groups of plants bearing edible fruit have been developed from species of Musa . In English, fruits which are sweet and used for dessert are usually called "bananas", whereas starchier varieties used for cooking are called "plantains", but these terms do not have any botanical significance. By far
4386-419: The nomenclature codes , which allow each species a single unique name that, for animals (including protists ), plants (also including algae and fungi ) and prokaryotes ( bacteria and archaea ), is Latin and binomial in form; this contrasts with common or vernacular names , which are non-standardized, can be non-unique, and typically also vary by country and language of usage. Except for viruses ,
4488-404: The platypus belongs to the genus Ornithorhynchus although George Shaw named it Platypus in 1799 (these two names are thus synonyms ) . However, the name Platypus had already been given to a group of ambrosia beetles by Johann Friedrich Wilhelm Herbst in 1793. A name that means two different things is a homonym . Since beetles and platypuses are both members of the kingdom Animalia,
4590-464: The sea urchin Strongylocentrotus purpuratus (female) and the sand dollar Dendraster excentricus (male). When two distinct types of organisms breed with each other, the resulting hybrids typically have intermediate traits (e.g., one plant parent has red flowers, the other has white, and the hybrid, pink flowers). Commonly, hybrids also combine traits seen only separately in one parent or
4692-438: The "suture region". It is at these regions that the subspecies were formed. Other hybrid zones have formed between described species of plants and animals. From the point of view of genetics, several different kinds of hybrid can be distinguished. A genetic hybrid carries two different alleles of the same gene , where for instance one allele may code for a lighter coat colour than the other. A structural hybrid results from
4794-468: The 1940s and 1950s, it became clear to botanists that the cultivated bananas and plantains could not usefully be assigned Linnean binomials, but were better given cultivar names. As for the word banana , it came to English from Spanish and Portuguese, which had apparently obtained it from a West African language, possibly Wolof (Senegal). The Wolof form might itself be a loanword from Arabic banān mawz (< banān 'finger'). Musa sections have
4896-625: The Americas. Modern studies examining the properties of the fruits have found diversity of bioactive compounds among genotypes compared with commercially grown cultivars . When the Linnaean binomial system was abandoned for cultivated bananas, an alternate genome-based system for the nomenclature of edible bananas in section Musa was devised. Thus, the plant previously known by the "species" name Musa cavendishii became Musa (AAA Group) 'Dwarf Cavendish'. The "new" name shows clearly that 'Dwarf Cavendish'
4998-503: The F1 generation is also phenotypically homogeneous, producing offspring that are all similar to each other. Double cross hybrids result from the cross between two different F1 hybrids (i.e., there are four unrelated grandparents). Three-way cross hybrids result from the cross between an F1 hybrid and an inbred line. Triple cross hybrids result from the crossing of two different three-way cross hybrids. Top cross (or "topcross") hybrids result from
5100-482: The banana, although Pliny says that a single fruit was sufficient to satisfy four persons. During the late Middle Ages , international trade brought bananas to Europe, which created the need for a name. In the 11th century, Medieval Latin innovated a term musa : this was most likely the latinization of the Arabic name for the fruit, mauz (موز). Thus, the 11th-century Arabic encyclopedia The Canon of Medicine , which
5202-442: The base for higher taxonomic ranks, such as the family name Canidae ("Canids") based on Canis . However, this does not typically ascend more than one or two levels: the order to which dogs and wolves belong is Carnivora ("Carnivores"). The numbers of either accepted, or all published genus names is not known precisely; Rees et al., 2020 estimate that approximately 310,000 accepted names (valid taxa) may exist, out of
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#17330923368915304-511: The butterfly Limenitis arthemis has two major subspecies in North America, L. a. arthemis (the white admiral) and L. a. astyanax (the red-spotted purple). The white admiral has a bright, white band on its wings, while the red-spotted purple has cooler blue-green shades. Hybridization occurs between a narrow area across New England, southern Ontario, and the Great Lakes,
