168-465: Samuel Goodere (1687–1741) was a captain in the British Royal Navy , who was the third son of Sir Edward Goodere, 1st Baronet . He was convicted and hanged at Bristol for the murder of his elder brother, Sir John Dineley Goodere, 2nd Baronet , in 1741. Samuel Goodere's mother -- Eleanor Goodere (née Dineley) -- was the daughter and heiress of Sir Edward Dineley of Charlton, Worcestershire , and
336-412: A basal lamina , the upper layer of the basement membrane , the lower layer is the reticular lamina lying next to the connective tissue in the extracellular matrix secreted by the epithelial cells. There are many different types of epithelium, modified to suit a particular function. In the respiratory tract there is a type of ciliated epithelial lining; in the small intestine there are microvilli on
504-450: A copulatory organ present in most species. The eggs are surrounded by amniotic membranes which prevents them from drying out and are laid on land, or develop internally in some species. The bladder is small as nitrogenous waste is excreted as uric acid . Turtles are notable for their protective shells. They have an inflexible trunk encased in a horny carapace above and a plastron below. These are formed from bony plates embedded in
672-481: A microscope . Human anatomy, physiology and biochemistry are complementary basic medical sciences, which are generally taught to medical students in their first year at medical school. Human anatomy can be taught regionally or systemically; that is, respectively, studying anatomy by bodily regions such as the head and chest, or studying by specific systems, such as the nervous or respiratory systems. The major anatomy textbook, Gray's Anatomy , has been reorganized from
840-470: A sessile lifestyle). Most animals have bodies differentiated into separate tissues and these animals are also known as eumetazoans . They have an internal digestive chamber, with one or two openings; the gametes are produced in multicellular sex organs, and the zygotes include a blastula stage in their embryonic development . Metazoans do not include the sponges , which have undifferentiated cells. Unlike plant cells , animal cells have neither
1008-402: A "treatise on the heart", with vessels carrying all the body's fluids to or from every member of the body. Ancient Greek anatomy and physiology underwent great changes and advances throughout the early medieval world. Over time, this medical practice expanded due to a continually developing understanding of the functions of organs and structures in the body. Phenomenal anatomical observations of
1176-460: A Royal Navy captain, has been responsible for the former duty whilst Commander Task Force Iraqi Maritime, a Royal Navy commodore, was responsible for the latter. The mission ended in May 2011. The financial costs attached to nuclear deterrence, including Trident missile upgrades and replacements, have become an increasingly significant issue for the navy. HMS Raleigh at Torpoint, Cornwall ,
1344-528: A cell wall nor chloroplasts . Vacuoles, when present, are more in number and much smaller than those in the plant cell. The body tissues are composed of numerous types of cells, including those found in muscles, nerves and skin. Each typically has a cell membrane formed of phospholipids , cytoplasm and a nucleus . All of the different cells of an animal are derived from the embryonic germ layers . Those simpler invertebrates which are formed from two germ layers of ectoderm and endoderm are called diploblastic and
1512-438: A charge of misconduct against Goodere, alleging that an attack on San Sebastián on 23 June had failed as a consequence of Goodere's actions. Goodere was tried by court-martial on 24 December 1719, was found guilty of "having been very much wanting in the performance of his duty", and was dismissed from his ship, which, in time of peace, was equivalent to being dismissed from the service. In the opinion of John Knox Laughton , it
1680-433: A class of arachnids have four pairs of legs; a body of two segments—a cephalothorax and an abdomen . Spiders have no wings and no antennae. They have mouthparts called chelicerae which are often connected to venom glands as most spiders are venomous. They have a second pair of appendages called pedipalps attached to the cephalothorax. These have similar segmentation to the legs and function as taste and smell organs. At
1848-552: A coat of glycoproteins . In more advanced animals, many glands are formed of epithelial cells. Muscle cells (myocytes) form the active contractile tissue of the body. Muscle tissue functions to produce force and cause motion, either locomotion or movement within internal organs. Muscle is formed of contractile filaments and is separated into three main types; smooth muscle , skeletal muscle and cardiac muscle . Smooth muscle has no striations when examined microscopically. It contracts slowly but maintains contractibility over
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#17328910007032016-593: A comprehensive transformation, brought about by steam propulsion , metal ship construction, and explosive munitions. Despite having to completely replace its war fleet, the Navy managed to maintain its overwhelming advantage over all potential rivals. Owing to British leadership in the Industrial Revolution , the country enjoyed unparalleled shipbuilding capacity and financial resources, which ensured that no rival could take advantage of these revolutionary changes to negate
2184-456: A daughter Eleanor Goodere born at Shoulden, Kent (who married William Wyborn). He later married a widow, Elizabeth Watts of Monmouthshire in or before 1729, and by her left three daughters and twin sons. Of the daughters two died unmarried; the third, Anne, married John Willyams, a commander in the navy, and was the mother of the Rev. Cooper Willyams . The eldest of the twin sons, Edward, who inherited
2352-457: A few species have no limbs and resemble snakes. Lizards have moveable eyelids, eardrums are present and some species have a central parietal eye. Snakes are closely related to lizards, having branched off from a common ancestral lineage during the Cretaceous period, and they share many of the same features. The skeleton consists of a skull, a hyoid bone, spine and ribs though a few species retain
2520-472: A fleet of technologically sophisticated ships, submarines, and aircraft, including 2 aircraft carriers , 2 amphibious transport docks , 4 ballistic missile submarines (which maintain the nuclear deterrent ), 6 nuclear fleet submarines , 6 guided missile destroyers , 9 frigates , 7 mine-countermeasure vessels and 26 patrol vessels . As of May 2024, there are 66 commissioned ships (including submarines as well as one historic ship, HMS Victory ) in
2688-800: A force multiplier for the Royal Navy, often doing patrols that frigates used to do. The Royal Navy is part of His Majesty's Naval Service, which also includes the Royal Marines and the Royal Fleet Auxiliary. The professional head of the Naval Service is the First Sea Lord who is an admiral and member of the Defence Council of the United Kingdom . The Defence Council delegates management of
2856-471: A gradual decline in its fleet size in accordance with the changed strategic environment it operated in. While new and more capable ships are continually brought into service, such as the Queen Elizabeth -class aircraft carriers , Astute -class submarines , and Type 45 destroyers , the total number of ships and submarines operated has continued to steadily reduce. This has caused considerable debate about
3024-469: A heavy reliance upon impressed labour. This would result in problems countering large, well-armed United States Navy frigates which outgunned Royal Naval vessels in single-opponent actions, as well as United States privateers, when the American War of 1812 broke out concurrent with the war against Napoleonic France and its allies. The Royal Navy still enjoyed a numerical advantage over the former colonists on
3192-513: A high metabolic rate , a light skeletal system and powerful muscles . The long bones are thin, hollow and very light. Air sac extensions from the lungs occupy the centre of some bones. The sternum is wide and usually has a keel and the caudal vertebrae are fused. There are no teeth and the narrow jaws are adapted into a horn-covered beak. The eyes are relatively large, particularly in nocturnal species such as owls. They face forwards in predators and sideways in ducks. The feathers are outgrowths of
3360-606: A lieutenant through the War of the Spanish Succession . On 12 January 1719 was appointed first lieutenant of HMS Preston with Captain Robert Johnson. Both Goodere and Johnson transferred to HMS Weymouth on 28 February 1719, and saw service in operations on the north coast of Spain in mid-1719. Later that year, on 6 November 1719, again with Johnson, he transferred to HMS Deptford . A few weeks later, however, Johnson preferred
3528-536: A male heir "of their bodies," then Samuel and his first born son would be next in line to inherit the bulk of the Dineley Estate. By the time of the death of John's son and then John himself his brother George should have inherited but he had either died or failed to have a son, therefore Samuel's son Edward (a lunatic) should have inherited the fortune, with Samuel's second born son John (Edward's twin) being next in line. Sir Edward Dineley's grandson John Goodere broke
