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128-461: The Sacramento NWR Complex was created in an attempt to resolve the conflict between the needs of migrating birds using the Pacific Flyway , and those of agriculture . The Sacramento NWR Complex headquarters and visitor center is located in the 10,819-acre (4378 ha) Sacramento National Wildlife Refuge off of 99W, and features a wildlife exhibit, bookstore, and Discovery Room. Visitors can enjoy

256-494: A white stork was found in the German state of Mecklenburg with an arrow made from central African hardwood, which provided some of the earliest evidence of long-distance stork migration. This bird was referred to as a Pfeilstorch , German for "Arrow stork". Since then, around 25 Pfeilstörche have been documented. Migration is the regular seasonal movement, often north and south, undertaken by many species of birds. Migration

384-536: A 'strongman' was often the best solution. Athens fell under a tyranny in the second half of the 6th century BC. When this tyranny was ended, the Athenians founded the world's first democracy as a radical solution to prevent the aristocracy regaining power. A citizens' assembly (the Ecclesia ), for the discussion of city policy, had existed since the reforms of Draco in 621 BC; all citizens were permitted to attend after

512-1235: A barrier, and detours avoiding such barriers are observed. For example, brent geese Branta bernicla bernicla migrating between the Taymyr Peninsula and the Wadden Sea travel via low-lying coastal feeding-areas on the White Sea and the Baltic Sea rather than directly across the Arctic Ocean and the Scandinavian mainland. Great snipes make non-stop flights of 4,000–7,000 km, lasting 60–90 h, during which they change their average cruising heights from 2,000 m (above sea level) at night to around 4,000 m during daytime. A similar situation occurs with waders (called shorebirds in North America). Many species, such as dunlin Calidris alpina and western sandpiper Calidris mauri , undertake long movements from their Arctic breeding grounds to warmer locations in

640-616: A chick on the Farne Islands in Northumberland off the British east coast, reached Melbourne , Australia in just three months from fledging, a sea journey of over 22,000 km (12,000 nmi), while another also from the Farne Islands with a light level geolocator tag 'G82' covered a staggering 96,000 km (52,000 nmi) in just 10 months from the end of one breeding season to

768-495: A coalition of 31 Greek city states, including Athens and Sparta, determined to resist the Persian invaders. At the same time, Greek Sicily was invaded by a Carthaginian force. In 480 BC, the first major battle of the invasion was fought at Thermopylae , where a small rearguard of Greeks, led by three hundred Spartans, held a crucial pass guarding the heart of Greece for several days; at the same time Gelon , tyrant of Syracuse, defeated

896-596: A council of elders (the Gerousia ) and magistrates specifically appointed to watch over the kings (the Ephors ). Only free, land-owning, native born men could be citizens entitled to the full protection of the law in a city-state. In most city-states, unlike the situation in Rome , social prominence did not allow special rights. Sometimes families controlled public religious functions, but this ordinarily did not give any extra power in

1024-514: A decisive victory, and in 447 lost Boeotia again. Athens and Sparta signed the Thirty Years' Peace in the winter of 446/5, ending the conflict. Despite the treaty, Athenian relations with Sparta declined again in the 430s, and in 431 BC the Peloponnesian War began. The first phase of the war saw a series of fruitless annual invasions of Attica by Sparta, while Athens successfully fought

1152-609: A good view of the birds while minimising the risk of disturbance to them. There is a visitor center at the Sacramento NWR , and routes for car tours on the Sacramento and Colusa sites, though visitors are allowed to leave their cars only at selected sites. There are walking trails at both those sites and also at the Sacramento River site. Limited hunting is permitted on some of the sites. Over 300 species of birds and mammals use

1280-436: A group of city-states allied themselves to defend Greece, the vast majority of poleis remained neutral, and after the Persian defeat, the allies quickly returned to infighting. Thus, the major peculiarities of the ancient Greek political system were its fragmented nature (and that this does not particularly seem to have tribal origin), and the particular focus on urban centers within otherwise tiny states. The peculiarities of

1408-483: A man's story about swallows being found in a chalk cliff collapse "while he was a schoolboy at Brighthelmstone", though the man denied being an eyewitness. However, he writes that "as to swallows being found in a torpid state during the winter in the Isle of Wight or any part of this country, I never heard any such account worth attending to", and that if early swallows "happen to find frost and snow they immediately withdraw for