5406-426: The chromosomes. A few animal species are the result of hybridization. The Lonicera fly is a natural hybrid. The American red wolf appears to be a hybrid of the gray wolf and the coyote , although its taxonomic status has been a subject of controversy. The European edible frog is a semi-permanent hybrid between pool frogs and marsh frogs ; its population requires the continued presence of at least one of
5508-539: The chromosomes. Chromosome duplication allows orderly meiosis and so viable seed can be produced. Plant hybrids are generally given names that include an "×" (not in italics), such as Platanus × hispanica for the London plane, a natural hybrid of P. orientalis (oriental plane) and P. occidentalis (American sycamore). The parent's names may be kept in their entirety, as seen in Prunus persica × Prunus americana , with
5610-481: The common pheasant and hen golden pheasant ( Chrysolophus pictus ). Spurs are absent in hybrids of the former type, although present in both parents. Hybridization is greatly influenced by human impact on the environment, through effects such as habitat fragmentation and species introductions. Such impacts make it difficult to conserve the genetics of populations undergoing introgressive hybridization . Humans have introduced species worldwide to environments for
5712-400: The crossing of a top quality or pure-bred male and a lower quality female, intended to improve the quality of the offspring, on average. Population hybrids result from the crossing of plants or animals in one population with those of another population. These include interspecific hybrids or crosses between different breeds. In biology, the result of crossing of two populations is called
5814-444: The developing embryo . Some act before fertilization and others after it. Similar barriers exist in plants, with differences in flowering times, pollen vectors, inhibition of pollen tube growth, somatoplastic sterility, cytoplasmic-genic male sterility and the structure of the chromosomes. A few animal species and many plant species, however, are the result of hybrid speciation , including important crop plants such as wheat , where
5916-407: The diverse Heliconius butterflies , but that is disputed. The two closely related harvester ant species Pogonomyrmex barbatus and Pogonomyrmex rugosus have evolved to depend on hybridization. When a queen fertilizes her eggs with sperm from males of her own species, the offspring is always new queens. And when she fertilizes the eggs with sperm from males of the other species, the offspring
6018-413: The doubling of chromosome sets, causing immediate genetic isolation. Hybridization may be important in speciation in some plant groups. However, homoploid hybrid speciation (not increasing the number of sets of chromosomes) may be rare: by 1997, only eight natural examples had been fully described. Experimental studies suggest that hybridization offers a rapid route to speciation, a prediction confirmed by
6120-611: The environment has resulted in an increase in the interbreeding between regional species, and the proliferation of introduced species worldwide has also resulted in an increase in hybridization. This has been referred to as genetic pollution out of concern that it may threaten many species with extinction. Similarly, genetic erosion from monoculture in crop plants may be damaging the gene pools of many species for future breeding. The conservation impacts of hybridization between species are highly debated. While hybridization could potentially threaten rare species or lineages by "swamping"
6222-429: The fact that early generation hybrids and ancient hybrid species have matching genomes, meaning that once hybridization has occurred, the new hybrid genome can remain stable. Many hybrid zones are known where the ranges of two species meet, and hybrids are continually produced in great numbers. These hybrid zones are useful as biological model systems for studying the mechanisms of speciation. Recently DNA analysis of
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#17330923368916324-465: The first formal designation of Musa sections. To do so, he named three subgenera that almost paralleled the sections that had been described by Sagot. These sections were: After this classification, in 1947, Cheeseman reclassified the taxa based on morphological features and chromosome number. This project proposed four sections: The addition of another Musa section came in 1976 by G.C.G. Ardent. The added section, M . sect. Ingentimusa , Ardent
