Misplaced Pages

#413586

146-538: Salting is used because most bacteria , fungi and other potentially pathogenic organisms cannot survive in a highly salty environment, due to the hypertonic nature of salt. Any living cell in such an environment will become dehydrated through osmosis and die or become temporarily inactivated. Fine grained salts were more expensive but also absorbed moisture faster than coarse salt. Salting could be combined with smoking to produce bacon in peasant homes. Instructions for preserving (salting) freshly killed venison in

292-574: A communist-led revolution in Afghanistan, Soviet forces invaded the country, ultimately starting the Soviet–Afghan War . In May 1988, the Soviets started to withdraw from Afghanistan , due to international opposition, persistent anti-Soviet guerrilla warfare, and a lack of support by Soviet citizens. From 1985 onwards, the last Soviet leader Mikhail Gorbachev , who sought to enact liberal reforms in

438-543: A potential difference analogous to a battery. The general lack of internal membranes in bacteria means these reactions, such as electron transport , occur across the cell membrane between the cytoplasm and the outside of the cell or periplasm . However, in many photosynthetic bacteria, the plasma membrane is highly folded and fills most of the cell with layers of light-gathering membrane. These light-gathering complexes may even form lipid-enclosed structures called chlorosomes in green sulfur bacteria . Bacteria do not have

584-404: A terminal electron acceptor in a redox reaction . Chemotrophs are further divided by the types of compounds they use to transfer electrons. Bacteria that derive electrons from inorganic compounds such as hydrogen, carbon monoxide , or ammonia are called lithotrophs , while those that use organic compounds are called organotrophs . Still, more specifically, aerobic organisms use oxygen as

730-626: A troika was designated to take charge. Eventually Joseph Stalin , the General Secretary of the Communist Party , managed to suppress all opposition factions and consolidate power in his hands to become the country's dictator by the 1930s. Leon Trotsky , the main proponent of world revolution , was exiled from the Soviet Union in 1929, and Stalin's idea of Socialism in One Country became

876-595: A Varangian from the Rus' people , named Rurik , was elected ruler of Novgorod in 862. In 882, his successor Oleg ventured south and conquered Kiev , which had been previously paying tribute to the Khazars . Rurik's son Igor and Igor's son Sviatoslav subsequently subdued all local East Slavic tribes to Kievan rule, destroyed the Khazar Khaganate, and launched several military expeditions to Byzantium and Persia . In

1022-744: A bacterial strain. However, liquid growth media are used when the measurement of growth or large volumes of cells are required. Growth in stirred liquid media occurs as an even cell suspension, making the cultures easy to divide and transfer, although isolating single bacteria from liquid media is difficult. The use of selective media (media with specific nutrients added or deficient, or with antibiotics added) can help identify specific organisms. Most laboratory techniques for growing bacteria use high levels of nutrients to produce large amounts of cells cheaply and quickly. However, in natural environments, nutrients are limited, meaning that bacteria cannot continue to reproduce indefinitely. This nutrient limitation has led

1168-501: A crisis. By 1991, economic and political turmoil began to boil over as the Baltic states chose to secede from the Soviet Union. On 17 March, a referendum was held, in which the vast majority of participating citizens voted in favour of changing the Soviet Union into a renewed federation . In June 1991, Boris Yeltsin became the first directly elected President in Russian history when he

1314-707: A decisive role for the Allies in World War II by leading large-scale efforts on the Eastern Front . With the onset of the Cold War , it competed with the United States for ideological dominance and international influence . The Soviet era of the 20th century saw some of the most significant Russian technological achievements , including the first human-made satellite and the first human expedition into outer space . In 1991,

1460-424: A disorganised slime layer of extracellular polymeric substances to a highly structured capsule . These structures can protect cells from engulfment by eukaryotic cells such as macrophages (part of the human immune system ). They can also act as antigens and be involved in cell recognition, as well as aiding attachment to surfaces and the formation of biofilms. The assembly of these extracellular structures

1606-413: A few micrometres in thickness to up to half a metre in depth, and may contain multiple species of bacteria, protists and archaea. Bacteria living in biofilms display a complex arrangement of cells and extracellular components, forming secondary structures, such as microcolonies , through which there are networks of channels to enable better diffusion of nutrients. In natural environments, such as soil or

SECTION 10

#1733084612414

1752-415: A few species are visible to the unaided eye—for example, Thiomargarita namibiensis is up to half a millimetre long, Epulopiscium fishelsoni reaches 0.7 mm, and Thiomargarita magnifica can reach even 2 cm in length, which is 50 times larger than other known bacteria. Among the smallest bacteria are members of the genus Mycoplasma , which measure only 0.3 micrometres, as small as

1898-426: A fixed size and then reproduce through binary fission , a form of asexual reproduction . Under optimal conditions, bacteria can grow and divide extremely rapidly, and some bacterial populations can double as quickly as every 17 minutes. In cell division, two identical clone daughter cells are produced. Some bacteria, while still reproducing asexually, form more complex reproductive structures that help disperse

2044-639: A global superpower. After World War II, according to the Potsdam Conference , the Red Army occupied parts of Eastern and Central Europe, including East Germany and the eastern regions of Austria . Dependent communist governments were installed in the Eastern Bloc satellite states. After becoming the world's second nuclear power , the Soviet Union established the Warsaw Pact alliance, and entered into

2190-400: A growing problem. Bacteria are important in sewage treatment and the breakdown of oil spills , the production of cheese and yogurt through fermentation , the recovery of gold, palladium , copper and other metals in the mining sector ( biomining , bioleaching ), as well as in biotechnology , and the manufacture of antibiotics and other chemicals. Once regarded as plants constituting

2336-778: A large confederacy, which was subsequently taken over by the Cumans and the Kipchaks . The ancestors of Russians are among the Slavic tribes that separated from the Proto-Indo-Europeans, who appeared in the northeastern part of Europe c.  1500  years ago. The East Slavs gradually settled western Russia (approximately between modern Moscow and Saint-Petersburg ) in two waves: one moving from Kiev towards present-day Suzdal and Murom and another from Polotsk towards Novgorod and Rostov . Prior to Slavic migration, that territory

2482-497: A member state of the G20 , SCO , BRICS , APEC , OSCE , and WTO ; and the leading member state of post-Soviet organisations such as CIS , CSTO , and EAEU/EEU . It possesses the largest stockpile of nuclear weapons and has the third-highest military expenditure . Russia is generally considered a great power and is a regional power . Internationally, Russia ranks very low in measurements of democracy , human rights and freedom of

2628-408: A membrane-bound nucleus, and their genetic material is typically a single circular bacterial chromosome of DNA located in the cytoplasm in an irregularly shaped body called the nucleoid . The nucleoid contains the chromosome with its associated proteins and RNA . Like all other organisms , bacteria contain ribosomes for the production of proteins, but the structure of the bacterial ribosome

2774-533: A million inhabitants. Its capital and largest city is Moscow . Saint Petersburg is Russia's second-largest city and its cultural capital . The East Slavs emerged as a recognised group in Europe between the 3rd and 8th centuries CE. The first East Slavic state, Kievan Rus' , arose in the 9th century, and in 988, it adopted Orthodox Christianity from the Byzantine Empire . Kievan Rus' ultimately disintegrated;