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1536-467: A powerful influence on ancient Rome , which carried a version of it throughout the Mediterranean and much of Europe. For this reason, Classical Greece is generally considered the cradle of Western civilization , the seminal culture from which the modern West derives many of its founding archetypes and ideas in politics, philosophy, science, and art. Classical antiquity in the Mediterranean region

1664-745: A result of Epaminondas ' liberation of Messenia from Spartan rule, the helot system there came to an end and the helots won their freedom. However, it did continue to persist in Laconia until the 2nd century BC. For most of Greek history, education was private, except in Sparta. During the Hellenistic period, some city-states established public schools . Only wealthy families could afford a teacher. Boys learned how to read, write and quote literature. They also learned to sing and play one musical instrument and were trained as athletes for military service. They studied not for

1792-480: A seasonal movement of birds between breeding and wintering grounds that occurs twice a year. It is typically from north to south or from south to north. Migration is inherently risky, due to predation and mortality. The Arctic tern holds the long-distance migration record for birds, travelling between Arctic breeding grounds and the Antarctic each year. Some species of tubenoses , such as albatrosses , circle

1920-465: A six-mile auto tour with 3 viewing areas and two walking trails. Before the arrival of European settlers in the 19th century, much of the Sacramento valley was taken up by seasonal wetlands and grasslands . By the beginning of the 20th century, much of this had been replaced by farmland, particularly for the growing of rice , and the rivers no longer create new wetlands because their flow is controlled by levees and irrigation schemes. Less than 10% of

2048-516: A special type of slaves called helots . Helots were Messenians enslaved en masse during the Messenian Wars by the state and assigned to families where they were forced to stay. Helots raised food and did household chores so that women could concentrate on raising strong children while men could devote their time to training as hoplites . Their masters treated them harshly, and helots revolted against their masters several times. In 370/69 BC, as

2176-414: A time—a circumstance this much more in favour of hiding than migration", since he doubts they would "return for a week or two to warmer latitudes". Only at the end of the eighteenth century was migration accepted as an explanation for the winter disappearance of birds from northern climes. Thomas Bewick 's A History of British Birds (Volume 1, 1797) mentions a report from "a very intelligent master of

2304-464: A timing mechanism, instead moving in response to local weather conditions. Thus mountain and moorland breeders, such as wallcreeper Tichodroma muraria and white-throated dipper Cinclus cinclus , may move only altitudinally to escape the cold higher ground. Other species such as merlin Falco columbarius and Eurasian skylark Alauda arvensis move further, to the coast or towards the south. Species like

2432-555: A vessel" who, "between the islands of Menorca and Majorca , saw great numbers of Swallows flying northward", and states the situation in Britain as follows: Swallows frequently roost at night, after they begin to congregate, by the sides of rivers and pools, from which circumstance it has been erroneously supposed that they retire into the water. Bewick then describes an experiment that succeeded in keeping swallows alive in Britain for several years, where they remained warm and dry through

2560-690: A war with the Roman Republic in the late 3rd century. Although the First Macedonian War was inconclusive, the Romans, in typical fashion, continued to fight Macedon until it was completely absorbed into the Roman Republic (by 149 BC). In the east, the unwieldy Seleucid Empire gradually disintegrated, although a rump survived until 64 BC, whilst the Ptolemaic Kingdom continued in Egypt until 30 BC when it too

2688-669: A year, the Thirty had been overthrown. The first half of the fourth century saw the major Greek states attempt to dominate the mainland; none were successful, and their resulting weakness led to a power vacuum which would eventually be filled by Macedon under Philip II and then Alexander the Great. In the immediate aftermath of the Peloponnesian war, Sparta attempted to extend their own power, leading Argos, Athens, Corinth, and Thebes to join against them. Aiming to prevent any single Greek state gaining

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2816-627: Is a much more complex phenomenon that may include both endogenous programs as well as learning. The primary physiological cue for migration is the changes in the day length. These changes are related to hormonal changes in the birds. In the period before migration, many birds display higher activity or Zugunruhe ( German : migratory restlessness ), first described by Johann Friedrich Naumann in 1795, as well as physiological changes such as increased fat deposition. The occurrence of Zugunruhe even in cage-raised birds with no environmental cues (e.g. shortening of day and falling temperature) has pointed to

2944-483: Is alterable with selective breeding. Many migration routes of long-distance migratory birds are circuitous due to evolutionary history: the breeding range of Northern wheatears Oenanthe oenanthe has expanded to cover the entire Northern Hemisphere, but the species still migrates up to 14,500 km to reach ancestral wintering grounds in sub-Saharan Africa rather than establish new wintering grounds closer to breeding areas. A migration route often does not follow

