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95-526: Anatomically modern humans (i.e. Homo sapiens ) are believed to have emerged in Africa around 300,000 years ago. It has been argued by some that their ways of life changed relatively little from that of archaic humans of the Middle Paleolithic , until about 50,000 years ago, when there was a marked increase in the diversity of artefacts found associated with modern human remains. This period coincides with

190-487: A gaggle ) but for theoretical discussions of behavioural ecology , the generic term herd can be used for all such kinds of assemblage. The word herd , as a noun, can also refer to one who controls, possesses and has care for such groups of animals when they are domesticated. Examples of herds in this sense include shepherds (who tend to sheep), goatherds (who tend to goats), and cowherds (who tend to cattle). When an association of animals (or, by extension, people)

285-1027: A 2017 study to be between 350 and 260,000 years ago, compatible with the estimated age of early H. sapiens . The study states that the deep split-time estimation of 350 to 260 thousand years ago is consistent with the archaeological estimate for the onset of the Middle Stone Age across sub-Saharan Africa and coincides with archaic H. sapiens in southern Africa represented by, for example, the Florisbad skull dating to 259 (± 35) thousand years ago. H. s. idaltu , found at Middle Awash in Ethiopia, lived about 160,000 years ago, and H. sapiens lived at Omo Kibish in Ethiopia about 233,000-195,000 years ago. Two fossils from Guomde, Kenya, dated to at least (and likely more than) 180,000 years ago and (more precisely) to 300–270,000 years ago, have been tentatively assigned to H. sapiens and similarities have been noted between them and

380-459: A comparatively homogeneous single species of H. sapiens from more diverse varieties of archaic humans (all of which were descended from the early dispersal of H. erectus some 1.8 million years ago) was debated in terms of two competing models during the 1980s: " recent African origin " postulated the emergence of H. sapiens from a single source population in Africa, which expanded and led to

475-468: A convention popular in the 1990s, Neanderthals were classified as a subspecies of H. sapiens , as H. s. neanderthalensis , while AMH (or European early modern humans , EEMH) was taken to refer to " Cro-Magnon " or H. s. sapiens . Under this nomenclature (Neanderthals considered H. sapiens ), the term "anatomically modern Homo sapiens " (AMHS) has also been used to refer to EEMH ("Cro-Magnons"). It has since become more common to designate Neanderthals as

570-407: A coordinated group where individuals have distinct roles. Many human groupings, such as army detachments or sports teams, show such coordination and differentiation of roles, but so do some animal groupings such as those of eusocial insects, which are coordinated through pheromones and other forms of animal communication. A herd is, by definition, relatively unstructured. However, there may be two or

665-408: A few animals which tend to be imitated by the bulk of the herd more than others. An animal in this role is called a "control animal", since its behaviour will predict that of the herd as a whole. It cannot be assumed, however, that the control animal is deliberately taking a leadership role; control animals are not necessarily socially dominant in conflict situations, though they often are. Group size

760-464: A group of Spinner dolphins is able to corral fish into a smaller volume, which makes catching them easier, as there is less opportunity for the fish to escape. Furthermore, large groups are able to monopolize resources and defend them against solitary animals or smaller groups of the same or different species. It has been shown that larger groups of lions tend to be more successful in protecting prey from hyenas than smaller ones. Being able to communicate

855-406: A higher forehead, and reduced brow ridge . Early modern people and some living people do however have quite pronounced brow ridges, but they differ from those of archaic forms by having both a supraorbital foramen or notch, forming a groove through the ridge above each eye. This splits the ridge into a central part and two distal parts. In current humans, often only the central section of the ridge

950-757: A lineage-based ( cladistic ) definition of H. sapiens has been suggested, in which H. sapiens would by definition refer to the modern human lineage following the split from the Neanderthal lineage. Such a cladistic definition would extend the age of H. sapiens to over 500,000 years. Estimates for the split between the Homo sapiens line and combined Neanderthal / Denisovan line range from between 503,000 and 565,000 years ago; between 550,000 and 765,000 years ago; and (based on rates of dental evolution) possibly more than 800,000 years ago. Extant human populations have historically been divided into subspecies , but since around

1045-503: A number of physiological details which can be taken as reliably differentiating the physiology of Neanderthals vs. anatomically modern humans. The term "anatomically modern humans" (AMH) is used with varying scope depending on context, to distinguish "anatomically modern" Homo sapiens from archaic humans such as Neanderthals and Middle and Lower Paleolithic hominins with transitional features intermediate between H. erectus , Neanderthals and early AMH called archaic Homo sapiens . In

