The Tem (also known as the Temba or Kotokoliare ) an ethnic group of West Africa, mainly living in Togo , Benin , Burkina Faso , and Ghana . There is reported to be about 417,000 of the Tem, with 339,000 in Togo, 60,000 in Ghana and 18,000 in Benin. They speak the Tem language .
95-476: The Tem are referred to by a number of terms: Chaucho, Cotocoli, Cotokol, Kiamba, Kotokoli, Kotokolis, Kotokol, Temba, Tems, Timn, Tim, and Timu. Historically, they were also referred to as lions (gouni), named after the local area "Lion Forest" (Gounilawou). Little is known about the history of the Tem before the colonisation period. The Tem originated as a coalition of Gurma chiefdoms who settled around Sokodé during
190-417: A buildup of scent attracting the attention of predators, the lioness moves her cubs to a new den site several times a month, carrying them one-by-one by the nape of the neck. Usually, the mother does not integrate herself and her cubs back into the pride until the cubs are six to eight weeks old. Sometimes the introduction to pride life occurs earlier, particularly if other lionesses have given birth at about
285-540: A closer relationship with P. leo melanochaita . Lion samples from some parts of the Ethiopian Highlands cluster genetically with those from Cameroon and Chad, while lions from other areas of Ethiopia cluster with samples from East Africa. Researchers, therefore, assume Ethiopia is a contact zone between the two subspecies. Genome -wide data of a wild-born historical lion sample from Sudan showed that it clustered with P. l. leo in mtDNA-based phylogenies, but with
380-540: A common ancestor who evolved to function as a nocturnal species, decreasing their eyesight in favor of a better sense of smell. The anomaly to this theory were anthropoids , who appeared to have the most divergence from nocturnality of all organisms examined. While most mammals did not exhibit the morphological characteristics expected of a nocturnal creature, reptiles and birds fit in perfectly. A larger cornea and pupil correlated well with whether these two classes of organisms were nocturnal or not. Being active at night
475-471: A constant lookout for circling vultures, which indicate the death or distress of an animal. Most carrion on which both hyenas and lions feed upon are killed by hyenas rather than lions. Carrion is thought to provide a large part of lion diet. Lions and spotted hyenas occupy a similar ecological niche and compete for prey and carrion; a review of data across several studies indicates a dietary overlap of 58.6%. Lions typically ignore hyenas unless they are on
570-485: A decrease in mate calls and continued to move around instead of waiting for a potential mate to arrive. This hurts the overall fitness of the species, which is concerning considering the overall decrease in amphibian populations. Predation Some nocturnal predator-prey relationships are interrupted by artificial lighting. Bats that are fast-moving are often at an advantage with insects being drawn to light; they are fast enough to escape any predators also attracted to
665-482: A dry climate, water is not plentiful, and their houses are often made of flammable materials. The festival begins by preparing a bonfire to the sound of drums. Performers begin as children and practice during the festivals, which are held several times per year. At the end of the festival, the fire is put out with bare feet. The Tem language is a Gur language belonging to the Niger-Congo language family . The language
760-425: A favoured position in the group, either stalking prey on the "wing", then attacking, or moving a smaller distance in the centre of the group and capturing prey fleeing from other lionesses. Males attached to prides do not usually participate in group hunting. Some evidence suggests, however, that males are just as successful as females; they are typically solo hunters who ambush prey in small bushland. They may join in
855-470: A few reported cases of old and wounded lions falling prey to wild dogs. Most lionesses reproduce by the time they are four years of age. Lions do not mate at a specific time of year and the females are polyestrous . Like those of other cats, the male lion's penis has spines that point backward. During withdrawal of the penis, the spines rake the walls of the female's vagina, which may cause ovulation . A lioness may mate with more than one male when she
950-548: A heavier mane. On average, Asiatic lions have sparser manes than African lions. This feature likely evolved to signal the fitness of males to females. Males with darker manes appear to have greater reproductive success and are more likely to remain in a pride for longer. They have longer and thicker hair and higher testosterone levels, but they are also more vulnerable to heat stress. The core body temperature does apparently not increase regardless of sex, season, feeding time, length and colour of mane, but only surface temperature
1045-590: A high affinity to P. l. melanochaita . This result suggested that the taxonomic position of lions in Central Africa may require revision. Other lion subspecies or sister species to the modern lion existed in prehistoric times: The Panthera lineage is estimated to have genetically diverged from the common ancestor of the Felidae around 9.32 to 4.47 million years ago to 11.75 to 0.97 million years ago . Results of analyses differ in
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#17332020376491140-473: A key species sought for exhibition in zoological gardens across the world since the late 18th century. Cultural depictions of lions were prominent in Ancient Egypt , and depictions have occurred in virtually all ancient and medieval cultures in the lion's historic and current range. The English word lion is derived via Anglo-Norman liun from Latin leōnem (nominative: leō ), which in turn