3072-567: Is an example of leap-frog migration . Many fully migratory species show leap-frog migration (birds that nest at higher latitudes spend the winter at lower latitudes), and many show the alternative, chain migration, where populations 'slide' more evenly north and south without reversing the order. Within a population, it is common for different ages and/or sexes to have different patterns of timing and distance. Female chaffinches Fringilla coelebs in Eastern Fennoscandia migrate earlier in

3200-695: Is clockwise migration, where birds flying North tend to be further West, and flying South tend to shift Eastwards. Many, if not most, birds migrate in flocks. For larger birds, flying in flocks reduces the energy cost. Geese in a V formation may conserve 12–20% of the energy they would need to fly alone. Red knots Calidris canutus and dunlins Calidris alpina were found in radar studies to fly 5 km/h (2.5 kn) faster in flocks than when they were flying alone. Birds fly at varying altitudes during migration. An expedition to Mt. Everest found skeletons of northern pintail Anas acuta and black-tailed godwit Limosa limosa at 5,000 m (16,000 ft) on

3328-557: Is commonly considered to have begun in the 8th century BC (around the time of the earliest recorded poetry of Homer) and ended in the 6th century AD. Classical antiquity in Greece was preceded by the Greek Dark Ages ( c.  1200 – c.  800 BC ), archaeologically characterised by the protogeometric and geometric styles of designs on pottery. Following the Dark Ages was

3456-465: Is debated. Herodotus was succeeded by authors such as Thucydides , Xenophon , Demosthenes , Plato and Aristotle . Most were either Athenian or pro-Athenian, which is why far more is known about the history and politics of Athens than of many other cities. Their scope is further limited by a focus on political, military and diplomatic history, ignoring economic and social history. The archaic period, lasting from approximately 800 to 500 BC, saw

3584-651: Is evidence that this enables the migrants to obtain more of their preferred foods such as fruits. Altitudinal migration is common on mountains worldwide, such as in the Himalayas and the Andes . Dusky grouse in Colorado migrate less than a kilometer away from their summer grounds to winter sites which may be higher or lower by about 400 m in altitude than the summer sites. Many bird species in arid regions across southern Australia are nomadic; they follow water and food supply around

3712-503: Is known as "partial migration". Partial migration is very common in the southern continents; in Australia, 44% of non-passerine birds and 32% of passerine species are partially migratory. In some species, the population at higher latitudes tends to be migratory and will often winter at lower latitude. The migrating birds bypass the latitudes where other populations may be sedentary, where suitable wintering habitats may already be occupied. This

3840-499: Is marked by its annual seasonality and movement between breeding and non-breeding areas. Nonmigratory bird movements include those made in response to environmental changes including in food availability, habitat, or weather. Sometimes, journeys are not termed "true migration" because they are irregular (nomadism, invasions, irruptions) or in only one direction (dispersal, movement of young away from natal area). Non-migratory birds are said to be resident or sedentary. Approximately 1,800 of

3968-452: Is mountainous, and as a result, ancient Greece consisted of many smaller regions, each with its own dialect, cultural peculiarities, and identity. Regionalism and regional conflicts were prominent features of ancient Greece. Cities tended to be located in valleys between mountains, or on coastal plains, and dominated a certain area around them. In the south lay the Peloponnese , consisting of

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4096-475: Is unclear exactly how this change occurred. For instance, in Athens, the kingship had been reduced to a hereditary, lifelong chief magistracy ( archon ) by c. 1050 BC; by 753 BC this had become a decennial, elected archonship; and finally by 683 BC an annually elected archonship. Through each stage, more power would have been transferred to the aristocracy as a whole, and away from a single individual. Inevitably,

4224-407: Is widely known as the "father of history": his Histories are eponymous of the entire field . Written between the 450s and 420s BC, Herodotus' work reaches about a century into the past, discussing 6th century BC historical figures such as Darius I of Persia , Cambyses II and Psamtik III , and alluding to some 8th century BC persons such as Candaules . The accuracy of Herodotus' works

4352-650: The Archaic Period , beginning around the 8th century BC, which saw early developments in Greek culture and society leading to the Classical Period from the Persian invasion of Greece in 480 BC until the death of Alexander the Great in 323 BC. The Classical Period is characterized by a "classical" style, i.e. one which was considered exemplary by later observers, most famously in the Parthenon of Athens. Politically,