2920-474: A number of conflicts in former Soviet states and other countries , including its war with Georgia in 2008 and its war with Ukraine since 2014, which has involved the internationally unrecognised annexations of Ukrainian territory including Crimea in 2014 and four other regions in 2022 during an ongoing invasion . Russia is a permanent member of the United Nations Security Council ;

3066-528: A particular organism or group of organisms ( syntrophy ). Bacterial growth follows four phases. When a population of bacteria first enter a high-nutrient environment that allows growth, the cells need to adapt to their new environment. The first phase of growth is the lag phase , a period of slow growth when the cells are adapting to the high-nutrient environment and preparing for fast growth. The lag phase has high biosynthesis rates, as proteins necessary for rapid growth are produced. The second phase of growth

SECTION 20

#1733084612414

3212-415: A process called transformation . Many bacteria can naturally take up DNA from the environment, while others must be chemically altered in order to induce them to take up DNA. The development of competence in nature is usually associated with stressful environmental conditions and seems to be an adaptation for facilitating repair of DNA damage in recipient cells. Second, bacteriophages can integrate into

3358-563: A secret protocol within the Molotov–Ribbentrop Pact with Nazi Germany . The Soviet Union later invaded Finland , and occupied and annexed the Baltic states , as well as parts of Romania . On 22 June 1941, Germany invaded the Soviet Union , opening the Eastern Front , the largest theater of World War II. Eventually, some 5 million Red Army troops were captured by the Nazis;

3504-425: A single continuous stretch of DNA. Although several different types of introns do exist in bacteria, these are much rarer than in eukaryotes. Bacteria, as asexual organisms, inherit an identical copy of the parent's genome and are clonal . However, all bacteria can evolve by selection on changes to their genetic material DNA caused by genetic recombination or mutations . Mutations arise from errors made during

3650-427: A single linear chromosome, while some Vibrio species contain more than one chromosome. Some bacteria contain plasmids , small extra-chromosomal molecules of DNA that may contain genes for various useful functions such as antibiotic resistance , metabolic capabilities, or various virulence factors . Bacteria genomes usually encode a few hundred to a few thousand genes. The genes in bacterial genomes are usually

3796-649: A source of electrons and a substrate for carbon anabolism . In many ways, bacterial metabolism provides traits that are useful for ecological stability and for human society. For example, diazotrophs have the ability to fix nitrogen gas using the enzyme nitrogenase . This trait, which can be found in bacteria of most metabolic types listed above, leads to the ecologically important processes of denitrification , sulfate reduction , and acetogenesis , respectively. Bacterial metabolic processes are important drivers in biological responses to pollution ; for example, sulfate-reducing bacteria are largely responsible for

3942-458: A struggle for global dominance, known as the Cold War , with the rivalling United States and NATO . After Stalin's death in 1953 and a short period of collective rule , the new leader Nikita Khrushchev denounced Stalin and launched the policy of de-Stalinization , releasing many political prisoners from the Gulag labour camps. The general easement of repressive policies became known later as

4088-785: A thick peptidoglycan cell wall like a Gram-positive bacterium, but also a second outer layer of lipids. In many bacteria, an S-layer of rigidly arrayed protein molecules covers the outside of the cell. This layer provides chemical and physical protection for the cell surface and can act as a macromolecular diffusion barrier . S-layers have diverse functions and are known to act as virulence factors in Campylobacter species and contain surface enzymes in Bacillus stearothermophilus . Flagella are rigid protein structures, about 20 nanometres in diameter and up to 20 micrometres in length, that are used for motility . Flagella are driven by

4234-417: A three- dimensional random walk . Bacterial species differ in the number and arrangement of flagella on their surface; some have a single flagellum ( monotrichous ), a flagellum at each end ( amphitrichous ), clusters of flagella at the poles of the cell ( lophotrichous ), while others have flagella distributed over the entire surface of the cell ( peritrichous ). The flagella of a group of bacteria,

4380-480: Is caused by a toxin released by the bacteria that grow from the spores. Clostridioides difficile infection , a common problem in healthcare settings, is caused by spore-forming bacteria. Bacteria exhibit an extremely wide variety of metabolic types. The distribution of metabolic traits within a group of bacteria has traditionally been used to define their taxonomy , but these traits often do not correspond with modern genetic classifications. Bacterial metabolism

4526-412: Is classified into nutritional groups on the basis of three major criteria: the source of energy , the electron donors used, and the source of carbon used for growth. Phototrophic bacteria derive energy from light using photosynthesis , while chemotrophic bacteria breaking down chemical compounds through oxidation , driving metabolism by transferring electrons from a given electron donor to

Salting (food) - Misplaced Pages Continue

4672-536: Is dependent on bacterial secretion systems . These transfer proteins from the cytoplasm into the periplasm or into the environment around the cell. Many types of secretion systems are known and these structures are often essential for the virulence of pathogens, so are intensively studied. Some genera of Gram-positive bacteria, such as Bacillus , Clostridium , Sporohalobacter , Anaerobacter , and Heliobacterium , can form highly resistant, dormant structures called endospores . Endospores develop within

4818-534: Is determined by the bacterial cell wall and cytoskeleton and is important because it can influence the ability of bacteria to acquire nutrients, attach to surfaces, swim through liquids and escape predators . Multicellularity . Most bacterial species exist as single cells; others associate in characteristic patterns: Neisseria forms diploids (pairs), streptococci form chains, and staphylococci group together in "bunch of grapes" clusters. Bacteria can also group to form larger multicellular structures, such as

4964-441: Is different from that of eukaryotes and archaea. Some bacteria produce intracellular nutrient storage granules, such as glycogen , polyphosphate , sulfur or polyhydroxyalkanoates . Bacteria such as the photosynthetic cyanobacteria , produce internal gas vacuoles , which they use to regulate their buoyancy, allowing them to move up or down into water layers with different light intensities and nutrient levels. Around

5110-414: Is essential to the survival of many bacteria, and the antibiotic penicillin (produced by a fungus called Penicillium ) is able to kill bacteria by inhibiting a step in the synthesis of peptidoglycan. There are broadly speaking two different types of cell wall in bacteria, that classify bacteria into Gram-positive bacteria and Gram-negative bacteria . The names originate from the reaction of cells to

5256-529: Is made of about 20 proteins, with approximately another 30 proteins required for its regulation and assembly. The flagellum is a rotating structure driven by a reversible motor at the base that uses the electrochemical gradient across the membrane for power. Bacteria can use flagella in different ways to generate different kinds of movement. Many bacteria (such as E. coli ) have two distinct modes of movement: forward movement (swimming) and tumbling. The tumbling allows them to reorient and makes their movement

5402-468: Is made primarily of phospholipids . This membrane encloses the contents of the cell and acts as a barrier to hold nutrients, proteins and other essential components of the cytoplasm within the cell. Unlike eukaryotic cells , bacteria usually lack large membrane-bound structures in their cytoplasm such as a nucleus , mitochondria , chloroplasts and the other organelles present in eukaryotic cells. However, some bacteria have protein-bound organelles in

5548-541: Is more common in Kosher Shechita (where it is all but required) than in Halal Dhabiha (as in most cases, draining alone will suffice). The American Institute for Cancer Research (AICR) and World Cancer Research Fund International (WCRF) have stated that there is strong evidence that consuming salted foods including salt-preserved fish and salt-preserved foods in general increases risk of stomach cancer and that

