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120-510: The yellow-headed temple turtle ( Heosemys annandalii ) is a large species of turtle in the family Geoemydidae . The species is native to Southeast Asia . The common name, "yellow-headed temple turtle", is derived from the fact that it is often found near Buddhist temples within its range. The specific name , annandalii , is in honor of Scottish herpetologist Nelson Annandale . H. annandalii may grow to over 20 in (51 cm) in straight carapace length. H. annandalii

240-423: A mutation–selection balance . It is predicted that a viral quasispecies at a low but evolutionarily neutral and highly connected (that is, flat) region in the fitness landscape will outcompete a quasispecies located at a higher but narrower fitness peak in which the surrounding mutants are unfit, "the quasispecies effect" or the "survival of the flattest". There is no suggestion that a viral quasispecies resembles

360-495: A species binomial (estimated range: 1.5–8 million). Less than 1% of all species that have been described beyond simply noting its existence. From these figures, the IUCN reports that 23% of vertebrates , 5% of invertebrates and 70% of plants that have been evaluated are designated as endangered or threatened . Better knowledge is being constructed by The Plant List for actual numbers of species. Systematic conservation planning

480-515: A taxonomic rank of an organism, as well as a unit of biodiversity . Other ways of defining species include their karyotype , DNA sequence, morphology , behaviour, or ecological niche . In addition, paleontologists use the concept of the chronospecies since fossil reproduction cannot be examined. The most recent rigorous estimate for the total number of species of eukaryotes is between 8 and 8.7 million. About 14% of these had been described by 2011. All species (except viruses ) are given

600-542: A two-part name , called a "binomial". The first part of a binomial is the genus to which the species belongs. The second part is called the specific name or the specific epithet (in botanical nomenclature , also sometimes in zoological nomenclature ). For example, Boa constrictor is one of the species of the genus Boa , with constrictor being the species' epithet. While the definitions given above may seem adequate at first glance, when looked at more closely they represent problematic species concepts. For example,

720-776: A wildlife sanctuary was founded on the Farne Islands by St Cuthbert in response to his religious beliefs. Natural history was a major preoccupation in the 18th century, with grand expeditions and the opening of popular public displays in Europe and North America . By 1900 there were 150 natural history museums in Germany , 250 in Great Britain , 250 in the United States , and 300 in France . Preservationist or conservationist sentiments are

840-400: A "classical" method of determining species, such as with Linnaeus, early in evolutionary theory. However, different phenotypes are not necessarily different species (e.g. a four-winged Drosophila born to a two-winged mother is not a different species). Species named in this manner are called morphospecies . In the 1970s, Robert R. Sokal , Theodore J. Crovello and Peter Sneath proposed

960-424: A 'smallest clade' idea" (a phylogenetic species concept). Mishler and Wilkins and others concur with this approach, even though this would raise difficulties in biological nomenclature. Wilkins cited the ichthyologist Charles Tate Regan 's early 20th century remark that "a species is whatever a suitably qualified biologist chooses to call a species". Wilkins noted that the philosopher Philip Kitcher called this

1080-428: A connected series of neighbouring populations, each of which can sexually interbreed with adjacent related populations, but for which there exist at least two "end" populations in the series, which are too distantly related to interbreed, though there is a potential gene flow between each "linked" population. Such non-breeding, though genetically connected, "end" populations may co-exist in the same region thus closing

1200-510: A conservation project, it is important to understand how an ecosystem functions and what role different species and abiotic factors have within the system. It is important to have a precise reason for why ecological monitoring is implemented; within the context of conservation, this reasoning is often to track changes before, during, or after conservation measures are put in place to help a species or habitat recover from degradation and/or maintain integrity. Another benefit of ecological monitoring

1320-417: A development of the late 18th to early 20th centuries. Before Charles Darwin set sail on HMS Beagle , most people in the world, including Darwin, believed in special creation and that all species were unchanged. George-Louis Leclerc was one of the first naturalist that questioned this belief. He proposed in his 44 volume natural history book that species evolve due to environmental influences. Erasmus Darwin

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1440-432: A different species from its ancestors. Viruses have enormous populations, are doubtfully living since they consist of little more than a string of DNA or RNA in a protein coat, and mutate rapidly. All of these factors make conventional species concepts largely inapplicable. A viral quasispecies is a group of genotypes related by similar mutations, competing within a highly mutagenic environment, and hence governed by

1560-518: A faster rate than other species. Amphibians spend parts of their time in the water and on land, making them susceptible to changes in both environments. They also have very permeable skin that allows them to breath and intake water, which means they also take any air or water-soluble pollutants in as well. Birds often cover a wide range in habitat types annually, and also generally revisit the same nesting site each year. This makes it easier for researchers to track ecological effects at both an individual and

1680-508: A genetic boundary suitable for defining a species concept is present. DNA barcoding has been proposed as a way to distinguish species suitable even for non-specialists to use. One of the barcodes is a region of mitochondrial DNA within the gene for cytochrome c oxidase . A database, Barcode of Life Data System , contains DNA barcode sequences from over 190,000 species. However, scientists such as Rob DeSalle have expressed concern that classical taxonomy and DNA barcoding, which they consider

1800-465: A lineage should be divided into multiple chronospecies , or when populations have diverged to have enough distinct character states to be described as cladistic species. Species and higher taxa were seen from the time of Aristotle until the 18th century as categories that could be arranged in a hierarchy, the great chain of being . In the 19th century, biologists grasped that species could evolve given sufficient time. Charles Darwin 's 1859 book On