1235-495: A kill or are being harassed, while the latter tend to visibly react to the presence of lions with or without the presence of food. In the Ngorongoro crater , lions subsist largely on kills stolen from hyenas, causing them to increase their kill rate. In Botswana's Chobe National Park, the situation is reversed as hyenas there frequently challenge lions and steal their kills, obtaining food from 63% of all lion kills. When confronted on
1330-505: A kill, hyenas may either leave or wait patiently at a distance of 30–100 m (98–328 ft) until the lions have finished. Hyenas may feed alongside lions and force them off a kill. The two species attack one another even when there is no food involved for no apparent reason. Lions can account for up to 71% of hyena deaths in Etosha National Park . Hyenas have adapted by frequently mobbing lions that enter their home ranges. When
1425-519: A kill. Crippled lions and cubs may fall victim to hyenas and leopards or be trampled by buffalo or elephants. Careless lions may be maimed when hunting prey. Nile crocodiles may also kill and eat lions, evidenced by the occasional lion claw found in crocodile stomachs. Ticks commonly infest the ears, neck and groin regions of the lions. Adult forms of several tapeworm species of the genus Taenia have been isolated from lion intestines, having been ingested as larvae in antelope meat. Lions in
1520-558: A lion makes when sniffing chemical signals and involves an open mouth with bared teeth, raised muzzle, wrinkled nose, closed eyes and relaxed ears. Lions also use chemical and visual marking; males spray urine and scrape plots of ground and objects within the territory. The lion's repertoire of vocalisations is large; variations in intensity and pitch appear to be central to communication. Most lion vocalisations are variations of growling , snarling , meowing and roaring. Other sounds produced include puffing, bleating and humming. Roaring
1615-473: A longer evolutionary history than genetically less diverse lion samples from Asia and West and Central Africa. A whole genome-wide sequence of lion samples showed that samples from West Africa shared alleles with samples from Southern Africa, and samples from Central Africa shared alleles with samples from Asia. This phenomenon indicates that Central Africa was a melting pot of lion populations after they had become isolated, possibly migrating through corridors in
1710-399: A loose pride while females associate with up to 12 other females, forming a stronger pride together with their cubs. Female and male lions associate only when mating. Coalitions of males hold territory for a longer time than single lions. Males in coalitions of three or four individuals exhibit a pronounced hierarchy, in which one male dominates the others and mates more frequently. The lion
1805-418: A major effect on nocturnal animals, as well as diurnal species. The causes of these can be traced to distinct, sometimes overlapping areas: light pollution and spatial disturbance. Light pollution is a major issue for nocturnal species, and the impact continues to increase as electricity reaches parts of the world that previously had no access. Species in the tropics are generally more affected by this due to
1900-433: A nomad is a "range". Males associated with a pride patrol the fringes. Both males and females defend the pride against intruders, but the male lion is better-suited for this purpose due to its stockier, more powerful build. Some individuals consistently lead the defense against intruders, while others lag behind. Lions tend to assume specific roles in the pride; slower-moving individuals may provide other valuable services to
1995-544: A nomadic phase before gaining residence in a pride. A study undertaken in the Serengeti National Park revealed that nomadic coalitions gain residency at between 3.5 and 7.3 years of age. In Kruger National Park, dispersing male lions move more than 25 km (16 mi) away from their natal pride in search of their own territory. Female lions stay closer to their natal pride. Therefore, female lions in an area are more closely related to each other than male lions in
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#17332020376492090-495: A number of threats to the different endangered species. Adults are likely to stay away from artificially lit beaches that they might prefer to lay eggs on, as there is less cover against predators. Additionally, baby sea turtles that hatch from eggs on artificially lit beaches often get lost, heading towards the light sources as opposed to the ocean. Rhythmic behaviors are affected by light pollution both seasonally and daily patterns. Migrating birds or mammals might have issues with
2185-425: A reed-bed, a cave, or some other sheltered area, usually away from the pride. She will often hunt alone while the cubs are still helpless, staying relatively close to the den. Lion cubs are born blind, their eyes opening around seven days after birth. They weigh 1.2–2.1 kg (2.6–4.6 lb) at birth and are almost helpless, beginning to crawl a day or two after birth and walking around three weeks of age. To avoid
2280-470: A relatively similar spatial habitat as they did before. In comparison, herbivorous prey tend to stay in areas where human disturbance is low, limiting both resources and their spatial habitat. This leads to an imbalance in favor of predators, who increase in population and come out more often at night. In zoos , nocturnal animals are usually kept in special night-illumination enclosures to invert their normal sleep-wake cycle and to keep them active during