5694-496: Is motile in liquid or solid media. Several Listeria and Shigella species move inside host cells by usurping the cytoskeleton , which is normally used to move organelles inside the cell. By promoting actin polymerisation at one pole of their cells, they can form a kind of tail that pushes them through the host cell's cytoplasm. A few bacteria have chemical systems that generate light. This bioluminescence often occurs in bacteria that live in association with fish, and

5840-444: Is the stationary phase and is caused by depleted nutrients. The cells reduce their metabolic activity and consume non-essential cellular proteins. The stationary phase is a transition from rapid growth to a stress response state and there is increased expression of genes involved in DNA repair , antioxidant metabolism and nutrient transport . The final phase is the death phase where

5986-440: Is the logarithmic phase , also known as the exponential phase. The log phase is marked by rapid exponential growth . The rate at which cells grow during this phase is known as the growth rate ( k ), and the time it takes the cells to double is known as the generation time ( g ). During log phase, nutrients are metabolised at maximum speed until one of the nutrients is depleted and starts limiting growth. The third phase of growth

Salting (food) - Misplaced Pages Continue

6132-676: Is the plural of the Neo-Latin bacterium , which is the Latinisation of the Ancient Greek βακτήριον ( baktḗrion ), the diminutive of βακτηρία ( baktēría ), meaning "staff, cane", because the first ones to be discovered were rod-shaped . The ancestors of bacteria were unicellular microorganisms that were the first forms of life to appear on Earth, about 4 billion years ago. For about 3 billion years, most organisms were microscopic, and bacteria and archaea were

6278-606: The Oxford English Dictionary , the English name Russia first appeared in the 14th century, borrowed from Medieval Latin : Russia , used in the 11th century and frequently in 12th-century British sources, in turn derived from Russi , 'the Russians'; and the suffix -ia . In modern historiography, this state is usually denoted as Kievan Rus' after its capital city. Another Medieval Latin name for Rus'

6424-543: The Battle of Kursk in the summer of 1943. Another German failure was the Siege of Leningrad , in which the city was fully blockaded on land between 1941 and 1944 by German and Finnish forces, and suffered starvation and more than a million deaths, but never surrendered. Soviet forces steamrolled through Eastern and Central Europe in 1944–1945 and captured Berlin in May 1945. In August 1945,

6570-731: The Brusilov Offensive of the Imperial Russian Army almost completely destroyed the Austro-Hungarian Army . However, the already-existing public distrust of the regime was deepened by the rising costs of war, high casualties , and rumors of corruption and treason. All this formed the climate for the Russian Revolution of 1917, carried out in two major acts. In early 1917, Nicholas II was forced to abdicate ; he and his family were imprisoned and later executed during

6716-470: The Congress of Vienna , which defined the map of post-Napoleonic Europe. The officers who pursued Napoleon into Western Europe brought ideas of liberalism back to Russia, and attempted to curtail the tsar's powers during the abortive Decembrist revolt of 1825. At the end of the conservative reign of Nicholas I (1825–1855), a zenith period of Russia's power and influence in Europe, was disrupted by defeat in

6862-677: The Crimean War . Nicholas's successor Alexander II (1855–1881) enacted significant changes throughout the country, including the emancipation reform of 1861 . These reforms spurred industrialisation, and modernised the Imperial Russian Army, which liberated much of the Balkans from Ottoman rule in the aftermath of the 1877–1878 Russo-Turkish War . During most of the 19th and early 20th century, Russia and Britain colluded over Afghanistan and its neighbouring territories in Central and South Asia;

7008-468: The Gram stain , a long-standing test for the classification of bacterial species. Gram-positive bacteria possess a thick cell wall containing many layers of peptidoglycan and teichoic acids . In contrast, Gram-negative bacteria have a relatively thin cell wall consisting of a few layers of peptidoglycan surrounded by a second lipid membrane containing lipopolysaccharides and lipoproteins . Most bacteria have

7154-574: The Grand Duchy of Moscow led the unification of Russian lands, leading to the proclamation of the Tsardom of Russia in 1547. By the early 18th century, Russia had vastly expanded through conquest, annexation, and the efforts of Russian explorers , developing into the Russian Empire , which remains the third-largest empire in history . However, with the Russian Revolution in 1917, Russia's monarchic rule

7300-600: The Great Northern War (1700–1721), securing Russia's access to the sea and sea trade. In 1703, on the Baltic Sea, Peter founded Saint Petersburg as Russia's new capital. Throughout his rule, sweeping reforms were made , which brought significant Western European cultural influences to Russia. He was succeeded by Catherine I (1725–1727), followed by Peter II (1727–1730), and Anna . The reign of Peter I's daughter Elizabeth in 1741–1762 saw Russia's participation in

7446-672: The Khrushchev Thaw . At the same time, Cold War tensions reached its peak when the two rivals clashed over the deployment of the United States Jupiter missiles in Turkey and Soviet missiles in Cuba . In 1957, the Soviet Union launched the world's first artificial satellite , Sputnik 1 , thus starting the Space Age . Russian cosmonaut Yuri Gagarin became the first human to orbit

SECTION 50

#1733084612414

7592-681: The Oldowan period in the early Lower Paleolithic . About 2 million years ago, representatives of Homo erectus migrated to the Taman Peninsula in southern Russia. Flint tools, some 1.5 million years old, have been discovered in the North Caucasus . Radiocarbon dated specimens from Denisova Cave in the Altai Mountains estimate the oldest Denisovan specimen lived 195–122,700 years ago. Fossils of Denny , an archaic human hybrid that

7738-556: The Rurik dynasty that ruled Kievan Rus' collectively. Kiev's dominance waned, to the benefit of Vladimir-Suzdal in the north-east, the Novgorod Republic in the north, and Galicia-Volhynia in the south-west. By the 12th century, Kiev lost its pre-eminence and Kievan Rus' had fragmented into different principalities. Prince Andrey Bogolyubsky sacked Kiev in 1169 and made Vladimir his base, leading to political power being shifted to

7884-786: The Russian Civil War . The monarchy was replaced by a shaky coalition of political parties that declared itself the Provisional Government , and proclaimed the Russian Republic . On 19 January [ O.S. 6 January], 1918, the Russian Constituent Assembly declared Russia a democratic federal republic (thus ratifying the Provisional Government's decision). The next day the Constituent Assembly

8030-611: The Seven Years' War (1756–1763). During the conflict, Russian troops overran East Prussia , reaching Berlin. However, upon Elizabeth's death, all these conquests were returned to the Kingdom of Prussia by pro-Prussian Peter III of Russia . Catherine II ("the Great"), who ruled in 1762–1796, presided over the Russian Age of Enlightenment . She extended Russian political control over

8176-810: The Siberian River Routes , and by the mid-17th century, there were Russian settlements in eastern Siberia, on the Chukchi Peninsula , along the Amur River , and on the coast of the Pacific Ocean. In 1648, Semyon Dezhnyov became the first European to navigate through the Bering Strait . Under Peter the Great , Russia was proclaimed an empire in 1721, and established itself as one of the European great powers. Ruling from 1682 to 1725, Peter defeated Sweden in

