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94-566: (Redirected from Trolltindene ) Trolltinden ( singular ) or Trolltindane (plural) are the names of several summits or mountain ranges in Norway: Innlandet county [ edit ] Trolltinden (Innlandet) (2,018 m) in Rondane, previously called Sagtinden Møre og Romsdal county [ edit ] Trolltindane, a mountain ridge in Rauma, on

188-1054: A dagger (†); groups of uncertain placement are labelled with a question mark (?) and dashed lines (- - - - -). Jawless fishes (118 species: hagfish , lampreys ) [REDACTED] † Thelodonti , † Conodonta , † Anaspida [REDACTED] [REDACTED] [REDACTED] † Galeaspida [REDACTED] † Osteostraci [REDACTED] † Placodermi [REDACTED] † Acanthodii [REDACTED]  (>1,100 species: sharks , rays , chimaeras ) [REDACTED]  (2 species: coelacanths ) [REDACTED] Dipnoi (6 species: lungfish ) [REDACTED] Tetrapoda (>38,000 species, not considered fish: amphibians, reptiles, birds, mammals) [REDACTED]  (14 species: bichirs , reedfish ) [REDACTED]  (27 species: sturgeons , paddlefish ) [REDACTED] Ginglymodi (7 species: gars , alligator gars ) [REDACTED] Halecomorphi (2 species: bowfin , eyetail bowfin ) [REDACTED]  (>32,000 species) [REDACTED] Fishes (without tetrapods) are

282-579: A paraphyletic group and for this reason, the class Pisces seen in older reference works is no longer used in formal classifications. Traditional classification divides fish into three extant classes (Agnatha, Chondrichthyes, and Osteichthyes), and with extinct forms sometimes classified within those groups, sometimes as their own classes. Fish account for more than half of vertebrate species. As of 2016, there are over 32,000 described species of bony fish, over 1,100 species of cartilaginous fish, and over 100 hagfish and lampreys. A third of these fall within

376-728: A facultative trial, like in Ngan'gi . Most languages with a trial are in the Austronesian family, and most non-Austronesian languages with a trial are nearby in Oceania. The latter category includes the Austronesian-influenced English creole languages of Tok Pisin , Bislama , and Pijin . In Australia, the trial can also be found in Aboriginal languages of many different language families. In Indonesia, trial pronouns are common in

470-487: A group of 100,000 referred to in the plural. Much like the dual, it is crosslinguistically variable which words and parts of speech may be marked with the paucal. Baiso has the paucal only for nouns and not pronouns, whereas Yimas has the paucal only for pronouns and not nouns. In Meryam Mir , the paucal is mostly marked on the verbs. Avar has the paucal for only about 90 specific nouns, including brush, spade, snake, and daughter-in-law (the only kin term that can take

564-452: A large number of something, and has been called the plural of abundance. In other languages like Kaytetye , it can refer to all of something in existence, and has been called the global plural. Like some other grammatical numbers, languages also vary as to which cases the greater plural may be used in. The greater plural is more common in nouns than in pronouns. Accordingly, in Kaytetye,

658-436: A more spherical lens . Their retinas generally have both rods and cones (for scotopic and photopic vision ); many species have colour vision , often with three types of cone. Teleosts can see polarized light ; some such as cyprinids have a fourth type of cone that detects ultraviolet . Amongst jawless fish , the lamprey has well-developed eyes, while the hagfish has only primitive eyespots. Hearing too

752-615: A number distinction is pronouns. An example of a personal pronoun system distinguishing singular and plural is that of Wayoró : Like the singular denotes exactly one item, the dual number denotes exactly two items. For example, in Camsá : In languages with a singular/dual/plural paradigm, the exact meaning of plural depends on whether the dual is obligatory or facultative (optional). In contrast to English and other singular/plural languages where plural means two or more, in languages with an obligatory dual, plural strictly means three or more. This

846-558: A number larger than and beyond greater plural. It has also been called the "even greater plural". For example, in Warekena : A similar system is found in Banyun , where the greater plural represents unlimitedness, and the greatest plural represents "a higher degree of unlimitedness". Linguist Daniel Harbour has represented the paucal, greater paucal, plural, greater plural, and greatest plural as collectively definable by "cuts" that divide

940-529: A paucal. Similar things have been said about trial pronouns in Larike and Anejom̃ . Russian has what has variably been called paucal numerals, the count form, the adnumerative, or the genitive of quantification. When a noun in the nominative case has a numeral added to quantify it, the noun becomes genitive singular with 2, 3, or 4, but genitive plural with 5 or above. Many linguists have described these as paucal constructions. However, some have disagreed on