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#17328910007033696-746: A mammal. Humans have a head, neck, trunk (which includes the thorax and abdomen ), two arms and hands, and two legs and feet. Generally, students of certain biological sciences , paramedics , prosthetists and orthotists, physiotherapists , occupational therapists , nurses , podiatrists , and medical students learn gross anatomy and microscopic anatomy from anatomical models, skeletons, textbooks, diagrams, photographs, lectures and tutorials and in addition, medical students generally also learn gross anatomy through practical experience of dissection and inspection of cadavers . The study of microscopic anatomy (or histology ) can be aided by practical experience examining histological preparations (or slides) under
3864-774: A national institution and not a possession of the Crown as it had been before. Following the Glorious Revolution of 1688, England joined the War of the Grand Alliance which marked the end of France 's brief pre-eminence at sea and the beginning of an enduring British supremacy which would help with the creation of the British Empire. In 1707, the Scottish navy was united with the English Royal Navy. On Scottish men-of-war,
4032-732: A navy, this meant the French landed at Sandwich unopposed in April 1216. John's flight to Winchester and his death later that year left the Earl of Pembroke as regent, and he was able to marshal ships to fight the French in the Battle of Sandwich in 1217 – one of the first major English battles at sea. The outbreak of the Hundred Years War emphasised the need for an English fleet. French plans for an invasion of England failed when Edward III of England destroyed
4200-425: A note to Smith, saying that, having heard his brother was there, he would be glad to meet him if Smith would allow him to come in. Accordingly, in the evening he went to Smith's house, and the two brothers smoked and drank together, (John declined alcohol and would take only water) and to all appearance made up their quarrel. But, as John was walking towards his lodgings, he was seized by Samuel's orders, carried down to
4368-538: A peak of efficiency, dominating the navies of all Britain's adversaries, which spent most of the war blockaded in port. Under Lord Nelson , the navy defeated the combined Franco-Spanish fleet at Trafalgar (1805). Ships of the line and even frigates, as well as manpower, were prioritised for the naval war in Europe, however, leaving only smaller vessels on the North America Station and other less active stations, and
4536-420: A pump action in which air is first drawn into the buccopharyngeal region through the nostrils. These are then closed and the air is forced into the lungs by contraction of the throat. They supplement this with gas exchange through the skin which needs to be kept moist. In frogs the pelvic girdle is robust and the hind legs are much longer and stronger than the forelimbs. The feet have four or five digits and
4704-424: A range of vessels, including so-called "motherships" planned for procurement by the Royal Fleet Auxiliary. The Hunt-class vessels combine the separate roles of the traditional minesweeper and the active minehunter in one hull. If required, the vessels can take on the role of offshore patrol vessels. A fleet of eight River-class offshore patrol vessels are in service with the Royal Navy. The three Batch 1 ships of
4872-421: A rope while Samuel stood sentry at the door with a drawn sword to prevent any interference. A man and his wife in the adjoining cabin had heard and witnessed through gaps in the ship's timbers part of what had occurred. Goodere had apparently intended to put to sea at once, but Smith became suspicious and having had information the previous night that a gentleman resembling his guest had been taken prisoner on board
5040-465: A single force. During the early 17th century, England's relative naval power deteriorated until Charles I undertook a major programme of shipbuilding. His methods of financing the fleet contributed to the outbreak of the English Civil War , and the abolition of the monarchy . The Commonwealth of England replaced many names and symbols in the new Commonwealth Navy, associated with royalty and
5208-543: A skeleton to support or protect the body. An exoskeleton is a thickened, rigid cuticle which is stiffened by mineralization , as in crustaceans or by the cross-linking of its proteins as in insects . An endoskeleton is internal and present in all developed animals, as well as in many of those less developed. Epithelial tissue is composed of closely packed cells, bound to each other by cell adhesion molecules , with little intercellular space. Epithelial cells can be squamous (flat), cuboidal or columnar and rest on
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5376-448: A small permanent core of warships in peacetime. England's naval organisation was haphazard and the mobilisation of fleets when war broke out was slow. Control of the sea only became critical to Anglo-Saxon kings in the 10th century. In the 11th century, Aethelred II had a large fleet built by a national levy. During the period of Danish rule in the 11th century, authorities maintained a standing fleet by taxation, and this continued for
5544-497: A systems format to a regional format, in line with modern teaching methods. A thorough working knowledge of anatomy is required by physicians, especially surgeons and doctors working in some diagnostic specialties, such as histopathology and radiology . Academic anatomists are usually employed by universities, medical schools or teaching hospitals. They are often involved in teaching anatomy, and research into certain systems, organs, tissues or cells. Invertebrates constitute
5712-785: A time under Edward the Confessor , who frequently commanded fleets in person. After the Norman Conquest , English naval power waned and England suffered large naval raids from the Vikings. In 1069, this allowed for the invasion and ravaging of England by Jarl Osborn, brother of King Svein Estridsson , and his sons. The lack of an organised navy came to a head during the First Barons' War , in which Prince Louis of France invaded England in support of northern barons. With King John unable to organise
5880-431: A vast array of living organisms ranging from the simplest unicellular eukaryotes such as Paramecium to such complex multicellular animals as the octopus , lobster and dragonfly . They constitute about 95% of the animal species. By definition, none of these creatures has a backbone. The cells of single-cell protozoans have the same basic structure as those of multicellular animals but some parts are specialized into
6048-479: A vertebrate is the vertebral column , formed in the development of the segmented series of vertebrae . In most vertebrates the notochord becomes the nucleus pulposus of the intervertebral discs . However, a few vertebrates, such as the sturgeon and the coelacanth , retain the notochord into adulthood. Jawed vertebrates are typified by paired appendages, fins or legs, which may be secondarily lost. The limbs of vertebrates are considered to be homologous because
6216-693: A vestige of the pelvis and rear limbs in the form of pelvic spurs . The bar under the second fenestra has also been lost and the jaws have extreme flexibility allowing the snake to swallow its prey whole. Snakes lack moveable eyelids, the eyes being covered by transparent "spectacle" scales. They do not have eardrums but can detect ground vibrations through the bones of their skull. Their forked tongues are used as organs of taste and smell and some species have sensory pits on their heads enabling them to locate warm-blooded prey. Crocodilians are large, low-slung aquatic reptiles with long snouts and large numbers of teeth. The head and trunk are dorso-ventrally flattened and
6384-594: A wide range of stretch lengths. It is found in such organs as sea anemone tentacles and the body wall of sea cucumbers . Skeletal muscle contracts rapidly but has a limited range of extension. It is found in the movement of appendages and jaws. Obliquely striated muscle is intermediate between the other two. The filaments are staggered and this is the type of muscle found in earthworms that can extend slowly or make rapid contractions. In higher animals striated muscles occur in bundles attached to bone to provide movement and are often arranged in antagonistic sets. Smooth muscle
6552-480: Is constantly evolving as discoveries are made. In recent years, there has been a significant increase in the use of advanced imaging techniques, such as MRI and CT scans , which allow for more detailed and accurate visualizations of the body's structures. The discipline of anatomy is divided into macroscopic and microscopic parts. Macroscopic anatomy , or gross anatomy, is the examination of an animal's body parts using unaided eyesight. Gross anatomy also includes
6720-577: Is definitely wrong since his uncle was certainly the second baronet. In the opinion of John Knox Laughton , it is probable "that the baronetcy became extinct in 1741, on the sentence of Samuel Goodere, though the twins may have been allowed the title by courtesy". [REDACTED] This article incorporates text from a publication now in the public domain : Stephen, Leslie ; Lee, Sidney , eds. (1890). " Goodere, Samuel ". Dictionary of National Biography . Vol. 22. London: Smith, Elder & Co. Royal Navy The Royal Navy ( RN )
6888-529: Is doubtful whether Goodere served again at sea until November 1733, when he was posted to HMS Antelope for a brief period of two weeks "apparently on some electioneering job" rather than as an active officer. He may have been living with his father, now nearing 80, who was on bad terms with his eldest surviving son, John. John was described as "rough, uncouth, and of no education". Samuel was also on bad terms with his brother; and these became worse when John, who had divorced his wife for adultery, found that she too