8322-629: The Soviet Union , by joining the Russian SFSR into a single state with the Byelorussian , Transcaucasian , and Ukrainian republics. Eventually internal border changes and annexations during World War II created a union of 15 republics ; the largest in size and population being the Russian SFSR, which dominated the union politically, culturally, and economically. Following Lenin's death in 1924,

8468-601: The gut , though there are many on the skin. Most of the bacteria in and on the body are harmless or rendered so by the protective effects of the immune system , and many are beneficial , particularly the ones in the gut. However, several species of bacteria are pathogenic and cause infectious diseases , including cholera , syphilis , anthrax , leprosy , tuberculosis , tetanus and bubonic plague . The most common fatal bacterial diseases are respiratory infections . Antibiotics are used to treat bacterial infections and are also used in farming, making antibiotic resistance

8614-444: The spirochaetes , are found between two membranes in the periplasmic space. They have a distinctive helical body that twists about as it moves. Two other types of bacterial motion are called twitching motility that relies on a structure called the type IV pilus , and gliding motility , that uses other mechanisms. In twitching motility, the rod-like pilus extends out from the cell, binds some substrate, and then retracts, pulling

8760-496: The vacuum and radiation of outer space , leading to the possibility that bacteria could be distributed throughout the universe by space dust , meteoroids , asteroids , comets , planetoids , or directed panspermia . Endospore-forming bacteria can cause disease; for example, anthrax can be contracted by the inhalation of Bacillus anthracis endospores, and contamination of deep puncture wounds with Clostridium tetani endospores causes tetanus , which, like botulism ,

8906-649: The 10th to 11th centuries, Kievan Rus' became one of the largest and most prosperous states in Europe. The reigns of Vladimir the Great (980–1015) and his son Yaroslav the Wise (1019–1054) constitute the Golden Age of Kiev, which saw the acceptance of Orthodox Christianity from Byzantium , and the creation of the first East Slavic written legal code , the Russkaya Pravda . The age of feudalism and decentralisation had come, marked by constant in-fighting between members of

SECTION 60

#1733084612414

9052-666: The 14th century involved covering the animal with bracken as soon as possible and carrying it to a place where it could be butchered, boiled in brine, and dry salted for long term preservation in a barrel . People in the 14th century could also put salt on vegetables for taste. Salted meat was a staple of the mariner's diet in the Age of Sail . It was stored in barrels, and often had to last for months at sea. The basic Royal Navy diet consisted of salted beef, salted pork , ship's biscuit , and oatmeal , supplemented with smaller quantities of peas, cheese and butter. Even in 1938, Eric Newby found

9198-554: The 16th century, Russia expanded east of the Ural Mountains . However, the Tsardom was weakened by the long and unsuccessful Livonian War against the coalition of the Kingdom of Poland and the Grand Duchy of Lithuania (later the united Polish–Lithuanian Commonwealth ), the Kingdom of Sweden , and Denmark–Norway for access to the Baltic coast and sea trade. In 1572, an invading army of Crimean Tatars were thoroughly defeated in

9344-583: The Byzantine double-headed eagle his own, and eventually Russia's, coat-of-arms. Vasili III united all of Russia by annexing the last few independent Russian states in the early 16th century. In development of the Third Rome ideas, the grand duke Ivan IV ("the Terrible") was officially crowned the first tsar of Russia in 1547. The tsar promulgated a new code of laws ( Sudebnik of 1550 ), established

9490-406: The Earth, aboard the Vostok 1 crewed spacecraft on 12 April 1961 . Following the ousting of Khrushchev in 1964, another period of collective rule ensued, until Leonid Brezhnev became the leader. The era of the 1970s and the early 1980s was later designated as the Era of Stagnation . The 1965 Kosygin reform aimed for partial decentralisation of the Soviet economy . In 1979, after

9636-675: The Gram-negative cell wall, and only members of the Bacillota group and actinomycetota (previously known as the low G+C and high G+C Gram-positive bacteria, respectively) have the alternative Gram-positive arrangement. These differences in structure can produce differences in antibiotic susceptibility; for instance, vancomycin can kill only Gram-positive bacteria and is ineffective against Gram-negative pathogens , such as Haemophilus influenzae or Pseudomonas aeruginosa . Some bacteria have cell wall structures that are neither classically Gram-positive or Gram-negative. This includes clinically important bacteria such as mycobacteria which have

9782-415: The Polish–Lithuanian Commonwealth and annexed most of its territories into Russia , making it the most populous country in Europe. In the south, after the successful Russo-Turkish Wars against the Ottoman Empire , Catherine advanced Russia's boundary to the Black Sea, by dissolving the Crimean Khanate , and annexing Crimea . As a result of victories over Qajar Iran through the Russo-Persian Wars , by

9928-692: The Provisional Government and gave full governing power to the soviets, leading to the creation of the world's first socialist state . The Russian Civil War broke out between the anti-communist White movement and the Bolsheviks with its Red Army . In the aftermath of signing the Treaty of Brest-Litovsk that concluded hostilities with the Central Powers of World War I ; Bolshevist Russia surrendered most of its western territories, which hosted 34% of its population, 54% of its industries, 32% of its agricultural land, and roughly 90% of its coal mines. The Allied powers launched an unsuccessful military intervention in support of anti-communist forces. In

10074-406: The Red Army invaded Manchuria and ousted the Japanese from Northeast Asia, contributing to the Allied victory over Japan. The 1941–1945 period of World War II is known in Russia as the Great Patriotic War . The Soviet Union, along with the United States, the United Kingdom and China were considered the Big Four of Allied powers in World War II, and later became the Four Policemen , which

10220-436: The Russian SFSR emerged from the dissolution of the Soviet Union as the Russian Federation. A new constitution was adopted, which established a federal semi-presidential system . Since the turn of the century, Russia's political system has been dominated by Vladimir Putin , under whom the country has experienced democratic backsliding and become an authoritarian dictatorship . Russia has been militarily involved in

10366-403: The Russian autocratic state. The destruction of Kievan Rus' saw the eventual rise of the Grand Duchy of Moscow , initially a part of Vladimir-Suzdal . While still under the domain of the Mongol - Tatars and with their connivance, Moscow began to assert its influence in the region in the early 14th century, gradually becoming the leading force in the "gathering of the Russian lands". When

10512-476: The Russian parliament culminated in a constitutional crisis which ended violently through military force. During the crisis, Yeltsin was backed by Western governments, and over 100 people were killed. In December, a referendum was held and approved, which introduced a new constitution, giving the president enormous powers. The 1990s were plagued by armed conflicts in the North Caucasus , both local ethnic skirmishes and separatist Islamist insurrections. From

10658-408: The Soviet Union led Russia into a deep and prolonged depression. During and after the disintegration of the Soviet Union, wide-ranging reforms including privatisation and market and trade liberalisation were undertaken, including radical changes along the lines of " shock therapy ". The privatisation largely shifted control of enterprises from state agencies to individuals with inside connections in

10804-571: The Soviet Union. The transitional disorganisation of the country's agriculture, combined with the harsh state policies and a drought, led to the Soviet famine of 1932–1933 ; which killed 5.7 to 8.7 million, 3.3 million of them in the Russian SFSR. The Soviet Union, ultimately, made the costly transformation from a largely agrarian economy to a major industrial powerhouse within a short span of time. The Soviet Union entered World War II on 17 September 1939 with its invasion of Poland , in accordance with