1920-492: A misnomer, need to be reconciled, as they delimit species differently. Genetic introgression mediated by endosymbionts and other vectors can further make barcodes ineffective in the identification of species. A phylogenetic or cladistic species is "the smallest aggregation of populations (sexual) or lineages (asexual) diagnosable by a unique combination of character states in comparable individuals (semaphoronts)". The empirical basis – observed character states – provides

2040-517: A multitude of other variables such as loss of biodiversity as a function of the rate of habitat loss and site occupancy to obtain such estimates. The Theory of Island Biogeography is possibly the most significant contribution toward the scientific understanding of both the process and how to measure the rate of species extinction. The current background extinction rate is estimated to be one species every few years. Actual extinction rates are estimated to be orders of magnitudes higher. While this

2160-449: A particular species, including which genus (and higher taxa) it is placed in, is a hypothesis about the evolutionary relationships and distinguishability of that group of organisms. As further information comes to hand, the hypothesis may be corroborated or refuted. Sometimes, especially in the past when communication was more difficult, taxonomists working in isolation have given two distinct names to individual organisms later identified as

2280-400: A perfect model of life, it is still a useful tool to scientists and conservationists for studying life on Earth, regardless of the theoretical difficulties. If species were fixed and clearly distinct from one another, there would be no problem, but evolutionary processes cause species to change. This obliges taxonomists to decide, for example, when enough change has occurred to declare that

2400-458: A population level for the species. Many conservation researchers believe that having a long-term ecological monitoring program should be a priority for conservation projects, protected areas, and regions where environmental harm mitigation is used. Conservation biologists are interdisciplinary researchers that practice ethics in the biological and social sciences. Chan states that conservationists must advocate for biodiversity and can do so in

2520-415: A protected existence that halts interference from the humans. In this regard, conservationists differ from preservationists in the social dimension, as conservation biology engages society and seeks equitable solutions for both society and ecosystems. Some preservationists emphasize the potential of biodiversity in a world without humans. Ecological monitoring is the systematic collection of data relevant to

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2640-481: A scientifically ethical manner by not promoting simultaneous advocacy against other competing values. A conservationist may be inspired by the resource conservation ethic , which seeks to identify what measures will deliver "the greatest good for the greatest number of people for the longest time." In contrast, some conservation biologists argue that nature has an intrinsic value that is independent of anthropocentric usefulness or utilitarianism . Aldo Leopold

2760-400: A short way of saying that something applies to many species within a genus, but not to all. If scientists mean that something applies to all species within a genus, they use the genus name without the specific name or epithet. The names of genera and species are usually printed in italics . However, abbreviations such as "sp." should not be italicised. When a species' identity is not clear,

2880-404: A specialist may use "cf." before the epithet to indicate that confirmation is required. The abbreviations "nr." (near) or "aff." (affine) may be used when the identity is unclear but when the species appears to be similar to the species mentioned after. With the rise of online databases, codes have been devised to provide identifiers for species that are already defined, including: The naming of

3000-523: A species as groups of actually or potentially interbreeding natural populations, which are reproductively isolated from other such groups. It has been argued that this definition is a natural consequence of the effect of sexual reproduction on the dynamics of natural selection. Mayr's use of the adjective "potentially" has been a point of debate; some interpretations exclude unusual or artificial matings that occur only in captivity, or that involve animals capable of mating but that do not normally do so in

3120-400: A species as determined by a taxonomist. A typological species is a group of organisms in which individuals conform to certain fixed properties (a type), so that even pre-literate people often recognise the same taxon as do modern taxonomists. The clusters of variations or phenotypes within specimens (such as longer or shorter tails) would differentiate the species. This method was used as

3240-488: A species. All species definitions assume that an organism acquires its genes from one or two parents very like the "daughter" organism, but that is not what happens in HGT. There is strong evidence of HGT between very dissimilar groups of prokaryotes , and at least occasionally between dissimilar groups of eukaryotes , including some crustaceans and echinoderms . The evolutionary biologist James Mallet concludes that there

3360-685: A species. Generally the term includes the unknown element of a distinct act of creation. Many authors have argued that a simple textbook definition, following Mayr's concept, works well for most multi-celled organisms , but breaks down in several situations: Species identification is made difficult by discordance between molecular and morphological investigations; these can be categorised as two types: (i) one morphology, multiple lineages (e.g. morphological convergence , cryptic species ) and (ii) one lineage, multiple morphologies (e.g. phenotypic plasticity , multiple life-cycle stages). In addition, horizontal gene transfer (HGT) makes it difficult to define

3480-519: A taxonomic decision at the discretion of cognizant specialists, is not governed by the Codes of Zoological or Botanical Nomenclature, in contrast to the PhyloCode , and contrary to what is done in several other fields, in which the definitions of technical terms, like geochronological units and geopolitical entities, are explicitly delimited. The nomenclatural codes that guide the naming of species, including

3600-506: A traditional biological species. The International Committee on Taxonomy of Viruses has since 1962 developed a universal taxonomic scheme for viruses; this has stabilised viral taxonomy. Most modern textbooks make use of Ernst Mayr 's 1942 definition, known as the Biological Species Concept as a basis for further discussion on the definition of species. It is also called a reproductive or isolation concept. This defines