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7056-473: Is found in the walls of the uterus , bladder , intestines , stomach , oesophagus , respiratory airways , and blood vessels . Cardiac muscle is found only in the heart , allowing it to contract and pump blood round the body. Nervous tissue is composed of many nerve cells known as neurons which transmit information. In some slow-moving radially symmetrical marine animals such as ctenophores and cnidarians (including sea anemones and jellyfish ),
7224-422: Is generated, both over immediate and long-term timescales. Anatomy and physiology , which study the structure and function of organisms and their parts respectively, make a natural pair of related disciplines, and are often studied together. Human anatomy is one of the essential basic sciences that are applied in medicine, and is often studied alongside physiology . Anatomy is a complex and dynamic field that
7392-564: Is one of the largest ships in the Navy. As of 2018, the newly commissioned HMS Magpie also undertakes survey duties at sea. The Royal Fleet Auxiliary plans to introduce two new Multi-Role Ocean Surveillance Ships , in part to protect undersea cables and gas pipelines and partly to compensate for the withdrawal of all ocean-going survey vessels from Royal Navy service. The first of these vessels, RFA Proteus , entered service in October 2023. The Royal Fleet Auxiliary (RFA) provides support to
7560-464: Is opened and its organs studied, and endoscopy , in which a video camera -equipped instrument is inserted through a small incision in the body wall and used to explore the internal organs and other structures. Angiography using X-rays or magnetic resonance angiography are methods to visualize blood vessels. The term "anatomy" is commonly taken to refer to human anatomy . However, substantially similar structures and tissues are found throughout
7728-562: Is primarily designed for anti-aircraft and anti-missile warfare and the Royal Navy describe the destroyer's mission as "to shield the Fleet from air attack". They are equipped with the PAAMS (also known as Sea Viper) integrated anti-aircraft warfare system which incorporates the sophisticated SAMPSON and S1850M long range radars and the Aster 15 and 30 missiles. Sixteen Type 23 frigates were delivered to
7896-489: Is startlingly rapid". Between 1275 and 1326, the anatomists Mondino de Luzzi , Alessandro Achillini and Antonio Benivieni at Bologna carried out the first systematic human dissections since ancient times. Mondino's Anatomy of 1316 was the first textbook in the medieval rediscovery of human anatomy. It describes the body in the order followed in Mondino's dissections, starting with the abdomen, thorax, head, and limbs. It
8064-579: Is the naval warfare force of the United Kingdom , British Overseas Territories and Crown Dependencies , and a component of His Majesty's Naval Service . Although warships were used by English and Scottish kings from the early medieval period, the first major maritime engagements were fought in the Hundred Years' War against France . The modern Royal Navy traces its origins to the English Navy of
8232-752: Is the submarine based element of the Royal Navy. It is sometimes referred to as the " Silent Service ", as the submarines are generally required to operate undetected. Founded in 1901, the service made history in 1982 when, during the Falklands War, HMS Conqueror became the first nuclear-powered submarine to sink a surface ship, ARA General Belgrano . Today, all of the Royal Navy's submarines are nuclear-powered . The Royal Navy operates four Vanguard -class ballistic missile submarines displacing nearly 16,000 tonnes and equipped with Trident II missiles (armed with nuclear weapons ) and heavyweight Spearfish torpedoes , to carry out Operation Relentless,
8400-752: Is the basic training facility for newly enlisted ratings. Britannia Royal Naval College in Dartmouth, Devon is the initial officer training establishment for the Royal Navy. Personnel are divided into a warfare branch , which includes Warfare Officers (previously named seamen officers) and Naval Aviators , as well other branches including the Royal Naval Engineers , Royal Navy Medical Branch , and Logistics Officers (previously named Supply Officers ). Present-day officers and ratings have several different uniforms ; some are designed to be worn aboard ship, others ashore or in ceremonial duties. Women began to join
8568-403: Is the study of the structures that make up a discrete body system—that is, a group of structures that work together to perform a unique body function, such as the digestive system. Anatomy can be studied using both invasive and non-invasive methods with the goal of obtaining information about the structure and organization of organs and systems. Methods used include dissection , in which a body
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#17328910007038736-468: Is the traditional workhorse of the Navy. As of May 2024 there are six Type 45 destroyers and 9 Type 23 frigates in commission. Among their primary roles is to provide escort for the larger capital ships—protecting them from air, surface and subsurface threats. Other duties include undertaking the Royal Navy's standing deployments across the globe, which often consists of: counter-narcotics, anti-piracy missions and providing humanitarian aid. The Type 45
8904-411: Is the vertebral column, composed of articulating vertebrae which are lightweight yet strong. The ribs attach to the spine and there are no limbs or limb girdles. The main external features of the fish, the fins , are composed of either bony or soft spines called rays, which with the exception of the caudal fins , have no direct connection with the spine. They are supported by the muscles which compose
9072-463: Is to conduct amphibious warfare, they have also been deployed for humanitarian aid missions. Both vessels were in reserve as of 2024 and in November 2024, the newly elected Labour government indicated that they would in fact be retired from service completely by March 2025. While second-line amphibious capabilities remained within the Royal Fleet Auxiliary, the future of the navy's amphibious capability
9240-590: The Hippocratic Corpus , an Ancient Greek medical work written by unknown authors. Aristotle described vertebrate anatomy based on animal dissection . Praxagoras identified the difference between arteries and veins . Also in the 4th century BCE, Herophilos and Erasistratus produced more accurate anatomical descriptions based on vivisection of criminals in Alexandria during the Ptolemaic period . In
9408-565: The American War of Independence . The United States was allied to France , and the Netherlands and Spain were also at war with Britain. In the Battle of the Chesapeake , the British fleet failed to lift the French blockade, resulting in the surrender of an entire British army at Yorktown . The French Revolutionary and Napoleonic Wars (1793–1801, 1803–1814 & 1815) saw the Royal Navy reach
9576-628: The Battle of Britain in order to neutralise the Home Fleet , but faced stiff resistance from the Royal Air Force . The Luftwaffe bombing offensive during the Kanalkampf phase of the battle targeted naval convoys and bases in order to lure large concentrations of RAF fighters into attrition warfare . At Taranto , Admiral Cunningham commanded a fleet that launched the first all-aircraft naval attack in history. The Royal Navy suffered heavy losses in
9744-574: The College of Arms , wrote that Goodere's sons Edward and John succeeded their murdered uncle rather than their father. But Burke thought that the baronetcy descended to Samuel and to his sons after him. In his Baronetage of 1741, Arthur Collins wrote that the baronetcy was extinct, while Thomas Wotton ( Baronetage , 1771) thought the baronetcy was " attainted ". Treadway Russell Nash in his History of Worcestershire (volume i, page 972) says that Sir Edward Dineley-Goodere succeeded his grandfather, which
9912-711: The Invergordon Mutiny took place in the Atlantic Fleet over the National Government 's proposed 25% pay cut, which was eventually reduced to 10%. International tensions increased in the mid-1930s and the re-armament of the Royal Navy was well under way by 1938. In addition to new construction, several existing old battleships , battlecruisers and heavy cruisers were reconstructed, and anti-aircraft weaponry reinforced, while new technologies, such as ASDIC , Huff-Duff and hydrophones , were developed. At
10080-682: The Kingdom of England had possessed less-organised naval forces for centuries prior to this. The Royal Scots Navy (or Old Scots Navy) had its origins in the Middle Ages until its merger with the English Royal Navy per the Acts of Union 1707 . During much of the medieval period, fleets or "king's ships" were often established or gathered for specific campaigns or actions, and these would disperse afterwards. These were generally merchant ships enlisted into service. Unlike some European states, England did not maintain
10248-575: The Middle East , especially Iraq . The Royal Navy played an historic role in several great global explorations of science and discovery. Beginning in the 18th century many great voyages were commissioned often in co-operation with the Royal Society , such as the Northwest Passage expedition of 1741 . James Cook led three great voyages, with goals such as discovering Terra Australis , observing