4480-705: The Archaic period , the Greek population grew beyond the capacity of the limited arable land of Greece proper, resulting in the large-scale establishment of colonies elsewhere: according to one estimate, the population of the widening area of Greek settlement increased roughly tenfold from 800 BC to 400 BC, from 800,000 to as many as 7 + 1 ⁄ 2 -10 million. This was not simply for trade, but also to found settlements. These Greek colonies were not, as Roman colonies were, dependent on their mother-city, but were independent city-states in their own right. Greeks settled outside of Greece in two distinct ways. The first

4608-670: The Battle of Aegospotami , and began to blockade Athens' harbour; driven by hunger, Athens sued for peace, agreeing to surrender their fleet and join the Spartan-led Peloponnesian League. Following the Athenian surrender, Sparta installed an oligarchic regime, the Thirty Tyrants , in Athens, one of a number of Spartan-backed oligarchies which rose to power after the Peloponnesian war. Spartan predominance did not last: after only

4736-474: The Bay of Fundy and Delaware Bay . Some bar-tailed godwits Limosa lapponica baueri have the longest known non-stop flight of any migrant, flying 11,000 km from Alaska to their New Zealand non-breeding areas. Prior to migration, 55 percent of their bodyweight is stored as fat to fuel this uninterrupted journey. Seabird migration is similar in pattern to those of the waders and waterfowl. Some, such as

4864-688: The Bosphorus at migration times. More common species, such as the European honey buzzard Pernis apivorus , can be counted in hundreds of thousands in autumn. Other barriers, such as mountain ranges, can cause funnelling, particularly of large diurnal migrants, as in the Central American migratory bottleneck. The Batumi bottleneck in the Caucasus is one of the heaviest migratory funnels on earth, created when hundreds of thousands of soaring birds avoid flying over

4992-564: The Delian League gradually transformed from a defensive alliance of Greek states into an Athenian empire, as Athens' growing naval power intimidated the other league states. Athens ended its campaigns against Persia in 450, after a disastrous defeat in Egypt in 454, and the death of Cimon in action against the Persians on Cyprus in 450. As the Athenian fight against the Persian empire waned, conflict grew between Athens and Sparta. Suspicious of

5120-729: The Early Middle Ages and the Byzantine period. Three centuries after the decline of Mycenaean Greece during the Bronze Age Collapse , Greek urban poleis began to form in the 8th century BC, ushering in the Archaic period and the colonization of the Mediterranean Basin. This was followed by the age of Classical Greece , from the Greco-Persian Wars to the death of Alexander the Great in 323 BC, and which included

5248-671: The Golden Age of Athens and the Peloponnesian War . The unification of Greece by Macedon under Philip II and subsequent conquest of the Achaemenid Empire by Alexander the Great spread Hellenistic civilization across the Middle East. The Hellenistic Period is considered to have ended in 30 BC, when the last Hellenistic kingdom, Ptolemaic Egypt , was annexed by the Roman Republic . Classical Greek culture , especially philosophy, had

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5376-519: The Khumbu Glacier . Bar-headed geese Anser indicus have been recorded by GPS flying at up to 6,540 m (21,460 ft) while crossing the Himalayas, at the same time engaging in the highest rates of climb to altitude for any bird. Anecdotal reports of them flying much higher have yet to be corroborated with any direct evidence. Seabirds fly low over water but gain altitude when crossing land, and

5504-710: The Paeonians due north, the Thracians to the northeast, and the Illyrians , with whom the Macedonians were frequently in conflict, to the northwest. Chalcidice was settled early on by southern Greek colonists and was considered part of the Greek world, while from the late 2nd millennium BC substantial Greek settlement also occurred on the eastern shores of the Aegean , in Anatolia . During

5632-510: The black guillemot Cepphus grylle and some gulls , are quite sedentary; others, such as most terns and auks breeding in the temperate northern hemisphere, move varying distances south in the northern winter. The Arctic tern Sterna paradisaea has the longest-distance migration of any bird, and sees more daylight than any other, moving from its Arctic breeding grounds to the Antarctic non-breeding areas. One Arctic tern, ringed (banded) as

5760-778: The dark-eyed junco migrates from subarctic and arctic climates to the contiguous United States and the American goldfinch from taiga to wintering grounds extending from the American South northwestward to Western Oregon . Some ducks, such as the garganey Anas querquedula , move completely or partially into the tropics. The European pied flycatcher Ficedula hypoleuca follows this migratory trend, breeding in Asia and Europe and wintering in Africa. Migration routes and wintering grounds are both genetically and traditionally determined depending on