10950-455: The Soviet system, introduced the policies of glasnost (openness) and perestroika (restructuring) in an attempt to end the period of economic stagnation and to democratise the government . This, however, led to the rise of strong nationalist and separatist movements across the country. Prior to 1991, the Soviet economy was the world's second-largest, but during its final years, it went into

11096-594: The Ukrainian leader, Bohdan Khmelnytsky , offered to place Ukraine under the protection of the Russian tsar, Alexis ; whose acceptance of this offer led to another Russo-Polish War . Ultimately, Ukraine was split along the Dnieper , leaving the eastern part, ( Left-bank Ukraine and Kiev ) under Russian rule. In the east, the rapid Russian exploration and colonisation of vast Siberia continued, hunting for valuable furs and ivory. Russian explorers pushed eastward primarily along

11242-801: The Volga-Dnieper region of southern Russia and Ukraine as the urheimat of the Proto-Indo-Europeans . Early Indo-European migrations from the Pontic–Caspian steppe of Ukraine and Russia spread Yamnaya ancestry and Indo-European languages across large parts of Eurasia. Nomadic pastoralism developed in the Pontic–Caspian steppe beginning in the Chalcolithic . Remnants of these steppe civilizations were discovered in places such as Ipatovo , Sintashta , Arkaim , and Pazyryk , which bear

11388-405: The absence of refrigeration . In more recent times, freeze-drying , water binding humectants , and fully automated equipment with temperature and humidity control have been added. Often a combination of these techniques is used. Jewish and Muslim dietary laws require the removal of blood from freshly slaughtered meat. Salt and brine are used for the purpose in both traditions, but salting

11534-502: The air, soil, water, acidic hot springs , radioactive waste , and the deep biosphere of Earth's crust . Bacteria play a vital role in many stages of the nutrient cycle by recycling nutrients and the fixation of nitrogen from the atmosphere . The nutrient cycle includes the decomposition of dead bodies ; bacteria are responsible for the putrefaction stage in this process. In the biological communities surrounding hydrothermal vents and cold seeps , extremophile bacteria provide

11680-425: The archaeal/eukaryotic lineage. The most recent common ancestor (MRCA) of bacteria and archaea was probably a hyperthermophile that lived about 2.5 billion–3.2 billion years ago. The earliest life on land may have been bacteria some 3.22 billion years ago. Bacteria were also involved in the second great evolutionary divergence, that of the archaea and eukaryotes. Here, eukaryotes resulted from

11826-455: The atmosphere and one cubic metre of air holds around one hundred million bacterial cells. The oceans and seas harbour around 3 x 10 bacteria which provide up to 50% of the oxygen humans breathe. Only around 2% of bacterial species have been fully studied. Size . Bacteria display a wide diversity of shapes and sizes. Bacterial cells are about one-tenth the size of eukaryotic cells and are typically 0.5–5.0  micrometres in length. However,

11972-520: The bacteria have come into contact with in the past, which allows them to block virus replication through a form of RNA interference . Third, bacteria can transfer genetic material through direct cell contact via conjugation . In ordinary circumstances, transduction, conjugation, and transformation involve transfer of DNA between individual bacteria of the same species, but occasionally transfer may occur between individuals of different bacterial species, and this may have significant consequences, such as

12118-444: The bacteria perform separate tasks; for example, about one in ten cells migrate to the top of a fruiting body and differentiate into a specialised dormant state called a myxospore, which is more resistant to drying and other adverse environmental conditions. Biofilms . Bacteria often attach to surfaces and form dense aggregations called biofilms and larger formations known as microbial mats . These biofilms and mats can range from

12264-401: The bacteria run out of nutrients and die. Most bacteria have a single circular chromosome that can range in size from only 160,000 base pairs in the endosymbiotic bacteria Carsonella ruddii , to 12,200,000 base pairs (12.2 Mbp) in the soil-dwelling bacteria Sorangium cellulosum . There are many exceptions to this; for example, some Streptomyces and Borrelia species contain

12410-412: The bacterial chromosome, introducing foreign DNA in a process known as transduction . Many types of bacteriophage exist; some infect and lyse their host bacteria, while others insert into the bacterial chromosome. Bacteria resist phage infection through restriction modification systems that degrade foreign DNA and a system that uses CRISPR sequences to retain fragments of the genomes of phage that

12556-576: The capital Moscow. In 1612, the Poles were forced to retreat by the Russian volunteer corps, led by merchant Kuzma Minin and prince Dmitry Pozharsky . The Romanov dynasty acceded to the throne in 1613 by the decision of the Zemsky Sobor, and the country started its gradual recovery from the crisis. Russia continued its territorial growth through the 17th century, which was the age of the Cossacks . In 1654,

12702-437: The cell forward. Motile bacteria are attracted or repelled by certain stimuli in behaviours called taxes : these include chemotaxis , phototaxis , energy taxis , and magnetotaxis . In one peculiar group, the myxobacteria, individual bacteria move together to form waves of cells that then differentiate to form fruiting bodies containing spores. The myxobacteria move only when on solid surfaces, unlike E. coli , which

12848-576: The class Schizomycetes ("fission fungi"), bacteria are now classified as prokaryotes . Unlike cells of animals and other eukaryotes , bacterial cells do not contain a nucleus and rarely harbour membrane -bound organelles . Although the term bacteria traditionally included all prokaryotes, the scientific classification changed after the discovery in the 1990s that prokaryotes consist of two very different groups of organisms that evolved from an ancient common ancestor . These evolutionary domains are called Bacteria and Archaea . The word bacteria

12994-530: The control of the Golden Horde and consolidated the whole of northern Rus' under Moscow's dominion, and was the first Russian ruler to take the title "Grand Duke of all Rus'". After the fall of Constantinople in 1453, Moscow claimed succession to the legacy of the Eastern Roman Empire . Ivan III married Sophia Palaiologina , the niece of the last Byzantine emperor Constantine XI , and made

13140-834: The country was more often referred to by its inhabitants as Rus ' , the Russian land ( Russkaia zemlia ), or the Muscovite state ( Moskovskoe gosudarstvo ), among other variations. In 1721, Peter the Great changed the name of the state from Tsardom of Russia ( Russian : Русское царство , romanized :  Russkoye tsarstvo ) or Tsardom of Muscovy ( Russian : Московское царство , romanized :  Moskovskoye tsarstvo ) to Russian Empire ( Rossiiskaia imperiia ). There are several words in Russian which translate to "Russians" in English. The noun and adjective русский , russkiy refers to ethnic Russians . The adjective российский , rossiiskiy denotes Russian citizens regardless of ethnicity. The same applies to

13286-446: The crucial Battle of Molodi . The death of Ivan's sons marked the end of the ancient Rurik dynasty in 1598, and in combination with the disastrous famine of 1601–1603 , led to a civil war, the rule of pretenders, and foreign intervention during the Time of Troubles in the early 17th century. The Polish–Lithuanian Commonwealth , taking advantage, occupied parts of Russia, extending into