2375-565: A short, rounded head, a reduced neck, and round ears; males have broader heads. The fur varies in colour from light buff to silvery grey, yellowish red, and dark brown. The colours of the underparts are generally lighter. A new-born lion has dark spots , which fade as the cub reaches adulthood, although faint spots may still be seen on the legs and underparts. The tail of all lions ends in a dark, hairy tuft that, in some lions, conceals an approximately 5 mm (0.20 in)-long, hard "spine" or "spur" composed of dermal papillae. The functions of
2470-429: A year old and begin to hunt effectively when nearing the age of two. Single lions are capable of bringing down zebra and wildebeest, while larger prey like buffalo and giraffe are riskier. In Chobe National Park , large prides have been observed hunting African bush elephants up to around 15 years old in exceptional cases, with the victims being calves, juveniles, and even subadults. In typical hunts, each lioness has
2565-404: Is in heat . Lions of both sexes may be involved in group homosexual and courtship activities. Males will also head-rub and roll around with each other before mounting each other. Generation length of the lion is about seven years. The average gestation period is around 110 days; the female gives birth to a litter of between one and four cubs in a secluded den, which may be a thicket,
2660-698: Is pollination - nocturnal pollinators such as moths, beetles, thrips, and bats have a lower risk of being seen by predators, and the plants evolved temporal scent production and ambient heat to attract nocturnal pollination. Like with predators hunting the same prey, some plants such as apples can be pollinated both during the day and at night. Nocturnality is a form of crypsis , an adaptation to avoid or enhance predation . Although lions are cathemeral , and may be active at any time of day or night, they prefer to hunt at night because many of their prey species ( zebra , antelope , impala, wildebeest , etc.) have poor night vision . Many species of small rodents, such as
2755-666: Is a behavior in some non-human animals characterized by being active during the night and sleeping during the day. The common adjective is " nocturnal ", versus diurnal meaning the opposite. Nocturnal creatures generally have highly developed senses of hearing , smell , and specially adapted eyesight . Some animals, such as cats and ferrets , have eyes that can adapt to both low-level and bright day levels of illumination (see metaturnal ). Others, such as bushbabies and (some) bats , can function only at night. Many nocturnal creatures including tarsiers and some owls have large eyes in comparison with their body size to compensate for
2850-905: Is a generalist hypercarnivore and is considered to be both an apex and keystone predator due to its wide prey spectrum. Its prey consists mainly of medium-sized to large ungulates , particularly blue wildebeest , plains zebra , African buffalo , gemsbok and giraffe . It also frequently takes common warthog despite it being much smaller. In India, chital and sambar deer are the most common wild prey, while livestock contributes significantly to lion kills outside protected areas. It usually avoids fully grown adult elephants , rhinoceros and hippopotamus and small prey like dik-dik , hyraxes , hares and monkeys . Unusual prey include porcupines and small reptiles. Lions kill other predators but seldom consume them. Young lions first display stalking behaviour at around three months of age, although they do not participate in hunting until they are almost
2945-450: Is a form of niche differentiation , where a species' niche is partitioned not by the amount of resources but by the amount of time (i.e. temporal division of the ecological niche ). Hawks and owls can hunt the same field or meadow for the same rodents without conflict because hawks are diurnal and owls are nocturnal. This means they are not in competition for each other's prey. Another niche that being nocturnal lessens competition within
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3040-532: Is a large cat of the genus Panthera , native to Africa and India . It has a muscular, broad-chested body ; a short, rounded head; round ears; and a dark, hairy tuft at the tip of its tail. It is sexually dimorphic ; adult male lions are larger than females and have a prominent mane. It is a social species , forming groups called prides. A lion's pride consists of a few adult males, related females, and cubs. Groups of female lions usually hunt together, preying mostly on medium-sized and large ungulates . The lion
3135-543: Is a mixture of dry savannah forest and very dry, deciduous scrub forest . In Africa, the range of the lion originally spanned most of the central African rainforest zone and the Sahara desert. In the 1960s, it became extinct in North Africa, except in the southern part of Sudan. During the mid- Holocene , around 8,000-6,000 years ago, the range of lions expanded into Southeastern and Eastern Europe, partially re-occupying
3230-924: Is affected. Unlike in other felid species, female lions consistently interact with multiple males at once. Another hypothesis suggests that the mane also serves to protect the neck in fights, but this is disputed. During fights, including those involving maneless females and adolescents, the neck is not targeted as much as the face, back, and hindquarters. Injured lions also begin to lose their manes. Almost all male lions in Pendjari National Park are either maneless or have very short manes. Maneless lions have also been reported in Senegal, in Sudan 's Dinder National Park and in Tsavo East National Park , Kenya. Castrated lions often have little to no mane because