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#173289100070310416-457: The Ministry of Defence published figures showing that the Royal Navy and Royal Marines had 29,090 full-time trained personnel compared with a target of 30,600. In 2023, it was reported that the Royal Navy was experiencing significant recruiting challenges with a net drop of some 1,600 personnel (4 percent of the force) from mid-2022 to mid-2023. This was posing a significant problem in the ability of
10584-727: The North America and West Indies Station . After the First World War, this formation assumed responsibility for the eastern Pacific Ocean and the western South Atlantic and was known as the America and West Indies Station until 1956. In 1921, due to the ambitions of Imperial Japan and the threat of the Imperial Japanese Navy , it was decided to construct the Singapore Naval Base . During this period, naval warfare underwent
10752-487: The Ruby , applied to the mayor for an investigation. Goodere and his accomplices were apprehended. All four were tried on 26 March, found guilty, and sentenced to death. They were hanged on 15 April 1741. His body was sent for anatomical dissection , but was afterwards interred with his ancestors at Hereford . Papers relating to the murder and trial are held at Bristol Archives (Ref. 14754) ( online catalogue ). Samuel had
10920-555: The Transit of Venus and searching for the elusive North-West Passage , these voyages are considered to have contributed to world knowledge and science. In the late 18th century, during a four year voyage Captain George Vancouver made detailed maps of the western coastline of North America . In the 19th century, Charles Darwin made further contributions to science during the second voyage of HMS Beagle . The Ross expedition to
11088-506: The Triassic period. There is one living species, Sphenodon punctatus . The skull has two openings (fenestrae) on either side and the jaw is rigidly attached to the skull. There is one row of teeth in the lower jaw and this fits between the two rows in the upper jaw when the animal chews. The teeth are merely projections of bony material from the jaw and eventually wear down. The brain and heart are more primitive than those of other reptiles, and
11256-692: The Type 26 Frigate , with the incremental retirement of the remaining Type 23s commencing in 2021. The Strategic Defence and Security Review 2015 reduced the procurement of Type 26 to eight with five Type 31e frigates also to be procured. There are two classes of MCMVs in the Royal Navy: one Sandown -class minehunter and six Hunt-class mine countermeasures vessels . All the Sandown-class vessels are to be withdrawn from service by 2025 and are being replaced by autonomous systems that are planned to operate from
11424-589: The War of the Spanish Succession (1702–1714) granted Britain Gibraltar and Menorca , providing the Navy with Mediterranean bases. The expansion of the Royal Navy would encourage the British colonisation of the Americas , with British (North) America becoming a vital source of timber for the Royal Navy. There was a defeat during the frustrated siege of Cartagena de Indias in 1741. A new French attempt to invade Britain
11592-400: The epidermis and are found in localized bands from where they fan out over the skin. Large flight feathers are found on the wings and tail, contour feathers cover the bird's surface and fine down occurs on young birds and under the contour feathers of water birds. The only cutaneous gland is the single uropygial gland near the base of the tail. This produces an oily secretion that waterproofs
11760-529: The extracellular matrix . Often called fascia (from the Latin "fascia," meaning "band" or "bandage"), connective tissues give shape to organs and holds them in place. The main types are loose connective tissue, adipose tissue , fibrous connective tissue, cartilage and bone. The extracellular matrix contains proteins , the chief and most abundant of which is collagen . Collagen plays a major part in organizing and maintaining tissues. The matrix can be modified to form
11928-534: The high church , and expanded it to become the most powerful in the world. The fleet was quickly tested in the First Anglo-Dutch War (1652–1654) and the Anglo-Spanish War (1654–1660) , which saw the British conquest of Jamaica and successful attacks on Spanish treasure fleets . The 1660 Restoration saw Charles II rename the Royal Navy again, and started use of the prefix HMS . The Navy remained
12096-459: The increasing cost of weapons systems . In 1981, Defence Secretary John Nott had advocated and initiated a series of cutbacks to the Navy . The Falklands War however proved a need for the Royal Navy to regain an expeditionary and littoral capability which, with its resources and structure at the time, would prove difficult. At the beginning of the 1980s, the Royal Navy was a force focused on blue-water anti-submarine warfare . Its purpose
12264-461: The mouthparts . The thorax has three pairs of segmented legs , one pair each for the three segments that compose the thorax and one or two pairs of wings . The abdomen is composed of eleven segments, some of which may be fused and houses the digestive , respiratory , excretory and reproductive systems. There is considerable variation between species and many adaptations to the body parts, especially wings, legs, antennae and mouthparts. Spiders
12432-400: The stomach . All vertebrates have a similar basic body plan and at some point in their lives, mostly in the embryonic stage, share the major chordate characteristics: a stiffening rod, the notochord ; a dorsal hollow tube of nervous material, the neural tube ; pharyngeal arches ; and a tail posterior to the anus. The spinal cord is protected by the vertebral column and is above
12600-420: The 2nd century, Galen of Pergamum , an anatomist, clinician , writer, and philosopher, wrote the final and highly influential anatomy treatise of ancient times. He compiled existing knowledge and studied anatomy through the dissection of animals. He was one of the first experimental physiologists through his vivisection experiments on animals. Galen's drawings, based mostly on dog anatomy, became effectively
12768-574: The Antarctic made several important discoveries in biology and zoology . Several of the Royal Navy's voyages ended in disaster such as those of Franklin and Scott . Between 1872 and 1876 HMS Challenger undertook the first global marine research expedition, the Challenger expedition . During World War I , the Royal Navy's strength was mostly deployed at home in the Grand Fleet , confronting
12936-697: The Atlantic Ocean and Mediterranean Sea. Control of military forces in Nova Scotia passed to the new Government of Canada after the 1867 Confederation of Canada and control of the naval dockyard in Halifax, Nova Scotia was transferred to the Government of Canada in 1905, five years prior to the establishment of the Royal Canadian Navy . Prior to the 1920s, it was presumed that the only navies that could challenge
13104-809: The Atlantic, and from its base in Bermuda it blockaded the Atlantic seaboard of the United States throughout the war and carried out (with Royal Marines, Colonial Marines , British Army , and Board of Ordnance military corps units) various amphibious operations, most notably the Chesapeake campaign . On the Great Lakes , however, the United States Navy established an advantage. In 1860, Albert, Prince Consort , wrote to
13272-802: The British Army and the Board of Ordnance military corps were cutback, weakening garrisons around the Empire, the Militia became a paper tiger, and the Volunteer Force and Fencible units disbanded, though the Yeomanry was maintained as a back-up to the police). Britain relied, throughout the 19th century and the first half of the 20th century, on imperial fortress colonies (originally Bermuda , Gibraltar, Halifax (Nova Scotia) , and Malta ). These areas permitted Britain to control
13440-753: The British advantage in ship numbers. In 1889, Parliament passed the Naval Defence Act , which formally adopted the 'two-power standard', which stipulated that the Royal Navy should maintain a number of battleships at least equal to the combined strength of the next two largest navies. The end of the 19th century saw structural changes and older vessels were scrapped or placed into reserve, making funds and manpower available for newer ships. The launch of HMS Dreadnought in 1906 rendered all existing battleships obsolete. The transition at this time from coal to fuel-oil for boiler firing would encourage Britain to expand their foothold in former Ottoman territories in
13608-599: The Foreign Secretary John Russell, 1st Earl Russell with his concern about "a perfect disgrace to our country, and particularly to the Admiralty ". The stated shipbuilding policy of the British monarchy was to take advantage of technological change and so be able to deploy a new weapons system that could defend British interests before other national and imperial resources are reasonably mobilized. Nevertheless, British taxpayers scrutinized progress in modernizing
13776-488: The French fleet in the Battle of Sluys in 1340. England's naval forces could not prevent frequent raids on the south-coast ports by the French and their allies. Such raids halted only with the occupation of northern France by Henry V . A Scottish fleet existed by the reign of William the Lion . In the early 13th century there was a resurgence of Viking naval power in the region. The Vikings clashed with Scotland over control of