13432-629: The cytoplasm of the cell; generally, a single endospore develops in each cell. Each endospore contains a core of DNA and ribosomes surrounded by a cortex layer and protected by a multilayer rigid coat composed of peptidoglycan and a variety of proteins. Endospores show no detectable metabolism and can survive extreme physical and chemical stresses, such as high levels of UV light , gamma radiation , detergents , disinfectants , heat, freezing, pressure, and desiccation . In this dormant state, these organisms may remain viable for millions of years. Endospores even allow bacteria to survive exposure to

13578-419: The cytoplasm which compartmentalise aspects of bacterial metabolism, such as the carboxysome . Additionally, bacteria have a multi-component cytoskeleton to control the localisation of proteins and nucleic acids within the cell, and to manage the process of cell division . Many important biochemical reactions, such as energy generation, occur due to concentration gradients across membranes, creating

13724-550: The diet on the tall ship Moshulu to consist almost entirely of salted meat. Moshulu's lack of refrigeration left little choice as the ship made voyages which could exceed 100 days passage between ports. It was discovered in the 19th century that salt mixed with nitrates (such as saltpeter ) would color meats red, rather than grey, and consumers at that time then strongly preferred the red-colored meat. The food hence preserved stays healthy and fresh for days avoiding bacterial decay. Salting, either with dry salt or brine ,

13870-411: The dominant forms of life. Although bacterial fossils exist, such as stromatolites , their lack of distinctive morphology prevents them from being used to examine the history of bacterial evolution, or to date the time of origin of a particular bacterial species. However, gene sequences can be used to reconstruct the bacterial phylogeny , and these studies indicate that bacteria diverged first from

14016-559: The earliest known traces of horses in warfare . The genetic makeup of speakers of the Uralic language family in northern Europe was shaped by migration from Siberia that began at least 3,500 years ago. In the 3rd to 4th centuries CE, the Gothic kingdom of Oium existed in southern Russia, which was later overrun by Huns . Between the 3rd and 6th centuries CE, the Bosporan Kingdom , which

14162-526: The elongated filaments of Actinomycetota species, the aggregates of Myxobacteria species, and the complex hyphae of Streptomyces species. These multicellular structures are often only seen in certain conditions. For example, when starved of amino acids, myxobacteria detect surrounding cells in a process known as quorum sensing , migrate towards each other, and aggregate to form fruiting bodies up to 500 micrometres long and containing approximately 100,000 bacterial cells. In these fruiting bodies,

14308-501: The energy released by the transfer of ions down an electrochemical gradient across the cell membrane. Fimbriae (sometimes called " attachment pili ") are fine filaments of protein, usually 2–10 nanometres in diameter and up to several micrometres in length. They are distributed over the surface of the cell, and resemble fine hairs when seen under the electron microscope . Fimbriae are believed to be involved in attachment to solid surfaces or to other cells, and are essential for

14454-597: The entering of ancient bacteria into endosymbiotic associations with the ancestors of eukaryotic cells, which were themselves possibly related to the Archaea. This involved the engulfment by proto-eukaryotic cells of alphaproteobacterial symbionts to form either mitochondria or hydrogenosomes , which are still found in all known Eukarya (sometimes in highly reduced form , e.g. in ancient "amitochondrial" protozoa). Later, some eukaryotes that already contained mitochondria also engulfed cyanobacteria -like organisms, leading to

14600-641: The evolution of different growth strategies (see r/K selection theory ). Some organisms can grow extremely rapidly when nutrients become available, such as the formation of algal and cyanobacterial blooms that often occur in lakes during the summer. Other organisms have adaptations to harsh environments, such as the production of multiple antibiotics by Streptomyces that inhibit the growth of competing microorganisms. In nature, many organisms live in communities (e.g., biofilms ) that may allow for increased supply of nutrients and protection from environmental stresses. These relationships can be essential for growth of

14746-474: The first Europeans to reach and colonise Alaska . In 1803–1806, the first Russian circumnavigation was made. In 1820, a Russian expedition discovered the continent of Antarctica . During the Napoleonic Wars , Russia joined alliances with various European powers, and fought against France. The French invasion of Russia at the height of Napoleon's power in 1812 reached Moscow, but eventually failed as

14892-549: The first Russian feudal representative body (the Zemsky Sobor ), revamped the military, curbed the influence of the clergy, and reorganised local government. During his long reign, Ivan nearly doubled the already large Russian territory by annexing the three Tatar khanates: Kazan and Astrakhan along the Volga , and the Khanate of Sibir in southwestern Siberia. Ultimately, by the end of

15038-571: The first half of the 19th century, Russia also conquered the Caucasus . Catherine's successor, her son Paul , was unstable and focused predominantly on domestic issues . Following his short reign, Catherine's strategy was continued with Alexander I's (1801–1825) wresting of Finland from the weakened Sweden in 1809, and of Bessarabia from the Ottomans in 1812. In North America, the Russians became

15184-566: The formation of chloroplasts in algae and plants. This is known as primary endosymbiosis . Bacteria are ubiquitous, living in every possible habitat on the planet including soil, underwater, deep in Earth's crust and even such extreme environments as acidic hot springs and radioactive waste. There are thought to be approximately 2×10 bacteria on Earth, forming a biomass that is only exceeded by plants. They are abundant in lakes and oceans, in arctic ice, and geothermal springs where they provide

15330-485: The government, which led to the rise of Russian oligarchs . Many of the newly rich moved billions in cash and assets outside of the country in an enormous capital flight . The depression of the economy led to the collapse of social services—the birth rate plummeted while the death rate skyrocketed, and millions plunged into poverty; while extreme corruption, as well as criminal gangs and organised crime rose significantly. In late 1993, tensions between Yeltsin and

15476-515: The growth in cell population. Russia Russia , or the Russian Federation , is a country spanning Eastern Europe and North Asia . It is the largest country in the world by area , extending across eleven time zones and sharing land borders with fourteen countries . It is the world's ninth-most populous country and Europe's most populous country . Russia is a highly urbanised country including 16 population centres with over

15622-421: The high salt content of processed meat may result in damage to the stomach mucosal lining. The American Cancer Society have stated that "stomach cancer risk is increased in people whose diets include large amounts of foods preserved by salting, such as salted fish and meat and pickled vegetables." Additionally, an excessive intake of salt has a dose-response relationship with elevated blood pressure , increasing

15768-405: The large corns or grains of salt used (see wiktionary:corn ). Corned beef retains this name, although it is typically brined today. Salt inhibits the growth of microorganisms by drawing water out of microbial cells through osmosis . Concentrations of salt up to 20% are required to kill most species of unwanted bacteria . Smoking , often used in the process of curing meat, adds chemicals to

15914-627: The largest viruses . Some bacteria may be even smaller, but these ultramicrobacteria are not well-studied. Shape . Most bacterial species are either spherical, called cocci ( singular coccus , from Greek kókkos , grain, seed), or rod-shaped, called bacilli ( sing . bacillus, from Latin baculus , stick). Some bacteria, called vibrio , are shaped like slightly curved rods or comma-shaped; others can be spiral-shaped, called spirilla , or tightly coiled, called spirochaetes . A small number of other unusual shapes have been described, such as star-shaped bacteria. This wide variety of shapes