3325-465: Is an apex and keystone predator ; although some lions scavenge when opportunities occur and have been known to hunt humans , lions typically do not actively seek out and prey on humans. The lion inhabits grasslands , savannahs , and shrublands . It is usually more diurnal than other wild cats, but when persecuted, it adapts to being active at night and at twilight . During the Neolithic period,
3420-410: Is avoiding the heat of the day. This is especially true in arid biomes like deserts , where nocturnal behavior prevents creatures from losing precious water during the hot, dry daytime. This is an adaptation that enhances osmoregulation . One of the reasons that ( cathemeral ) lions prefer to hunt at night is to conserve water. Hamiltons Frog , found on Stephens and Maud islands, stays hidden for
3515-487: Is currently spoken by over 300,000. French is also sometimes used. Tem literature predominantly consists of tales passed down orally. Several compilations of these tales have been created, notably that of Silvano Galli , who, in the 2000s, compiled stories of approximately 100 authors and translated them into French and Italian. Tales are told at festive assemblies, attended by renowned storytellers such as Seybou Sebabe Giafago and Aboulaye Djibirila . Traditionally, once
3610-399: Is generally mutual and the recipient appears to express pleasure. The head and neck are the most common parts of the body licked; this behaviour may have arisen out of utility because lions cannot lick these areas themselves. Lions have an array of facial expressions and body postures that serve as visual gestures. A common facial expression is the "grimace face" or flehmen response , which
3705-521: Is no evidence for gene flow between the two lineages, indicating that they did not share the same geographic area. The Eurasian and American cave lions became extinct at the end of the last glacial period without mitochondrial descendants on other continents. The modern lion was probably widely distributed in Africa during the Middle Pleistocene and started to diverge in sub-Saharan Africa during
3800-429: Is not needed anymore due to the evolution of compensatory sensory systems, such as a heightened sense of smell and more astute auditory systems. In a recent study, recently extinct elephant birds and modern day nocturnal kiwi bird skulls were examined to recreate their likely brain and skull formation. They indicated that olfactory bulbs were much larger in comparison to their optic lobes , indicating they both have
3895-415: Is short and powerful; it attempts to catch prey with a fast rush and final leap, usually pulls it down by the rump, and kills with a clamping bite to the throat or muzzle . It can hold the prey's throat for up to 13 minutes, until the prey stops moving. It has a bite force from 1593.8 to 1768 Newtons at the canine tip and up 4167.6 Newtons at the carnassial notch. Lions typically consume prey at
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3990-586: Is the Tsavo lion pride that always has just one adult male. Prides act as fission–fusion societies , and members will split into subgroups that keep in contact with roars . Nomadic lions range widely and move around sporadically, either in pairs or alone. Pairs are more frequent among related males. A lion may switch lifestyles; nomads can become residents and vice versa. Interactions between prides and nomads tend to be hostile, although pride females in estrus allow nomadic males to approach them. Males spend years in
4085-615: Is the most social of all wild felid species, living in groups of related individuals with their offspring. Such a group is called a " pride ". Groups of male lions are called "coalitions". Females form the stable social unit in a pride and do not tolerate outside females. The majority of females remain in their birth prides while all males and some females will disperse . The average pride consists of around 15 lions, including several adult females and up to four males and their cubs of both sexes. Large prides, consisting of up to 30 individuals, have been observed. The sole exception to this pattern
4180-475: Is typically brownish and tinged with yellow, rust, and black hairs. Mutations in the genes microphthalmia-associated transcription factor and tyrosinase are possibly responsible for the colour of manes. It starts growing when lions enter adolescence, when testosterone levels increase, and reach their full size at around four years old. Cool ambient temperatures in European and North American zoos may result in
4275-426: Is unable to consume all of the kill, it rests for a few hours before continuing to eat. On hot days, the pride retreats to shade with one or two males standing guard. Lions defend their kills from scavengers such as vultures and hyenas. Lions scavenge on carrion when the opportunity arises, scavenging animals dead from natural causes such as disease or those that were killed by other predators. Scavenging lions keep
4370-563: Is used to advertise its presence. Lions most often roar at night, a sound that can be heard from a distance of 8 kilometres (5 mi). They tend to roar in a very characteristic manner starting with a few deep, long roars that subside into grunts. The lion is listed as Vulnerable on the IUCN Red List . The Indian population is listed on CITES Appendix I and the African population on CITES Appendix II . Nocturnality Nocturnality
