Mongolian is the principal language of the Mongolic language family that originated in the Mongolian Plateau . It is spoken by ethnic Mongols and other closely related Mongolic peoples who are native to modern Mongolia and surrounding parts of East and North Asia . Mongolian is the official language of Mongolia and Inner Mongolia and a recognized language of Xinjiang and Qinghai .
150-427: Przewalski's horse ( /( p ) ʃ ə ˈ v ɑː l s k iː z , ˌ p ɜːr ʒ ə -/ (p)shə- VAHL -skeez, PUR -zhə- ; Russian: [prʐɨˈvalʲskʲɪj] ( Пржевальский ); Polish: [pʂɛˈvalskʲi] ; Equus ferus przewalskii or Equus przewalskii ), also called the takhi ( Mongolian : Тахь ), Mongolian wild horse or Dzungarian horse , is a rare and endangered horse originally native to
300-480: A fission of a large chromosome ancestral to domestic horse chromosome 5 to produce Przewalski's horse chromosomes 23 and 24, though conversely, a Robertsonian translocation that fused two chromosomes ancestral to those seen in Przewalski's horse to produce the single large domestic horse chromosome has also been proposed. Many smaller inversions , insertions and other rearrangements were observed between
450-544: A geneticist who developed the idea of cryopreserving genetic material from species considered to be endangered. His ideas led to creating the Frozen Zoo as a genetic library. In 2021, Kurt was relocated to the breeding herd at the San Diego Zoo Safari Park . In order to integrate him into the existing herd, Kurt was partnered with a young female named Holly, a few months older than him, in order to allow him to learn
600-741: A literary standard for Mongolian in whose grammar is said to be based on the Standard Mongolian of Inner Mongolia and whose pronunciation is based on the Chakhar dialect as spoken in the Plain Blue Banner . Dialectologically, however, western Mongolian dialects in Inner Mongolia are closer to Khalkha than they are to eastern Mongolian dialects in Inner Mongolia: e.g. Chakhar is closer to Khalkha than to Khorchin. Juha Janhunen (2003: 179) lists
750-464: A tree of life arose from ancient notions of a ladder-like progression from lower into higher forms of life (such as in the Great Chain of Being ). Early representations of "branching" phylogenetic trees include a "paleontological chart" showing the geological relationships among plants and animals in the book Elementary Geology , by Edward Hitchcock (first edition: 1840). Charles Darwin featured
900-407: A vasectomy on a Przewalski horse in 2007—the first operation of its kind on this species, and possibly the first ever on any endangered species. While normally, a vasectomy may be performed on an endangered animal under limited circumstances, particularly if an individual has already produced many offspring and its genes are overrepresented in the population, scientists realized the animal in question
1050-428: A Przewlaski horse mithocondrial clade. Mongolian language The number of speakers across all its dialects may be 5–6 million, including the vast majority of the residents of Mongolia and many of the ethnic Mongol residents of the Inner Mongolia of China . In Mongolia , Khalkha Mongolian is predominant, and is currently written in both Cyrillic and the traditional Mongolian script . In Inner Mongolia, it
1200-814: A combination of genes that come from different genomic sources (e.g., from mitochondrial or plastid vs. nuclear genomes), or genes that would be expected to evolve under different selective regimes, so that homoplasy (false homology ) would be unlikely to result from natural selection. When extinct species are included as terminal nodes in an analysis (rather than, for example, to constrain internal nodes), they are considered not to represent direct ancestors of any extant species. Extinct species do not typically contain high-quality DNA . The range of useful DNA materials has expanded with advances in extraction and sequencing technologies. Development of technologies able to infer sequences from smaller fragments, or from spatial patterns of DNA degradation products, would further expand
1350-620: A diagrammatic evolutionary "tree" in his 1859 book On the Origin of Species . Over a century later, evolutionary biologists still use tree diagrams to depict evolution because such diagrams effectively convey the concept that speciation occurs through the adaptive and semirandom splitting of lineages. The term phylogenetic , or phylogeny , derives from the two ancient greek words φῦλον ( phûlon ), meaning "race, lineage", and γένεσις ( génesis ), meaning "origin, source". A rooted phylogenetic tree (see two graphics at top)
1500-433: A domestic horse was bred back into the captive Przewalski's horse population. However, recent studies have shown only minimal genetic contribution of this domestic horse to the captive population. The situation was improved when the exchange of breeding animals among facilities increased genetic diversity and there was a consequent improvement in fertility, but the population experienced another genetic bottleneck when many of
1650-590: A few decades later in 1750, a large hunt with thousands of beaters organized by the Manchurian emperor killed between two and three hundred of these horses. The species is named after a Russian colonel of Polish descent, Nikolai Przhevalsky (1839–1888) (Nikołaj Przewalski in Polish). An explorer and naturalist, he obtained the skull and hide of an animal shot in 1878 in the Gobi near today's China–Mongolia border. He would travel to
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#17328842315801800-526: A few weeks but are not weaned for 8–13 months after birth. They reach sexual maturity at two years of age. Przewalski's-type wild horses appear in European cave art dating as far back as 20,000 years ago, but genetic investigation of a 35,870-year-old specimen from one such cave instead showed an affinity with extinct Iberian horse lineage and the modern domestic horse, suggesting that it was not Przewalski's horse being depicted in this art. Horse skeletons dating to
1950-427: A function of the number of tips. For 10 tips, there are more than 34 × 10 6 {\displaystyle 34\times 10^{6}} possible bifurcating trees, and the number of multifurcating trees rises faster, with ca. 7 times as many of the latter as of the former. A dendrogram is a general name for a tree, whether phylogenetic or not, and hence also for the diagrammatic representation of
2100-402: A lone filly captured. Although local herdsmen reported seeing as many as 50 to 100 takhis grazing in small groups then, there were only sporadic sightings of single groups of two or three animals after that, mostly near natural wells. Two scientific expeditions in 1955 and 1962 failed to find any. After herders and naturalists reported single harem groups in 1966 and 1967, the last observation of
2250-609: A more suitable metaphor than the tree . Indeed, phylogenetic corals are useful for portraying past and present life, and they have some advantages over trees ( anastomoses allowed, etc.). Phylogenetic trees composed with a nontrivial number of input sequences are constructed using computational phylogenetics methods. Distance-matrix methods such as neighbor-joining or UPGMA , which calculate genetic distance from multiple sequence alignments , are simplest to implement, but do not invoke an evolutionary model. Many sequence alignment methods such as ClustalW also create trees by using
2400-574: A much broader "Mongolian language" consisting of a Central dialect (Khalkha, Chakhar, Ordos), an Eastern dialect (Kharchin, Khorchin), a Western dialect (Oirat, Kalmyk), and a Northern dialect (consisting of two Buryat varieties). Additionally, the Language Policy in the People's Republic of China: Theory and Practice Since 1949 , states that Mongolian can be classified into four dialects: the Khalkha dialect in
2550-693: A much-needed boost from this new source, with the spread of her bloodline through the inbred captive groups leading to their increased reproductive success, and by 1965, there were more than 130 animals spread among thirty-two zoos and parks. In 1977, the Foundation for the Preservation and Protection of the Przewalski Horse was founded in Rotterdam , the Netherlands, by Jan and Inge Bouman. The foundation started
2700-477: A new study indicated ancient horses of the Botai culture are related to takhis, not to domestic horses as was previously thought. Specifically, the Botai horses appeared to be ancestral to the modern takhi, because all seven takhis nested within the phylogenetic tree of the 20 Botai horses. No comparison was made to definitively wild early takhis. The authors posit that modern Przewalski's horses are feral descendants of