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1140-620: A semi-technical usage in behavioral finance to describe the largest group of market investors or market speculators who tend to "move with the market", or "follow the general market trend". This is at least a plausible example of genuine herding, though according to some researchers it results from rational decisions through processes such as information cascade and rational expectations . Other researchers, however, ascribe it to non-rational process such as mimicry , fear and greed contagion . "Contrarians" or contrarian investors are those who deliberately choose to invest or speculate counter to

1235-481: A separate species, H. neanderthalensis , so that AMH in the European context refers to H. sapiens , but the question is by no means resolved. In this more narrow definition of H. sapiens , the subspecies Homo sapiens idaltu , discovered in 2003, also falls under the umbrella of "anatomically modern". The recognition of H. sapiens idaltu as a valid subspecies of the anatomically modern human lineage would justify

1330-508: A smaller, more receded dentary, making the rest of the jaw-line stand out, giving an often quite prominent chin. The central part of the mandible forming the chin carries a triangularly shaped area forming the apex of the chin called the mental trigon , not found in archaic humans. Particularly in living populations, the use of fire and tools requires fewer jaw muscles, giving slender, more gracile jaws. Compared to archaic people, modern humans have smaller, lower faces. The body skeletons of even

1425-480: A specific purpose. The early modern humans who expanded into Europe, commonly referred to as the Cro-Magnons , left many sophisticated stone tools, carved and engraved pieces on bone, ivory and antler , cave paintings and Venus figurines . The Neanderthals continued to use Mousterian stone tool technology and possibly Châtelperronian technology. These tools disappeared from the archeological record at around

1520-573: A study published in 2020, there are indications that 2% to 19% (or about ≃6.6 and ≃7.0%) of the DNA of four West African populations may have come from an unknown archaic hominin which split from the ancestor of humans and Neanderthals between 360 kya to 1.02 mya. Generally, modern humans are more lightly built (or more "gracile") than the more "robust" archaic humans . Nevertheless, contemporary humans exhibit high variability in many physiological traits , and may exhibit remarkable "robustness". There are still

1615-491: A virtual skull shape of the last common human ancestor to modern humans/ H. sapiens , representative of the earliest modern humans, and suggested that modern humans arose between 350,000 and 260,000 years ago through a merging of populations in East and South Africa while North-African fossils may represent a population which introgressed into Neandertals during the LMP. According to

1710-513: A way to convey seasonal behavioural information about hunted animals. Lines (|) and dots (•) were apparently used interchangeably to denote lunar months, while the (Y) sign apparently signified "To give birth". These characters were seemingly combined to convey the breeding period of hunted animals. The climate of the period in Europe saw dramatic changes, and included the Last Glacial Maximum ,

1805-420: Is (scramble or direct) competition with other group members. In general, it is clear that the amount of resources available for each individual decreases with group size. If the resource availability is critical, competition within the group may get so intense, that animals no longer experience benefits from living in groups. However, only the relative importance of within- and between-group competition determines

1900-427: Is a social group of certain animals of the same species, either wild or domestic . The form of collective animal behavior associated with this is called herding . These animals are known as gregarious animals. The term herd is generally applied to mammals, and most particularly to the grazing ungulates that classically display this behaviour. Different terms are used for similar groupings in other species; in

1995-510: Is also found from a 125,000 years old artefacts in Buya , Eritrea and in other places such as Blombos cave in South Africa . More complex social groupings emerged, supported by more varied and reliable food sources and specialized tool types. This probably contributed to increasing group identification or ethnicity . The peopling of Australia most likely took place before c. 60 ka . Europe

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2090-401: Is an important characteristic of the social environment of gregarious species. The reason why animals form herds can not always be stated easily, since the underlying mechanisms are diverse and complex. Understanding the social behaviour of animals and the formation of groups has been a fundamental goal in the field of sociobiology and behavioural ecology . Theoretical framework is focused on

2185-676: Is associated with the African megadroughts during MIS 5 , beginning 130,000 years ago. A 2011 study located the origin of basal population of contemporary human populations at 130,000 years ago, with the Khoi-San representing an "ancestral population cluster" located in southwestern Africa (near the coastal border of Namibia and Angola ). While early modern human expansion in Sub-Saharan Africa before 130 kya persisted, early expansion to North Africa and Asia appears to have mostly disappeared by

2280-568: Is based on a cave lion skeleton found in Seigsdorf, Germany which has hunting lesions. 14,000 BP Fertile Crescent : Europe : Africa : Siberia : The Upper Paleolithic in the Franco-Cantabrian region : Anatomically modern humans Early modern human ( EMH ), or anatomically modern human ( AMH ), are terms used to distinguish Homo sapiens (the only extant Hominina species) that are anatomically consistent with