6426-413: The five sections of Musa defined by morphology (and listed above) were not monophyletic. Based on the incorrect section grouping, Markku Häkkinen proposed another reclassification of the Musa sections in 2013. Using a multitude of genetic evidence and markers from other studies, Häkkinen suggested the reduction of five Musa sections into two: Musa and Callimusa . Unlike sectional classifications of
6528-446: The form "author, year" in zoology, and "standard abbreviated author name" in botany. Thus in the examples above, the genus Canis would be cited in full as " Canis Linnaeus, 1758" (zoological usage), while Hibiscus , also first established by Linnaeus but in 1753, is simply " Hibiscus L." (botanical usage). Each genus should have a designated type , although in practice there is a backlog of older names without one. In zoology, this
6630-560: The fusion of gametes that have differing structure in at least one chromosome, as a result of structural abnormalities . A numerical hybrid results from the fusion of gametes having different haploid numbers of chromosomes . A permanent hybrid results when only the heterozygous genotype occurs, as in Oenothera lamarckiana , because all homozygous combinations are lethal. In the early history of genetics, Hugo de Vries supposed these were caused by mutation . Genetic complementation
6732-727: The generic name (or its abbreviated form) still forms the leading portion of the scientific name, for example, Canis lupus lupus for the Eurasian wolf subspecies, or as a botanical example, Hibiscus arnottianus ssp. immaculatus . Also, as visible in the above examples, the Latinised portions of the scientific names of genera and their included species (and infraspecies, where applicable) are, by convention, written in italics . The scientific names of virus species are descriptive, not binomial in form, and may or may not incorporate an indication of their containing genus; for example,
6834-507: The genetic relationships between ducks are further complicated by the widespread gene flow between wild and domestic mallards. One of the most common interspecific hybrids in geese occurs between Greylag and Canada geese ( Anser anser x Branta canadensis ). One potential mechanism for the occurrence of hybrids in these geese is interspecific nest parasitism , where an egg is laid in the nest of another species to be raised by non-biological parents. The chick imprints upon and eventually seeks
6936-630: The genetically "pure" individuals with hybrids, hybridization could also save a rare lineage from extinction by introducing genetic diversity. It has been proposed that hybridization could be a useful tool to conserve biodiversity by allowing organisms to adapt, and that efforts to preserve the separateness of a "pure" lineage could harm conservation by lowering the organisms' genetic diversity and adaptive potential, particularly in species with low populations. While endangered species are often protected by law, hybrids are often excluded from protection, resulting in challenges to conservation. The term hybrid
7038-564: The genome of edible bananas from section Musa , combinations such as AA, BB, ABB, BBB and even AAAB can be found. No such nomenclature system has been developed for the group of edible bananas derived from section Callimusa . This group is known generally as the "Fe'i" or "Fehi" bananas, and numerous cultivars are found in the South Pacific region. They are very distinctive plants with upright fruit bunches, featuring in three of Paul Gauguin 's paintings. The flesh can be cooked before eating and
7140-401: The hybrids are more fit and have breeding advantages over the indigenous ecotype or species. These hybridization events can result from the introduction of non-native genotypes by humans or through habitat modification, bringing previously isolated species into contact. Genetic mixing can be especially detrimental for rare species in isolated habitats, ultimately affecting the population to such
7242-407: The individual parentage. In genetics , attention is focused on the numbers of chromosomes . In taxonomy, a key question is how closely related the parent species are. Species are reproductively isolated by strong barriers to hybridization, which include genetic and morphological differences, differing times of fertility, mating behaviors and cues, and physiological rejection of sperm cells or
7344-465: The largest and now the most widely distributed group of cultivated bananas is derived from section Musa , particularly M. acuminata and M. balbisiana , either alone or in various hybrid combinations. The next but much smaller group is derived from members of section Callimusa (previously classified as Australimusa ) and is restricted in importance to Polynesia . Of even more restricted importance are small groups of hybrids from Papua New Guinea ;