2850-409: A nucleus collected from a cultured Przewalski's horse fibroblast. The resulting embryo was induced to begin division. It was cultured until it reached the blastocyst stage, then implanted into a domestic horse surrogate mare, which carried the embryo to term and delivered a foal with the Przewalski horse DNA of the long-deceased stallion. The cloned horse was named Kurt, after Dr. Kurt Benirschke ,
3000-536: A number of different formats, all of which must represent the nested structure of a tree. They may or may not encode branch lengths and other features. Standardized formats are critical for distributing and sharing trees without relying on graphics output that is hard to import into existing software. Commonly used formats are Although phylogenetic trees produced on the basis of sequenced genes or genomic data in different species can provide evolutionary insight, these analyses have important limitations. Most importantly,
3150-605: A number that had grown by the early 1990s to over 1,500. While dozens of zoos worldwide have Przewalski's horses in small numbers, specialized reserves are also dedicated primarily to the species. The world's largest captive-breeding program for Przewalski's horses is at the Askania Nova preserve in Ukraine. From 1998, thirty-one horses were also released in the unenclosed Chernobyl Exclusion Zone in Ukraine and Belarus . People evacuated
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#17328842315803300-534: A parameter called ATR ( advanced tongue root ); the groups are −ATR, +ATR, and neutral. This alignment seems to have superseded an alignment according to oral backness. However, some scholars still describe Mongolian as being characterized by a distinction between front vowels and back vowels, and the front vowel spellings 'ö' and 'ü' are still often used in the West to indicate two vowels which were historically front. The Mongolian vowel system also has rounding harmony. Length
3450-441: A phylogenetic tree. A cladogram only represents a branching pattern; i.e., its branch lengths do not represent time or relative amount of character change, and its internal nodes do not represent ancestors. A phylogram is a phylogenetic tree that has branch lengths proportional to the amount of character change. A chronogram is a phylogenetic tree that explicitly represents time through its branch lengths. A Dahlgrenogram
3600-530: A postalveolar or palatalized consonant will be followed by an epenthetic [i] , as in [ˈatʃĭɮ] . Stress in Mongolian is nonphonemic (does not distinguish different meanings) and thus is considered to depend entirely on syllable structure. But scholarly opinions on stress placement diverge sharply. Most native linguists, regardless of which dialect they speak, claim that stress falls on the first syllable. Between 1941 and 1975, several Western scholars proposed that
3750-432: A program of exchange between captive populations in zoos worldwide to reduce inbreeding and later began its own breeding program. As a result of such efforts, the extant herd has retained a far greater genetic diversity than its genetic bottleneck made likely. By 1979, when this concerted program of population management to maximize genetic diversity was begun, there were almost four hundred horses in sixteen facilities,
3900-469: A program to reintroduce Przewalski's horses that were bred in France into Mongolia. Instrumental to that 2004 reintroduction was Claudia Feh , a Swiss equine specialist and conservation biologist, Feh led an effort to bring together animals that zoos had conserved to create a breeding population in southern France. Then, after it was established, three family groups were relocated to Khovd in western Mongolia. At
4050-491: A report on sports composed in Mongolian script on stone, which is most often dated at 1224 or 1225. The Mongolian- Armenian wordlist of 55 words compiled by Kirakos of Gandzak (13th century) is the first written record of Mongolian words. From the 13th to the 15th centuries, Mongolian language texts were written in four scripts (not counting some vocabulary written in Western scripts): Uyghur Mongolian (UM) script (an adaptation of
4200-426: A short first syllable are stressed on the second syllable. But if their first syllable is long, then the data for different acoustic parameters seems to support conflicting conclusions: intensity data often seems to indicate that the first syllable is stressed, while F0 seems to indicate that it is the second syllable that is stressed. The grammar in this article is also based primarily on Khalkha Mongolian. Unlike
4350-579: A site on the northern edge of the Gobi Desert , Feh worked in cooperation with local people to ensure the horses survived and flourished. For this work, Feh received a Rolex Award in 2004. In 2004 and 2005, 22 horses were released by the Association Takh to a third reintroduction site in the buffer zone of the Khar Us Nuur National Park , in the northern edge of the Gobi ecoregion. In
4500-583: A system of vowel harmony : For historical reasons, these have been traditionally labeled as "front" vowels and "back" vowels, as /o/ and /u/ developed from /ø/ and /y/, while /ɔ/ and /ʊ/ developed from /o/ and /u/ in Middle Mongolian. Indeed, in Mongolian romanizations , the vowels /o/ and /u/ are often conventionally rendered as ⟨ö⟩ and ⟨ü⟩ , while the vowels /ɔ/ and /ʊ/ are expressed as ⟨o⟩ and ⟨u⟩ . However, for modern Mongolian phonology, it
4650-445: A tree shape, defines a topology only. Some sequence-based trees built from a small genomic locus, such as Phylotree, feature internal nodes labeled with inferred ancestral haplotypes. The number of possible trees for a given number of leaf nodes depends on the specific type of tree, but there are always more labeled than unlabeled trees, more multifurcating than bifurcating trees, and more rooted than unrooted trees. The last distinction
Przewalski's horse - Misplaced Pages Continue
4800-446: A typical Przewalski's horse environment, including: Elymus repens , Carex spp., Fabaceae , and Asteraceae . Looking at the species' diet overall, Przewalski's horses most often eat E. repens , Trifolium pratense , Vicia cracca , Poa trivialis , Dactylis glomerata , and Bromus inermis . While the horses eat a variety of different plant species, they tend to favor different species at different times of year. In
4950-417: A variety of the domesticated horse ( E. caballus ). Przewalski's horse is stockily built, smaller, and shorter than its domesticated relatives. Typical height is about 12–14 hands (48–56 inches, 122–142 cm), and the average weight is around 300 kg (660 lb). They have a dun coat with pangaré features and often have dark primitive markings . Przewalski's horse
5100-508: Is Classical Mongolian , which is dated from the 17th to the 19th century. This is a written language with a high degree of standardization in orthography and syntax that sets it quite apart from the subsequent Modern Mongolian. The most notable documents in this language are the Mongolian Kangyur and Tengyur as well as several chronicles. In 1686, the Soyombo alphabet ( Buddhist texts )
5250-433: Is determined according to phonotactic requirements. The following table lists the consonants of Khalkha Mongolian. The consonants enclosed in parentheses occur only in loanwords. The occurrence of palatalized consonant phonemes, except /tʃ/ /tʃʰ/ /ʃ/ /j/ , is restricted to words with [−ATR] vowels. A rare feature among the world's languages, Mongolian has neither a voiced lateral approximant, such as [l] , nor
5400-705: Is dialectally more diverse and written in the traditional Mongolian script. However, Mongols in both countries often use the Latin script for convenience on the Internet. In the discussion of grammar to follow, the variety of Mongolian treated is the standard written Khalkha formalized in the writing conventions and in grammar as taught in schools, but much of it is also valid for vernacular (spoken) Khalkha and other Mongolian dialects, especially Chakhar Mongolian . Some classify several other Mongolic languages like Buryat and Oirat as varieties of Mongolian, but this classification