13944-588: The German High Seas Fleet across the North Sea. Several inconclusive clashes took place between them, chiefly the Battle of Jutland in 1916. The British fighting advantage proved insurmountable, leading the High Seas Fleet to abandon any attempt to challenge British dominance. The Royal Navy under John Jellicoe also tried to avoid combat and remained in port at Scapa Flow for much of the war. This
14112-519: The King's Dock, rowed out to the man o' war Ruby , and confined in a spare cabin. Goodere told the men on deck to ignore Sir John's cries for help, as he was out of his mind, and would have to be watched to prevent his attempting his own life. Three men were chosen to attend the prisoner, and these three men, after being well primed with brandy, and on the promise of large rewards, went into the cabin early next morning (19 January 1741), and strangled Sir John with
14280-774: The Naval Service to the Admiralty Board , chaired by the Secretary of State for Defence . The Royal Navy operates from three bases in Britain where commissioned ships and submarines are based: Portsmouth , Clyde and Devonport , the last being the largest operational naval base in Western Europe, as well as two naval air stations, RNAS Yeovilton and RNAS Culdrose where maritime aircraft are based. The Royal Navy stated its six major roles in umbrella terms in 2017 as. The Royal Navy protects British interests at home and abroad, executing
14448-592: The Pacific and had begun building large, modern fleets which went to war with each other in 1904. Britain's reliance on Malta, via the Suez Canal, as the nearest Imperial fortress was improved, relying on amity and common interests that developed between Britain and the United States during and after World War I, by the completion of the Panama Canal in 1914, allowing the cruisers based in Bermuda to more easily and rapidly reach
14616-663: The Queen's ships in highly profitable raids against Spanish commerce and colonies. The Royal Navy was then used in 1588 to repulse the Spanish Armada , but the English Armada was lost the next year. In 1603, the Union of the Crowns created a personal union between England and Scotland. While the two remained distinct sovereign states for a further century, the two navies increasingly fought as
14784-526: The Royal Navy at sea in several capacities. For fleet replenishment, it deploys one Fleet Solid Support Ship and six fleet tankers (three of which are maintained in reserve). The RFA also has one aviation training and casualty reception vessel, which also operates as a Littoral Strike Ship . Three amphibious transport docks are also incorporated within its fleet. These are known as the Bay-class landing ships, of which four were introduced in 2006–2007, but one
14952-604: The Royal Navy belonged to nations on the Atlantic Ocean or its connected seas, despite the growth of the Imperial Russian and United States Pacific fleets during the latter half of the 19th Century. Britain relied on Malta, in the Mediterranean Sea, to project power to the Indian Ocean and western Pacific Ocean via the Suez Canal after its completion in 1869. It relied on friendship and common interests between Britain and
15120-512: The Royal Navy in 1917 with the formation of the Women's Royal Naval Service (WRNS), which was disbanded after the end of the First World War in 1919. It was revived in 1939, and the WRNS continued until disbandment in 1993, as a result of the decision to fully integrate women into the structures of the Royal Navy. Women now serve in all sections of the Royal Navy including the Royal Marines . In August 2019,
15288-530: The Royal Navy so as to ensure, that taypayers' money is not wasted. Between 1815 and 1914, the Royal Navy saw little serious action, owing to the absence of any opponent strong enough to challenge its dominance, though it did not suffer the drastic cutbacks the various military forces underwent in the period of economic austerity that followed the end of the Napoleonic Wars and the American War of 1812 (when
15456-665: The Royal Navy, plus 13 ships of the Royal Fleet Auxiliary (RFA). There are also four Point-class sealift ships from the Merchant Navy available to the RFA under a private finance initiative , while the civilian Marine Services operate auxiliary vessels which further support the Royal Navy in various capacities. The RFA replenishes Royal Navy warships at sea, and augments the Royal Navy's amphibious warfare capabilities through its three Bay-class landing ship vessels. It also works as
15624-530: The Royal Navy, with the final vessel, HMS St Albans , commissioned in June 2002. However, the 2004 Delivering Security in a Changing World review announced that three frigates would be paid off as part of a cost-cutting exercise, and these were subsequently sold to the Chilean Navy . The 2010 Strategic Defence and Security Review announced that the remaining 13 Type 23 frigates would eventually be replaced by
15792-610: The Royal Navy. In December 2019, the modified Batch 1 River-class vessel, HMS Clyde , was decommissioned, with the Batch 2 HMS Forth taking over duties as the Falkland Islands patrol ship. HMS Protector is a dedicated Antarctica patrol ship that fulfils the nation's mandate to provide support to the British Antarctic Survey (BAS). HMS Scott is an ocean survey vessel and at 13,500 tonnes
15960-825: The STOVL variant of the F-35 Lightning II . Queen Elizabeth began sea trials in June 2017, was commissioned later that year, and entered service in 2020, while the second, HMS Prince of Wales , began sea trials on 22 September 2019, was commissioned in December 2019 and was declared operational as of October 2021. The aircraft carriers form a central part of the UK Carrier Strike Group alongside escorts and support ships. Amphibious warfare ships in current service include two landing platform docks ( HMS Albion and HMS Bulwark ). While their primary role
16128-410: The United Kingdom's Continuous At Sea Deterrent (CASD). The UK government has committed to replace these submarines with four new Dreadnought -class submarines , which will enter service in the "early 2030s" to maintain this capability. Anatomy Anatomy (from Ancient Greek ἀνατομή ( anatomḗ ) ' dissection ') is the branch of morphology concerned with the study of
16296-647: The United States (which controlled transit through the Panama Canal , completed in 1914) during and after the First World War, and on Bermuda, to project power the length of the western Atlantic, including the Caribbean Sea and Gulf of Mexico. The area controlled from Bermuda (and Halifax until 1905) had been part of the North America Station, until the 1820s, which then absorbed the Jamaica Station to become
16464-425: The appearance and position of the various parts, the materials from which they are composed, and their relationships with other parts. Anatomy is quite distinct from physiology and biochemistry , which deal respectively with the functions of those parts and the chemical processes involved. For example, an anatomist is concerned with the shape, size, position, structure, blood supply and innervation of an organ such as
16632-531: The arts and sciences from the Ptolemaic dynasty of Egypt helped raise Alexandria up, further rivalling other Greek states' cultural and scientific achievements. Some of the most striking advances in early anatomy and physiology took place in Hellenistic Alexandria. Two of the most famous anatomists and physiologists of the third century were Herophilus and Erasistratus . These two physicians helped pioneer human dissection for medical research, using
16800-490: The atria were parts of the heart. Herophilus's knowledge of the human body has provided vital input towards understanding the brain, eye, liver, reproductive organs, and nervous system and characterizing the course of the disease. Erasistratus accurately described the structure of the brain, including the cavities and membranes, and made a distinction between its cerebrum and cerebellum During his study in Alexandria, Erasistratus
16968-526: The baronetcy on his father's execution, died a lunatic and unmarried in March 1761 at Clapton, London . The other, John , died in 1809 at Windsor Castle where he lived as "one of the Poor Knights of Windsor ". Although apparently entitled to inherit the baronetcy from his brother, he did not assume the title, and was referred to as "Mr Goodere" throughout his trial. Ralph Bigland , in his manuscript collections in
17136-518: The basis of sense organs and there is a central nervous system (brain and spinal cord) and a peripheral nervous system . The latter consists of sensory nerves that transmit information from sense organs and motor nerves that influence target organs. The peripheral nervous system is divided into the somatic nervous system which conveys sensation and controls voluntary muscle , and the autonomic nervous system which involuntarily controls smooth muscle , certain glands and internal organs, including
17304-560: The beginning of the war without the loss of a single life. The Royal Navy nevertheless remained active in other theatres, most notably in the Mediterranean Sea , where they waged the Dardanelles and Gallipoli campaigns in 1914 and 1915. British cruisers hunted down German commerce raiders across the world's oceans in 1914 and 1915, including the battles of Coronel , Falklands Islands , Cocos , and Rufiji Delta , among others. At
17472-462: The body and they swim by undulating their body from side to side. Reptiles are a class of animals comprising turtles , tuataras , lizards , snakes and crocodiles . They are tetrapods , but the snakes and a few species of lizard either have no limbs or their limbs are much reduced in size. Their bones are better ossified and their skeletons stronger than those of amphibians. The teeth are conical and mostly uniform in size. The surface cells of