5888-501: The poleis grouped themselves into leagues, membership of which was in a constant state of flux. Later in the Classical period, the leagues would become fewer and larger, be dominated by one city (particularly Athens , Sparta and Thebes ); and often poleis would be compelled to join under threat of war (or as part of a peace treaty). Even after Philip II of Macedon conquered the heartlands of ancient Greece, he did not attempt to annex

6016-478: The stork in the heavens knows its seasons, and the turtle dove , the swift and the crane keep the time of their arrival." In the Pacific, traditional land-finding techniques used by Micronesians and Polynesians suggest that bird migration was observed and interpreted for more than 3,000 years. In Samoan tradition, for example, Tagaloa sent his daughter Sina to Earth in the form of a bird, Tuli, to find dry land,

6144-523: The 'tubenose' order Procellariiformes , are great wanderers, and the albatrosses of the southern oceans may circle the globe as they ride the " Roaring Forties " outside the breeding season. The tubenoses spread widely over large areas of open ocean, but congregate when food becomes available. Many are among the longest-distance migrants; sooty shearwaters Puffinus griseus nesting on the Falkland Islands migrate 14,000 km (7,600 nmi) between

6272-507: The 12th–9th centuries BC to the end of classical antiquity ( c.  600 AD ), that comprised a loose collection of culturally and linguistically related city-states and other territories. Prior to the Roman period , most of these regions were officially unified once under the Kingdom of Macedon from 338 to 323 BC. In Western history , the era of classical antiquity was immediately followed by

6400-460: The 146 BC conquest of Greece after the Battle of Corinth. Macedonia became a Roman province while southern Greece came under the surveillance of Macedonia's prefect ; however, some Greek poleis managed to maintain a partial independence and avoid taxation. The Aegean Islands were added to this territory in 133 BC. Athens and other Greek cities revolted in 88 BC, and the peninsula was crushed by

6528-521: The 1940s onward, additional refuges were created, so that the Sacramento NWR Complex now includes the following refuges, located between 80 and 145 kilometres (50 and 90 mi) north of the city of Sacramento : The total area of the refuges is about 140 square kilometres (35,000 acres)s). The water flows in the refuge have to be controlled artificially, and the vegetation has to be managed actively through irrigation and burning, to ensure that

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6656-840: The Black Sea surface and across high mountains. Birds of prey such as honey buzzards which migrate using thermals lose only 10 to 20% of their weight during migration, which may explain why they forage less during migration than do smaller birds of prey with more active flight such as falcons, hawks and harriers. From observing the migration of eleven soaring bird species over the Strait of Gibraltar, species which did not advance their autumn migration dates were those with declining breeding populations in Europe. Many long-distance migrants appear to be genetically programmed to respond to changing day length. Species that move short distances, however, may not need such

6784-401: The Carthaginian invasion at the Battle of Himera . The Persians were decisively defeated at sea by a primarily Athenian naval force at the Battle of Salamis , and on land in 479 BC at the Battle of Plataea . The alliance against Persia continued, initially led by the Spartan Pausanias but from 477 by Athens, and by 460 Persia had been driven out of the Aegean. During this long campaign,

6912-416: The Classical Period was dominated by Athens and the Delian League during the 5th century, but displaced by Spartan hegemony during the early 4th century BC, before power shifted to Thebes and the Boeotian League and finally to the League of Corinth led by Macedon . This period was shaped by the Greco-Persian Wars , the Peloponnesian War , and the Rise of Macedon . Following the Classical period

7040-466: The Corinthian empire in northwest Greece and defended its own empire, despite a plague which killed the leading Athenian statesman Pericles . The war turned after Athenian victories led by Cleon at Pylos and Sphakteria , and Sparta sued for peace, but the Athenians rejected the proposal. The Athenian failure to regain control of Boeotia at Delium and Brasidas ' successes in northern Greece in 424 improved Sparta's position after Sphakteria. After

7168-488: The Earth, flying over the southern oceans, while others such as Manx shearwaters migrate 14,000 km (8,700 mi) between their northern breeding grounds and the southern ocean. Shorter migrations are common, while longer ones are not. The shorter migrations include altitudinal migrations on mountains, including the Andes and Himalayas . The timing of migration seems to be controlled primarily by changes in day length. Migrating birds navigate using celestial cues from