3720-447: A variation on the morphological species concept, a phenetic species, defined as a set of organisms with a similar phenotype to each other, but a different phenotype from other sets of organisms. It differs from the morphological species concept in including a numerical measure of distance or similarity to cluster entities based on multivariate comparisons of a reasonably large number of phenotypic traits. A mate-recognition species

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3840-515: A variety of reasons. Viruses are a special case, driven by a balance of mutation and selection , and can be treated as quasispecies . Biologists and taxonomists have made many attempts to define species, beginning from morphology and moving towards genetics . Early taxonomists such as Linnaeus had no option but to describe what they saw: this was later formalised as the typological or morphological species concept. Ernst Mayr emphasised reproductive isolation, but this, like other species concepts,

3960-470: A whole is good, then every part is good, whether we understand it or not. If the biota, in the course of aeons, has built something we like but do not understand, then who but a fool would discard seemingly useless parts? To keep every cog and wheel is the first precaution of intelligent tinkering. Extinction rates are measured in a variety of ways. Conservation biologists measure and apply statistical measures of fossil records , rates of habitat loss , and

4080-438: Is "an entity composed of organisms which maintains its identity from other such entities through time and over space, and which has its own independent evolutionary fate and historical tendencies". This differs from the biological species concept in embodying persistence over time. Wiley and Mayden stated that they see the evolutionary species concept as "identical" to Willi Hennig 's species-as-lineages concept, and asserted that

4200-400: Is a group of sexually reproducing organisms that recognise one another as potential mates. Expanding on this to allow for post-mating isolation, a cohesion species is the most inclusive population of individuals having the potential for phenotypic cohesion through intrinsic cohesion mechanisms; no matter whether populations can hybridise successfully, they are still distinct cohesion species if

4320-458: Is a set of organisms adapted to a particular set of resources, called a niche, in the environment. According to this concept, populations form the discrete phenetic clusters that we recognise as species because the ecological and evolutionary processes controlling how resources are divided up tend to produce those clusters. A genetic species as defined by Robert Baker and Robert Bradley is a set of genetically isolated interbreeding populations. This

4440-537: Is an effective way to seek and identify efficient and effective types of reserve design to capture or sustain the highest priority biodiversity values and to work with communities in support of local ecosystems. Margules and Pressey identify six interlinked stages in the systematic planning approach: Conservation biologists regularly prepare detailed conservation plans for grant proposals or to effectively coordinate their plan of action and to identify best management practices (e.g. ). Systematic strategies generally employ

4560-492: Is an interdisciplinary network with professional alliances in the biological as well as social sciences. Those dedicated to the cause and profession advocate for a global response to the current biodiversity crisis based on morals , ethics , and scientific reason. Organizations and citizens are responding to the biodiversity crisis through conservation action plans that direct research, monitoring, and education programs that engage concerns at local through global scales. There

4680-725: Is aquatic, and is generally herbivorous . The Convention on International Trade in Endangered Species of Wild Fauna and Flora ( CITES ) suspended trade of yellow-headed temple turtles in July 2012. H. annandalii is found in Cambodia , Laos , Malaysia , Thailand , Vietnam , and possibly Myanmar . The preferred natural habitats of H. annandalii are wet forests and freshwater wetlands . H. annandalii may live in captivity for up to 35 years. Two turtles were hatched in November 2019 at

4800-490: Is based on a new philosophy or leadership theory steering away from historical notions of power, authority, and dominance. Adaptive conservation leadership is reflective and more equitable as it applies to any member of society who can mobilize others toward meaningful change using communication techniques that are inspiring, purposeful, and collegial. Adaptive conservation leadership and mentoring programs are being implemented by conservation biologists through organizations such as

4920-712: Is based on the findings of conservation biology. The term conservation biology and its conception as a new field originated with the convening of "The First International Conference on Research in Conservation Biology" held at the University of California, San Diego in La Jolla, California, in 1978 led by American biologists Bruce A. Wilcox and Michael E. Soulé with a group of leading university and zoo researchers and conservationists including Kurt Benirschke , Sir Otto Frankel , Thomas Lovejoy , and Jared Diamond . The meeting

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5040-414: Is called speciation . Charles Darwin was the first to describe the role of natural selection in speciation in his 1859 book The Origin of Species . Speciation depends on a measure of reproductive isolation , a reduced gene flow. This occurs most easily in allopatric speciation, where populations are separated geographically and can diverge gradually as mutations accumulate. Reproductive isolation

5160-423: Is debate on which metrics that money, time and personnel should be dedicated to for the best chance of making a positive impact. One specific general discussion topic is whether monitoring should happen where there is little human impact (to understand a system that has not been degraded by humans), where there is human impact (so the effects from humans can be investigated), or where there is data deserts and little

5280-403: Is described formally, in a publication that assigns it a unique scientific name. The description typically provides means for identifying the new species, which may not be based solely on morphology (see cryptic species ), differentiating it from other previously described and related or confusable species and provides a validly published name (in botany) or an available name (in zoology) when

5400-671: Is further weakened by the existence of microspecies , groups of organisms, including many plants, with very little genetic variability, usually forming species aggregates . For example, the dandelion Taraxacum officinale and the blackberry Rubus fruticosus are aggregates with many microspecies—perhaps 400 in the case of the blackberry and over 200 in the dandelion, complicated by hybridisation , apomixis and polyploidy , making gene flow between populations difficult to determine, and their taxonomy debatable. Species complexes occur in insects such as Heliconius butterflies, vertebrates such as Hypsiboas treefrogs, and fungi such as