17640-399: The brain, appreciated the division between cerebellum and cerebrum and recognized that the brain was the "seat of intellect" and not a "cooling chamber" as propounded by Aristotle Herophilus is also credited with describing the optic, oculomotor, motor division of the trigeminal, facial, vestibulocochlear and hypoglossal nerves. Incredible feats were made during the third century BCE in both
17808-401: The branch of superficial anatomy . Microscopic anatomy involves the use of optical instruments in the study of the tissues of various structures, known as histology , and also in the study of cells . The history of anatomy is characterized by a progressive understanding of the functions of the organs and structures of the human body. Methods have also improved dramatically, advancing from
17976-516: The cadavers of condemned criminals, which was considered taboo until the Renaissance—Herophilus was recognized as the first person to perform systematic dissections. Herophilus became known for his anatomical works, making impressive contributions to many branches of anatomy and many other aspects of medicine. Some of the works included classifying the system of the pulse, the discovery that human arteries had thicker walls than veins, and that
18144-653: The class serve in U.K. waters in a sovereignty and fisheries protection role while the five Batch 2 ships are forward-deployed on a long-term basis to Gibraltar, the Caribbean, the Falkland Islands and the Indo-Pacific region. The vessel MV Grampian Frontier is leased from Scottish-based North Star Shipping for patrol duties around the British Indian Ocean Territory . However, she is not in commission with
18312-652: The cross of St Andrew was replaced with the Union Jack. On English ships, the red, white, or blue ensigns had the St George's Cross of England removed from the canton, and the combined crosses of the Union flag put in its place. Throughout the 18th and 19th centuries, the Royal Navy was the largest maritime force in the world, maintaining superiority in financing, tactics, training, organisation, social cohesion, hygiene, logistical support and warship design. The peace settlement following
18480-418: The dermis which are overlain by horny ones and are partially fused with the ribs and spine. The neck is long and flexible and the head and the legs can be drawn back inside the shell. Turtles are vegetarians and the typical reptile teeth have been replaced by sharp, horny plates. In aquatic species, the front legs are modified into flippers. Tuataras superficially resemble lizards but the lineages diverged in
18648-402: The digestive and reproductive systems. Herophilus discovered and described not only the salivary glands but also the small intestine and liver. He showed that the uterus is a hollow organ and described the ovaries and uterine tubes. He recognized that spermatozoa were produced by the testes and was the first to identify the prostate gland. The anatomy of the muscles and skeleton is described in
18816-686: The early 16th century; the oldest of the UK's armed services , it is consequently known as the Senior Service . From the 19th century until the Second World War , it was the world's most powerful navy. The Royal Navy played a key part in establishing and defending the British Empire , and four Imperial fortress colonies and a string of imperial bases and coaling stations secured the Royal Navy's ability to assert naval superiority. Following World War I , it
18984-566: The eastern Pacific Ocean (after the war, the Royal Navy's Bermuda-based North America and West Indies Station was consequently re-designated the America and West Indies station , including a South American division. The rising power and increasing belligerence of the Japanese Empire after World War I, however, resulted in the construction of the Singapore Naval Base , which was completed in 1938, less than four years before hostilities with Japan did commence during World War II . In 1932,
19152-402: The egg-laying monotremes , the platypus and the echidnas of Australia. Most other mammals have a placenta through which the developing foetus obtains nourishment, but in marsupials , the foetal stage is very short and the immature young is born and finds its way to its mother's pouch where it latches on to a teat and completes its development. Humans have the overall body plan of
19320-523: The end of World War I, the Royal Navy remained by far the world's most powerful navy, larger than the U.S. Navy and French Navy combined, and over twice as large as the Imperial Japanese Navy and Royal Italian Navy combined. Its former primary competitor, the Imperial German Navy, was destroyed at the end of the war . In the inter-war period , the Royal Navy was stripped of much of its power. The Washington and London Naval Treaties imposed
19488-523: The end of each male pedipalp is a spoon-shaped cymbium that acts to support the copulatory organ . In 1600 BCE, the Edwin Smith Papyrus , an Ancient Egyptian medical text , described the heart and its vessels, as well as the brain and its meninges and cerebrospinal fluid , and the liver , spleen , kidneys , uterus and bladder . It showed the blood vessels diverging from the heart. The Ebers Papyrus ( c. 1550 BCE ) features
19656-467: The epidermis are modified into horny scales which create a waterproof layer. Reptiles are unable to use their skin for respiration as do amphibians and have a more efficient respiratory system drawing air into their lungs by expanding their chest walls. The heart resembles that of the amphibian but there is a septum which more completely separates the oxygenated and deoxygenated bloodstreams. The reproductive system has evolved for internal fertilization, with
19824-413: The epidermis is normally formed of epithelial cells and secretes an extracellular matrix which provides support to the organism. An endoskeleton derived from the mesoderm is present in echinoderms , sponges and some cephalopods . Exoskeletons are derived from the epidermis and is composed of chitin in arthropods (insects, spiders, ticks, shrimps, crabs, lobsters). Calcium carbonate constitutes
19992-429: The epithelial lining and in the large intestine there are intestinal villi . Skin consists of an outer layer of keratinized stratified squamous epithelium that covers the exterior of the vertebrate body. Keratinocytes make up to 95% of the cells in the skin . The epithelial cells on the external surface of the body typically secrete an extracellular matrix in the form of a cuticle . In simple animals this may just be
20160-610: The equivalent of tissues and organs. Locomotion is often provided by cilia or flagella or may proceed via the advance of pseudopodia , food may be gathered by phagocytosis , energy needs may be supplied by photosynthesis and the cell may be supported by an endoskeleton or an exoskeleton . Some protozoans can form multicellular colonies. Metazoans are a multicellular organism, with different groups of cells serving different functions. The most basic types of metazoan tissues are epithelium and connective tissue, both of which are present in nearly all invertebrates. The outer surface of
20328-527: The examination of animals by dissection of carcasses and cadavers (corpses) to 20th-century medical imaging techniques, including X-ray , ultrasound , and magnetic resonance imaging . Derived from the Greek ἀνατομή anatomē "dissection" (from ἀνατέμνω anatémnō "I cut up, cut open" from ἀνά aná "up", and τέμνω témnō "I cut"), anatomy is the scientific study of the structure of organisms including their systems, organs and tissues . It includes
20496-405: The father of microscopical anatomy, discovered that plants had tubules similar to those he saw in insects like the silk worm. He observed that when a ring-like portion of bark was removed on a trunk a swelling occurred in the tissues above the ring, and he unmistakably interpreted this as growth stimulated by food coming down from the leaves, and being captured above the ring. Arthropods comprise
20664-435: The feathers when the bird preens . There are scales on the legs, feet and claws on the tips of the toes. Mammals are a diverse class of animals, mostly terrestrial but some are aquatic and others have evolved flapping or gliding flight. They mostly have four limbs, but some aquatic mammals have no limbs or limbs modified into fins, and the forelimbs of bats are modified into wings. The legs of most mammals are situated below
20832-443: The features of ancient fish. They have a bony skeleton, are generally laterally flattened, have five pairs of gills protected by an operculum , and a mouth at or near the tip of the snout. The dermis is covered with overlapping scales . Bony fish have a swim bladder which helps them maintain a constant depth in the water column, but not a cloaca. They mostly spawn a large number of small eggs with little yolk which they broadcast into
21000-402: The first two years of the war. Over 3,000 people were lost when the converted troopship Lancastria was sunk in June 1940, the greatest maritime disaster in Britain's history. The Navy's most critical struggle was the Battle of the Atlantic defending Britain's vital North American commercial supply lines against U-boat attack. A traditional convoy system was instituted from the start of
21168-436: The foreign and defence policies of His Majesty's Government through the exercise of military effect, diplomatic activities and other activities in support of these objectives. It is also a key element of the British contribution to NATO, with a number of ships or aircraft allocated to NATO tasks at any time. In 2007 core capabilities were described as: The English Royal Navy was formally founded in 1546 by Henry VIII , though