16060-525: The latter deliberately starved to death or otherwise killed 3.3 million Soviet POWs , and a vast number of civilians, as the " Hunger Plan " sought to fulfil Generalplan Ost . Although the Wehrmacht had considerable early success, their attack was halted in the Battle of Moscow . Subsequently, the Germans were dealt major defeats first at the Battle of Stalingrad in the winter of 1942–1943, and then in

16206-716: The light probably serves to attract fish or other large animals. Bacteria often function as multicellular aggregates known as biofilms , exchanging a variety of molecular signals for intercell communication and engaging in coordinated multicellular behaviour. The communal benefits of multicellular cooperation include a cellular division of labour , accessing resources that cannot effectively be used by single cells, collectively defending against antagonists, and optimising population survival by differentiating into distinct cell types. For example, bacteria in biofilms can have more than five hundred times increased resistance to antibacterial agents than individual "planktonic" bacteria of

16352-669: The meantime, both the Bolsheviks and White movement carried out campaigns of deportations and executions against each other, known respectively as the Red Terror and White Terror . By the end of the violent civil war, Russia's economy and infrastructure were heavily damaged, and as many as 10 million perished during the war, mostly civilians. Millions became White émigrés , and the Russian famine of 1921–1922 claimed up to five million victims. On 30 December 1922, Lenin and his aides formed

16498-596: The more recently coined noun россиянин , rossiianyn , "Russian" in the sense of citizen of the Russian state. According to the Primary Chronicle , the word Rus' is derived from the Rus' people , who were a Swedish tribe, and where the three original members of the Rurikid dynasty came from. The Finnish word for Swedes, ruotsi , has the same origin. Later archeological studies mostly confirmed this theory. The first human settlement on Russia dates back to

16644-417: The newly formed daughter cells. Examples include fruiting body formation by myxobacteria and aerial hyphae formation by Streptomyces species, or budding. Budding involves a cell forming a protrusion that breaks away and produces a daughter cell. In the laboratory, bacteria are usually grown using solid or liquid media. Solid growth media , such as agar plates , are used to isolate pure cultures of

16790-625: The north-east. Led by Prince Alexander Nevsky , Novgorodians repelled the invading Swedes in the Battle of the Neva in 1240, as well as the Germanic crusaders in the Battle on the Ice in 1242. Kievan Rus' finally fell to the Mongol invasion of 1237–1240, which resulted in the sacking of Kiev and other cities, as well as the death of a major part of the population. The invaders, later known as Tatars , formed

16936-473: The nutrients needed to sustain life by converting dissolved compounds, such as hydrogen sulphide and methane , to energy. They live on and in plants and animals. Most do not cause diseases, are beneficial to their environments, and are essential for life. The soil is a rich source of bacteria and a few grams contain around a thousand million of them. They are all essential to soil ecology, breaking down toxic waste and recycling nutrients. They are even found in

17082-517: The nutrients needed to sustain life by converting dissolved compounds, such as hydrogen sulphide and methane , to energy. Bacteria also live in mutualistic , commensal and parasitic relationships with plants and animals. Most bacteria have not been characterised and there are many species that cannot be grown in the laboratory. The study of bacteria is known as bacteriology , a branch of microbiology . Like all animals, humans carry vast numbers (approximately 10 to 10 ) of bacteria. Most are in

17228-601: The obstinate resistance in combination with the bitterly cold Russian winter led to a disastrous defeat of invaders, in which the pan-European Grande Armée faced utter destruction. Led by Mikhail Kutuzov and Michael Andreas Barclay de Tolly , the Imperial Russian Army ousted Napoleon and drove throughout Europe in the War of the Sixth Coalition , ultimately entering Paris. Alexander I controlled Russia's delegation at

17374-643: The official line. The continued internal struggle in the Bolshevik party culminated in the Great Purge . Under Stalin's leadership, the government launched a command economy , industrialisation of the largely rural country , and collectivisation of its agriculture . During this period of rapid economic and social change, millions of people were sent to penal labour camps , including many political convicts for their suspected or real opposition to Stalin's rule; and millions were deported and exiled to remote areas of

17520-475: The outside of the cell membrane is the cell wall . Bacterial cell walls are made of peptidoglycan (also called murein), which is made from polysaccharide chains cross-linked by peptides containing D- amino acids . Bacterial cell walls are different from the cell walls of plants and fungi , which are made of cellulose and chitin , respectively. The cell wall of bacteria is also distinct from that of achaea, which do not contain peptidoglycan. The cell wall

17666-407: The press ; the country also has high levels of perceived corruption . As of 2024, Russia has a high-income economy which ranks eleventh in the world by nominal GDP and fourth at purchasing power parity , relying on its vast mineral and energy resources ; the world's second-largest for oil production and natural gas production . Russia is home to 32 UNESCO World Heritage Sites . According to

17812-612: The production of the highly toxic forms of mercury ( methyl- and dimethylmercury ) in the environment. Nonrespiratory anaerobes use fermentation to generate energy and reducing power, secreting metabolic by-products (such as ethanol in brewing) as waste. Facultative anaerobes can switch between fermentation and different terminal electron acceptors depending on the environmental conditions in which they find themselves. Unlike in multicellular organisms, increases in cell size ( cell growth ) and reproduction by cell division are tightly linked in unicellular organisms. Bacteria grow to

17958-560: The replication of DNA or from exposure to mutagens . Mutation rates vary widely among different species of bacteria and even among different clones of a single species of bacteria. Genetic changes in bacterial genomes emerge from either random mutation during replication or "stress-directed mutation", where genes involved in a particular growth-limiting process have an increased mutation rate. Some bacteria transfer genetic material between cells. This can occur in three main ways. First, bacteria can take up exogenous DNA from their environment in

18104-492: The risk of several cardiovascular diseases . Bacteria See § Phyla Bacteria ( / b æ k ˈ t ɪər i ə / ; sg. : bacterium) are ubiquitous, mostly free-living organisms often consisting of one biological cell . They constitute a large domain of prokaryotic microorganisms . Typically a few micrometres in length, bacteria were among the first life forms to appear on Earth , and are present in most of its habitats . Bacteria inhabit

18250-573: The rivalry between the two major European empires came to be known as the Great Game . The late 19th century saw the rise of various socialist movements in Russia. Alexander II was assassinated in 1881 by revolutionary terrorists. The reign of his son Alexander III (1881–1894) was less liberal but more peaceful. Under last Russian emperor, Nicholas II (1894–1917), the Revolution of 1905

18396-530: The same species. One type of intercellular communication by a molecular signal is called quorum sensing , which serves the purpose of determining whether the local population density is sufficient to support investment in processes that are only successful if large numbers of similar organisms behave similarly, such as excreting digestive enzymes or emitting light. Quorum sensing enables bacteria to coordinate gene expression and to produce, release, and detect autoinducers or pheromones that accumulate with

18542-845: The seat of the Metropolitan of the Russian Orthodox Church moved to Moscow in 1325, its influence increased. Moscow's last rival, the Novgorod Republic , prospered as the chief fur trade centre and the easternmost port of the Hanseatic League . Led by Prince Dmitry Donskoy of Moscow, the united army of Russian principalities inflicted a milestone defeat on the Mongol-Tatars in the Battle of Kulikovo in 1380. Moscow gradually absorbed its parent duchy and surrounding principalities, including formerly strong rivals such as Tver and Novgorod . Ivan III ("the Great") threw off