4465-471: The Large Japanese Field Mouse , are active at night because most of the dozen or so birds of prey that hunt them are diurnal. There are many diurnal species that exhibit some nocturnal behaviors. For example, many seabirds and sea turtles only gather at breeding sites or colonies at night to reduce the risk of predation to themselves and/or their offspring. Nocturnal species take advantage of
4560-459: The Mesozoic , many ancestors of modern-day mammals evolved nocturnal characteristics in order to avoid contact with the numerous diurnal predators. A recent study attempts to answer the question as to why so many modern day mammals retain these nocturnal characteristics even though they are not active at night. The leading answer is that the high visual acuity that comes with diurnal characteristics
4655-505: The Narmada River in central India. Lions spend much of their time resting; they are inactive for about twenty hours per day. Although lions can be active at any time, their activity generally peaks after dusk with a period of socialising, grooming, and defecating. Intermittent bursts of activity continue until dawn, when hunting most often takes place. They spend an average of two hours a day walking and fifty minutes eating. The lion
4750-478: The Nile Basin during the early Holocene . In zoos, lions have been bred with tigers to create hybrids for the curiosity of visitors or for scientific purpose. The liger is bigger than a lion and a tiger, whereas most tigons are relatively small compared to their parents because of reciprocal gene effects. The leopon is a hybrid between a lion and leopard. The lion is a muscular, broad-chested cat with
4845-661: The 17th or 18th century, founding the Tchaoudjo Kingdom . The Tem originated in Burkina Faso, settling along the White Volta before most migrated to Togo and Benin. Over several centuries, many people groups such as the Hausa , Yoruba , and Mandinka have integrated into the Tem people group. The Tem converted to Islam during the 19th century via the influence of Chakosi merchants. The majority of Tem profess Islam today, and have
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#17332020376494940-749: The Asiatic lion population. In 2017, the Cat Classification Task Force of the Cat Specialist Group revised lion taxonomy , and recognises two subspecies based on results of several phylogeographic studies on lion evolution , namely: However, there seems to be some degree of overlap between both groups in northern Central Africa. DNA analysis from a more recent study indicates that Central African lions are derived from both northern and southern lions, as they cluster with P. leo leo in mtDNA-based phylogenies whereas their genomic DNA indicates
5035-609: The Late Pleistocene. Lion populations in East and Southern Africa became separated from populations in West and North Africa when the equatorial rainforest expanded 183,500 to 81,800 years ago. They shared a common ancestor probably between 98,000 and 52,000 years ago. Due to the expansion of the Sahara between 83,100 and 26,600 years ago, lion populations in West and North Africa became separated. As
5130-558: The Ngorongoro Crater were afflicted by an outbreak of stable fly ( Stomoxys calcitrans ) in 1962, resulting in lions becoming emaciated and covered in bloody, bare patches. Lions sought unsuccessfully to evade the biting flies by climbing trees or crawling into hyena burrows; many died or migrated and the local population dropped from 70 to 15 individuals. A more recent outbreak in 2001 killed six lions. Captive lions have been infected with canine distemper virus (CDV) since at least
5225-691: The adjacent Timbavati Private Game Reserve in eastern South Africa. They were removed from the wild in the 1970s, thus decreasing the white lion gene pool . Nevertheless, 17 births have been recorded in five prides between 2007 and 2015. White lions are selected for breeding in captivity. They have reportedly been bred in camps in South Africa for use as trophies to be killed during canned hunts . African lions live in scattered populations across sub-Saharan Africa. The lion prefers grassy plains and savannahs , scrub bordering rivers, and open woodlands with bushes. It rarely enters closed forests. On Mount Elgon ,
5320-424: The age of two. When a new male lion takes over a pride, adolescents both male and female may be evicted. Lions may live 12–17 years in the wild. Although adult lions have no natural predators, evidence suggests most die violently from attacks by humans or other lions. Lions often inflict serious injuries on members of other prides they encounter in territorial disputes or members of the home pride when fighting at
5415-544: The animal's expressive movements are highly developed. The most common peaceful, tactile gestures are head rubbing and social licking , which have been compared with the role of allogrooming among primates. Head rubbing, nuzzling the forehead, face and neck against another lion appears to be a form of greeting and is seen often after an animal has been apart from others or after a fight or confrontation. Males tend to rub other males, while cubs and females rub females. Social licking often occurs in tandem with head rubbing; it
5510-408: The artificial lighting. Insects are the most obvious example, who are attracted by the lighting and are usually killed by either the heat or electrical current. Some species of frogs are blinded by the quick changes in light, while nocturnal migratory birds may be disoriented, causing them to lose direction, tire out, or be captured by predators. Sea turtles are particularly affected by this, adding to