21336-445: The granddaughter of Lewis Watson, 1st Baron Rockingham . He had several brothers, the eldest of whom was killed in a duel. On the death of Sir Edward Dineley, the next eldest surviving brother, John, inherited the extensive Dineley estate. Sir Edward Dineley's Will made bequests to his grandsons George Goodere, Samuel Goodere and Henry Goodere, and also John Goodere. Since the first born Goodere grandson had been killed before this Will
21504-482: The head. The dermis is covered with separate dermal placoid scales . They have a cloaca into which the urinary and genital passages open, but not a swim bladder . Cartilaginous fish produce a small number of large, yolky eggs. Some species are ovoviviparous and the young develop internally but others are oviparous and the larvae develop externally in egg cases. The bony fish lineage shows more derived anatomical traits, often with major evolutionary changes from
21672-512: The human body were made, which contributed to the understanding of the brain, eye, liver, reproductive organs, and nervous system. The Hellenistic Egyptian city of Alexandria was the stepping-stone for Greek anatomy and physiology. Alexandria not only housed the biggest library for medical records and books of the liberal arts in the world during the time of the Greeks but was also home to many medical practitioners and philosophers. Great patronage of
21840-403: The internal structure of organisms and their parts. Anatomy is a branch of natural science that deals with the structural organization of living things. It is an old science, having its beginnings in prehistoric times. Anatomy is inherently tied to developmental biology , embryology , comparative anatomy , evolutionary biology , and phylogeny , as these are the processes by which anatomy
22008-463: The isles though Alexander III was ultimately successful in asserting Scottish control. The Scottish fleet was of particular import in repulsing English forces in the early 14th century. A standing "Navy Royal", with its own secretariat, dockyards and a permanent core of purpose-built warships, emerged during the reign of Henry VIII. Under Elizabeth I , England became involved in a war with Spain , which saw privately owned vessels combining with
22176-492: The largest phylum of invertebrates in the animal kingdom with over a million known species. Insects possess segmented bodies supported by a hard-jointed outer covering, the exoskeleton , made mostly of chitin . The segments of the body are organized into three distinct parts, a head, a thorax and an abdomen . The head typically bears a pair of sensory antennae , a pair of compound eyes , one to three simple eyes ( ocelli ) and three sets of modified appendages that form
22344-399: The length of the sides of fish, and these respond to nearby movements and to changes in water pressure. Sharks and rays are basal fish with numerous primitive anatomical features similar to those of ancient fish, including skeletons composed of cartilage. Their bodies tend to be dorso-ventrally flattened, they usually have five pairs of gill slits and a large mouth set on the underside of
22512-439: The liver; while a physiologist is interested in the production of bile , the role of the liver in nutrition and the regulation of bodily functions. The discipline of anatomy can be subdivided into a number of branches, including gross or macroscopic anatomy and microscopic anatomy. Gross anatomy is the study of structures large enough to be seen with the naked eye, and also includes superficial anatomy or surface anatomy,
22680-437: The lungs have a single chamber and lack bronchi . The tuatara has a well-developed parietal eye on its forehead. Lizards have skulls with only one fenestra on each side, the lower bar of bone below the second fenestra having been lost. This results in the jaws being less rigidly attached which allows the mouth to open wider. Lizards are mostly quadrupeds, with the trunk held off the ground by short, sideways-facing legs, but
22848-400: The main part of the trunk. The heart has two chambers and pumps the blood through the respiratory surfaces of the gills and on round the body in a single circulatory loop. The eyes are adapted for seeing underwater and have only local vision. There is an inner ear but no external or middle ear . Low frequency vibrations are detected by the lateral line system of sense organs that run along
23016-477: The more developed animals whose structures and organs are formed from three germ layers are called triploblastic . All of a triploblastic animal's tissues and organs are derived from the three germ layers of the embryo, the ectoderm , mesoderm and endoderm . Animal tissues can be grouped into four basic types: connective , epithelial , muscle and nervous tissue . Connective tissues are fibrous and made up of cells scattered among inorganic material called
23184-737: The navy to meet its commitments. In December 2019 the First Sea Lord , Admiral Tony Radakin , outlined a proposal to reduce the number of Rear-Admirals at Navy Command by five. The fighting arms (excluding Commandant General Royal Marines ) would be reduced to commodore (1-star) rank and the surface flotillas would be combined. Training would be concentrated under the Fleet Commander . The Royal Navy has two Queen Elizabeth -class aircraft carriers. Each carrier cost £ 3 billion and displaces 65,000 tonnes (64,000 long tons; 72,000 short tons). The first, HMS Queen Elizabeth , commenced flight trials in 2018. Both are intended to operate
23352-475: The nerves form a nerve net , but in most animals they are organized longitudinally into bundles. In simple animals, receptor neurons in the body wall cause a local reaction to a stimulus. In more complex animals, specialized receptor cells such as chemoreceptors and photoreceptors are found in groups and send messages along neural networks to other parts of the organism. Neurons can be connected together in ganglia . In higher animals, specialized receptors are
23520-416: The notochord, and the gastrointestinal tract is below it. Nervous tissue is derived from the ectoderm , connective tissues are derived from mesoderm , and gut is derived from the endoderm . At the posterior end is a tail which continues the spinal cord and vertebrae but not the gut. The mouth is found at the anterior end of the animal, and the anus at the base of the tail. The defining characteristic of
23688-428: The only anatomical textbook for the next thousand years. His work was known to Renaissance doctors only through Islamic Golden Age medicine until it was translated from Greek sometime in the 15th century. Anatomy developed little from classical times until the sixteenth century; as the historian Marie Boas writes, "Progress in anatomy before the sixteenth century is as mysteriously slow as its development after 1500
23856-430: The property to one of the sons of his sister Eleanor, the mother of Samuel Foote the comedian. In November 1740, Samuel Goodere was appointed to the command of HMS Ruby , then lying off Bristol . On Sunday, 18 January 1741, Samuel was on shore and learned that his brother, Sir John, who was on his way to Bath in the hope of curing his ill health, was dining with an attorney in the city named Jarrett Smith. Samuel sent
24024-419: The rest of the animal kingdom, and the term also includes the anatomy of other animals. The term zootomy is also sometimes used to specifically refer to non-human animals. The structure and tissues of plants are of a dissimilar nature and they are studied in plant anatomy . The kingdom Animalia contains multicellular organisms that are heterotrophic and motile (although some have secondarily adopted
24192-493: The same underlying skeletal structure was inherited from their last common ancestor. This is one of the arguments put forward by Charles Darwin to support his theory of evolution . The body of a fish is divided into a head, trunk and tail, although the divisions between the three are not always externally visible. The skeleton, which forms the support structure inside the fish, is either made of cartilage, in cartilaginous fish , or bone in bony fish . The main skeletal element
24360-661: The scrapping of some capital ships and limitations on new construction. The lack of an imperial fortress in the region of Asia , the Indian Ocean , and the Pacific Ocean was always to be a weakness throughout the 19th century as the former North American colonies that had become the United States of America had multiplied towards the Pacific Coast of North America, and the Russian Empire and Japanese Empire both had ports on
24528-670: The shells of molluscs , brachiopods and some tube-building polychaete worms and silica forms the exoskeleton of the microscopic diatoms and radiolaria . Other invertebrates may have no rigid structures but the epidermis may secrete a variety of surface coatings such as the pinacoderm of sponges, the gelatinous cuticle of cnidarians ( polyps , sea anemones , jellyfish ) and the collagenous cuticle of annelids . The outer epithelial layer may include cells of several types including sensory cells, gland cells and stinging cells. There may also be protrusions such as microvilli , cilia, bristles, spines and tubercles . Marcello Malpighi ,
24696-411: The size of the Royal Navy. A 2013 report found that the Royal Navy was already too small, and that Britain would have to depend on her allies if her territories were attacked. The Royal Navy was responsible for training the fledgling Iraqi Navy and securing Iraq's oil terminals following the cessation of hostilities in the country. The Iraqi Training and Advisory Mission (Navy) ( Umm Qasr ), headed by