1034-472: A plural, leaving the former plural with a greater plural meaning. A different four-way distinction of singular, paucal, plural, and greater plural can be found in some verbs of Hualapai . A more complex system is found in Mele-Fila : pronouns distinguish singular, dual, plural, and greater plural, but articles attached to nouns distinguish singular, paucal, and plural. The result is that for full sentences, there

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1128-419: A protective bony cover or operculum . They are able to oxygenate their gills using muscles in the head. Some 400 species of fish in 50 families can breathe air, enabling them to live in oxygen-poor water or to emerge on to land. The ability of fish to do this is potentially limited by their single-loop circulation, as oxygenated blood from their air-breathing organ will mix with deoxygenated blood returning to

1222-481: A quadral or a quintal. Linguist Susan McBurney has contended that American Sign Language has a true dual, but that the trial, quadral, and quintal should instead be classified as numeral incorporation rather than grammatical number. This is motivated by the dual marker handshape being distinct from the handshape for the numeral two, in contrast to higher number markers; the ability to also incorporate these numerals into other words, including those for times and amounts; and

1316-479: A quadral; the final 2016 reference grammar of Marshallese by Byron W. Bender , a linguist with expertise in the language, still refers to it as having a quadral. Besides singular, dual, trial, and quadral or paucal, Marshallese additionally has two different plural forms, one for five or more and one for two or more (referred to as multiple and plural absolute respectively), creating a partially overlapping six-way number distinction. Kove has been recorded as having

1410-408: A rare pronoun form for exactly six people. Some American Sign Language speakers have incorporated numerals up to nine into inclusive pronouns upon solicitation. Israeli Sign Language theoretically has the grammatical ability to incorporate numerals up to ten into pronouns. Greater plural is a number larger than and beyond plural. In various forms across different languages, it has also been called

1504-441: A sense of touch and of hearing . Blind cave fish navigate almost entirely through the sensations from their lateral line system. Some fish, such as catfish and sharks, have the ampullae of Lorenzini , electroreceptors that detect weak electric currents on the order of millivolt. Vision is an important sensory system in fish. Fish eyes are similar to those of terrestrial vertebrates like birds and mammals, but have

1598-464: A similar pronoun system as Marshallese, with one addition: the plural (2+) is split between two categories, one for members of the same family and one for members of different families, creating a seven-way distinction. A few other languages have also been claimed to have quadral pronouns. Robert Blust and others have said they exist in some of the Austronesian Kenyah languages , specifically

1692-501: A spoken language with the trial (in both pronouns and verbs) outside of Oceania is Muklom Tangsa , spoken in northeast India. The paucal number represents 'a few', a small inexactly numbered group of items. For example, in Motuna : Almost all languages with a paucal also have a dual. However, this is not universal. Nouns in Mocoví only have singular, paucal, and plural. On the other hand,

1786-456: A system of paucal, greater paucal, plural. Other examples can be found in the related languages of Northern Gumuz and Daatsʼiin . Northern Gumuz is said to mark the plural and greater plural on verbs, and Daatsʼiin is said to mark "three degrees of plurality" (plural, greater plural, and greatest plural) on verbs. In both languages though, the "plural" is often actually a paucal, understood to mean about two to four. However, in neither language

1880-485: A tail fin, jaws, skin covered with scales , and lays eggs. Each criterion has exceptions, creating a wide diversity in body shape and way of life. For example, some fast-swimming fish are warm-blooded, while some slow-swimming fish have abandoned streamlining in favour of other body shapes. Fish species are roughly divided equally between freshwater and marine (oceanic) ecosystems; there are some 15,200 freshwater species and around 14,800 marine species. Coral reefs in

1974-585: A true "land fish" as this worm-like catfish strictly lives among waterlogged leaf litter . Cavefish of multiple families live in underground lakes , underground rivers or aquifers . Like other animals, fish suffer from parasitism . Some species use cleaner fish to remove external parasites. The best known of these are the bluestreak cleaner wrasses of coral reefs in the Indian and Pacific oceans. These small fish maintain cleaning stations where other fish congregate and perform specific movements to attract

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2068-852: A true quadral did exist, but it has since morphed into a plural form. It has thus been hypothesized that the quadral existed in Proto-Oceanic and Proto-Southern Vanuatu. The quintal number denotes exactly five items. Apparent examples of its use can mostly only be found in pronouns of sign languages. Like the quadral, its existence has been contested, and only some classifications accept it. Like trial and quadral forms, rare quintal forms of pronouns have been said to be attested in Tok Pisin and Bislama. These languages insert numerals to represent exact numbers of referents. For example, in Bislama, the numerals tu (two) and tri (three) are contained within