7446-628: The largest component, with 23,236 ± 5,379 accepted genus names, of which 20,845 ± 4,494 are angiosperms (superclass Angiospermae). By comparison, the 2018 annual edition of the Catalogue of Life (estimated >90% complete, for extant species in the main) contains currently 175,363 "accepted" genus names for 1,744,204 living and 59,284 extinct species, also including genus names only (no species) for some groups. The number of species in genera varies considerably among taxonomic groups. For instance, among (non-avian) reptiles , which have about 1180 genera,
7548-414: The largest extant herbaceous plants, some reaching up to 9 m (30 ft) in height or 18 m (59 ft) in the case of Musa ingens . The large herb is composed of a modified underground stem ( rhizome ), a false trunk or pseudostem formed by the basal parts of tightly rolled leaves, a network of roots, and a large flower spike. A single leaf is divided into a leaf sheath, a contracted part called
7650-531: The lizard genus Anolis has been suggested to be broken down into 8 or so different genera which would bring its ~400 species to smaller, more manageable subsets. Hybrid (biology) In biology , a hybrid is the offspring resulting from combining the qualities of two organisms of different varieties , subspecies , species or genera through sexual reproduction . Generally, it means that each cell has genetic material from two different organisms, whereas an individual where some cells are derived from
7752-400: The loss of genetic diversity . Since the indigenous breeds are often well-adapted to local extremes in climate and have immunity to local pathogens, this can be a significant genetic erosion of the gene pool for future breeding. Therefore, commercial plant geneticists strive to breed "widely adapted" cultivars to counteract this tendency. Familiar examples of equid hybrids are the mule,
7854-403: The most (>300) have only 1 species, ~360 have between 2 and 4 species, 260 have 5–10 species, ~200 have 11–50 species, and only 27 genera have more than 50 species. However, some insect genera such as the bee genera Lasioglossum and Andrena have over 1000 species each. The largest flowering plant genus, Astragalus , contains over 3,000 species. Which species are assigned to a genus
7956-428: The name could not be used for both. Johann Friedrich Blumenbach published the replacement name Ornithorhynchus in 1800. However, a genus in one kingdom is allowed to bear a scientific name that is in use as a generic name (or the name of a taxon in another rank) in a kingdom that is governed by a different nomenclature code. Names with the same form but applying to different taxa are called "homonyms". Although this
8058-452: The number of chromosomes has been doubled. A form of often intentional human-mediated hybridization is the crossing of wild and domesticated species. This is common in both traditional horticulture and modern agriculture ; many commercially useful fruits, flowers, garden herbs, and trees have been produced by hybridization. One such flower, Oenothera lamarckiana , was central to early genetics research into mutationism and polyploidy. It
8160-420: The only remaining evidence of prior species, they need to be conserved as well. Regionally developed ecotypes can be threatened with extinction when new alleles or genes are introduced that alter that ecotype. This is sometimes called genetic mixing. Hybridization and introgression, which can happen in natural and hybrid populations, of new genetic material can lead to the replacement of local genotypes if
8262-438: The other (e.g., a bird hybrid might combine the yellow head of one parent with the orange belly of the other). Interspecific hybrids are bred by mating individuals from two species, normally from within the same genus. The offspring display traits and characteristics of both parents, but are often sterile , preventing gene flow between the species. Sterility is often attributed to the different number of chromosomes between
8364-573: The parent species), and hybrid swarms (highly variable populations with much interbreeding as well as backcrossing with the parent species). Depending on where a population falls along this continuum, the management plans for that population will change. Hybridization is currently an area of great discussion within wildlife management and habitat management. Global climate change is creating other changes such as difference in population distributions which are indirect causes for an increase in anthropogenic hybridization. Conservationists disagree on when