5520-726: Is hard or even impossible to test. Later biologists have tried to refine Mayr's definition with the recognition and cohesion concepts, among others. Many of the concepts are quite similar or overlap, so they are not easy to count: the biologist R. L. Mayden recorded about 24 concepts, and the philosopher of science John Wilkins counted 26. Wilkins further grouped the species concepts into seven basic kinds of concepts: (1) agamospecies for asexual organisms (2) biospecies for reproductively isolated sexual organisms (3) ecospecies based on ecological niches (4) evolutionary species based on lineage (5) genetic species based on gene pool (6) morphospecies based on form or phenotype and (7) taxonomic species,

5640-598: Is important, it's worth noting that there are no models in existence that account for the complexity of unpredictable factors like species movement, a non-analog climate, changing species interactions, evolutionary rates on finer time scales, and many other stochastic variables. The measure of ongoing species loss is made more complex by the fact that most of the Earth's species have not been described or evaluated. Estimates vary greatly on how many species actually exist (estimated range: 3,600,000–111,700,000) to how many have received

5760-411: Is increasing recognition that conservation is not just about what is achieved but how it is done. The conservation of natural resources is the fundamental problem. Unless we solve that problem, it will avail us little to solve all others. Conscious efforts to conserve and protect global biodiversity are a recent phenomenon. Natural resource conservation, however, has a history that extends prior to

5880-436: Is known about the habitats' and communities' response to human perturbations . The concept of bioindicators / indicator species can be applied to ecological monitoring as a way to investigate how pollution is affecting an ecosystem. Species like amphibians and birds are highly susceptible to pollutants in their environment due to their behaviours and physiological features that cause them to absorb pollutants at

6000-403: Is no easy way to tell whether related geographic or temporal forms belong to the same or different species. Species gaps can be verified only locally and at a point of time. One is forced to admit that Darwin's insight is correct: any local reality or integrity of species is greatly reduced over large geographic ranges and time periods. The botanist Brent Mishler argued that the species concept

6120-468: Is not valid, notably because gene flux decreases gradually rather than in discrete steps, which hampers objective delimitation of species. Indeed, complex and unstable patterns of gene flux have been observed in cichlid teleosts of the East African Great Lakes . Wilkins argued that "if we were being true to evolution and the consequent phylogenetic approach to taxa, we should replace it with

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6240-563: Is now directed at urban conservation biology. The Society for Conservation Biology originated in 1985. By 1992, most of the countries of the world had become committed to the principles of conservation of biological diversity with the Convention on Biological Diversity ; subsequently many countries began programmes of Biodiversity Action Plans to identify and conserve threatened species within their borders, as well as protect associated habitats. The late 1990s saw increasing professionalism in

6360-660: Is particularly relevant to practitioners in that it has the potential to generate cause-and-effect relationships and reveal the factors that contribute to population declines. The Society for Conservation Biology is a global community of conservation professionals dedicated to advancing the science and practice of conserving biodiversity. Conservation biology as a discipline reaches beyond biology, into subjects such as philosophy , law , economics , humanities , arts , anthropology , and education . Within biology, conservation genetics and evolution are immense fields unto themselves, but these disciplines are of prime importance to

6480-586: Is similar to Mayr's Biological Species Concept, but stresses genetic rather than reproductive isolation. In the 21st century, a genetic species could be established by comparing DNA sequences. Earlier, other methods were available, such as comparing karyotypes (sets of chromosomes ) and allozymes ( enzyme variants). An evolutionarily significant unit (ESU) or "wildlife species" is a population of organisms considered distinct for purposes of conservation. In palaeontology , with only comparative anatomy (morphology) and histology from fossils as evidence,

6600-513: Is similar to the medical profession advocating for healthy lifestyle options, both are beneficial to human well-being yet remain scientific in their approach. There is a movement in conservation biology suggesting a new form of leadership is needed to mobilize conservation biology into a more effective discipline that is able to communicate the full scope of the problem to society at large. The movement proposes an adaptive leadership approach that parallels an adaptive management approach. The concept

6720-407: Is sometimes an important source of genetic variation. Viruses can transfer genes between species. Bacteria can exchange plasmids with bacteria of other species, including some apparently distantly related ones in different phylogenetic domains , making analysis of their relationships difficult, and weakening the concept of a bacterial species. Conservation biology The conservation ethic

6840-493: Is the hard evidence it provides scientists to use for advising policy makers and funding bodies about conservation efforts. Not only is ecological monitoring data important for convincing politicians, funders, and the public why a conservation program is important to implement, but also to keep them convinced that a program should be continued to be supported. There is plenty of debate on how conservation resources can be used most efficiently; even within ecological monitoring, there

6960-594: Is threatened by hybridisation, but this can be selected against once a pair of populations have incompatible alleles of the same gene, as described in the Bateson–Dobzhansky–Muller model . A different mechanism, phyletic speciation, involves one lineage gradually changing over time into a new and distinct form (a chronospecies ), without increasing the number of resultant species. Horizontal gene transfer between organisms of different species, either through hybridisation , antigenic shift , or reassortment ,

7080-489: The Columbus Zoo and Aquarium . Heosemys annadali has been breed in captivity. The leech Placobdelloides siamensis is an ectoparasite of this turtle. Species A species ( pl. : species) is a population of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring , typically by sexual reproduction . It is the basic unit of classification and