5605-499: The attack. They take advantage of factors that reduce visibility; many kills take place near some form of cover or at night. One study in 2018 recorded a lion running at a top speed of 74.1 km/h (46.0 mph). The lion accelerates at the start of the chase by 9.5 m/s² , whereas zebras, wildebeest and Thomson's gazelle accelerate by 5 m/s², 5.6 m/s² and 4.5 m/s², respectively; acceleration appears to be more important than steady displacement speed in lion hunts. The lion's attack
5700-454: The cause of the decline is not fully understood, habitat loss and conflicts with humans are the greatest causes for concern. One of the most widely recognised animal symbols in human culture, the lion has been extensively depicted in sculptures and paintings, on national flags, and in literature and films. Lions have been kept in menageries since the time of the Roman Empire and have been
5795-428: The change in their relatively constant light patterns, but temperate species relying on day-night triggers for behavioral patterns are also affected as well. Many diurnal species see the benefit of a "longer day", allowing for a longer hunting period which is detrimental to their nocturnal prey trying to avoid them. Light pollution can disorient species that are used to darkness, as their adaptive eyes are not as used to
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#17332020376495890-428: The cost. The increasing amount of habitat destruction worldwide as a result of human expansion has given both advantages and disadvantages to different nocturnal animals. As a result of peak human activity in the daytime, more species are likely to be active at night in order to avoid the new disturbance in their habitat. Carnivorous predators however are less timid of the disturbance, feeding on human waste and keeping
5985-411: The cubs varies—one male could patiently let the cubs play with his tail or his mane, while another may snarl and bat the cubs away. Pride lionesses often synchronise their reproductive cycles and communal rearing and suckling of the young, which suckle indiscriminately from any or all of the nursing females in the pride. The synchronisation of births is advantageous because the cubs grow to being roughly
6080-399: The daytime. Crepuscular species, such as rabbits , skunks , tigers and hyenas , are often erroneously referred to as nocturnal. Cathemeral species, such as fossas and lions , are active both in the day and at night. While it is difficult to say which came first, nocturnality or diurnality, a hypothesis in evolutionary biology , the nocturnal bottleneck theory, postulates that in
6175-534: The group. Alternatively, there may be rewards associated with being a leader that fends off intruders; the rank of lionesses in the pride is reflected in these responses. The male or males associated with the pride must defend their relationship with the pride from outside males who may attempt to usurp them. Dominance hierarchies do not appear to exist among individuals of either sex in a pride. Asiatic lion prides differ in group composition. Male Asiatic lions are solitary or associate with up to three males, forming
6270-452: The highest percentage to do so in Togo. The Tem people are known for performing the famous "fire dance festival", during which performers dance with fire, and are able to bring fire in contact with their skin seemingly without being hurt. Some also appear to consume the flames. It is thought by some that resistance to fire was built up by the people group out of necessity - fires are common in such
6365-570: The hunting of large, slower-moving prey like buffalo; and even hunt them on their own. Moderately-sized hunting groups generally have higher success rates than lone females and larger groups. Lions are not particularly known for their stamina. For instance, a lioness's heart comprises only 0.57% of her body weight and a male's is about 0.45% of his body weight, whereas a hyena's heart comprises almost 1% of its body weight. Thus, lions run quickly only in short bursts at about 48–59 km/h (30–37 mph) and need to be close to their prey before starting
6460-399: The light, leaving slow-moving bats at a disadvantage. Another example is harbor seals eating juvenile salmon that moved down a river lit by nearby artificial lighting. Once the lights were turned off, predation levels decreased. Many diurnal prey species forced into being nocturnal are susceptible to nocturnal predators and those species with poor nocturnal eyesight often bear the brunt of
6555-705: The lion has been recorded up to an elevation of 3,600 m (11,800 ft) and close to the snow line on Mount Kenya . Savannahs with an annual rainfall of 300 to 1,500 mm (12 to 59 in) make up the majority of lion habitat in Africa, estimated at 3,390,821 km (1,309,203 sq mi) at most, but remnant populations are also present in tropical moist forests in West Africa and montane forests in East Africa. The Asiatic lion now survives only in and around Gir National Park in Gujarat, western India. Its habitat
6650-801: The lion population in Kenya's Masai Mara National Reserve declined, the spotted hyena population increased rapidly. Lions tend to dominate cheetahs and leopards, steal their kills and kill their cubs and even adults when given the chance. Cheetahs often lose their kills to lions or other predators. A study in the Serengeti ecosystem revealed that lions killed at least 17 of 125 cheetah cubs born between 1987 and 1990. Cheetahs avoid their competitors by hunting at different times and habitats. Leopards, by contrast, do not appear to be motivated by an avoidance of lions, as they use heavy vegetation regardless of whether lions are present in an area and both cats are active around