2162-422: A typical fish is adapted for efficient swimming by alternately contracting paired sets of muscles on either side of the backbone. These contractions form S-shaped curves that move down the body. As each curve reaches the tail fin, force is applied to the water, moving the fish forward. The other fins act as control surfaces like an aircraft's flaps, enabling the fish to steer in any direction. Since body tissue

2256-509: Is Nukna , which has only a single trial pronoun, nanggula , which can be either 2nd or 3rd person. The trial may also be marked on verbs, such as in Lenakel . While the dual can be obligatory or facultative, according to Greville Corbett there are no known cases of an obligatory trial, so the trial might always be facultative. However, languages may have both a facultative dual and a facultative trial, like in Larike, or an obligatory dual and

2350-452: Is a combined five-way distinction of singular, dual, paucal, plural, and greater plural. Singular and plural have straightforward number agreements, whereas dual has dual pronouns but paucal articles, paucal has plural pronouns but paucal articles, and greater plural has greater plural pronouns but plural articles. The exact meaning of and terminology for the greater plural differs between languages. In some languages like Miya , it represents

2444-435: Is a larger paucal category, for an inexactly numbered group that is larger in size than a smaller paucal. It can be found in the pronouns of the Austronesian language of Sursurunga , which exhibit a five-way distinction described as singular, dual, paucal, greater paucal, and plural. The Sursurunga paucal is used for smaller groups, usually of about three or four, or for nuclear families of any size. The Sursurunga greater paucal

2538-423: Is an aquatic , anamniotic , gill -bearing vertebrate animal with swimming fins and a hard skull , but lacking limbs with digits . Fish can be grouped into the more basal jawless fish and the more common jawed fish , the latter including all living cartilaginous and bony fish , as well as the extinct placoderms and acanthodians . Most fish are cold-blooded , their body temperature varying with

2632-465: Is an important sensory system in fish. Fish sense sound using their lateral lines and otoliths in their ears, inside their heads. Some can detect sound through the swim bladder. Some fish, including salmon, are capable of magnetoreception ; when the axis of a magnetic field is changed around a circular tank of young fish, they reorient themselves in line with the field. The mechanism of fish magnetoreception remains unknown; experiments in birds imply

2726-431: Is denser than water, fish must compensate for the difference or they will sink. Many bony fish have an internal organ called a swim bladder that allows them to adjust their buoyancy by increasing or decreasing the amount of gas it contains. The scales of fish provide protection from predators at the cost of adding stiffness and weight. Fish scales are often highly reflective; this silvering provides camouflage in

2820-426: Is different from Wikidata All article disambiguation pages All disambiguation pages Grammatical number The word "number" is also used in linguistics to describe the distinction between certain grammatical aspects that indicate the number of times an event occurs, such as the semelfactive aspect, the iterative aspect, etc. For that use of the term, see " Grammatical aspect ". Most languages of

2914-515: Is found particularly in the isolating languages of West Africa. One of the simplest number distinctions a language can make is singular and plural. Singular denotes exactly one referent, while plural denotes more than one referent. For example, in English: To mark number, English has different singular and plural forms for nouns and verbs (in the third person): "my dog watch es television" (singular) and "my dog s watch television" (plural). This

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3008-422: Is not universal: Wambaya marks number on nouns but not verbs, and Onondaga marks number on verbs but not nouns. Latin has different singular and plural forms for nouns, verbs, and adjectives, in contrast to English where adjectives do not change for number. Tundra Nenets can mark singular and plural on nouns, verbs, adjectives, adverbs, and postpositions . However, the most common part of speech to show

3102-447: Is obligatory for pronouns but facultative for nouns. In Comanche , it is obligatory when referring to humans, facultative for other animate nouns, and rarely used for inanimate nouns. There are also languages where use of the dual number is more restricted than singular and plural. In the possessive noun forms of Northern Sámi , the possessor can be in the dual number, but the noun possessed can only be singular or plural. Pronouns are

3196-419: Is rare for a language to mark the trial on nouns, and some sources even claim that trial marking on nouns does not exist. However, it has been recorded for a few languages; besides Awa, Arabana , Urama , and Angaataha have trial number. It is much more common for a language to have trial pronouns, the case for the Austronesian languages of Larike , Tolai , Raga , and Wamesa . A minimal example

3290-465: Is the biggest part of the brain; it is small in hagfish and lampreys , but very large in mormyrids , processing their electrical sense . The brain stem or myelencephalon controls some muscles and body organs, and governs respiration and osmoregulation . The lateral line system is a network of sensors in the skin which detects gentle currents and vibrations, and senses the motion of nearby fish, whether predators or prey. This can be considered both