8466-532: The parent species. Cave paintings indicate that the European bison is a natural hybrid of the aurochs and the steppe bison . Plant hybridization is more commonplace compared to animal hybridization. Many crop species are hybrids, including notably the polyploid wheats : some have four sets of chromosomes (tetraploid) or six (hexaploid), while other wheat species have (like most eukaryotic organisms) two sets ( diploid ), so hybridization events likely involved
8568-506: The parents' common ancestor living tens of millions of years ago. Among insects, so-called killer bees were accidentally created during an attempt to breed a strain of bees that would both produce more honey and be better adapted to tropical conditions. It was done by crossing a European honey bee and an African bee . The Colias eurytheme and C. philodice butterflies have retained enough genetic compatibility to produce viable hybrid offspring. Hybrid speciation may have produced
8670-701: The past, this hypothesis was based on genetic markers rather than morphological features or chromosome number. The two groups were generally formed by the clustering of the previously defined groups: The advance of genomic analysis technologies and further data on the relatedness of Musa species, formulated Häkkinen's two sections and later corroborated them as correct subcategories for the genus. The history of Musa sections provides an example of genomics superseding morphological evidence and thus classifications. The World Checklist of Selected Plant Families accepts 68 species and two primary hybrids, as of January 2013, which are listed below. The assignment to sections
8772-637: The point of view of taxonomy , hybrids differ according to their parentage. Hybrids between different subspecies (such as between the dog and Eurasian wolf ) are called intra-specific hybrids. Interspecific hybrids are the offspring from interspecies mating ; these sometimes result in hybrid speciation. Intergeneric hybrids result from matings between different genera, such as between sheep and goats . Interfamilial hybrids, such as between chickens and guineafowl or pheasants , are reliably described but extremely rare. Interordinal hybrids (between different orders) are few, but have been engineered between
8874-560: The presence of latex-producing cells, flowers with five connate tepals and one member of the inner whorl distinct, and a petiole with one row of air channels. Bananas have a green skin when unripe. While plantains remain green all the way, many varieties change their skin to a yellow, orange or reddish colour as they ripen. Pliny the Elder used the name ariena for the fruit of a tree found in India. It has been speculated that this might have been
8976-526: The provisions of the ICZN Code, e.g., incorrect original or subsequent spellings, names published only in a thesis, and generic names published after 1930 with no type species indicated. According to "Glossary" section of the zoological Code, suppressed names (per published "Opinions" of the International Commission of Zoological Nomenclature) remain available but cannot be used as the valid name for
9078-466: The same kind as other (analogous) genera. The term "genus" comes from Latin genus , a noun form cognate with gignere ('to bear; to give birth to'). The Swedish taxonomist Carl Linnaeus popularized its use in his 1753 Species Plantarum , but the French botanist Joseph Pitton de Tournefort (1656–1708) is considered "the founder of the modern concept of genera". The scientific name (or
9180-408: The scientific epithet) of a genus is also called the generic name ; in modern style guides and science, it is always capitalised. It plays a fundamental role in binomial nomenclature , the system of naming organisms , where it is combined with the scientific name of a species : see Botanical name and Specific name (zoology) . The rules for the scientific names of organisms are laid down in
9282-502: The section Musa bananas. Cultivars can be formally named, as e.g. Musa (Fe'i Group) 'Utafun'. In addition to the edible fruits, the flowers can be eaten cooked, and the heart of the plant (like heart of palm ) can be eaten raw or cooked. Additionally, the rootstocks and leaf sheaths of some species can be cooked and eaten. The leaves are used in several cultures as cooking wrappers, such as for Puerto Rican pasteles or Indonesian pepes , and for food plating. Musa textilis (abacá)
9384-497: The specific name particular to the wolf. A botanical example would be Hibiscus arnottianus , a particular species of the genus Hibiscus native to Hawaii. The specific name is written in lower-case and may be followed by subspecies names in zoology or a variety of infraspecific names in botany . When the generic name is already known from context, it may be shortened to its initial letter, for example, C. lupus in place of Canis lupus . Where species are further subdivided,