7296-416: The Greek city-states. It greatly widened the horizons of the Greeks and led to a steady emigration of the young and ambitious to the new Greek empires in the east. Many Greeks migrated to Alexandria, Antioch and the many other new Hellenistic cities founded in Alexander's wake, as far away as present-day Afghanistan and Pakistan , where the Greco-Bactrian Kingdom and the Indo-Greek Kingdom survived until

7424-460: The Greek colonies Syracusae ( Συράκουσαι ), Neapolis ( Νεάπολις ), Massalia ( Μασσαλία ) and Byzantion ( Βυζάντιον ). These colonies played an important role in the spread of Greek influence throughout Europe and also aided in the establishment of long-distance trading networks between the Greek city-states, boosting the economy of ancient Greece . Ancient Greece consisted of several hundred relatively independent city-states ( poleis ). This

7552-470: The Greek colony Sybaris in southern Italy, its allies, and the Serdaioi. In 499 BC, the Ionian city states under Persian rule rebelled against their Persian-supported tyrant rulers. Supported by troops sent from Athens and Eretria , they advanced as far as Sardis and burnt the city before being driven back by a Persian counterattack. The revolt continued until 494, when the rebelling Ionians were defeated. Darius did not forget that Athens had assisted

7680-420: The Greek system are further evidenced by the colonies that they set up throughout the Mediterranean , which, though they might count a certain Greek polis as their 'mother' (and remain sympathetic to her), were completely independent of the founding city. Inevitably smaller poleis might be dominated by larger neighbors, but conquest or direct rule by another city-state appears to have been quite rare. Instead

7808-660: The Greeks began 250 years of expansion, settling colonies in all directions. To the east, the Aegean coast of Asia Minor was colonized first, followed by Cyprus and the coasts of Thrace , the Sea of Marmara and south coast of the Black Sea . Eventually, Greek colonization reached as far northeast as present-day Ukraine and Russia ( Taganrog ). To the west the coasts of Illyria , Southern Italy (called " Magna Graecia ") were settled, followed by Southern France , Corsica , and even eastern Spain . Greek colonies were also founded in Egypt and Libya . Modern Syracuse , Naples , Marseille and Istanbul had their beginnings as

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7936-423: The Greeks were very aware of their tribal origins; Herodotus was able to extensively categorise the city-states by tribe. Yet, although these higher-level relationships existed, they seem to have rarely had a major role in Greek politics. The independence of the poleis was fiercely defended; unification was something rarely contemplated by the ancient Greeks. Even when, during the second Persian invasion of Greece,

8064-420: The Ionian revolt, and in 490 he assembled an armada to retaliate. Though heavily outnumbered, the Athenians—supported by their Plataean allies—defeated the Persian hordes at the Battle of Marathon , and the Persian fleet turned tail. Ten years later, a second invasion was launched by Darius' son Xerxes . The city-states of northern and central Greece submitted to the Persian forces without resistance, but

8192-430: The League of Corinth following the death of Philip, Alexander began his campaign against Persia in 334 BC. He conquered Persia, defeating Darius III at the Battle of Issus in 333 BC, and after the Battle of Gaugamela in 331 BC proclaimed himself king of Asia. From 329 BC he led expeditions to Bactria and then India; further plans to invade Arabia and North Africa were halted by his death in 323 BC. The period from

8320-471: The Roman general Sulla . The Roman civil wars devastated the land even further, until Augustus organized the peninsula as the province of Achaea in 27 BC. Greece was a key eastern province of the Roman Empire, as the Roman culture had long been in fact Greco-Roman . The Greek language served as a lingua franca in the East and in Italy , and many Greek intellectuals such as Galen would perform most of their work in Rome . The territory of Greece

8448-498: The Southern Hemisphere) to overwinter; In contrast, among (pelagic) seabirds, species of the Southern Hemisphere are more likely to migrate. This is because there is a large area of ocean in the Southern Hemisphere, and more islands suitable for seabirds to nest. The control of migration, its timing and response are genetically controlled and appear to be a primitive trait that is present even in non-migratory species of birds. The ability to navigate and orient themselves during migration

8576-468: The Sun and stars, the Earth's magnetic field, and mental maps. Writings of ancient Greeks recognized the seasonal comings and goings of birds. Aristotle wrote that birds transmuted into other birds or species like fish and animals, which explained their disappearance and reappearance. Aristotle thought many birds disappeared during cold weather because they were torpid , undetected in unseen environments like tree hollows or burrowing down in mud found at