24864-416: The start of World War II in 1939, the Royal Navy was still the largest in the world, with over 1,400 vessels. The Royal Navy provided critical cover during Operation Dynamo , the British evacuations from Dunkirk , and as the ultimate deterrent to a German invasion of Britain during the following four months. The Luftwaffe under Hermann Göring attempted to gain air supremacy over southern England in
25032-400: The study by sight of the external body features. Microscopic anatomy is the study of structures on a microscopic scale, along with histology (the study of tissues), and embryology (the study of an organism in its immature condition). Regional anatomy is the study of the interrelationships of all of the structures in a specific body region, such as the abdomen. In contrast, systemic anatomy
25200-741: The tail is laterally compressed. It undulates from side to side to force the animal through the water when swimming. The tough keratinized scales provide body armour and some are fused to the skull. The nostrils, eyes and ears are elevated above the top of the flat head enabling them to remain above the surface of the water when the animal is floating. Valves seal the nostrils and ears when it is submerged. Unlike other reptiles, crocodilians have hearts with four chambers allowing complete separation of oxygenated and deoxygenated blood. Birds are tetrapods but though their hind limbs are used for walking or hopping, their front limbs are wings covered with feathers and adapted for flight. Birds are endothermic , have
25368-464: The terms of the 1706 Will by leaving his Estate to his sister Eleanor Foote's son. John and Samuel Goodere both served as Mayors of Evesham. Their solicitor friend, Jarrett Smith, was unwittingly involved in the plot by Samuel to get rid of his brother John. Samuel entered the navy in 1705 as a volunteer on board HMS Ipswich , with Captain Kirktowne, and served in a subordinate rank and afterwards as
25536-401: The toes are often webbed for swimming or have suction pads for climbing. Frogs have large eyes and no tail. Salamanders resemble lizards in appearance; their short legs project sideways, the belly is close to or in contact with the ground and they have a long tail. Caecilians superficially resemble earthworms and are limbless. They burrow by means of zones of muscle contractions which move along
25704-538: The trunk, which is held well clear of the ground. The bones of mammals are well ossified and their teeth, which are usually differentiated, are coated in a layer of prismatic enamel . The teeth are shed once ( milk teeth ) during the animal's lifetime or not at all, as is the case in cetaceans . Mammals have three bones in the middle ear and a cochlea in the inner ear . They are clothed in hair and their skin contains glands which secrete sweat . Some of these glands are specialized as mammary glands , producing milk to feed
25872-442: The vertebrae. Their skulls are mostly broad and short, and are often incompletely ossified. Their skin contains little keratin and lacks scales, but contains many mucous glands and in some species, poison glands. The hearts of amphibians have three chambers, two atria and one ventricle . They have a urinary bladder and nitrogenous waste products are excreted primarily as urea . Amphibians breathe by means of buccal pumping ,
26040-502: The war, but German submarine tactics, based on group attacks by " wolf-packs ", were much more effective than in the previous war, and the threat remained serious for well over three years. After World War II, the decline of the British Empire and the economic hardships in Britain forced the reduction in the size and capability of the Royal Navy. The United States Navy instead took on the role of global naval power. Governments since have faced increasing budgetary pressures, partly due to
26208-437: The water column. Amphibians are a class of animals comprising frogs , salamanders and caecilians . They are tetrapods , but the caecilians and a few species of salamander have either no limbs or their limbs are much reduced in size. Their main bones are hollow and lightweight and are fully ossified and the vertebrae interlock with each other and have articular processes . Their ribs are usually short and may be fused to
26376-411: The young. Mammals breathe with lungs and have a muscular diaphragm separating the thorax from the abdomen which helps them draw air into the lungs. The mammalian heart has four chambers, and oxygenated and deoxygenated blood are kept entirely separate. Nitrogenous waste is excreted primarily as urea. Mammals are amniotes , and most are viviparous , giving birth to live young. Exceptions to this are
26544-606: Was acquired in 2023 to act as a mothership for autonomous minehunting systems. The Royal Navy also includes a number of smaller non-commissioned assets such as the Sea-class workboats . On 29 July 2022, the Royal Navy christened a new experimental ship, XV Patrick Blackett , which it aims to use as a testbed for autonomous systems. Whilst the ship flies the Blue Ensign , it is crewed by Royal Navy personnel and will participate in Royal Navy and NATO exercises. The Submarine Service
26712-425: Was also responsible for naming and describing the function of the epiglottis and the heart's valves, including the tricuspid. During the third century, Greek physicians were able to differentiate nerves from blood vessels and tendons and to realize that the nerves convey neural impulses. It was Herophilus who made the point that damage to motor nerves induced paralysis. Herophilus named the meninges and ventricles in
26880-628: Was contrary to widespread prewar expectations that in the event of a Continental conflict Britain would primarily provide naval support to the Entente Powers while sending at most only a small ground army. Nevertheless, the Royal Navy played an important role in securing the British Isles and the English Channel , notably ferrying the entire British Expeditionary Force to the Western Front at
27048-426: Was drawn up he is not named in the Will which is dated 12 November 1706 and bears a codicil. Sir Edward Dineley's Will stipulated that grandson John Goodere's first born son "of his body" should in turn inherit, or if John's first born son died, then the second son born of John's body should inherit. If John had no sons, his brother George Goodere and George's son should inherit. Should John and George fail to produce
27216-426: Was now in question. The Royal Navy clearance diving unit, the Fleet Diving Squadron, was reorganised and renamed the Diving and Threat Exploitation Group in 2022. The group consists of five squadrons: Alpha, Bravo, Charlie, Delta, and Echo. The Royal Navy has a separate unit with divers the special forces unit the Special Boat Service . The escort fleet comprises guided missile destroyers and frigates and
27384-421: Was particularly concerned with studies of the circulatory and nervous systems. He could distinguish the human body's sensory and motor nerves and believed air entered the lungs and heart, which was then carried throughout the body. His distinction between the arteries and veins—the arteries carrying the air through the body, while the veins carry the blood from the heart was a great anatomical discovery. Erasistratus
27552-407: Was significantly reduced in size. During the Cold War , the Royal Navy transformed into a primarily anti-submarine force , hunting for Soviet submarines and mostly active in the GIUK gap . Following the collapse of the Soviet Union , its focus has returned to expeditionary operations around the world and it remains one of the world's foremost blue-water navies . The Royal Navy maintains
27720-497: Was sold to the Royal Australian Navy in 2011. In November 2006, the First Sea Lord Admiral Sir Jonathon Band described the Royal Fleet Auxiliary vessels as "a major uplift in the Royal Navy's war fighting capability". In February 2023, a commercial vessel was also acquired to act as a Multi-Role Ocean Surveillance (MROS) Ship for the protection of critical seabed infrastructure and other tasks. She entered service as RFA Proteus . An additional vessel, RFA Stirling Castle ,
27888-445: Was supported against him by Samuel. On the death of their father Sir Edward Goodere on 29 March 1739, John inherited the baronetcy , but a share of the Goodere estate was left to Samuel. John thought Samuel's share was too generous, while Samuel thought it too little. The brothers quarrelled, and John, joining with his son who was of age, cut off the entail (bequest). Shortly after, his son died, and John announced his intention of leaving
28056-406: Was thwarted by the defeat of their escort fleet in the extraordinary Battle of Quiberon Bay in 1759, fought in dangerous conditions. In 1762, the resumption of hostilities with Spain led to the British capture of Manila and of Havana , along with a Spanish fleet sheltering there. British naval supremacy could however be challenged still in this period by coalitions of other nations, as seen in
28224-418: Was to search for and destroy Soviet submarines in the North Atlantic, and to operate the nuclear deterrent submarine force. The navy received its first nuclear weapons with the introduction of the first of the Resolution -class submarines armed with the Polaris missile . Following the conclusion of the collapse of the Soviet Union and the end of the Cold War in 1991, the Royal Navy began to experience
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