7200-523: The ICZN for animals and the ICN for plants, do not make rules for defining the boundaries of the species. Research can change the boundaries, also known as circumscription, based on new evidence. Species may then need to be distinguished by the boundary definitions used, and in such cases the names may be qualified with sensu stricto ("in the narrow sense") to denote usage in the exact meaning given by an author such as

7320-480: The ecology of a species or habitat at repeating intervals with defined methods. Long-term monitoring for environmental and ecological metrics is an important part of any successful conservation initiative. Unfortunately, long-term data for many species and habitats is not available in many cases. A lack of historical data on species populations , habitats, and ecosystems means that any current or future conservation work will have to make assumptions to determine if

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7440-441: The environment , that there was a civic duty to maintain the environment for future generations, and that scientific, empirically based methods should be applied to ensure this duty was carried out. Sir James Ranald Martin was prominent in promoting this ideology, publishing many medico-topographical reports that demonstrated the scale of damage wrought through large-scale deforestation and desiccation, and lobbying extensively for

7560-399: The fly agaric . Natural hybridisation presents a challenge to the concept of a reproductively isolated species, as fertile hybrids permit gene flow between two populations. For example, the carrion crow Corvus corone and the hooded crow Corvus cornix appear and are classified as separate species, yet they can hybridise where their geographical ranges overlap. A ring species is

7680-498: The jaguar ( Panthera onca ) of Latin America or the leopard ( Panthera pardus ) of Africa and Asia. In contrast, the scientific names of species are chosen to be unique and universal (except for some inter-code homonyms ); they are in two parts used together : the genus as in Puma , and the specific epithet as in concolor . A species is given a taxonomic name when a type specimen

7800-721: The mountain gorilla and establish Albert National Park (since renamed Virunga National Park ) in what is now Democratic Republic of Congo . By the 1970s, led primarily by work in the United States under the Endangered Species Act along with the Species at Risk Act (SARA) of Canada, Biodiversity Action Plans developed in Australia , Sweden , the United Kingdom , hundreds of species specific protection plans ensued. Notably

7920-406: The "cynical species concept", and arguing that far from being cynical, it usefully leads to an empirical taxonomy for any given group, based on taxonomists' experience. Other biologists have gone further and argued that we should abandon species entirely, and refer to the "Least Inclusive Taxonomic Units" (LITUs), a view that would be coherent with current evolutionary theory. The species concept

8040-594: The Aldo Leopold Leadership Program. Conservation may be classified as either in-situ conservation , which is protecting an endangered species in its natural habitat , or ex-situ conservation , which occurs outside the natural habitat. In-situ conservation involves protecting or restoring the habitat. Ex-situ conservation, on the other hand, involves protection outside of an organism's natural habitat, such as on reservations or in gene banks , in circumstances where viable populations may not be present in

8160-920: The Commons ". From this principle, conservation biologists can trace communal resource based ethics throughout cultures as a solution to communal resource conflict. For example, the Alaskan Tlingit peoples and the Haida of the Pacific Northwest had resource boundaries, rules, and restrictions among clans with respect to the fishing of sockeye salmon . These rules were guided by clan elders who knew lifelong details of each river and stream they managed. There are numerous examples in history where cultures have followed rules, rituals, and organized practice with respect to communal natural resource management. The Mauryan emperor Ashoka around 250 BC issued edicts restricting

8280-530: The Endangered Species Act (1966) and National Environmental Policy Act (1970), which together injected major funding and protection measures to large-scale habitat protection and threatened species research. Other conservation developments, however, have taken hold throughout the world. India, for example, passed the Wildlife Protection Act of 1972 . In 1980, a significant development was

8400-622: The Origin of Species explained how species could arise by natural selection . That understanding was greatly extended in the 20th century through genetics and population ecology . Genetic variability arises from mutations and recombination , while organisms themselves are mobile, leading to geographical isolation and genetic drift with varying selection pressures . Genes can sometimes be exchanged between species by horizontal gene transfer ; new species can arise rapidly through hybridisation and polyploidy ; and species may become extinct for

8520-623: The RSPB. The National Trust formed in 1895 with the manifesto to "...promote the permanent preservation, for the benefit of the nation, of lands, ... to preserve (so far practicable) their natural aspect." In May 1912, a month after the Titanic sank, banker and expert naturalist Charles Rothschild held a meeting at the Natural History Museum in London to discuss his idea for a new organisation to save

8640-542: The United Nations acted to conserve sites of outstanding cultural or natural importance to the common heritage of mankind. The programme was adopted by the General Conference of UNESCO in 1972. As of 2006, a total of 830 sites are listed: 644 cultural, 162 natural. The first country to pursue aggressive biological conservation through national legislation was the United States, which passed back to back legislation in

8760-405: The abbreviation "sp." in the singular or "spp." (standing for species pluralis , Latin for "multiple species") in the plural in place of the specific name or epithet (e.g. Canis sp.). This commonly occurs when authors are confident that some individuals belong to a particular genus but are not sure to which exact species they belong, as is common in paleontology . Authors may also use "spp." as

8880-400: The age of conservation. Resource ethics grew out of necessity through direct relations with nature. Regulation or communal restraint became necessary to prevent selfish motives from taking more than could be locally sustained, therefore compromising the long-term supply for the rest of the community. This social dilemma with respect to natural resource management is often called the " Tragedy of