3384-411: Is the case for Sanskrit , North Mansi , and Alutiiq . In languages with a facultative dual, two of something can be referred to using either the dual or the plural, and so plural means two or more. This is the case for modern Arabic dialects, at least some Inuktitut dialects, and Yandruwandha . In some languages, the dual is obligatory in certain cases but facultative in others. In Slovene , it

3478-417: Is this always the case. The Northern Gumuz paucal/plural may sometimes refer to "much greater than four". In some languages, the default form of a noun is not singular, but rather general, which does not specify number and could mean one or more than one. Singular and plural forms are marked from the general form. The general is used when the specific number is deemed irrelevant or unimportant. In this system,

3572-456: Is to categorize the apparent trial/quadral/quintal forms as "cardinal plurals", or forms of the grammatical plural number where the number of people is specified. Other authors have treated these concepts as perfectly equivalent, referring to pronoun numeral incorporation while still applying the terms quadral and quintal. There are also cases of sign language pronouns indicating specific numbers of referents above five. Ugandan Sign Language has

3666-605: Is used for groups of four or more (and must be used instead of the plural for a group of two or more dyads). There is thus some overlap between the two groups; a family of four can be referred to in Sursurunga by either of the paucals. This distinction is found both in Sursurunga's personal pronouns and in two different sets of possessive pronouns, one for edible things and one for non-edible things. The quadral number denotes exactly four items. Apparent examples of its use are almost entirely confined to pronouns, and specifically those in

3760-606: The Gnathostomata or (for bony fish) Osteichthyes , also contains the clade of tetrapods (four-limbed vertebrates, mostly terrestrial), which are usually not considered fish. Some tetrapods, such as cetaceans and ichthyosaurs , have secondarily acquired a fish-like body shape through convergent evolution . Fishes of the World comments that "it is increasingly widely accepted that tetrapods, including ourselves, are simply modified bony fishes, and so we are comfortable with using

3854-709: The Indo-Pacific constitute the center of diversity for marine fishes, whereas continental freshwater fishes are most diverse in large river basins of tropical rainforests , especially the Amazon , Congo , and Mekong basins. More than 5,600 fish species inhabit Neotropical freshwaters alone, such that Neotropical fishes represent about 10% of all vertebrate species on the Earth. Fish are abundant in most bodies of water. They can be found in nearly all aquatic environments, from high mountain streams (e.g., char and gudgeon ) to

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3948-661: The Silurian , with giant armoured placoderms such as Dunkleosteus . Jawed fish, too, appeared during the Silurian: the cartilaginous Chondrichthyes and the bony Osteichthyes . During the Devonian , fish diversity greatly increased, including among the placoderms, lobe-finned fishes, and early sharks, earning the Devonian the epithet "the age of fishes". Fishes are a paraphyletic group, since any clade containing all fish, such as

4042-647: The Sorbian languages . Indo-European languages that have long ago lost the dual still sometimes have residual traces of it, such as the English distinctions both vs. all , either vs. any , and neither vs. none . The Norwegian både , cognate with English both , has further evolved to be able to refer to more than two items, as in både epler, pærer, og druer , literally "both apples, pears, and grapes." The trial number denotes exactly three items. For example, in Awa : It

4136-469: The abyssal and even hadal depths of the deepest oceans (e.g., cusk-eels and snailfish ), although none have been found in the deepest 25% of the ocean. The deepest living fish in the ocean so far found is a cusk-eel, Abyssobrotula galatheae , recorded at the bottom of the Puerto Rico Trench at 8,370 m (27,460 ft). In terms of temperature, Jonah's icefish live in cold waters of

4230-440: The end-Devonian extinction wiped out the apex predators , the placoderms. Bony fish are further divided into the lobe-finned and ray-finned fish . About 96% of all living fish species today are teleosts , a crown group of ray-finned fish that can protrude their jaws . The tetrapods , a mostly terrestrial clade of vertebrates that have dominated the top trophic levels in both aquatic and terrestrial ecosystems since

4324-400: The intertidal zone , are facultative air breathers, able to breathe air when out of water, as may occur daily at low tide , and to use their gills when in water. Some coastal fish like rockskippers and mudskippers choose to leave the water to feed in habitats temporarily exposed to the air. Some catfish absorb air through their digestive tracts. The digestive system consists of a tube,

4418-405: The ostracoderms , had heavy bony plates that served as protective exoskeletons against invertebrate predators . The first fish with jaws , the placoderms, appeared in the Silurian and greatly diversified during the Devonian , the "Age of Fishes". Bony fish, distinguished by the presence of swim bladders and later ossified endoskeletons , emerged as the dominant group of fish after