5550-409: Is ellipsis . The rules governing the morphology of Mongolian case endings are intricate, and so the rules given below are only indicative. In many situations, further (more general) rules must also be taken into account in order to produce the correct form: these include the presence of an unstable nasal or unstable velar, as well as the rules governing when a penultimate vowel should be deleted from
5700-466: Is CVVCCC, where the last C is a word-final suffix. A single short vowel rarely appears in syllable-final position . If a word was monosyllabic historically, *CV has become CVV. In native words, the following consonants do not occur word-initially: /w̜/ , /ɮ/ , /r/ , /w̜ʲ/ , /ɮʲ/ , /rʲ/ , /tʰʲ/ , and /tʲ/ . [ŋ] is restricted to codas (else it becomes [n] ), and /p/ and /pʲ/ do not occur in codas for historical reasons. For two-consonant clusters,
5850-399: Is a directed tree with a unique node — the root — corresponding to the (usually imputed ) most recent common ancestor of all the entities at the leaves of the tree. The root node does not have a parent node, but serves as the parent of all other nodes in the tree. The root is therefore a node of degree 2, while other internal nodes have a minimum degree of 3 (where "degree" here refers to
6000-438: Is a +ATR vowel, then every vowel of the word must be either /i/ or a +ATR vowel. In the case of suffixes, which must change their vowels to conform to different words, two patterns predominate. Some suffixes contain an archiphoneme /A/ that can be realized as /a, ɔ, e, o/ ; e.g. Other suffixes can occur in /U/ being realized as /ʊ, u/ , in which case all −ATR vowels lead to /ʊ/ and all +ATR vowels lead to /u/ ; e.g. If
6150-409: Is a branching diagram or a tree showing the evolutionary relationships among various biological species or other entities based upon similarities and differences in their physical or genetic characteristics. In evolutionary biology, all life on Earth is theoretically part of a single phylogenetic tree, indicating common ancestry . Phylogenetics is the study of phylogenetic trees. The main challenge
Przewalski's horse - Misplaced Pages Continue
6300-604: Is a diagram representing a cross section of a phylogenetic tree. A phylogenetic network is not strictly speaking a tree, but rather a more general graph , or a directed acyclic graph in the case of rooted networks. They are used to overcome some of the limitations inherent to trees. A spindle diagram, or bubble diagram, is often called a romerogram, after its popularisation by the American palaeontologist Alfred Romer . It represents taxonomic diversity (horizontal width) against geological time (vertical axis) in order to reflect
6450-447: Is about 2.1 m (6 ft 11 in). It weighs around 300 kilograms (660 lb). The coat is generally dun in color with pangaré features, varying from dark brown around the mane, to pale brown on the flanks, and yellowish-white on the belly, as well as around the muzzle. The legs of Przewalski's horse are often faintly striped, also typical of primitive markings . The mane stands erect and does not extend as far forward, while
6600-485: Is based primarily on the Khalkha dialect as spoken in Ulaanbaatar , Mongolia's capital. The phonologies of other varieties such as Ordos, Khorchin, and even Chakhar, differ considerably. This section discusses the phonology of Khalkha Mongolian with subsections on Vowels, Consonants, Phonotactics and Stress. The standard language has seven monophthong vowel phonemes. They are aligned into three vowel harmony groups by
6750-634: Is being established in the Iberian System in Spain, the first free-roaming Przewalski’s horses in Western Europe. In 2024, a Colorado rancher discovered what appears to be a critically endangered Przewalski's horse at a Kansas livestock auction, mistakenly identified as a mule. Another similar horse was found at a Utah sanctuary. Genetic tests suggest both are Przewalski's horses, raising concerns about how they ended up in U.S. auctions. The owners care for
6900-566: Is impeded by the fact that existing data for the major varieties is not easily arrangeable according to a common set of linguistic criteria. Such data might account for the historical development of the Mongolian dialect continuum , as well as for its sociolinguistic qualities. Though phonological and lexical studies are comparatively well developed, the basis has yet to be laid for a comparative morphosyntactic study, for example between such highly diverse varieties as Khalkha and Khorchin. In Juha Janhunen's book titled Mongolian , he groups
7050-410: Is more appropriate to instead characterize the two vowel-harmony groups by the dimension of tongue root position. There is also one neutral vowel, /i/ , not belonging to either group. All the vowels in a non compound word, including all its suffixes, must belong to the same group. If the first vowel is −ATR, then every vowel of the word must be either /i/ or a −ATR vowel. Likewise, if the first vowel
7200-506: Is most true of genetic material that is subject to lateral gene transfer and recombination , where different haplotype blocks can have different histories. In these types of analysis, the output tree of a phylogenetic analysis of a single gene is an estimate of the gene's phylogeny (i.e. a gene tree) and not the phylogeny of the taxa (i.e. species tree) from which these characters were sampled, though ideally, both should be very close. For this reason, serious phylogenetic studies generally use
7350-427: Is not in line with the current international standard. Mongolian is a language with vowel harmony and a complex syllabic structure compared to other Mongolic languages, allowing clusters of up to three consonants syllable-finally. It is a typical agglutinative language that relies on suffix chains in the verbal and nominal domains. While there is a basic word order, subject–object–verb , ordering among noun phrases
7500-1261: Is phonemic for vowels, and except short [e], which has merged into short [i], at least in Ulaanbaatar dialect, each of the other six phonemes occurs both short and long. Phonetically, short /o/ has become centralised to the central vowel [ɵ] . In the following table, the seven vowel phonemes, with their length variants, are arranged and described phonetically. The vowels in the Mongolian Cyrillic alphabet are: Khalkha also has four diphthongs : historically /ui, ʊi, ɔi, ai/ but are pronounced more like [ʉe̯, ʊe̯, ɞe̯, æe̯] ; e.g. ой in нохой ( nohoi ) [nɔ̙ˈχɞe̯] 'dog', ай in далай ( dalai ) [taˈɮæe̯] sea', уй in уйлах ( uilah ) [ˈʊe̯ɮɐχ] 'to cry', үй in үйлдвэр ( üildver ) [ˈʉe̯ɮtw̜ɘr] 'factory', эй in хэрэгтэй ( heregtei ) [çiɾɪxˈtʰe] 'necessary'. There are three additional rising diphthongs /ia/ (иа), /ʊa/ (уа) /ei/ (эй); e.g. иа in амиараа ( amiaraa ) [aˈmʲæɾa] 'individually', уа in хуаран ( huaran ) [ˈχʷaɾɐɴ] 'barracks'. This table below lists vowel allophones (short vowels allophones in non-initial positions are used interchangeably with schwa): Mongolian divides vowels into three groups in
7650-458: Is relatively free, as grammatical roles are indicated by a system of about eight grammatical cases . There are five voices . Verbs are marked for voice, aspect , tense and epistemic modality / evidentiality . In sentence linking, a special role is played by converbs . Modern Mongolian evolved from Middle Mongol , the language spoken in the Mongol Empire of the 13th and 14th centuries. In
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#17328842315807800-411: Is seen in modern domestic horses, indicating neither is an ancestor of the other. For example, Przewalski's horse has 33 chromosome pairs, compared to 32 for the domestic horse. Their ancestral lineages split from a common ancestor between 160,000 and 38,000 years ago, long before the domestication of the horse . Przewalski's horse was long considered the only remaining truly wild horse, in contrast with
7950-539: Is the most biologically relevant; it arises because there are many places on an unrooted tree to put the root. For bifurcating labeled trees, the total number of rooted trees is: For bifurcating labeled trees, the total number of unrooted trees is: Among labeled bifurcating trees, the number of unrooted trees with n {\displaystyle n} leaves is equal to the number of rooted trees with n − 1 {\displaystyle n-1} leaves. The number of rooted trees grows quickly as
8100-458: Is to find a phylogenetic tree representing optimal evolutionary ancestry between a set of species or taxa. Computational phylogenetics (also phylogeny inference) focuses on the algorithms involved in finding optimal phylogenetic tree in the phylogenetic landscape. Phylogenetic trees may be rooted or unrooted. In a rooted phylogenetic tree, each node with descendants represents the inferred most recent common ancestor of those descendants, and