9000-570: The amount of hybridisation is insufficient to completely mix their respective gene pools . A further development of the recognition concept is provided by the biosemiotic concept of species. In microbiology , genes can move freely even between distantly related bacteria, possibly extending to the whole bacterial domain. As a rule of thumb, microbiologists have assumed that members of Bacteria or Archaea with 16S ribosomal RNA gene sequences more similar than 97% to each other need to be checked by DNA–DNA hybridisation to decide if they belong to

9120-467: The area covered due to unplanned urbanization activities. Then they plant the new saplings of same tree families of that existing forest in the areas where the old forest has been lost and also plant those saplings to the barren areas connected to the forest. This maintains the density and area covered by the forest. Also, non-interference may be used, which is termed a preservationist method. Preservationists advocate for giving areas of nature and species

9240-862: The best places for wildlife in the British Isles. This meeting led to the formation of the Society for the Promotion of Nature Reserves, which later became the Wildlife Trusts . In the United States , the Forest Reserve Act of 1891 gave the President power to set aside forest reserves from the land in the public domain. John Muir founded the Sierra Club in 1892, and the New York Zoological Society

9360-474: The biological species concept, "the several versions" of the phylogenetic species concept, and the idea that species are of the same kind as higher taxa are not suitable for biodiversity studies (with the intention of estimating the number of species accurately). They further suggested that the concept works for both asexual and sexually-reproducing species. A version of the concept is Kevin de Queiroz 's "General Lineage Concept of Species". An ecological species

9480-505: The biological species concept, a cladistic species does not rely on reproductive isolation – its criteria are independent of processes that are integral in other concepts. Therefore, it applies to asexual lineages. However, it does not always provide clear cut and intuitively satisfying boundaries between taxa, and may require multiple sources of evidence, such as more than one polymorphic locus, to give plausible results. An evolutionary species, suggested by George Gaylord Simpson in 1951,

9600-428: The boundaries between closely related species become unclear with hybridisation , in a species complex of hundreds of similar microspecies , and in a ring species . Also, among organisms that reproduce only asexually , the concept of a reproductive species breaks down, and each clone is potentially a microspecies. Although none of these are entirely satisfactory definitions, and while the concept of species may not be

9720-421: The broad range of taxa (i.e. including microbes, plants, and animals). Physiology is considered in the broadest possible terms to include functional and mechanistic responses at all scales, and conservation includes the development and refinement of strategies to rebuild populations, restore ecosystems, inform conservation policy, generate decision-support tools, and manage natural resources. Conservation physiology

9840-433: The concept of a chronospecies can be applied. During anagenesis (evolution, not necessarily involving branching), some palaeontologists seek to identify a sequence of species, each one derived from the phyletically extinct one before through continuous, slow and more or less uniform change. In such a time sequence, some palaeontologists assess how much change is required for a morphologically distinct form to be considered

9960-662: The early 19th century biogeography was ignited through the efforts of Alexander von Humboldt , Charles Lyell and Charles Darwin . The 19th-century fascination with natural history engendered a fervor to be the first to collect rare specimens with the goal of doing so before they became extinct by other such collectors. Although the work of many 18th and 19th century naturalists were to inspire nature enthusiasts and conservation organizations , their writings, by modern standards, showed insensitivity towards conservation as they would kill hundreds of specimens for their collections. The modern roots of conservation biology can be found in

10080-611: The emergence of the urban conservation movement. A local organization was established in Birmingham , UK, a development followed in rapid succession in cities across the UK, then overseas. Although perceived as a grassroots movement , its early development was driven by academic research into urban wildlife. Initially perceived as radical, the movement's view of conservation being inextricably linked with other human activity has now become mainstream in conservation thought. Considerable research effort

10200-435: The evidence to support hypotheses about evolutionarily divergent lineages that have maintained their hereditary integrity through time and space. Molecular markers may be used to determine diagnostic genetic differences in the nuclear or mitochondrial DNA of various species. For example, in a study done on fungi , studying the nucleotide characters using cladistic species produced the most accurate results in recognising

10320-712: The first conservation societies was the Royal Society for the Protection of Birds , founded in 1889 in Manchester as a protest group campaigning against the use of great crested grebe and kittiwake skins and feathers in fur clothing . Originally known as "the Plumage League", the group gained popularity and eventually amalgamated with the Fur and Feather League in Croydon, and formed

10440-475: The institutionalization of forest conservation activities in British India through the establishment of Forest Departments . The Madras Board of Revenue started local conservation efforts in 1842, headed by Alexander Gibson , a professional botanist who systematically adopted a forest conservation program based on scientific principles. This was the first case of state conservation management of forests in

10560-549: The late 18th-century Enlightenment period particularly in England and Scotland . Thinkers including Lord Monboddo described the importance of "preserving nature"; much of this early emphasis had its origins in Christian theology . Scientific conservation principles were first practically applied to the forests of British India . The conservation ethic that began to evolve included three core principles: that human activity damaged

10680-479: The maintenance, loss, and restoration of biodiversity and the science of sustaining evolutionary processes that engender genetic , population , species , and ecosystem diversity. The concern stems from estimates suggesting that up to 50% of all species on the planet will disappear within the next 50 years, which will increase poverty and starvation, and will reset the course of evolution on this planet. Researchers acknowledge that projections are difficult, given