9486-412: The standard format for a species name comprises the generic name, indicating the genus to which the species belongs, followed by the specific epithet, which (within that genus) is unique to the species. For example, the gray wolf 's scientific name is Canis lupus , with Canis ( Latin for 'dog') being the generic name shared by the wolf's close relatives and lupus (Latin for 'wolf') being
9588-631: The stem (the rachis) continues growth downward, where terminal male flowers grow. The leaves originate from a pseudostem and unroll to show a leaf blade with two lamina halves. The lamina can be as much as 7 m (23 ft) long in the case of M. acuminata subsp. truncata (syn. M. truncata ) of the Malay Peninsula ). Musa species reproduce by both sexual (seed) and asexual (suckers) processes, using asexual means when producing sterile (unseeded) fruits. Further qualities to distinguish Musa include spirally arranged leaves, fruits as berries,
9690-403: The taxon is termed a synonym ; some authors also include unavailable names in lists of synonyms as well as available names, such as misspellings, names previously published without fulfilling all of the requirements of the relevant nomenclatural code, and rejected or suppressed names. A particular genus name may have zero to many synonyms, the latter case generally if the genus has been known for
9792-470: The term negative heterosis refers to weaker or smaller hybrids. Heterosis is common in both animal and plant hybrids. For example, hybrids between a lion and a tigress (" ligers ") are much larger than either of the two progenitors, while " tigons " (lioness × tiger) are smaller. Similarly, the hybrids between the common pheasant ( Phasianus colchicus ) and domestic fowl ( Gallus gallus ) are larger than either of their parents, as are those produced between
9894-400: The two parental mutant organisms are defective in different genes. If the hybrid organism displays a distinctly mutant phenotype, the two mutant parental organisms are considered to be defective in the same gene. However, in some cases the hybrid organism may display a phenotype that is only weakly (or partially) wild-type, and this may reflect intragenic (interallelic) complementation. From
9996-462: The two species. For example, donkeys have 62 chromosomes , horses have 64 chromosomes, and mules or hinnies have 63 chromosomes. Mules, hinnies, and other normally sterile interspecific hybrids cannot produce viable gametes, because differences in chromosome structure prevent appropriate pairing and segregation during meiosis , meiosis is disrupted, and viable sperm and eggs are not formed. However, fertility in female mules has been reported with
10098-566: The values quoted are the mean of "accepted" names alone (all "uncertain" names treated as unaccepted) and "accepted + uncertain" names (all "uncertain" names treated as accepted), with the associated range of uncertainty indicating these two extremes. Within Animalia, the largest phylum is Arthropoda , with 151,697 ± 33,160 accepted genus names, of which 114,387 ± 27,654 are insects (class Insecta). Within Plantae, Tracheophyta (vascular plants) make up
10200-429: The virus species " Salmonid herpesvirus 1 ", " Salmonid herpesvirus 2 " and " Salmonid herpesvirus 3 " are all within the genus Salmonivirus ; however, the genus to which the species with the formal names " Everglades virus " and " Ross River virus " are assigned is Alphavirus . As with scientific names at other ranks, in all groups other than viruses, names of genera may be cited with their authorities, typically in
10302-496: Was based on a single species, Musa ingens . This designation put the number of sections in Musa at five: Eumusa , Rhodochlamys , Callimusa , Australimusa , and Ingentimusa . In the 21st century, genomics have become cheaper, more efficient, and more accurate, and Musa genetic research has increased exponentially. Research was conducted around a diversity of genomic markers (cpDNA, nrDNA , rDNA, introns, various spacers, etc.). The results of many of these studies suggested that
10404-615: Was translated to Latin in medieval times and well known in Europe, shows a correspondence between Arabic mauz and Latin musa . Muz is also the Turkish, Persian, and Somali name for the fruit. According to linguist Mark Donohue and archaeologist Tim Denham, the ultimate origin of the Latinized form musa is in the Trans–New Guinea languages , where certain cultivars of bananas are known under
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