8250-402: Is word-final, it gets stressed anyway. In cases where there is only one phonemic short word-initial syllable, even this syllable can get the stress: More recently, the most extensive collection of phonetic data so far in Mongolian studies has been applied to a partial account of stress placement in the closely related Chakhar dialect. The conclusion is drawn that di- and trisyllabic words with
8400-516: The Altaic language family and contrasted with the Mainland Southeast Asia linguistic area . However, instead of a common genetic origin, Clauson, Doerfer, and Shcherbak proposed that Turkic, Mongolic and Tungusic languages form a language Sprachbund , rather than common origin. Mongolian literature is well attested in written form from the 13th century but has earlier Mongolic precursors in
8550-559: The Great Gobi B Strictly Protected Area continued. as of 2011, an estimated total of almost 400 horses existed in three free-ranging populations in the wild. Prague Zoo has transported horses to Mongolia in several rounds in cooperation with partners (Czech Air Force, European Breeding Programme for Przewalski's Horses, Association pour le cheval de Przewalski: Takh, Czech Development Agency, Czech Embassy in Mongolia, and others). The zoo has
8700-479: The Great Gobi B Strictly Protected Area . The ranges of harems are separated, but slightly overlapping. They have few modern predators, but one of the few is the Himalayan wolf . Horses maintain visual contact with their family and herd at all times, and have a host of ways to communicate with one another, including vocalizations, scent marking , and a wide range of visual and tactile signals. Each kick, groom, tilt of
8850-501: The Kalmyk variety ) and Buryat, both of which are spoken in Russia, Mongolia, and China; and Ordos , spoken around Inner Mongolia's Ordos City . The influential classification of Sanžeev (1953) proposed a "Mongolian language" consisting of just the three dialects Khalkha, Chakhar, and Ordos, with Buryat and Oirat judged to be independent languages. On the other hand, Luvsanvandan (1959) proposed
9000-642: The Shuluun Huh/Zhènglán Banner , and is written in the traditional Mongolian script . The number of Mongolian speakers in China is still larger than in the state of Mongolia, where the majority of Mongolians in China speak one of the Khorchin dialects , or rather more than two million of them speak the Khorchin dialect itself as their mother tongue, so that the Khorchin dialect group has about as many speakers as
9150-627: The Uyghur alphabet), 'Phags-pa script (Ph) (used in decrees), Chinese (SM) ( The Secret History of the Mongols ), and Arabic (AM) (used in dictionaries). While they are the earliest texts available, these texts have come to be called " Middle Mongol " in scholarly practice. The documents in UM script show some distinct linguistic characteristics and are therefore often distinguished by terming their language "Preclassical Mongolian". The Yuan dynasty referred to
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#17328842315809300-723: The steppes of Central Asia . It is named after the Russian geographer and explorer Nikolay Przhevalsky . Once extinct in the wild , since the 1990s it has been reintroduced to its native habitat in Mongolia in the Khustain Nuruu National Park , Takhin Tal Nature Reserve, and Khomiin Tal, as well as several other locales in Central Asia and Eastern Europe . Several genetic characteristics of Przewalski's horse differ from what
9450-553: The syllable 's position in the word. In word-initial syllables, there is a phonemic contrast in vowel length . A long vowel has about 208% the length of a short vowel. In word-medial and word-final syllables, formerly long vowels are now only 127% as long as short vowels in initial syllables, but they are still distinct from initial-syllable short vowels. Short vowels in noninitial syllables differ from short vowels in initial syllables by being only 71% as long and by being centralized in articulation. As they are nonphonemic, their position
9600-438: The 1980s had already substantially increased the genetic diversity of the captive population after he was discovered to have more unique alleles than any other horse living at the time, including otherwise lost genetic material from two of the original captive founders. To produce the clone, frozen skin fibroblasts were thawed, and grown in cell culture . An oocyte was collected from a domestic horse, and its nucleus replaced by
9750-474: The American mustang and the Australian brumby , which are instead feral horses descended from domesticated animals. That status was called into question when domestic horses of the 5,000-year-old Botai culture of Central Asia were found to be more closely related to Przewalski's horses than to E. f. caballus . The study raised the possibility that modern Przewalski's horses could be the feral descendants of
9900-667: The Dzungarian Basin to observe it in the wild. In 1881, the horse received a formal scientific description and was named Equus przevalskii by Ivan Semyonovich Polyakov, based on Przewalski's collection and description, while in 1884, the sole exemplar of the horse in Europe was a preserved specimen in the Museum of the Russian Academy of Sciences in St. Petersburg . This was supplemented in 1894 when
10050-588: The Inner Mongolia since September, which caused widespread protests among ethnic Mongol communities. These protests were quickly suppressed by the Chinese government. Mandarin has been deemed the only language of instruction for all subjects as of September 2023. Mongolian belongs to the Mongolic languages . The delimitation of the Mongolian language within Mongolic is a much disputed theoretical problem, one whose resolution
10200-461: The Khalkha dialect group in the State of Mongolia. Nevertheless, the Chakhar dialect, which today has only about 100,000 native speakers and belongs to the Khalkha dialect group, is the basis of standard Mongolian in China. The characteristic differences in the pronunciation of the two standard varieties include the umlauts in Inner Mongolia and the palatalized consonants in Mongolia (see below) as well as
10350-812: The Mongolian language in Chinese as "Guoyu" ( Chinese : 國語 ), which means "National language", a term also used by other non-Han dynasties to refer to their languages such as the Manchu language during the Qing dynasty , the Jurchen language during the Jin dynasty (1115–1234) , the Khitan language during the Liao dynasty , and the Xianbei language during the Northern Wei period. The next distinct period
10500-497: The Mongolic language family into four distinct linguistic branches: The Common Mongolic branch is grouped in the following way: There is no disagreement that the Khalkha dialect of the Mongolian state is Mongolian. However, the status of certain varieties in the Common Mongolic group—whether they are languages distinct from Mongolian or just dialects of it—is disputed. There are at least three such varieties: Oirat (including
10650-467: The United States. Many facilities failed in their attempts at captive breeding, but a few programs were established. However, by the mid-1930s, inbreeding had caused reduced fertility, and the captive population experienced a genetic bottleneck , with the surviving captive breeding stock descended from only 11 of the founder captives. In addition, in at least one instance, the progeny of interbreeding with
10800-426: The accusative, while it must take the nominative if it is indefinite . In addition to case, a number of postpositions exist that usually govern the genitive, dative-locative, comitative and privative cases, including a marked form of the nominative (which can itself then take further case forms). There is also a possible attributive case (when a noun is used attributively ), which is unmarked in most nouns but takes
10950-407: The age of five. Stallions assemble groups of mares or challenge the leader of another group for dominance. Females are able to give birth at the age of three and have a gestation period of 11–12 months. Foals are able to stand about an hour after birth. The rate of infant mortality among foals is 25%, with 83.3% of these deaths resulting from leading stallion infanticide. Foals begin grazing within