8704-528: The ancient Greeks did not think in terms of race . Most families owned slaves as household servants and laborers, and even poor families might have owned a few slaves. Owners were not allowed to beat or kill their slaves. Owners often promised to free slaves in the future to encourage slaves to work hard. Unlike in Rome, freedmen did not become citizens. Instead, they were mixed into the population of metics , which included people from foreign countries or other city-states who were officially allowed to live in

8832-405: The autumn passage of southbound passerine migrants, which it feeds to its young. A similar strategy is adopted by the greater noctule bat , which preys on nocturnal passerine migrants. The higher concentrations of migrating birds at stopover sites make them prone to parasites and pathogens, which require a heightened immune response. Within a species not all populations may be migratory; this

8960-659: The autumn than males do and the European tits of genera Parus and Cyanistes only migrate their first year. Most migrations begin with the birds starting off in a broad front. Often, this front narrows into one or more preferred routes termed flyways . These routes typically follow mountain ranges or coastlines, sometimes rivers, and may take advantage of updrafts and other wind patterns or avoid geographical barriers such as large stretches of open water. The specific routes may be genetically programmed or learned to varying degrees. The routes taken on forward and return migration are often different. A common pattern in North America

9088-474: The bottom of ponds, then reemerging months later. Still, Aristotle recorded that cranes traveled from the steppes of Scythia to marshes at the headwaters of the Nile , an observation repeated by Pliny the Elder in his Historia Naturalis . Two books of the Bible may address avian migration. The Book of Job notes migrations with the inquiry: "Is it by your insight that the hawk hovers, spreads its wings southward?" The Book of Jeremiah comments: "Even

9216-669: The breeding colony and the North Atlantic Ocean off Norway . Some Manx shearwaters Puffinus puffinus do this same journey in reverse. As they are long-lived birds, they may cover enormous distances during their lives; one record-breaking Manx shearwater is calculated to have flown 8 million kilometres (4.5 million nautical miles) during its over-50-year lifespan. Some large broad-winged birds rely on thermal columns of rising hot air to enable them to soar. These include many birds of prey such as vultures , eagles , and buzzards , but also storks . These birds migrate in

9344-646: The center, while in the east lay Boeotia , Attica , and Megaris . Northeast lay Thessaly , while Epirus lay to the northwest. Epirus stretched from the Ambracian Gulf in the south to the Ceraunian Mountains and the Aoos river in the north, and consisted of Chaonia (north), Molossia (center), and Thesprotia (south). In the northeast corner was Macedonia , originally consisting Lower Macedonia and its regions, such as Elimeia , Pieria , and Orestis . Around

9472-483: The chaffinch are much less migratory in Britain than those of continental Europe, mostly not moving more than 5 km in their lives. Short-distance passerine migrants have two evolutionary origins. Those that have long-distance migrants in the same family, such as the common chiffchaff Phylloscopus collybita , are species of southern hemisphere origins that have progressively shortened their return migration to stay in

9600-455: The composition of a migrating flock, and can sometimes encode the sex of a migrating individual, and to avoid collision in the air. Nocturnal migration can be monitored using weather radar data, allowing ornithologists to estimate the number of birds migrating on a given night, and the direction of the migration. Future research includes the automatic detection and identification of nocturnally calling migrant birds. Nocturnal migrants land in

9728-498: The country in an irregular pattern, unrelated to season but related to rainfall. Several years may pass between visits to an area by a particular species. Sometimes circumstances such as a good breeding season followed by a food source failure the following year lead to irruptions in which large numbers of a species move far beyond the normal range. Bohemian waxwings Bombycilla garrulus well show this unpredictable variation in annual numbers, with five major arrivals in Britain during

9856-455: The course of the archaic period. Already in the seventh century, the right of all citizen men to attend the assembly appears to have been established. After a failed coup led by Cylon of Athens around 636 BC, Draco was appointed to establish a code of laws in 621. This failed to reduce the political tension between the poor and the elites, and in 594 Solon was given the authority to enact another set of reforms, which attempted to balance

9984-473: The course of the eighth and seventh century. According to Spartan tradition, this constitution was established by the legendary lawgiver Lycurgus . Over the course of the first and second Messenian wars , Sparta subjugated the neighbouring region of Messenia , enserfing the population. In the sixth century, Greek city-states began to develop formal relationships with one another, where previously individual rulers had relied on personal relationships with

10112-574: The culmination of political and social developments which had begun in the Greek dark age, with the polis (city-state) becoming the most important unit of political organisation in Greece. The absence of powerful states in Greece after the collapse of Mycenaean power, and the geography of Greece, where many settlements were separated from their neighbours by mountainous terrain, encouraged the development of small independent city-states. Several Greek states saw tyrants rise to power in this period, most famously at Corinth from 657 BC. The period also saw