2375-426: Is beneficial. Whenever groups, such as shoals of fish, synchronize their movements, it becomes harder for predators to focus on particular individuals. However, animals that are weak and slower or on the periphery are preferred by predators, so that certain positions within the group are better than others (see selfish herd theory ). For fit animals, being in a group with such vulnerable individuals may thus decrease

2470-435: Is described as a herd, the implication is that the group tends to act together (for example, all moving in the same direction at a given time), but that this does not occur as a result of planning or coordination. Rather, each individual is choosing behaviour in correspondence with most other members, possibly through imitation or possibly because all are responding to the same external circumstances. A herd can be contrasted with

2565-479: Is estimated as having taken place over 500,000 years ago (marking the split of the H. sapiens lineage from ancestors shared with other known archaic hominins). But the oldest split among modern human populations (such as the Khoisan split from other groups) has been recently dated to between 350,000 and 260,000 years ago, and the earliest known examples of H. sapiens fossils also date to about that period, including

2660-577: Is estimated to have occurred in Africa roughly 500,000 years ago. The earliest fossil evidence of early modern humans appears in Africa around 300,000 years ago, with the earliest genetic splits among modern people, according to some evidence, dating to around the same time. Sustained archaic human admixture with modern humans is known to have taken place both in Africa and (following the recent Out-Of-Africa expansion ) in Eurasia, between about 100,000 and 30,000 years ago. The binomial name Homo sapiens

2755-553: Is evidence that the characteristic human brain development, especially the prefrontal cortex, was due to "an exceptional acceleration of metabolome evolution ... paralleled by a drastic reduction in muscle strength. The observed rapid metabolic changes in brain and muscle, together with the unique human cognitive skills and low muscle performance, might reflect parallel mechanisms in human evolution." The Schöningen spears and their correlation of finds are evidence that complex technological skills already existed 300,000 years ago, and are

2850-445: Is harsh and cold: By staying close together animals experience better thermoregulation, because their overall surface to volume ratio is reduced. Consequently, maintaining adequate body temperatures becomes less energetically costly. The collective force of a group mobbing predators can reduce risk of predation significantly. Flocks of raven are able to actively defend themselves against eagles and baboons collectively mob lions, which

2945-541: Is impossible for individuals alone. This behaviour may be based on reciprocal altruism, meaning animals are more likely to help each other if their conspecifics did so earlier. Animals living in groups are more likely to find mates than those living in solitary and are also able to compare potential partners in order to optimize genetic quality for their offspring. Domestic animal herds are assembled by humans for practicality in raising them and controlling them. Their behaviour may be quite different from that of wild herds of

Upper Paleolithic - Misplaced Pages Continue

3040-470: Is of the order of about 1% to 4% in Europeans and East Asians, and highest among Melanesians (the last also having Denisova hominin admixture at 4% to 6% in addition to neanderthal admixture). Cumulatively, about 20% of the Neanderthal genome is estimated to remain present spread in contemporary populations. In September 2019, scientists reported the computerized determination, based on 260 CT scans , of

3135-934: Is positively correlated to their fitness. If their energy requirement is fixed and additional energy is not increasing fitness, they will use as little time for foraging as possible (time minimizers). If on the other hand time allocated to foraging is fixed, an animal's gain in fitness is related to the quantity and quality of resources it feeds on (Energy maximizers). Since foraging may be energetically costly (searching, hunting, handling, etc.) and may induce risk of predation , animals in groups may have an advantage, since their combined effort in locating and handling food will reduce time needed to forage sufficiently. Thus, animals in groups may have shorter searching and handling times as well as an increased chance of finding (or monopolizing) highly profitable food, which makes foraging in groups beneficial for time minimizers and energy maximizers alike. The obvious disadvantage of foraging in groups

3230-477: Is preserved (if it is preserved at all). This contrasts with archaic humans, where the brow ridge is pronounced and unbroken. Modern humans commonly have a steep, even vertical forehead whereas their predecessors had foreheads that sloped strongly backwards. According to Desmond Morris , the vertical forehead in humans plays an important role in human communication through eyebrow movements and forehead skin wrinkling. Brain size in both Neanderthals and AMH

3325-411: Is primarily a direct effect of group size instead of the position within a herd. Greater group sizes result in higher visibility and detection rates for predators, but this relation is not directly proportional and saturates at some point, while the risk of being attacked for an individual is directly proportional to group size. Thus, the net effect for an individual in a group concerning its predation risk

3420-512: Is significantly larger on average (but overlapping in range) than brain size in H. erectus . Neanderthal and AMH brain sizes are in the same range, but there are differences in the relative sizes of individual brain areas, with significantly larger visual systems in Neanderthals than in AMH. Compared to archaic people, anatomically modern humans have smaller, differently shaped teeth. This results in