4512-425: The stout infantfish . Swimming performance varies from fish such as tuna, salmon , and jacks that can cover 10–20 body-lengths per second to species such as eels and rays that swim no more than 0.5 body-lengths per second. A typical fish is cold-blooded , has a streamlined body for rapid swimming, extracts oxygen from water using gills, has two sets of paired fins, one or two dorsal fins, an anal fin and

4606-457: The English sentences below: The quantity of apples is marked on the noun—"apple" singular number (one item) vs. "apples" plural number (more than one item)—on the demonstrative, that/those , and on the verb, is/are . In the second sentence, all this information is redundant , since quantity is already indicated by the numeral two . A language has grammatical number when its noun forms are subdivided into morphological classes according to

4700-467: The Kiwaian languages, but it is now recognized that many actually have a paucal instead. Linguist Michael Cysouw has suggested that most languages reported to have trials in fact have mislabelled paucals, and that true trials are very rare. On the other hand, Luise Hercus stated in her published grammar of Arabana that the language's trial (which can be marked on nouns) is a true trial which cannot act as

4794-610: The Late Paleozoic , evolved from lobe-finned fish during the Carboniferous , developing air-breathing lungs homologous to swim bladders. Despite the cladistic lineage, tetrapods are usually not considered to be fish, making "fish" a paraphyletic group. Fish have been an important natural resource for humans since prehistoric times, especially as food . Commercial and subsistence fishers harvest fish in wild fisheries or farm them in ponds or in breeding cages in

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4888-452: The Melanesian pidgins of Tok Pisin, Bislama, and Pijin. However, while these are grammatically possible, they are rare, and plural forms are almost always used in their place. Many different sign languages have been explicitly described as having quadral pronoun forms. Estonian Sign Language has even been described as having the quadral for nouns. Marshallese has been said to have

4982-594: The Southern Ocean, including under the Filchner–Ronne Ice Shelf at a latitude of 79°S, while desert pupfish live in desert springs, streams, and marshes, sometimes highly saline, with water temperatures as high as 36 C. A few fish live mostly on land or lay their eggs on land near water. Mudskippers feed and interact with one another on mudflats and go underwater to hide in their burrows. A single undescribed species of Phreatobius has been called

5076-556: The attention of the cleaners. Cleaning behaviors have been observed in a number of fish groups, including an interesting case between two cichlids of the same genus, Etroplus maculatus , the cleaner, and the much larger E. suratensis . Fish occupy many trophic levels in freshwater and marine food webs . Fish at the higher levels are predatory , and a substantial part of their prey consists of other fish. In addition, mammals such as dolphins and seals feed on fish, alongside birds such as gannets and cormorants . The body of

5170-550: The body, and produce a concentrated urine. The reverse happens in freshwater fish : they tend to gain water osmotically, and produce a dilute urine. Some fish have kidneys able to operate in both freshwater and saltwater. Fish have small brains relative to body size compared with other vertebrates, typically one-fifteenth the brain mass of a similarly sized bird or mammal. However, some fish have relatively large brains, notably mormyrids and sharks , which have brains about as large for their body weight as birds and marsupials . At

5264-554: The dual can only be used by an adult male speaking to another adult male. Dual number existed in all nouns and adjectives of Proto-Indo-European around 4000 BCE, and was inherited in some form in many of its prehistoric , protohistoric , ancient , and medieval descendents. Only rarely has it persisted in Indo-European languages to the modern day. It survived in Proto-Germanic in the first and second person pronouns, where it

5358-561: The exact root is unknown; some authorities reconstruct a Proto-Indo-European root * peysk- , attested only in Italic , Celtic , and Germanic . About 530 million years ago during the Cambrian explosion , fishlike animals with a notochord and eyes at the front of the body, such as Haikouichthys , appear in the fossil record . During the late Cambrian , other jawless forms such as conodonts appear. Jawed vertebrates appear in

5452-486: The existence of multiple plural categories may blur the line between paucal and plural. For example, Mele-Fila is said to have a paucal, plural, and greater plural. However, the transition between plural and greater plural occurs around 15 to 20. This puts the Mele-Fila "plural" in range of some larger "paucals" described in other languages. Thus the distinction is muddied between a system of paucal, plural, greater plural, and

5546-405: The front of the brain are the olfactory lobes , a pair of structures that receive and process signals from the nostrils via the two olfactory nerves . Fish that hunt primarily by smell, such as hagfish and sharks, have very large olfactory lobes. Behind these is the telencephalon , which in fish deals mostly with olfaction. Together these structures form the forebrain. Connecting the forebrain to