11100-453: The ancient Botai domesticated animals, rather than representing a surviving population of never-domesticated horses. Another geneticist pointed out that Przewalski's horses may have simply descended from the same wild population that the Botai horses came from, which would still be compatible with the findings of the study. In 2021, William Taylor and Christina Barron-Ortiz disputed the evidence for domestication of Przewalski's horse. Their case
11250-579: The animal was changed from "extinct in the wild" to " endangered " in 2005, while on the IUCN Red List they were reclassified from "extinct in the wild" to " critically endangered ", after a reassessment in 2008, and from "critically endangered" to "endangered" after a 2011 reassessment. In 2011, Prague Zoo started a new project, Return of the Wild Horses . With the support of public and many strategic partners, yearly transports of captive-bred horses into
11400-666: The animals but hope they can eventually join professional conservation programs. The Przewalski's Horse Reintroduction Project of China was initiated in 1985 when 11 wild horses were imported from overseas. After more than two decades of effort, the Xinjiang Wild Horse Breeding Centre has bred a large number of horses, 55 of which were released into the Kalamely Mountain area. The animals quickly adapted to their new environment. In 1988, six foals were born and survived, and by 2001, over 100 horses were at
11550-425: The brothers Grum-Grzhimailo returned several hides and skulls to St. Petersburg and described the horse's behavior in the wild. A number of these horses were captured around 1900 by Carl Hagenbeck and placed in zoos, and these, along with one later captive, reproduced to give rise to today's population. After 1903, there were no reports of the wild population until 1947, when several isolated groups were observed and
11700-514: The central problem remains the question of how to classify Chakhar, Khalkha, and Khorchin in relation to each other and in relation to Buryat and Oirat. The split of [tʃ] into [tʃ] before *i and [ts] before all other reconstructed vowels, which is found in Mongolia but not in Inner Mongolia, is often cited as a fundamental distinction, for example Proto-Mongolic *tʃil , Khalkha /tʃiɮ/ , Chakhar /tʃil/ 'year' versus Proto-Mongolic *tʃøhelen , Khalkha /tsoːɮəŋ/ , Chakhar /tʃoːləŋ/ 'few'. On
11850-535: The centre. As of 2013, the center hosted 127 horses divided into 13 breeding herds and three bachelor herds. Reintroductions organized by Western European countries started in the 1990s. Several populations have now been released into the wild. A cooperative venture between the Zoological Society of London and Mongolian scientists has successfully reintroduced these horses from zoos into their natural habitat in Mongolia. In 1992, 16 horses were released into
12000-470: The chromosomes of domestic and Przewalski's horses, while there was much lower hetero zygosity in Przewalski's horses, with extensive segments devoid of genetic diversity, a consequence of the recent severe bottleneck of the captive Przewalski's horse population. In comparison, the chromosomal differences between domestic horses and zebras include numerous large-scale translocations , fusions, inversions, and centromere repositioning. Przewalski's horse has
12150-400: The domestic Botai horses. However, it remains possible that both the Botai horses and the modern Przewalski's horses descend separately from the same ancient wild Przewalski's horse population. Its taxonomic position is still debated, with some taxonomists treating Przewalski's horse as a species , E. przewalskii , others as a subspecies of wild horse ( E. ferus przewalskii ) or
12300-526: The ear, or other contact with another horse is a means of communicating. This constant communication leads to complex social behaviors among Przewalski's horses. The historical population was said to have lived in the "wildest parts of the desert" with a preference for "especially saline districts". They were observed mostly during spring and summer at natural wells, migrating to them by crossing valleys rather than by way of higher mountains. Przewalski horse's diet consists of vegetation. Many plant species are in
12450-477: The earlier decades of captivity, the insular breeding by individual zoos led to inbreeding and reduced fertility. In 1979, several American zoos began a collaborative breeding-exchange program to maximize genetic diversity. Recent advances in equine reproductive science have also been used to preserve and expand the gene pool. Scientists at the Smithsonian Institution 's National Zoo successfully reversed
12600-455: The edge lengths in some trees may be interpreted as time estimates. Each node is called a taxonomic unit. Internal nodes are generally called hypothetical taxonomic units, as they cannot be directly observed. Trees are useful in fields of biology such as bioinformatics , systematics , and phylogenetics . Unrooted trees illustrate only the relatedness of the leaf nodes and do not require the ancestral root to be known or inferred. The idea of
12750-506: The end of the 1950s, only 12 individual horses were left in the world's zoos. A wild-caught mare captured as a foal a decade earlier was introduced into the Ukrainian captive population in 1957. This would prove the last wild-caught horse, and with the presumed extinction of the wild population, last sighted in Mongolia in the late 1960s, the captive population became the sole representatives of Przewalski's horse. Genetic diversity received
12900-412: The examples given above, the words are phonetically [ˈxɔjɔ̆r] , [ˈatʃĭɮ] , and [ˈsaːrmăɢ] . The phonetic form of the epenthetic vowel follows from vowel harmony triggered by the vowel in the preceding syllable. Usually it is a centralized version of the same sound, with the following exceptions: preceding /u/ produces [e] ; /i/ will be ignored if there is a nonneutral vowel earlier in the word; and
13050-402: The extinct tarpan E. f. ferus ); or even a subpopulation of the domestic horse. The American Society of Mammalogists considers Przewalski's horse and the tarpan both to be subspecies of Equus ferus , and classifies the domestic horse as a separate species, Equus caballus . Genetic analysis shows that the takhi and the domestic horse differ significantly, with neither ancestral to
13200-449: The family group until they are no longer dependent, usually at two or three years old. Young females join other harems, while bachelor stallions as well as old stallions who have lost their harems join bachelor groups. Family groups can join to form a herd that moves together. The patterns of their daily lives exhibit horse behavior similar to that of feral horse herds. Stallions herd, drive, and defend all members of their family, while
13350-611: The fifth to the third millennia BCE, found in Central Asia, with a range extending to the southern Urals and the Altai , belong to the genetic lineage of Przewalski's horse. Of particular note are the horses of this lineage found in the archaeological sites of the Chalcolithic Botai culture. Sites dating from the mid-fourth-millennium BCE show evidence of horse domestication. Analysis of ancient DNA from Botai horse specimens from about 3000 BCE reveals them to have DNA markers consistent with
13500-525: The first European sightings of the horses in the journal recounting his trip to Mongolia as a prisoner of the Mongol Khan . Another was recorded as a gift to the Manchurian emperor around 1630, its value as a gift suggesting a difficulty in obtaining them. John Bell , a Scottish doctor in service to Peter the Great from 1719 to 1722, observed a horse in Russia's Tomsk Oblast that was apparently this species, and
13650-612: The following Mongol dialects, most of which are spoken in Inner Mongolia . There are two standard varieties of Mongolian. Standard Mongolian in the state of Mongolia is based on the northern Khalkha Mongolian dialects, which include the dialect of Ulaanbaatar , and is written in the Mongolian Cyrillic script . Standard Mongolian in Inner Mongolia is based on the Chakhar Mongolian of the Khalkha dialect group, spoken in
13800-444: The following restrictions obtain: Clusters that do not conform to these restrictions will be broken up by an epenthetic nonphonemic vowel in a syllabification that takes place from right to left. For instance, hoyor 'two', azhil 'work', and saarmag 'neutral' are, phonemically, /xɔjr/ , /atʃɮ/ , and /saːrmɡ/ respectively. In such cases, an epenthetic vowel is inserted to prevent disallowed consonant clusters. Thus, in