3515-468: Is still unclear. However, some animals have found countermeasures such as propolis in beehives or grooming in social animals. Staying together in groups often brings energetic advantages. Birds flying together in a flock use aerodynamic effects to reduce energetic costs, e.g. by positioning themselves in a V-shaped formation. A similar effect can be observed when fish swim together in fixed formations. Another benefit of group living occurs when climate

3610-445: Is taken to reflect a process towards a smaller and more fine-boned skeleton beginning around 50,000–30,000 years ago. The cranium lacks a pronounced occipital bun in the neck, a bulge that anchored considerable neck muscles in Neanderthals. Modern humans, even the earlier ones, generally have a larger fore-brain than the archaic people, so that the brain sits above rather than behind the eyes. This will usually (though not always) give

3705-570: The Arctic . Both Neanderthal and EEMH had somewhat larger cranial volumes on average than modern Europeans, suggesting the relaxation of selection pressures for larger brain volume after the end of the LGM. Examples for still later adaptations related to agriculture and animal domestication including East Asian types of ADH1B associated with rice domestication , or lactase persistence , are due to recent selection pressures. Herd A herd

3800-749: The Balkans , parts of the Iberian Peninsula and areas around the Black Sea . This period saw cultures such as the Solutrean in France and Spain. Human life may have continued on top of the ice sheet, but we know next to nothing about it, and very little about the human life that preceded the European glaciers. In the early part of the period, up to about 30 kya, the Mousterian Pluvial made northern Africa, including

3895-727: The Florisbad site in South Africa, dating to about 259,000 years ago, and the Jebel Irhoud site in Morocco, dated about 315,000 years ago. Extinct species of the genus Homo include Homo erectus (extant from roughly 2 to 0.1 million years ago) and a number of other species (by some authors considered subspecies of either H. sapiens or H. erectus ). The divergence of the lineage leading to H. sapiens out of ancestral H. erectus (or an intermediate species such as Homo antecessor )

Upper Paleolithic - Misplaced Pages Continue

3990-594: The Jebel Irhoud remains from Morocco (ca. 300,000 or 350–280,000 years ago), the Florisbad Skull from South Africa (ca. 259,000 years ago), and the Omo remains from Ethiopia (ca. 195,000, or, as more recently dated, ca. 233,000 years ago). An mtDNA study in 2019 proposed an origin of modern humans in Botswana (and a Khoisan split) of around 200,000 years. However, this proposal has been widely criticized by scholars, with

4085-558: The Mesolithic and the Neolithic , due to increased selection pressures and due to founder effects associated with migration . Alleles predictive of light skin have been found in Neanderthals , but the alleles for light skin in Europeans and East Asians, associated with KITLG and ASIP , are (as of 2012 ) thought to have not been acquired by archaic admixture but recent mutations since

4180-815: The Sahara , well-watered and with lower temperatures than today; after the end of the Pluvial the Sahara became arid. The Last Glacial Maximum was followed by the Allerød oscillation , a warm and moist global interstadial that occurred around 13.5 to 13.8 kya. Then there was a very rapid onset, perhaps within as little as a decade, of the cold and dry Younger Dryas climate period, giving sub-arctic conditions to much of northern Europe. The Preboreal rise in temperatures also began sharply around 10.3 kya, and by its end around 9.0 kya had brought temperatures nearly to present day levels, although

4275-459: The brain case is quite rounded and distinct from that of the Neanderthals and is similar to the brain case of modern humans. It is uncertain whether the robust traits of some of the early modern humans like Skhul V reflects mixed ancestry or retention of older traits. The "gracile" or lightly built skeleton of anatomically modern humans has been connected to a change in behavior, including increased cooperation and "resource transport". There

4370-438: The fish hook , the oil lamp , rope , and the eyed needle . Fishing of pelagic fish species and navigating the open ocean is evidenced by sites from Timor and Buka ( Solomon Islands ). The changes in human behavior have been attributed to changes in climate, encompassing a number of global temperature drops. These led to a worsening of the already bitter cold of the last glacial period (popularly but incorrectly called

4465-522: The lipid catabolic process . A 2017 study found correlation of Neanderthal admixture in phenotypic traits in modern European populations. Physiological or phenotypical changes have been traced to Upper Paleolithic mutations, such as the East Asian variant of the EDAR gene, dated to c. 35,000 years ago. Recent divergence of Eurasian lineages was sped up significantly during the Last Glacial Maximum (LGM),

4560-542: The range of phenotypes seen in contemporary humans, from extinct archaic human species. This distinction is useful especially for times and regions where anatomically modern and archaic humans co-existed, for example, in Paleolithic Europe . Among the oldest known remains of Homo sapiens are those found at the Omo-Kibish I archaeological site in south-western Ethiopia , dating to about 233,000 to 196,000 years ago,