6745-427: The lion ranged throughout Africa and Eurasia, from Southeast Europe to India, but it has been reduced to fragmented populations in sub-Saharan Africa and one population in western India . It has been listed as Vulnerable on the IUCN Red List since 1996 because populations in African countries have declined by about 43% since the early 1990s. Lion populations are untenable outside designated protected areas. Although
6840-551: The location of the hunt but sometimes drag large prey into cover. They tend to squabble over kills, particularly the males. Cubs suffer most when food is scarce but otherwise all pride members eat their fill, including old and crippled lions, which can live on leftovers. Large kills are shared more widely among pride members. An adult lioness requires an average of about 5 kg (11 lb) of meat per day while males require about 7 kg (15 lb). Lions gorge themselves and eat up to 30 kg (66 lb) in one session. If it
6935-424: The lower light levels at night. More specifically, they have been found to have a larger cornea relative to their eye size than diurnal creatures to increase their visual sensitivity : in the low-light conditions. Nocturnality helps wasps , such as Apoica flavissima , avoid hunting in intense sunlight. Diurnal animals, including humans (except for night owls ), squirrels and songbirds, are active during
7030-408: The majority of the day when temperatures are warmer and are mainly active at night. They will only come out during the day if there are humid and cool conditions. Many plant species native to arid biomes have adapted so that their flowers only open at night when the sun's intense heat cannot wither and destroy their moist, delicate blossoms. These flowers are pollinated by bats, another creature of
7125-481: The mid-1970s. CDV is spread by domestic dogs and other carnivores; a 1994 outbreak in Serengeti National Park resulted in many lions developing neurological symptoms such as seizures. During the outbreak, several lions died from pneumonia and encephalitis . Feline immunodeficiency virus and lentivirus also affect captive lions. When resting, lion socialisation occurs through a number of behaviours;
7220-428: The night time to prey on species that are used to avoiding diurnal predators. Some nocturnal fish species will use the moonlight to prey on zooplankton species that come to the surface at night. Some species have developed unique adaptations that allow them to hunt in the dark. Bats are famous for using echolocation to hunt down their prey, using sonar sounds to capture them in the dark. Another reason for nocturnality
7315-428: The night. Climate-change and the change in global temperatures has led to an increasing amount of diurnal species to push their activity patterns closer towards crepuscular or fully nocturnal behavior. This adaptive measure allows species to avoid the heat of the day, without having to leave that particular habitat. The exponential increase in human expansion and technological advances in the last few centuries has had
7410-434: The phylogenetic relationship of the lion; it was thought to form a sister group with the jaguar that diverged 3.46 to 1.22 million years ago , but also with the leopard that diverged 3.1 to 1.95 million years ago to 4.32 to 0.02 million years ago . Hybridisation between lion and snow leopard ancestors possibly continued until about 2.1 million years ago. The lion-leopard clade
7505-612: The previous males associated with a pride, the victors often kill any existing young cubs , perhaps because females do not become fertile and receptive until their cubs mature or die. Females often fiercely defend their cubs from a usurping male but are rarely successful unless a group of three or four mothers within a pride join forces against the male. Cubs also die from starvation and abandonment, and predation by leopards, hyenas and wild dogs. Male cubs are excluded from their maternal pride when they reach maturity at around two or three years of age, while some females may leave when they reach
7600-579: The rainforest decreased and thus gave rise to more open habitats, lions moved from West to Central Africa. Lions from North Africa dispersed to southern Europe and Asia between 38,800 and 8,300 years ago. Extinction of lions in southern Europe, North Africa and the Middle East interrupted gene flow between lion populations in Asia and Africa. Genetic evidence revealed numerous mutations in lion samples from East and Southern Africa, which indicates that this group has
7695-457: The range of the now extinct cave lion . In Hungary , the modern lion was present from about 4,500 to 3,200 years Before Present . In Ukraine , the modern lion was present from about 6,400 to 2,000 years Before Present. In Greece, it was common, as reported by Herodotus in 480 BC; it was considered rare by 300 BC and extirpated by AD 100. In Asia the lion once ranged in regions where climatic conditions supported an abundance of prey. It
7790-488: The removal of the gonads inhibits testosterone production. Rarely, both wild and captive lionesses have manes. Increased testosterone may be the cause of maned lionesses reported in northern Botswana. The white lion is a rare morph with a genetic condition called leucism , which is caused by a double recessive allele . It is not albino; it has normal pigmentation in the eyes and skin. White lions have occasionally been encountered in and around Kruger National Park and
7885-429: The same area. The evolution of sociability in lions was likely driven both by high population density and the clumped resources of savannah habitats. The larger the pride, the more high-quality territory they can defend; "hotspots" being near river confluences , where the cats have better access to water, prey and shelter (via vegetation). The area occupied by a pride is called a "pride area" whereas that occupied by