13950-449: The form of an unrooted binary tree , a free tree with exactly three neighbors at each internal node. In contrast, a rooted multifurcating tree may have more than two children at some nodes and an unrooted multifurcating tree may have more than three neighbors at some nodes. Both rooted and unrooted trees can be either labeled or unlabeled. A labeled tree has specific values assigned to its leaves, while an unlabeled tree, sometimes called
14100-428: The free expression of natural Przewalski's horse behaviors. In 1993, eleven zoo-born horses were brought to Le Villaret. Horses born there are adapted to life in the wild, free to choose their mates, and required to forage independently. This was intended to produce individuals capable of being reintroduced into Mongolia. In 2012, 39 individuals were at Le Villaret. An intensely researched population of free-ranging animals
14250-605: The help of the San Diego Zoo Global Frozen Zoo. It is the first reported case of any endangered species having more than one clone successfully produced. This individual eventually joins Kurt and Holly at the San Diego Zoo Safari Park. Due to having been conceived through the transfer of a somatic cell nucleus into an egg cell obtained from a domestic horse donor, Kurt and Ollie both display the mithocondrial genome of domestic horses instead of belonging to
14400-546: The highest diploid chromosome number among all equine species. They can interbreed with the domestic horse and produce fertile offspring, with 65 chromosomes. The mitochondrial genome has 37 genes that are 99.63% identical to that of the domestic horse. Przewalski reported the horses forming troops of between five and fifteen members, consisting of a mature stallion, his mares and foals. Modern reintroduced populations similarly form family groups of one adult stallion, one to three mares, and their common offspring that stay in
14550-505: The horses failed to survive World War II. The most valuable group, in Askania Nova , Ukraine , was shot by German soldiers during World War II occupation, and the group in the United States had died out. Only two captive populations in zoos remained, in Munich and in Prague , and of the 31 remaining horses at war's end, only 9 became ancestors of the subsequent captive population. By
14700-512: The horses were released into the reserve proper in September. This introduction was intended to address the buildup of dense scrub caused by the decrease in traditional sheep grazing due to rural depopulation. The horses are intended to fill a niche similar to that of the extinct European wild horse and of contemporary domesticated herbivores by opening the landscape through low-intensity grazing and browsing, thereby enhancing biodiversity and lowering
14850-420: The ice and snow. Their winter diet is very similar to the winter diet of domestic horses, but differs from that revealed by isotope analysis of the historical (pre-captivity) population, which switched in winter to browsing shrubs, though the difference may be due to the extreme habitat pressure the historical population was under. In the wintertime, they eat their food more slowly than they do during other times of
15000-583: The independent words derived using verbal suffixes can roughly be divided into three classes: final verbs , which can only be used sentence-finally, i.e. ‑ na (mainly future or generic statements) or ‑ ö (second person imperative); participles (often called "verbal nouns"), which can be used clause-finally or attributively, i.e. ‑ san ( perfect - past ) or ‑ maar 'want to'; and converbs , which can link clauses or function adverbially , i.e. ‑ zh (qualifies for any adverbial function or neutrally connects two sentences ) or ‑ tal (the action of
15150-413: The language is spoken by roughly half of the country's 5.8 million ethnic Mongols (2005 estimate) However, the exact number of Mongolian speakers in China is unknown, as there is no data available on the language proficiency of that country's citizens. The use of Mongolian in Inner Mongolia has witnessed periods of decline and revival over the last few hundred years. The language experienced a decline during
15300-405: The late Qing period, a revival between 1947 and 1965, a second decline between 1966 and 1976, a second revival between 1977 and 1992, and a third decline between 1995 and 2012. However, in spite of the decline of the Mongolian language in some of Inner Mongolia's urban areas and educational spheres, the ethnic identity of the urbanized Chinese-speaking Mongols is most likely going to survive due to
15450-413: The latter was independent, involving a different wild population, from any possible domestication of Przewalski's horse by the Botai culture. Przewalski's horse is stockily built in comparison to domesticated horses, with shorter legs, and is much smaller and shorter than its domesticated relatives. Typical height is about 12–14 hands (48–56 inches, 122–142 cm), and length
15600-409: The leaf nodes without making assumptions about ancestry. They do not require the ancestral root to be known or inferred. Unrooted trees can always be generated from rooted ones by simply omitting the root. By contrast, inferring the root of an unrooted tree requires some means of identifying ancestry. This is normally done by including an outgroup in the input data so that the root is necessarily between
15750-418: The leftmost heavy syllable gets the stress. Yet other positions were taken in works published between 1835 and 1915. Walker (1997) proposes that stress falls on the rightmost heavy syllable unless this syllable is word-final: A "heavy syllable" is defined as one that is at least the length of a full vowel; short word-initial syllables are thereby excluded. If a word is bisyllabic and the only heavy syllable
15900-506: The lineage of modern Przewalski's horses. There are sporadic reports of Przewalski's horse in the historical record before its formal characterization. The Buddhist monk Bodowa wrote a description of what is thought to have been Przewalski's horse about AD 900, and an account from 1226 reports an incident involving wild horses during Genghis Khan 's campaign against the Tangut empire . In the fifteenth century, Johann Schiltberger recorded one of
16050-565: The literature of the Khitan and other Xianbei peoples. The Bugut inscription dated to 584 CE and the Inscription of Hüis Tolgoi dated to 604–620 CE appear to be the oldest substantial Mongolic or Para-Mongolic texts discovered. Writers such as Owen Lattimore referred to Mongolian as "the Mongol language". The earliest surviving Mongolian text may be the Stele of Yisüngge [ ru ] ,
16200-590: The longest uninterrupted history of breeding Przewalski's horses in the world and keeps the studbook of this species. The first reintroduction into the Orenburg region on the Russian steppe occurred in 2016. In May 2023, a herd of ten Przewalski's horses obtained from Monts D'Azur Biological Reserve in France was introduced by Rewilding Europe to the Iberian Highlands rewilding landscape in Spain, near Villanueva de Alcorón . Following an acclimatization period,
16350-421: The main clause takes place until the action expressed by the suffixed verb begins). Roughly speaking, Mongolian has between seven and nine cases : nominative ( unmarked ), genitive , dative - locative , accusative , ablative , instrumental , comitative , privative and directive , though the final two are not always considered part of the case paradigm. If a direct object is definite , it must take
16500-499: The mares often display leadership in the family. Stallions and mares stay with their preferred partners for years. While behavioral synchronization is high among mares, stallions other than the main harem stallion are generally less stable in this respect. Home range in the wild is little studied, but estimated as 1.2–24 km (0.46–9.3 sq mi) in the Hustai National Park and 150–825 km (58–319 sq mi) in
16650-635: The middle, the Horcin-Haracin dialect in the East, Oriat-Hilimag in the west, and Bargu–Buriyad in the north. Some Western scholars propose that the relatively well researched Ordos variety is an independent language due to its conservative syllable structure and phoneme inventory. While the placement of a variety like Alasha , which is under the cultural influence of Inner Mongolia but historically tied to Oirat, and of other border varieties like Darkhad would very likely remain problematic in any classification,