4655-839: The recent "out of Africa" migration , likely between 60,000 and 40,000 years ago. Recent admixture analyses have added to the complexity, finding that Eastern Neanderthals derive up to 2% of their ancestry from anatomically modern humans who left Africa some 100 kya . The extent of Neanderthal admixture (and introgression of genes acquired by admixture) varies significantly between contemporary racial groups, being absent in Africans, intermediate in Europeans and highest in East Asians. Certain genes related to UV-light adaptation introgressed from Neanderthals have been found to have been selected for in East Asians specifically from 45,000 years ago until around 5,000 years ago. The extent of archaic admixture

4750-474: The 1980s all extant groups have tended to be subsumed into a single species, H. sapiens , avoiding division into subspecies altogether. Some sources show Neanderthals ( H. neanderthalensis ) as a subspecies ( H. sapiens neanderthalensis ). Similarly, the discovered specimens of the H. rhodesiensis species have been classified by some as a subspecies ( H. sapiens rhodesiensis ), although it remains more common to treat these last two as separate species within

4845-626: The 2010s with the discovery of admixture events ( introgression ) of populations of H. sapiens with populations of archaic humans over the period of between roughly 100,000 and 30,000 years ago, both in Eurasia and in Sub-Saharan Africa. Neanderthal admixture , in the range of 1–4%, is found in all modern populations outside of Africa, including in Europeans, Asians, Papua New Guineans, Australian Aboriginals, Native Americans, and other non-Africans. This suggests that interbreeding between Neanderthals and anatomically modern humans took place after

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4940-493: The Americas by about 40,000–25,000 years ago. Evidence for the overwhelming contribution of this "recent" ( L3 -derived) expansion to all non-African populations was established based on mitochondrial DNA , combined with evidence based on physical anthropology of archaic specimens , during the 1990s and 2000s, and has also been supported by Y DNA and autosomal DNA . The assumption of complete replacement has been revised in

5035-597: The Americas by about 15 ka. In Western Eurasia, the Paleolithic eases into the so-called Epipaleolithic or Mesolithic from the end of the LGM, beginning 15 ka. The Holocene glacial retreat begins 11.7 ka ( 10th millennium BC ), falling well into the Old World Epipaleolithic, and marking the beginning of the earliest forms of farming in the Fertile Crescent . Both Homo erectus and Neanderthals used

5130-399: The LGM. Phenotypes associated with the " white " or " Caucasian " populations of Western Eurasian stock emerge during the LGM, from about 19,000 years ago. Average cranial capacity in modern human populations varies in the range of 1,200 to 1,450 cm for adult males. Larger cranial volume is associated with climatic region, the largest averages being found in populations of Siberia and

5225-438: The Neanderthal lineage, as "pre-Neanderthal" or "early Neanderthal", while the divergence time between the Neanderthal and modern lineages has been pushed back to before the emergence of H. heidelbergensis , to close to 800,000 years ago, the approximate time of disappearance of H. antecessor . The term Middle Paleolithic is intended to cover the time between the first emergence of H. sapiens (roughly 300,000 years ago) and

5320-714: The Omo Kibbish remains. Fossil evidence for modern human presence in West Asia is ascertained for 177,000 years ago, and disputed fossil evidence suggests expansion as far as East Asia by 120,000 years ago. In July 2019, anthropologists reported the discovery of 210,000 year old remains of a H. sapiens and 170,000 year old remains of a H. neanderthalensis in Apidima Cave , Peloponnese , Greece , more than 150,000 years older than previous H. sapiens finds in Europe. A significant dispersal event, within Africa and to West Asia,

5415-525: The age of Y-chromosomal Adam have been pushed back significantly with the discovery of an ancient Y-chromosomal lineage in 2013, to likely beyond 300,000 years ago. There have, however, been no reports of the survival of Y-chromosomal or mitochondrial DNA clearly deriving from archaic humans (which would push back the age of the most recent patrilinear or matrilinear ancestor beyond 500,000 years). Fossil teeth found at Qesem Cave (Israel) and dated to between 400,000 and 200,000 years ago have been compared to

5510-401: The case of birds, for example, the word is flocking , but flock may also be used for mammals, particularly sheep or goats . Large groups of carnivores are usually called packs , and in nature a herd is classically subject to predation from pack hunters . Special collective nouns may be used for particular taxa (for example a flock of geese, if not in flight, is sometimes called

5605-457: The chance of being preyed upon even further. The effect of collective vigilance in social groups has been widely studied within the framework of optimal foraging theory and animal decision making. While animals under the risk of predation are feeding or resting, they have to stay vigilant and watch for predators. It could be shown in many studies (especially for birds) that with increase in group size individual animals are less attentive, while