5640-412: The gills flows in the opposite direction to the water, resulting in efficient countercurrent exchange . The gills push the oxygen-poor water out through openings in the sides of the pharynx. Cartilaginous fish have multiple gill openings: sharks usually have five, sometimes six or seven pairs; they often have to swim to oxygenate their gills. Bony fish have a single gill opening on each side, hidden beneath

5734-532: The gills. Oxygen-rich blood then flows without further pumping, unlike in mammals, to the body tissues. Finally, oxygen-depleted blood returns to the heart. Fish exchange gases using gills on either side of the pharynx . Gills consist of comblike structures called filaments. Each filament contains a capillary network that provides a large surface area for exchanging oxygen and carbon dioxide . Fish exchange gases by pulling oxygen-rich water through their mouths and pumping it over their gills. Capillary blood in

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5828-558: The global plural, the remote plural, the plural of abundance, the unlimited plural, and the superplural. For example, in Tswana : The greater plural may also be a component of larger number systems. Nouns in Barngarla have a four-way distinction of singular, dual, plural, and greater plural. The same four-way distinction is found in Mokilese pronouns, where a former trial has evolved to become

5922-401: The greater plural exists only in nouns and not pronouns. Oppositely, Mokilese has the greater plural in pronouns but not nouns. Chamacoco has the greater plural only in first person inclusive pronouns, second person pronouns, and first person inclusive verb inflections. Tigre has the greater plural only in a single word, nälät , which means a large number of deer. Greatest plural is

6016-425: The grounds that a Russian noun cannot be declined to stand by itself and mean anywhere between 2 and 4. Similar constructions can be found in other Slavic languages , including Polish , Serbo-Croatian , and Slovene. Because Slovene also has a regular dual, there is a four-way distinction of nouns being singular with 1, dual with 2, plural with 3 or 4, and genitive plural with 5 or more. The greater paucal number

6110-447: The gut, leading from the mouth to the anus. The mouth of most fishes contains teeth to grip prey, bite off or scrape plant material, or crush the food. An esophagus carries food to the stomach where it may be stored and partially digested. A sphincter, the pylorus, releases food to the intestine at intervals. Many fish have finger-shaped pouches, pyloric caeca , around the pylorus, of doubtful function. The pancreas secretes enzymes into

6204-452: The heart from the rest of the body. Lungfish, bichirs, ropefish, bowfins, snakefish, and the African knifefish have evolved to reduce such mixing, and to reduce oxygen loss from the gills to oxygen-poor water. Bichirs and lungfish have tetrapod-like paired lungs, requiring them to surface to gulp air, and making them obligate air breathers. Many other fish, including inhabitants of rock pools and

6298-565: The highland Lepoʼ Sawa dialect spoken in Long Anap . There seems to be no other published sources of info on this dialect's pronouns, and an investigation into the lowland Lebo’ Vo’ dialect has revealed a paucal instead of a quadral. A quadral claim has also been made for the animate demonstrative pronouns in Nauruan . Outside the Austronesian family, Abun storytelling reportedly frequently contains quadral pronouns in addition to trial ones. Perhaps

6392-455: The intestine to digest the food; other enzymes are secreted directly by the intestine itself. The liver produces bile which helps to break up fat into an emulsion which can be absorbed in the intestine. Most fish release their nitrogenous wastes as ammonia . This may be excreted through the gills or filtered by the kidneys . Salt is excreted by the rectal gland. Saltwater fish tend to lose water by osmosis ; their kidneys return water to

6486-467: The lack of grammatical number with an extensive system of measure words . Joseph Greenberg has proposed a number category hierarchy as a linguistic universal : "No language has a trial number unless it has a dual. No language has a dual unless it has a plural." This hierarchy does not account for the paucal. Obligatory plural marking of all nouns is found throughout the languages of western and northern Eurasia and most parts of Africa . The rest of

6580-437: The languages of Oceania or in sign languages . It has been contested whether the quadral truly exists in natural language; some linguists have rejected it as an extant category, while others have accepted it. Some languages that have previously been described as having a quadral, like Sursurunga, have since been reanalyzed as having a paucal instead. Like trial forms, quadral forms of pronouns have been said to be attested in

6674-419: The midbrain is the diencephalon ; it works with hormones and homeostasis . The pineal body is just above the diencephalon; it detects light, maintains circadian rhythms, and controls color changes. The midbrain contains the two optic lobes . These are very large in species that hunt by sight, such as rainbow trout and cichlids . The hindbrain controls swimming and balance.The single-lobed cerebellum