16800-565: The onager, they were a steppe animal driven to this barren last refuge by the dual pressures of hunting and habitat loss to agricultural grazing. There were two distinct populations recognized by local Mongolians, a lighter steppe variety and a darker mountain one. This distinction is seen in early twentieth-century descriptions. Their mountainous habitat included the Takhiin Shar Nuruu (The Yellow Wild-Horse Mountain Range). In their last decades in
16950-491: The only vowel in the word stem is /i/ , the suffixes will use the +ATR suffix forms. Mongolian also has rounding harmony, which does not apply to close vowels. If a stem contains /o/ (or /ɔ/ ), a suffix that is specified for an open vowel will have [o] (or [ɔ] , respectively) as well. However, this process is blocked by the presence of /u/ (or /ʊ/ ) and /ei/ ; e.g. /ɔr-ɮɔ/ 'came in', but /ɔr-ʊɮ-ɮa/ 'inserted'. The pronunciation of long and short vowels depends on
17100-439: The optimal tree using many of these techniques is NP-hard , so heuristic search and optimization methods are used in combination with tree-scoring functions to identify a reasonably good tree that fits the data. Tree-building methods can be assessed on the basis of several criteria: Tree-building techniques have also gained the attention of mathematicians. Trees can also be built using T-theory . Trees can be encoded in
17250-608: The other hand, the split between the past tense verbal suffixes - /sŋ/ in the Central varieties v. - /dʒɛː/ in the Eastern varieties is usually seen as a merely stochastic difference. In Inner Mongolia, official language policy divides the Mongolian language into three dialects: Standard Mongolian of Inner Mongolia , Oirat, and Barghu-Buryat. The Standard Mongolian of Inner Mongolia is said to consist of Chakhar, Ordos, Baarin , Khorchin, Kharchin, and Alasha. The authorities have synthesized
17400-416: The other. The evolutionary divergence of the two populations was estimated to have occurred about 72,000–38,000 years ago, well before domestication, most likely due to climate, topography, or other environmental changes. According to a 2009 study, the earliest known domestic horses were found at settlements of the Botai culture, from about 5500 years ago. These horses were raised for meat and milk. In 2018,
17550-452: The outgroup and the rest of the taxa in the tree, or by introducing additional assumptions about the relative rates of evolution on each branch, such as an application of the molecular clock hypothesis . Both rooted and unrooted trees can be either bifurcating or multifurcating. A rooted bifurcating tree has exactly two descendants arising from each interior node (that is, it forms a binary tree ), and an unrooted bifurcating tree takes
17700-448: The phonology, most of what is said about morphology and syntax also holds true for Chakhar, while Khorchin is somewhat more diverse. Modern Mongolian is an agglutinative —almost exclusively suffixing—language, with the only exception being reduplication. Mongolian also does not have gendered nouns, or definite articles like "the". Most of the suffixes consist of a single morpheme . There are many derivational morphemes. For example,
17850-542: The preferential policies for minorities in education, healthcare, family planning, school admissions, the hiring and promotion, the financing and taxation of businesses, and regional infrastructural support given to ethnic minorities in China. In 2020, the Chinese government required three subjects—language and literature, politics, and history—to be taught in Mandarin in Mongolian-language primary and secondary schools in
18000-597: The presence of urban ethnic communities. The multilingual situation in Inner Mongolia does not appear to obstruct efforts by ethnic Mongols to preserve their language. Although an unknown number of Mongols in China, such as the Tumets, may have completely or partially lost the ability to speak their language, they are still registered as ethnic Mongols and continue to identify themselves as ethnic Mongols. The children of inter-ethnic Mongol-Chinese marriages also claim to be and are registered as ethnic Mongols so they can benefit from
18150-405: The range of DNA considered useful. Phylogenetic trees can also be inferred from a range of other data types, including morphology, the presence or absence of particular types of genes, insertion and deletion events – and any other observation thought to contain an evolutionary signal. Phylogenetic networks are used when bifurcating trees are not suitable, due to these complications which suggest
18300-472: The risk of forest fires . Future introductions are planned. In June 2024 six mares and a stallion were reintroduced to Kazakhstan from zoos in Europe, ten years after plans were announced to do so.The operation was organised by Prague Zoo , which selected horses from various programs in Europe, which were housed at Tierpark Berlin for some months before being transported to Kazakhstan in Czech army planes. In
18450-417: The simpler algorithms (i.e. those based on distance) of tree construction. Maximum parsimony is another simple method of estimating phylogenetic trees, but implies an implicit model of evolution (i.e. parsimony). More advanced methods use the optimality criterion of maximum likelihood , often within a Bayesian framework , and apply an explicit model of evolution to phylogenetic tree estimation. Identifying
18600-464: The social and communication behaviors of wild Przewalski's horses. On reaching maturity at three to four years of age, Kurt is intended to become the breeder stallion for the San Diego Zoo herd to pass Kuporovic's genes into the larger captive Przewalski's horse population and thereby increase the genetic variation of the species. In 2023, a genetic twin of Kurt, named Ollie, was born from cloning with
18750-596: The splitting of the Middle Mongol affricates * ʧ ( ᠴ č ) and * ʤ ( ᠵ ǰ ) into ʦ ( ц c ) and ʣ ( з z ) versus ʧ ( ч č ) and ʤ ( ж ž ) in Mongolia: Aside from these differences in pronunciation, there are also differences in vocabulary and language use: in the state of Mongolia more loanwords from Russian are being used, while in Inner Mongolia more loanwords from Chinese have been adopted. The following description
18900-530: The springtime, they favor Elymus repens , Corynephorus canescens , Festuca valesiaca , and Chenopodium album . In early summer, they favor Dactylis glomerata and Trifolium , and in late summer, they gravitate towards E. repens and Vicia cracca . In winter the horses eat Salix spp., Pyrus communis , Malus sylvatica , Pinus sylvestris , Rosa spp., and Alnus spp. Additionally, Przewalski's horses may dig for Festuca spp., Bromus inermis , and E. repens that grow beneath
19050-421: The stem with certain case endings (e.g. цэрэг ( tsereg ) → цэргийн ( tsergiin )). The additional morphological rules specific to loanwords are not covered. Phylogenetic tree A phylogenetic tree , phylogeny or evolutionary tree is a graphical representation which shows the evolutionary history between a set of species or taxa during a specific time. In other words, it
19200-430: The suffix ‑ н (‑ n ) when the stem has an unstable nasal. Nouns can also take a reflexive-possessive suffix , indicating that the marked noun is possessed by the subject of the sentence: bi najz-aa avar-san I friend- reflexive-possessive save- perfect "I saved my friend". However, there are also somewhat noun-like adjectives to which case suffixes seemingly cannot be attached directly unless there
19350-441: The tail is about 90 cm (35 in) long, with a longer dock and shorter hair than seen in domesticated horses. The hooves of Przewalski's horse are longer in the front and have significantly thicker sole horns than feral horses, an adaptation that improves hoof performance on terrain. The karyotype of Przewalski's horse differs from that of the domestic horse, having 33 chromosome pairs versus 32, apparently due to
19500-410: The total number of incoming and outgoing edges). The most common method for rooting trees is the use of an uncontroversial outgroup —close enough to allow inference from trait data or molecular sequencing, but far enough to be a clear outgroup. Another method is midpoint rooting, or a tree can also be rooted by using a non-stationary substitution model . Unrooted trees illustrate the relatedness of