5700-728: The climate was wetter. This period saw the Upper Paleolithic give way to the start of the following Mesolithic cultural period. As the glaciers receded sea levels rose; the English Channel , Irish Sea and North Sea were land at this time, and the Black Sea a fresh-water lake. In particular the Atlantic coastline was initially far out to sea in modern terms in most areas, though the Mediterranean coastline has retreated far less, except in

5795-452: The coldest phase of the last glacial period , which lasted from about 26.5 to 19 kya, being coldest at the end, before relatively rapid warming (all dates vary somewhat for different areas, and in different studies). During the Maximum, most of Northern Europe was covered by an ice-sheet , forcing human populations into the areas known as Last Glacial Maximum refugia , including modern Italy and

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5890-424: The competition for food can lead to the misuse of warning calls, as was observed for great tits: If food is scarce or monopolized by dominant birds, other birds (mainly subordinates) use antipredatory warning calls to induce an interruption of feeding and gain access to resources. Another study concerning a flock of geese suggested that the benefits of lower vigilance concerned only those in central positions, due to

5985-491: The contemporary height distribution measured among Malay people , for one. Following the peopling of Africa some 130,000 years ago, and the recent Out-of-Africa expansion some 70,000 to 50,000 years ago, some sub-populations of H. sapiens had been essentially isolated for tens of thousands of years prior to the early modern Age of Discovery . Combined with archaic admixture this has resulted in significant genetic variation , which in some instances has been shown to be

6080-440: The costs and benefits associated with living in groups in terms of the fitness of each individual compared to living solitarily. Living in groups evolved independently multiple times in various taxa and can only occur if its benefits outweigh the costs within an evolutionary timescale. Thus, animals form groups whenever this increases their fitness compared to living in solitary. The following includes an outline about some of

6175-647: The dental material from the younger (120,000–80,000 years ago) Skhul and Qafzeh hominins . Dispersal of early H. sapiens begins soon after its emergence, as evidenced by the North African Jebel Irhoud finds (dated to around 315,000 years ago). There is indirect evidence for H. sapiens presence in West Asia around 270,000 years ago. The Florisbad Skull from Florisbad, South Africa, dated to about 259,000 years ago, has also been classified as representing early H. sapiens . Scerri (2018) , pp. 582–594 In September 2019, scientists proposed that

6270-412: The description of contemporary humans with the subspecies name Homo sapiens sapiens . However, biological anthropologist Chris Stringer does not consider idaltu distinct enough within H. sapiens to warrant its own subspecies designation. A further division of AMH into "early" or "robust" vs. "post-glacial" or " gracile " subtypes has since been used for convenience. The emergence of "gracile AMH"

6365-623: The distal bones were shorter, usually thought to be an adaptation to cold climate. The same adaptation is found in some modern people living in the polar regions. Height ranges overlap between Neanderthals and AMH, with Neanderthal averages cited as 164 to 168 cm (65 to 66 in) and 152 to 156 cm (60 to 61 in) for males and females, respectively, which is largely identical to pre-industrial average heights for AMH. Contemporary national averages range between 158 to 184 cm (62 to 72 in) in males and 147 to 172 cm (58 to 68 in) in females. Neanderthal ranges approximate

6460-488: The earliest H. sapiens (and last common human ancestor to modern humans) arose between 350,000 and 260,000 years ago through a merging of populations in East and South Africa . Among extant populations, the Khoi-San (or " Capoid ") hunters-gatherers of Southern Africa may represent the human population with the earliest possible divergence within the group Homo sapiens sapiens . Their separation time has been estimated in

6555-437: The earliest and most robustly built modern humans were less robust than those of Neanderthals (and from what little we know from Denisovans), having essentially modern proportions. Particularly regarding the long bones of the limbs, the distal bones (the radius / ulna and tibia / fibula ) are nearly the same size or slightly shorter than the proximal bones (the humerus and femur ). In ancient people, particularly Neanderthals,

6650-572: The emergence of "anatomically modern humans". Since the 2000s, the discovery of older remains with comparable characteristics, and the discovery of ongoing hybridization between "modern" and "archaic" populations after the time of the Omo remains, have opened up a renewed debate on the age of H. sapiens in journalistic publications. H. s. idaltu , dated to 160,000 years ago, has been postulated as an extinct subspecies of H. sapiens in 2003. H. neanderthalensis , which became extinct about 40,000 years ago,

6745-433: The emergence of a much more detailed picture, intermediate between the two competing scenarios outlined above: The recent Out-of-Africa expansion accounts for the predominant part of modern human ancestry, while there were also significant admixture events with regional archaic humans. Since the 1970s, the Omo remains, originally dated to some 195,000 years ago, have often been taken as the conventional cut-off point for