18688-575: The state of the Golden Horde , which ruled over Russia for the next two centuries. Only the Novgorod Republic escaped foreign occupation after it agreed to pay tribute to the Mongols. Galicia-Volhynia would later be absorbed by Lithuania and Poland , while the Novgorod Republic continued to prosper in the north. In the northeast, the Byzantine-Slavic traditions of Kievan Rus' were adapted to form

18834-554: The surface of meat that reduce the concentration of salt required. Various types of salted meat are staples of the diets of people in North Africa , Southern China , Scandinavia , coastal Russia , and in the Arctic . Some of those salted meats (or foods that contain salted meat) are bacon , biltong , cecina , corned beef , ham , jamón , jerky , pastrami , and salt pork . Drying or salting, either with dry salt or with brine ,

18980-412: The surfaces of plants, the majority of bacteria are bound to surfaces in biofilms. Biofilms are also important in medicine, as these structures are often present during chronic bacterial infections or in infections of implanted medical devices , and bacteria protected within biofilms are much harder to kill than individual isolated bacteria. The bacterial cell is surrounded by a cell membrane , which

19126-457: The terminal electron acceptor, while anaerobic organisms use other compounds such as nitrate , sulfate , or carbon dioxide. Many bacteria, called heterotrophs , derive their carbon from other organic carbon . Others, such as cyanobacteria and some purple bacteria , are autotrophic , meaning they obtain cellular carbon by fixing carbon dioxide . In unusual circumstances, the gas methane can be used by methanotrophic bacteria as both

19272-523: The time Chechen separatists declared independence in the early 1990s, an intermittent guerrilla war was fought between the rebel groups and Russian forces. Terrorist attacks against civilians were carried out by Chechen separatists, claiming the lives of thousands of Russian civilians. After the dissolution of the Soviet Union, Russia assumed responsibility for settling the latter's external debts. In 1992, most consumer price controls were eliminated, causing extreme inflation and significantly devaluing

19418-447: The transfer of antibiotic resistance. In such cases, gene acquisition from other bacteria or the environment is called horizontal gene transfer and may be common under natural conditions. Many bacteria are motile (able to move themselves) and do so using a variety of mechanisms. The best studied of these are flagella , long filaments that are turned by a motor at the base to generate propeller-like movement. The bacterial flagellum

19564-495: The virulence of some bacterial pathogens. Pili ( sing . pilus) are cellular appendages, slightly larger than fimbriae, that can transfer genetic material between bacterial cells in a process called conjugation where they are called conjugation pili or sex pili (see bacterial genetics, below). They can also generate movement where they are called type IV pili . Glycocalyx is produced by many bacteria to surround their cells, and varies in structural complexity: ranging from

19710-504: Was Ruthenia . In Russian, the current name of the country, Россия ( Rossiya ), comes from the Byzantine Greek name for Rus', Ρωσία ( Rosía ). A new form of the name Rus ' , Росия ( Rosiya ), was borrowed from the Greek term and first attested in 1387. The name Rossiia appeared in Russian sources in the late 15th century, but until the end of the 17th century

19856-449: Was elected President of the Russian SFSR. In August 1991, a coup d'état attempt by members of Gorbachev's government, directed against Gorbachev and aimed at preserving the Soviet Union, instead led to the end of the Communist Party of the Soviet Union. On 25 December 1991, following the dissolution of the Soviet Union , along with contemporary Russia, fourteen other post-Soviet states emerged. The economic and political collapse of

20002-613: Was a Hellenistic polity that succeeded the Greek colonies, was also overwhelmed by nomadic invasions led by warlike tribes such as the Huns and Eurasian Avars . The Khazars , who were of Turkic origin , ruled the steppes between the Caucasus in the south, to the east past the Volga river basin, and west as far as Kyiv on the Dnieper river until the 10th century. After them came the Pechenegs who created

20148-447: Was a common method of preserving meat until the middle of the 20th century, becoming less popular after the advent of refrigeration. Meat that had been preserved in this way was frequently called "junk" or "salt horse". One early method of salt-curing meat was corning , or applying large, coarse pellets of salt, which were rubbed into the meat to keep it from spoiling and to preserve it. This term originates from Old English and references

20294-415: Was abolished and eventually replaced by the Russian SFSR —the world's first constitutionally socialist state . Following the Russian Civil War , the Russian SFSR established the Soviet Union with three other Soviet republics , within which it was the largest and principal constituent. At the expense of millions of lives , the Soviet Union underwent rapid industrialisation in the 1930s and later played

20440-639: Was dissolved by the All-Russian Central Executive Committee . An alternative socialist establishment co-existed, the Petrograd Soviet , wielding power through the democratically elected councils of workers and peasants, called soviets . The rule of the new authorities only aggravated the crisis in the country instead of resolving it, and eventually, the October Revolution , led by Bolshevik leader Vladimir Lenin , overthrew

20586-507: Was found at Kostyonki–Borshchyovo , and at Sungir , dating back to 34,600 years ago—both in western Russia . Humans reached Arctic Russia at least 40,000 years ago, in Mamontovaya Kurya . Ancient North Eurasian populations from Siberia genetically similar to Mal'ta–Buret' culture and Afontova Gora were an important genetic contributor to Ancient Native Americans and Eastern Hunter-Gatherers . The Kurgan hypothesis places

20732-568: Was half Neanderthal and half Denisovan, and lived some 90,000 years ago, was also found within the latter cave. Russia was home to some of the last surviving Neanderthals, from about 45,000 years ago, found in Mezmaiskaya cave . The first trace of an early modern human in Russia dates back to 45,000 years, in Western Siberia . The discovery of high concentration cultural remains of anatomically modern humans , from at least 40,000 years ago,

20878-567: Was populated by Finno-Ugrian peoples. From the 7th century onwards, the incoming East Slavs slowly assimilated the native Finno-Ugrians. The establishment of the first East Slavic states in the 9th century coincided with the arrival of Varangians , the Vikings who ventured along the waterways extending from the eastern Baltic to the Black and Caspian Seas. According to the Primary Chronicle ,

21024-474: Was the foundation of the United Nations Security Council . During the war, Soviet civilian and military death were about 26–27 million , accounting for about half of all World War II casualties . The Soviet economy and infrastructure suffered massive devastation, which caused the Soviet famine of 1946–1947 . However, at the expense of a large sacrifice, the Soviet Union emerged as

21170-488: Was the only widely available method of preserving fish until the 19th century. Dried fish and salted fish (or fish both dried and salted) are a staple of diets in the Caribbean , West Africa , North Africa , South Asia , Southeast Asia , Southern China , Scandinavia , parts of Canada including Newfoundland , coastal Russia , and in the Arctic . Like other salt-cured meats , it provides preserved animal protein even in

21316-681: Was triggered by the humiliating failure of the Russo-Japanese War . The uprising was put down, but the government was forced to concede major reforms ( Russian Constitution of 1906 ), including granting freedoms of speech and assembly , the legalisation of political parties, and the creation of an elected legislative body, the State Duma . In 1914, Russia entered World War I in response to Austria-Hungary 's declaration of war on Russia's ally Serbia , and fought across multiple fronts while isolated from its Triple Entente allies. In 1916,

#413586