10800-457: The management, mainly for economic reasons, of such natural resources as timber , fish, game, topsoil , pastureland , and minerals. In addition it referred to the preservation of forests ( forestry ), wildlife ( wildlife refuge ), parkland, wilderness , and watersheds . This period also saw the passage of the first conservation legislation and the establishment of the first nature conservation societies. The Sea Birds Preservation Act of 1869

10920-423: The metric must be able to capture the trend of the population or habitat as a whole. Long-term monitoring can include the continued measuring of many biological, ecological, and environmental metrics including annual breeding success, population size estimates, water quality, biodiversity (which can be measured in many way, i.e. Shannon Index ), and many other methods. When determining which metrics to monitor for

11040-602: The mid-20th century, efforts arose to target individual species for conservation, notably efforts in big cat conservation in South America led by the New York Zoological Society. In the early 20th century the New York Zoological Society was instrumental in developing concepts of establishing preserves for particular species and conducting the necessary conservation studies to determine the suitability of locations that are most appropriate as conservation priorities;

11160-456: The modern era of conservation science and policy . The inherent multidisciplinary basis for conservation biology has led to new subdisciplines including conservation social science, conservation behavior and conservation physiology. It stimulated further development of conservation genetics which Otto Frankel had originated first but is now often considered a subdiscipline as well. The rapid decline of established biological systems around

11280-581: The movement toward wildlife conservation . In the 21st century professional conservation officers have begun to collaborate with indigenous communities for protecting wildlife in Canada. Some conservation efforts are yet to fully take hold due to ecological neglect. For example in the USA, 21st century bowfishing of native fishes, which amounts to killing wild animals for recreation and disposing of them immediately afterwards, remains unregulated and unmanaged. In

11400-456: The natural habitat. The conservation of habitats like forest, water or soil in its natural state is crucial for any species depending in it to thrive. Instead of making the whole new environment looking alike the original habitat of wild animals is less effective than preserving the original habitats. An approach in Nepal named reforestation campaign has helped increase the density and area covered by

11520-478: The numerous fungi species of all the concepts studied. Versions of the phylogenetic species concept that emphasise monophyly or diagnosability may lead to splitting of existing species, for example in Bovidae , by recognising old subspecies as species, despite the fact that there are no reproductive barriers, and populations may intergrade morphologically. Others have called this approach taxonomic inflation , diluting

11640-418: The original forests which proved to be better than creating entirely new environment after original one is let to lost. Old Forests Store More Carbon than Young Ones as proved by latest researches, so it is more crucial to protect the old ones. The reforestation campaign launched by Himalayan Adventure Therapy in Nepal basically visits the old forests in periodic basis which are vulnerable to loss of density and

11760-585: The paper is accepted for publication. The type material is usually held in a permanent repository, often the research collection of a major museum or university, that allows independent verification and the means to compare specimens. Describers of new species are asked to choose names that, in the words of the International Code of Zoological Nomenclature , are "appropriate, compact, euphonious, memorable, and do not cause offence". Books and articles sometimes intentionally do not identify species fully, using

11880-674: The person who named the species, while the antonym sensu lato ("in the broad sense") denotes a wider usage, for instance including other subspecies. Other abbreviations such as "auct." ("author"), and qualifiers such as "non" ("not") may be used to further clarify the sense in which the specified authors delineated or described the species. Species are subject to change, whether by evolving into new species, exchanging genes with other species, merging with other species or by becoming extinct. The evolutionary process by which biological populations of sexually-reproducing organisms evolve to become distinct or reproductively isolated as species

12000-445: The practice and profession of conservation biology. Conservationists introduce bias when they support policies using qualitative description, such as habitat degradation , or healthy ecosystems . Conservation biologists advocate for reasoned and sensible management of natural resources and do so with a disclosed combination of science , reason , logic , and values in their conservation management plans. This sort of advocacy

12120-487: The result of misclassification leading to questions on whether there really are any ring species. The commonly used names for kinds of organisms are often ambiguous: "cat" could mean the domestic cat, Felis catus , or the cat family, Felidae . Another problem with common names is that they often vary from place to place, so that puma, cougar, catamount, panther, painter and mountain lion all mean Puma concolor in various parts of America, while "panther" may also mean

12240-400: The rich, soft soil has been carried off and only the bare framework of the district left." In the bible, through Moses, God commanded to let the land rest from cultivation every seventh year. Before the 18th century, however, much of European culture considered it a pagan view to admire nature. Wilderness was denigrated while agricultural development was praised. However, as early as AD 680

12360-573: The ring. Ring species thus present a difficulty for any species concept that relies on reproductive isolation. However, ring species are at best rare. Proposed examples include the herring gull – lesser black-backed gull complex around the North pole, the Ensatina eschscholtzii group of 19 populations of salamanders in America, and the greenish warbler in Asia, but many so-called ring species have turned out to be

12480-508: The same species. This concept was narrowed in 2006 to a similarity of 98.7%. The average nucleotide identity (ANI) method quantifies genetic distance between entire genomes , using regions of about 10,000 base pairs . With enough data from genomes of one genus, algorithms can be used to categorize species, as for Pseudomonas avellanae in 2013, and for all sequenced bacteria and archaea since 2020. Observed ANI values among sequences appear to have an "ANI gap" at 85–95%, suggesting that