19650-456: The transition, a major shift in the vowel-harmony paradigm occurred, long vowels developed, the case system changed slightly, and the verbal system was restructured. Mongolian is related to the extinct Khitan language . It was believed that Mongolian was related to Turkic , Tungusic , Korean and Japonic languages but this view is now seen as obsolete by a majority of (but not all) comparative linguists. These languages have been grouped under
19800-411: The tree before hybridisation takes place, and conserved sequences . Also, there are problems in basing an analysis on a single type of character, such as a single gene or protein or only on morphological analysis, because such trees constructed from another unrelated data source often differ from the first, and therefore great care is needed in inferring phylogenetic relationships among species. This
19950-529: The trees that they generate are not necessarily correct – they do not necessarily accurately represent the evolutionary history of the included taxa. As with any scientific result, they are subject to falsification by further study (e.g., gathering of additional data, analyzing the existing data with improved methods). The data on which they are based may be noisy ; the analysis can be confounded by genetic recombination , horizontal gene transfer , hybridisation between species that were not nearest neighbors on
20100-404: The variation of abundance of various taxa through time. A spindle diagram is not an evolutionary tree: the taxonomic spindles obscure the actual relationships of the parent taxon to the daughter taxon and have the disadvantage of involving the paraphyly of the parental group. This type of diagram is no longer used in the form originally proposed. Darwin also mentioned that the coral may be
20250-424: The voiceless velar plosive [k] ; instead, it has a voiced alveolar lateral fricative , /ɮ/ , which is often realized as voiceless [ɬ] . In word-final position, /n/ (if not followed by a vowel in historical forms) is realized as [ŋ] . Aspirated consonants are preaspirated in medial and word-final contexts, devoicing preceding consonants and vowels. Devoiced short vowels are often deleted. The maximal syllable
20400-462: The wild horse in its native habitat was of a single stallion in 1969. Expeditions after this failed to locate any horses, and the species would be designated "extinct in the wild" for over 30 years. Competition with livestock, hunting, capture of foals for zoological collections, military activities, and harsh winters recorded in 1945, 1948, and 1956 are considered to be main causes of the decline in Przewalski's horse population. The wild population
20550-593: The wild in Mongolia, followed by additional animals later. One of the areas to which they were reintroduced became Khustain Nuruu National Park in 1998. Another reintroduction site is Great Gobi B Strictly Protected Area , located at the fringes of the Gobi Desert . In 2001, Przewalski's horses were reintroduced into the Kalamaili Nature Reserve in Xinjiang , China. Since 2004, there has been
20700-461: The wild, the remnant population was limited to the small region between the Takhiin Shar Nuruu and Bajtag-Bogdo mountain ridges. Attempts to obtain specimens for exhibit and captive breeding were largely unsuccessful until 1902, when 28 captured foals were brought to Europe. These and a small number of additional captives would be distributed among zoos and breeding centers in Europe and
20850-452: The winter of 2009–2010, one of the worst dzud or snowy winter conditions ever hit Mongolia. The population of Przewalski's horse in the Great Gobi B SPA was drastically affected, providing clear evidence of the risks associated with reintroducing small and sequestered species in unpredictable and unfamiliar environments. After reintroduced horses had successfully reproduced, the status of
21000-547: The word baiguullagiinh consists of the root bai 'to be', an epenthetic ‑ g ‑, the causative ‑ uul ‑ (hence 'to found'), the derivative suffix ‑ laga that forms nouns created by the action (like - ation in organisation ) and the complex suffix ‑ iinh denoting something that belongs to the modified word (‑ iin would be genitive ). Nominal compounds are quite frequent. Some derivational verbal suffixes are rather productive , e.g. yarih 'to speak', yarilc 'to speak with each other'. Formally,
21150-472: The year. Przewalski's horses seasonally display a set of changes collectively characteristic of physiologic adaptation to starvation, with their basal metabolic rate in winter being half what it is during springtime. This is not a direct consequence of decreased nutrient intake, but rather a programmed response to predictable seasonal dietary fluctuation. Mating occurs in late spring or early summer. Mating stallions do not start looking for mating partners until
21300-571: The zone after the Chernobyl accident , so now it serves as a deserted de facto nature reserve. Though poaching has taken a toll on numbers, as of 2019 the estimated population in the Chernobyl zone was over 100 individuals. Le Villaret, located in the Cevennes National Park in southern France and run by the Association Takh, is a breeding site for Przewalski's horses that was created to allow
21450-505: Was a Przewalski horse stallion named Kuporovic, born in the UK in 1975 and relocated three years later to the US, where he died in 1998. Due to concerns over the loss of genetic variation in the captive Przewalski's horse population, and in anticipation of the development of new cloning techniques, tissue from the stallion was cryopreserved at the San Diego Zoo's Frozen Zoo . Breeding of this individual in
21600-428: Was already rare at its first scientific characterization. Przewalski reported seeing them only from a distance and may have instead sighted herds of local onager Mongolian wild asses. He was only able to obtain specimens of the type from Kirghiz hunters. The range of Przewalski's horse was limited to the arid Dzungarian Basin in the Gobi Desert . It has been suggested that this was not their natural habitat, but, like
21750-636: Was also introduced to the Hortobágy National Park puszta in Hungary; data on social structure, behavior, and diseases gathered from these animals are used to improve the Mongolian conservation effort. An additional breeding population of Przewalski's horses roams the former Döberitzer Heide military proving ground , now a nature reserve in Dallgow-Döberitz , Germany. Established in 2008, this population comprised 24 horses in 2019. Another population
21900-464: Was carried out by somatic cell nuclear transfer (SCNT), whereby a viable embryo is created by transplanting the DNA -containing nucleus of a somatic cell into an immature egg cell ( oocyte ) that has had its nucleus removed, producing offspring genetically identical to the somatic cell donor. Since the oocyte used was from a domestic horse, this was an example of interspecies SCNT. The somatic cell donor
22050-417: Was created, giving distinctive evidence on early classical Mongolian phonological peculiarities. Mongolian is the official national language of Mongolia, where it is spoken (but not always written) by nearly 3.6 million people (2014 estimate), and the official provincial language (both spoken and written forms) of Inner Mongolia, where there are at least 4.1 million ethnic Mongols. Across the whole of China,
22200-426: Was formally described as a novel species in 1881 by Ivan Semyonovich Polyakov . The taxonomic position of Przewalski's horse remains controversial, and no consensus exists about whether it is a full species (as Equus przewalskii ); a subspecies of Equus ferus the wild horse (as Equus ferus przewalskii in trinomial nomenclature , along with two other subspecies, the domestic horse E. f. caballus , and
22350-540: Was one of the most genetically valuable Przewalski's horses in the North American breeding program. The first birth by artificial insemination occurred on 27 July 2013 at the Smithsonian Conservation Biology Institute . In 2020, the first cloned Przewalski's horse was born, the result of a collaboration between San Diego Zoo Global , ViaGen Equine and Revive & Restore . The cloning
22500-462: Was rejected by Alan Outram and colleagues in a paper which was not dated or peer-reviewed. Taylor reiterated his arguments that Przewalski's horse had never been domesticated in an article in Scientific American in 2024. In any case, the Botai horses were found to have negligible genetic contribution to any of the ancient or modern domestic horses studied, indicating that the domestication of
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