6768-443: The nine largest families; from largest to smallest, these are Cyprinidae , Gobiidae , Cichlidae , Characidae , Loricariidae , Balitoridae , Serranidae , Labridae , and Scorpaenidae . About 64 families are monotypic , containing only one species. Fish range in size from the huge 16-metre (52 ft) whale shark to some tiny teleosts only 8-millimetre (0.3 in) long, such as the cyprinid Paedocypris progenetica and

6862-496: The ocean. Fish are caught for recreation , or raised by fishkeepers as ornaments for private and public exhibition in aquaria and garden ponds . Fish have had a role in human culture through the ages, serving as deities , religious symbols, and as the subjects of art, books and movies. The word fish is inherited from Proto-Germanic , and is related to German Fisch , the Latin piscis and Old Irish īasc , though

6956-433: The only known spoken language outside Oceania to have a claimed quadral is Apinayé of Brazil, recorded as having a third person pronominal prefix meaning "they four", although this has been little researched or described. In some Austronesian languages with a singular/dual/trial/plural pronoun system, the plural forms are etymologically related to the number four. This has led to suggestions or assertions that historically

7050-452: The only part of speech with a dual form in some Polynesian languages , including Samoan , Tuvaluan , and Māori . In Maltese , the dual only exists for about 30 specific nouns, of which it is obligatory for only 8 (hour, day, week, month, year, once, hundred, and thousand). Words that can take a facultative dual in Maltese include egg, branch, tear, and wicker basket. In Mezquital Otomi ,

7144-402: The open ocean. Because the water all around is the same colour, reflecting an image of the water offers near-invisibility. Fish have a closed-loop circulatory system . The heart pumps the blood in a single loop throughout the body; for comparison, the mammal heart has two loops, one for the lungs to pick up oxygen, one for the body to deliver the oxygen. In fish, the heart pumps blood through

7238-645: The paucal in Avar). Takivatan Bunun has a paucal only in its distal demonstratives used in reference to people. It is common for former trials to evolve in meaning to become paucals, and many Austronesian languages have paucal markers that are etymologically derived from the numeral three, indicating the old usage. It is less common for duals to evolve into paucals, but this has been observed in some dialects of Arabic. Paucals that are etymologically trials are sometimes incorrectly described as being trials. For example, trial pronouns were once described as being found in all

7332-418: The primary factor for using the paucal is not a specific number range, but the referents forming a single group; although the paucal is most common between 3 and 5, it has been used with more than 20. In Paamese , a major factor is relative group size compared to the plural, such that even though the paucal generally means 12 or fewer, a group of 2,000 people may be referred to in the paucal when contrasted with

7426-845: The pronouns in Mussau and Lihir have dual, trial, and paucal. The lower bound of the paucal is usually defined by what other number categories exist in the language. In singular/paucal/plural paradigms, use of the paucal begins at two, but with the addition of the dual, the paucal begins at three. There is usually no exact upper bound on how many paucal refers to, and its approximate range depends on both language and context. It has been recorded as going up to about 5 in Warndarrang , about 6 in Baiso , 10 in Arabic, and about 10 or 15 in Murrinh-patha . In Manam ,

7520-415: The quadral as a regular feature in its pronoun system. While the apparent Marshallese quadral can mean exactly four, it also has an alternate rhetorical use in speeches to larger groups in order to impart a sense of individual intimacy. According to Greville Corbett , this means it is better classified as a paucal. However, there is not consensus that this alternate use means Marshallese does not truly have

7614-761: The quantity they express, such that: This is partly true for English: every noun and pronoun form is singular or plural (a few, such as " fish ", " cannon " and " you ", can be either, according to context). Some modifiers of nouns—namely the demonstrative determiners—and finite verbs inflect to agree with the number of the noun forms they modify or have as subject: this car and these cars are correct, while * this cars and * these car are incorrect. However, adjectives do not inflect for and many verb forms do not distinguish between singular and plural ("She/They went", "She/They can go", "She/They had gone", "She/They will go"). Many languages distinguish between count nouns and mass nouns . Only count nouns can be freely used in

7708-525: The range of possible numbers into different sections. One low cut defines paucal and plural, and one high cut defines plural and greater plural. Two low cuts define paucal, greater paucal, and plural; one low cut and one high cut define paucal, plural, and greater plural; and two high cuts define plural, greater plural, and greatest plural. There does not appear to be any language with three such cuts, and so no language with three paucal categories and an "even greater paucal". Because they are inexactly defined,

7802-418: The same term [REDACTED] This disambiguation page lists articles associated with the title Trolltinden . If an internal link led you here, you may wish to change the link to point directly to the intended article. Retrieved from " https://en.wikipedia.org/w/index.php?title=Trolltinden&oldid=955631008 " Category : Disambiguation pages Hidden categories: Short description