6840-785: The end of MIS5 (75,000 years ago), and is known only from fossil evidence and from archaic admixture . Eurasia was re-populated by early modern humans in the so-called "recent out-of-Africa migration" post-dating MIS5, beginning around 70,000–50,000 years ago. In this expansion, bearers of mt-DNA haplogroup L3 left East Africa, likely reaching Arabia via the Bab-el-Mandeb , and in the Great Coastal Migration spread to South Asia, Maritime South Asia and Oceania between 65,000 and 50,000 years ago, while Europe , East and North Asia were reached by about 45,000 years ago. Some evidence suggests that an early wave of humans may have reached

6935-480: The extinction of all other human varieties, while the " multiregional evolution " model postulated the survival of regional forms of archaic humans, gradually converging into the modern human varieties by the mechanism of clinal variation , via genetic drift , gene flow and selection throughout the Pleistocene. Since the 2000s, the availability of data from archaeogenetics and population genetics has led to

7030-443: The fact that the possibly more vulnerable individuals in the flock's periphery have a greater need to stay attentive. This implies that the decrease in overall vigilance arises simply because the geese on the edge of the flock comprise a smaller group when groups get large. A special case of collective vigilance in groups is that of sentinels. Individuals take turn in keeping guard, while all others participate in other activities. Thus,

7125-486: The first obvious proof of an active (big game) hunt . H. heidelbergensis already had intellectual and cognitive skills like anticipatory planning, thinking and acting that so far have only been attributed to modern man. The ongoing admixture events within anatomically modern human populations make it difficult to estimate the age of the matrilinear and patrilinear most recent common ancestors of modern populations ( Mitochondrial Eve and Y-chromosomal Adam ). Estimates of

7220-438: The genus Homo rather than as subspecies within H. sapiens . All humans are considered to be a part of the subspecies H. sapiens sapiens , a designation which has been a matter of debate since a species is usually not given a subspecies category unless there is evidence of multiple distinct subspecies. The divergence of the lineage that would lead to H. sapiens out of archaic human varieties derived from H. erectus ,

7315-505: The last ice age ). Such changes may have reduced the supply of usable timber and forced people to look at other materials. In addition, flint becomes brittle at low temperatures and may not have functioned as a tool. Some notational signs, used next to images of animals, may have appeared as early as the Upper Palaeolithic in Europe circa 35,000 BCE, and may be the earliest proto-writing : several symbols were used in combination as

7410-500: The lineage of modern humans since the split from the lineage of Neanderthals , roughly 500,000 to 800,000 years ago. The time of divergence between archaic H. sapiens and ancestors of Neanderthals and Denisovans caused by a genetic bottleneck of the latter was dated at 744,000 years ago, combined with repeated early admixture events and Denisovans diverging from Neanderthals 300 generations after their split from H. sapiens , as calculated by Rogers et al. (2017). The derivation of

7505-402: The location and type of food to other group members may increase the chance for each individual to find profitable food sources, a mechanism which is known to be used by both bees (via a Waggle dance ) and several species of birds (using specific vocalisations to indicate food). In terms of Optimal foraging theory , animals always try to maximize their net energy gain when feeding, because this

7600-425: The major effects determining the trade-offs for living in groups. Perhaps the most studied effect of herds is the so-called dilution effect. The key argument is that the risk of being preyed upon for any particular individual is smaller within a larger group, strictly due to the fact that a predator has to decide which individual to attack. Although the dilution effect is influenced by so-called selfish herding , it

7695-435: The most common date assigned to expansion of modern humans from Africa throughout Asia and Eurasia, which contributed to the extinction of the Neanderthals . The Upper Paleolithic has the earliest known evidence of organized settlements , in the form of campsites, some with storage pits. Artistic work blossomed, with cave painting, petroglyphs , carvings and engravings on bone or ivory. The first evidence of human fishing

7790-580: The north of the Adriatic and the Aegean . The rise in sea levels continued until at least 7.5 kya ( 5500 BC ), so evidence of human activity along Europe's coasts in the Upper Paleolithic is mostly lost, though some traces have been recovered by fishing boats and marine archaeology , especially from Doggerland , the lost area beneath the North Sea. The first direct evidence for Neanderthals hunting cave lions . This

7885-467: The optimal group size and ultimately the decision of each individual whether or not to stay in the group. Since animals in groups stay near each other and interact frequently, infectious diseases and parasites spread much easier between them compared to solitary animals. Studies have shown a positive correlation between herd size and intensity of infections, but the extent to which this sometimes drastic reduction in fitness governs group size and structure