12600-529: The same species. When two species names are discovered to apply to the same species, the older species name is given priority and usually retained, and the newer name considered as a junior synonym, a process called synonymy . Dividing a taxon into multiple, often new, taxa is called splitting . Taxonomists are often referred to as "lumpers" or "splitters" by their colleagues, depending on their personal approach to recognising differences or commonalities between organisms. The circumscription of taxa, considered

12720-674: The sector, with the maturing of organisations such as the Institute of Ecology and Environmental Management and the Society for the Environment . Since 2000, the concept of landscape scale conservation has risen to prominence, with less emphasis being given to single-species or even single-habitat focused actions. Instead an ecosystem approach is advocated by most mainstream conservationists, although concerns have been expressed by those working to protect some high-profile species. Ecology has clarified

12840-546: The services of Geographic Information Systems to assist in the decision-making process. The SLOSS debate is often considered in planning. Conservation physiology was defined by Steven J. Cooke and colleagues as: An integrative scientific discipline applying physiological concepts, tools, and knowledge to characterizing biological diversity and its ecological implications; understanding and predicting how organisms, populations, and ecosystems respond to environmental change and stressors; and solving conservation problems across

12960-456: The slaughter of animals and certain kinds of birds, as well as opened veterinary clinics. Conservation ethics are also found in early religious and philosophical writings. There are examples in the Tao , Shinto , Hindu , Islamic and Buddhist traditions. In Greek philosophy, Plato lamented about pasture land degradation : "What is left now is, so to say, the skeleton of a body wasted by disease;

13080-502: The species concept and making taxonomy unstable. Yet others defend this approach, considering "taxonomic inflation" pejorative and labelling the opposing view as "taxonomic conservatism"; claiming it is politically expedient to split species and recognise smaller populations at the species level, because this means they can more easily be included as endangered in the IUCN red list and can attract conservation legislation and funding. Unlike

13200-540: The unknown potential impacts of many variables, including species introduction to new biogeographical settings and a non-analog climate. Conservation biologists research and educate on the trends and process of biodiversity loss , species extinctions , and the negative effect these are having on our capabilities to sustain the well-being of human society. Conservation biologists work in the field and office, in government, universities, non-profit organizations and industry. The topics of their research are diverse, because this

13320-537: The wild. It is difficult to define a species in a way that applies to all organisms. The debate about species concepts is called the species problem. The problem was recognised even in 1859, when Darwin wrote in On the Origin of Species : I was much struck how entirely vague and arbitrary is the distinction between species and varieties. He went on to write: No one definition has satisfied all naturalists; yet every naturalist knows vaguely what he means when he speaks of

13440-403: The work is having any effect on the population or ecosystem health. Ecological monitoring can provide early warning signals of deleterious effects (from human activities or natural changes in an environment) on an ecosystem and its species. In order for signs of negative trends in ecosystem or species health to be detected, monitoring methods must be carried out at appropriate time intervals, and

13560-458: The work of Henry Fairfield Osborn Jr., Carl E. Akeley , Archie Carr and his son Archie Carr III is notable in this era. Akeley for example, having led expeditions to the Virunga Mountains and observed the mountain gorilla in the wild, became convinced that the species and the area were conservation priorities. He was instrumental in persuading Albert I of Belgium to act in defense of

13680-420: The workings of the biosphere ; i.e., the complex interrelationships among humans, other species, and the physical environment. The burgeoning human population and associated agriculture , industry , and the ensuing pollution, have demonstrated how easily ecological relationships can be disrupted. The last word in ignorance is the man who says of an animal or plant: "What good is it?" If the land mechanism as

13800-419: The world means that conservation biology is often referred to as a "Discipline with a deadline". Conservation biology is tied closely to ecology in researching the population ecology ( dispersal , migration , demographics , effective population size , inbreeding depression , and minimum population viability ) of rare or endangered species . Conservation biology is concerned with phenomena that affect

13920-403: The world. Governor-General Lord Dalhousie introduced the first permanent and large-scale forest conservation program in the world in 1855, a model that soon spread to other colonies , as well the United States, where Yellowstone National Park was opened in 1872 as the world's first national park. The term conservation came into widespread use in the late 19th century and referred to

14040-410: Was also a naturalist who also suggested that species evolved. Erasmus Darwin noted that some species have vestigial structures which are anatomical structures that have no apparent function in the species currently but would have been useful for the species' ancestors. The thinking of these early 18th century naturalists helped to change the mindset and thinking of the early 19th century naturalists. By

14160-514: Was passed in Britain as the first nature protection law in the world after extensive lobbying from the Association for the Protection of Seabirds and the respected ornithologist Alfred Newton . Newton was also instrumental in the passage of the first Game laws from 1872, which protected animals during their breeding season so as to prevent the stock from being brought close to extinction. One of

14280-452: Was prompted due to concern over tropical deforestation, disappearing species, and eroding genetic diversity within species. The conference and proceedings that resulted sought to initiate the bridging of a gap between theory in ecology and evolutionary genetics on the one hand and conservation policy and practice on the other. Conservation biology and the concept of biological diversity ( biodiversity ) emerged together, helping crystallize

14400-517: Was set up in 1895. A series of national forests and preserves were established by Theodore Roosevelt from 1901 to 1909. The 1916 National Parks Act, included a 'use without impairment' clause, sought by John Muir, which eventually resulted in the removal of a proposal to build a dam in Dinosaur National Monument in 1959. In the 20th century, Canadian civil servants, including Charles Gordon Hewitt and James Harkin , spearheaded

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