7896-504: The second person pronouns yutufala (dual) and yutrifala (trial). These forms theoretically have no specific limit, but in practicality usually stop at three. Sign languages described as having a quintal in addition to the quadral include American Sign Language , Argentine Sign Language , British Sign Language , German Sign Language , Levantine Arabic Sign Language , and Ugandan Sign Language . The validity has been debated of categorizing sign language pronouns as having

7990-566: The singular and in the plural. Mass nouns, like "milk", "gold", and "furniture", are normally invariant. (In some cases, a normally mass noun X may be used as a count noun to collect several distinct kinds of X into an enumerable group; for example, a cheesemaker might speak of goat, sheep, and cow milk as milks .) Not all languages have number as a grammatical category. In those that do not, quantity must be expressed either directly, with numerals , or indirectly, through optional quantifiers . However, many of these languages compensate for

8084-757: The singular is often called the singulative, to distinguish it as derived from a different form. Similarly, the plural derived from the general has been called the plurative. For example, in Pular : However, some languages only have a two-way difference between general and plurative, like in Japanese : Less common is a two-way distinction between general and singulative. No language has this as its default number contrast, although some languages have specific nouns with this distinction. For example, in Sidama : Fish#⁘Fish⁘ or ⁘fishes⁘ A fish ( pl. : fish or fishes )

8178-554: The storytelling of Abun , a possible language isolate. In the Solomon Islands, trial pronouns are used very frequently in Touo , either a Central Solomon language or a language isolate. As a result, bilingual speakers of Touo and Pijin will use trial pronouns a lot more commonly in Pijin than other speakers, for whom the trial is usually a lot less common than the dual. A very rare example of

8272-547: The surrounding water, though some large active swimmers like white shark and tuna can hold a higher core temperature . Many fish can communicate acoustically with each other, such as during courtship displays . The study of fish is known as ichthyology . The earliest fish appeared during the Cambrian as small filter feeders ; they continued to evolve through the Paleozoic , diversifying into many forms. The earliest fish with dedicated respiratory gills and paired fins ,

8366-415: The taxon Osteichthyes as a clade, which now includes all tetrapods". The biodiversity of extant fish is unevenly distributed among the various groups; teleosts , bony fishes able to protrude their jaws , make up 96% of fish species. The cladogram shows the evolutionary relationships of all groups of living fishes (with their respective diversity ) and the tetrapods. Extinct groups are marked with

8460-537: The use of markers higher than the dual not being obligatory, with replacement by the plural being acceptable. There was not enough data available to McBurney to argue whether or not these reasons equally applied to other sign languages. Linguist Raquel Veiga Busto has argued they do not equally apply to Catalan Sign Language , and has applied the terms quadral and quintal to the language's pronouns for convenience without taking an official stance as to whether they are grammatical number or numeral incorporation. A third model

8554-795: The western side of Romsdalen valley, including: Store Trolltind (1,788 m), near Troll Wall in Rauma, part of Trolltindane in Rauma Trollveggen or Troll Wall in Rauma, a vertical rock face, part of Trolltindane Trollryggen a summit and pillar in Trolltindane range Trolltindane (890 m), mountains along the border of the municipalities Eide and Fræna Trolltinden (Møre og Romsdal) (1,170 m), also called Brustinden , in Vestnes Nordland county [ edit ] Trolltinden (Nordland) (426 m), in Andøy Topics referred to by

8648-464: The world have formal means to express differences of number. One widespread distinction, found in English and many other languages, involves a simple two-way contrast between singular and plural number ( car / cars , child / children , etc.). Discussion of other more elaborate systems of number appears below. Grammatical number is a morphological category characterized by the expression of quantity through inflection or agreement. As an example, consider

8742-680: The world's languages present a heterogeneous picture. Optional plural marking is common in Southeast and East Asia and Australian languages , and complete lack of plural marking is particularly found in New Guinea and Australian languages. In addition to the areal correlations , there also seems to be at least one correlation with morphological typology : isolating languages appear to favor no or non-obligatory plural marking. This can be seen particularly in Africa, where optionality or absence of plural marking

8836-563: Was then inherited by Old English , Old High German , Old Low German , Early Old Swedish , Old Norwegian , Old Icelandic , and Gothic . It continued in Icelandic until the 1700s, some dialects of Faroese until at least the late 1800s, and some dialects of North Frisian through the 1900s. From Proto-Greek it entered Ancient Greek , and from Proto-Indo-Iranian it entered Sanskrit. From Proto-Slavic , it still exists today in Slovene and

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