7980-410: The overall vigilance suffers little (many eyes effect). This means food intake and other activities related to fitness are optimized in terms of time allocation when animals stay in groups. However, some details about this concepts remain unclear. Being the first to detect predators and react accordingly can be advantageous, implying individuals may not fully be able to rely only on the group. Moreover,

8075-478: The period held by some to mark the emergence of full behavioral modernity (roughly by 50,000 years ago, corresponding to the start of the Upper Paleolithic ). Many of the early modern human finds, like those of Jebel Irhoud , Omo , Herto , Florisbad , Skhul , and Peștera cu Oase exhibit a mix of archaic and modern traits. Skhul V, for example, has prominent brow ridges and a projecting face. However,

8170-508: The recent evidence overall (genetic, fossil, and archaeological) supporting an origin for H. sapiens approximately 100,000 years earlier and in a broader region of Africa than the study proposes. In September 2019, scientists proposed that the earliest H. sapiens (and last common human ancestor to modern humans) arose between 350,000 and 260,000 years ago through a merging of populations in East and South Africa . An alternative suggestion defines H. sapiens cladistically as including

8265-769: The result of directional selection taking place over the past 15,000 years, i.e., significantly later than possible archaic admixture events. Some climatic adaptations, such as high-altitude adaptation in humans , are thought to have been acquired by archaic admixture. Introgression of genetic variants acquired by Neanderthal admixture have different distributions in European and East Asians , reflecting differences in recent selective pressures. A 2014 study reported that Neanderthal-derived variants found in East Asian populations showed clustering in functional groups related to immune and haematopoietic pathways , while European populations showed clustering in functional groups related to

8360-709: The same crude stone tools. Archaeologist Richard G. Klein , who has worked extensively on ancient stone tools, describes the stone tool kit of archaic hominids as impossible to categorize. He argues that almost everywhere, whether Asia , Africa or Europe , before 50,000 years ago all the stone tools are much alike and unsophisticated. Firstly among the artefacts of Africa, archeologists found they could differentiate and classify those of less than 50,000 years into many different categories, such as projectile points, engraving tools, knife blades, and drilling and piercing tools. These new stone-tool types have been described as being distinctly differentiated from each other; each tool had

8455-454: The same or related species, since both their composition (in terms of the distribution of age and sex within the herd) and their history (in terms of when and how the individuals joined the herd) are likely to be very different. The term herd is also applied metaphorically to human beings in social psychology , with the concept of herd behaviour . However both the term and concepts that underlie its use are controversial. The term has acquired

8550-420: The same time the Neanderthals themselves disappeared from the fossil record, about 40,000 cal BP. Settlements were often located in narrow valley bottoms, possibly associated with hunting of passing herds of animals. Some of them may have been occupied year round, though more commonly they appear to have been used seasonally; people moved between the sites to exploit different food sources at different times of

8645-439: The strength of social bonds and trust within these groups have to be much higher than in the former cases. Hunting together enables group-living predators, such as wolves and wild dogs, to catch large prey, which they are unable to achieve when hunting alone. Working together significantly improves foraging efficiency, meaning the net energy gain of each individual is increased when animals are feeding collectively. As an example,

8740-485: The year. Hunting was important, and caribou/wild reindeer "may well be the species of single greatest importance in the entire anthropological literature on hunting". Technological advances included significant developments in flint tool manufacturing, with industries based on fine blades rather than simpler and shorter flakes . Burins and racloirs were used to work bone, antler and hides . Advanced darts and harpoons also appear in this period, along with

8835-470: Was also at one point considered to be a subspecies, H. s. neanderthalensis . H. heidelbergensis , dated 600,000 to 300,000 years ago, has long been thought to be a likely candidate for the last common ancestor of the Neanderthal and modern human lineages. However, genetic evidence from the Sima de los Huesos fossils published in 2016 seems to suggest that H. heidelbergensis in its entirety should be included in

8930-458: Was coined by Linnaeus , 1758 . The Latin noun homō (genitive hominis ) means "human being", while the participle sapiēns means "discerning, wise, sensible". The species was initially thought to have emerged from a predecessor within the genus Homo around 300,000 to 200,000 years ago. A problem with the morphological classification of "anatomically modern" was that it would not have included certain extant populations. For this reason,

9025-483: Was peopled after c. 45 ka. Anatomically modern humans are known to have expanded northward into Siberia as far as the 58th parallel by about 45 ka ( Ust'-Ishim man ). The Upper Paleolithic is divided by the Last Glacial Maximum (LGM), from about 25 to 15 ka. The peopling of the Americas occurred during this time, with East and Central Asia populations reaching the Bering land bridge after about 35 ka, and expanding into

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