7980-420: The same size and have an equal chance of survival, and sucklings are not dominated by older cubs. Weaning occurs after six or seven months. Male lions reach maturity at about three years of age and at four to five years are capable of challenging and displacing adult males associated with another pride. They begin to age and weaken at between 10 and 15 years of age at the latest. When one or more new males oust
8075-406: The same time of day. In addition, there is no evidence that lions affect leopard abundance. Leopards take refuge in trees, though lionesses occasionally attempt to climb up and retrieve their kills. Lions similarly dominate African wild dogs , taking their kills and dispatching pups or adult dogs. Population densities of wild dogs are low in areas where lions are more abundant. However, there are
8170-420: The same time. When first introduced to the rest of the pride, lion cubs lack confidence when confronted with adults other than their mother. They soon begin to immerse themselves in the pride life, however, playing among themselves or attempting to initiate play with the adults. Lionesses with cubs of their own are more likely to be tolerant of another lioness's cubs than lionesses without cubs. Male tolerance of
8265-422: The spur are unknown. The tuft is absent at birth and develops at around 5 + 1 ⁄ 2 months of age. It is readily identifiable at the age of seven months. Its skull is very similar to that of the tiger, although the frontal region is usually more depressed and flattened and has a slightly shorter postorbital region and broader nasal openings than those of the tiger. Due to the amount of skull variation in
8360-573: The story is over, the audience thanks the storyteller by saying in unison "You and the Sky", to which the storyteller responds "You and the Earth". Tem literature is known across Togo, Benin, and even throughout West Africa. Many tales feature characters of domestic or savage animals who go on various adventures reflecting human society. This Togo -related article is a stub . You can help Misplaced Pages by expanding it . Lion The lion ( Panthera leo )
8455-413: The tiger in size. The size and weight of adult lions vary across its range and habitats. Accounts of a few individuals that were larger than average exist from Africa and India. The male lion's mane is the most recognisable feature of the species. It may have evolved around 320,000–190,000 years ago. It grows downwards and backwards, covering most of the head, neck, shoulders, and chest. The mane
8550-704: The timing of their movement for example. On a day-to-day basis, species can see significant changes in their internal temperatures, their general movement, feeding and body mass. These small scale changes can eventually lead to a population decline, as well as hurting local trophic levels and interconnecting species. Some typically diurnal species have even become crepuscular or nocturnal as a result of light pollution and general human disturbance. There have been documented effects of light pollution on reproductive cycles and factors in different species. It can affect mate choice , migration to breeding grounds, and nest site selection. In male green frogs , artificial light causes
8645-407: The two species, usually only the structure of the lower jaw can be used as a reliable indicator of species. The skeletal muscles of the lion make up 58.8% of its body weight and represent the highest percentage of muscles among mammals. The lion has a high concentration of fast twitch muscle fibers, giving them quick bursts of speed but less stamina. Among felids, the lion is second only to
8740-400: Was a borrowing from Ancient Greek λέων léōn . The Hebrew word לָבִיא lavi may also be related. The generic name Panthera is traceable to the classical Latin word 'panthēra' and the ancient Greek word πάνθηρ 'panther'. Felis leo was the scientific name used by Carl Linnaeus in 1758, who described the lion in his work Systema Naturae . The genus name Panthera
8835-593: Was coined by Lorenz Oken in 1816. Between the mid-18th and mid-20th centuries, 26 lion specimens were described and proposed as subspecies, of which 11 were recognised as valid in 2005. They were distinguished mostly by the size and colour of their manes and skins. In the 19th and 20th centuries, several lion type specimens were described and proposed as subspecies , with about a dozen recognised as valid taxa until 2017. Between 2008 and 2016, IUCN Red List assessors used only two subspecific names: P. l. leo for African lion populations, and P. l. persica for
8930-511: Was distributed in the Asian and African Palearctic since at least the early Pliocene . The earliest fossils recognisable as lions were found at Olduvai Gorge in Tanzania and are estimated to be up to 2 million years old. Estimates for the divergence time of the modern and cave lion lineages range from 529,000 to 392,000 years ago based on mutation rate per generation time of the modern lion. There
9025-910: Was present in the Caucasus until the 10th century. It lived in Palestine until the Middle Ages and in Southwest Asia until the late 19th century. By the late 19th century, it had been extirpated in most of Turkey. The last live lion in Iran was sighted in 1942, about 65 km (40 mi) northwest of Dezful , although the corpse of a lioness was found on the banks of the Karun river in Khuzestan province in 1944. It once ranged from Sind and Punjab in Pakistan to Bengal and
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