10240-495: The daytime. Migratory species in these groups have great difficulty crossing large bodies of water, since thermals only form over land, and these birds cannot maintain active flight for long distances. Mediterranean and other seas present a major obstacle to soaring birds, which must cross at the narrowest points. Massive numbers of large raptors and storks pass through areas such as the Strait of Messina , Gibraltar , Falsterbo , and

10368-461: The death of Alexander the Great in 323 BC until the death of Cleopatra , the last Macedonian ruler of Egypt, is known as the Hellenistic period. In the early part of this period, a new form of kingship developed based on Macedonian and Near Eastern traditions. The first Hellenistic kings were previously Alexander's generals, and took power in the period following his death, though they were not part of existing royal lineages and lacked historic claims to

10496-423: The deaths of Cleon and Brasidas, the strongest proponents of war on each side, a peace treaty was negoitiated in 421 by the Athenian general Nicias . The peace did not last, however. In 418 BC allied forces of Athens and Argos were defeated by Sparta at Mantinea . In 415 Athens launched an ambitious naval expedition to dominate Sicily; the expedition ended in disaster at the harbor of Syracuse , with almost

10624-657: The dominance that would allow it to challenge Persia, the Persian king initially joined the alliance against Sparta, before imposing the Peace of Antalcidas ("King's Peace") which restored Persia's control over the Anatolian Greeks. By 371 BC, Thebes was in the ascendancy, defeating Sparta at the Battle of Leuctra , killing the Spartan king Cleombrotus I , and invading Laconia. Further Theban successes against Sparta in 369 led to Messenia gaining independence; Sparta never recovered from

10752-409: The domination of politics and concomitant aggregation of wealth by small groups of families was apt to cause social unrest in many poleis . In many cities a tyrant (not in the modern sense of repressive autocracies), would at some point seize control and govern according to their own will; often a populist agenda would help sustain them in power. In a system wracked with class conflict , government by

10880-700: The east to the Indian king Chandragupta Maurya in exchange for war elephants, and later lost large parts of Persia to the Parthian Empire . By the mid-third century, the kingdoms of Alexander's successors was mostly stable, though there continued to be disputes over border areas. The great capitals of Hellenistic culture were Alexandria in the Ptolemaic Kingdom and Antioch in the Seleucid Empire . The conquests of Alexander had numerous consequences for

11008-537: The elites of other cities. Towards the end of the archaic period, Sparta began to build a series of alliances, the Peloponnesian League , with cities including Corinth , Elis , and Megara , isolating Messenia and reinforcing Sparta's position against Argos , the other major power in the Peloponnese. Other alliances in the sixth century included those between Elis and Heraea in the Peloponnese; and between

11136-648: The end of the first century BC. The city-states within Greece formed themselves into two leagues; the Achaean League (including Corinth and Argos) and the Aetolian League (including Sparta and Athens). For much of the period until the Roman conquest, these leagues were at war, often participating in the conflicts between the Diadochi (the successor states to Alexander's empire). The Antigonid Kingdom became involved in

11264-592: The entire army killed, and the ships destroyed. Soon after the Athenian defeat in Syracuse, Athens' Ionian allies began to rebel against the Delian league, while Persia began to once again involve itself in Greek affairs on the Spartan side. Initially the Athenian position continued relatively strong, with important victories at Cyzicus in 410 and Arginusae in 406. However, in 405 the Spartan Lysander defeated Athens in

11392-529: The exact borders of the Hellenistic kingdoms were not settled. Antigonus attempted to expand his territory by attacking the other successor kingdoms until they joined against him, and he was killed at the Battle of Ipsus in 301 BC. His son Demetrius spent many years in Seleucid captivity, and his son, Antigonus II , only reclaimed the Macedonian throne around 276. Meanwhile, the Seleucid kingdom gave up territory in

11520-412: The far south to support long-distance migration. The primary motivation for migration appears to be food; for example, some hummingbirds choose not to migrate if fed through the winter. In addition, the longer days of the northern summer provide extended time for breeding birds to feed their young. This helps diurnal birds to produce larger clutches than related non-migratory species that remain in

11648-464: The founding of Greek colonies around the Mediterranean, with Euboean settlements at Al-Mina in the east as early as 800 BC, and Ischia in the west by 775. Increasing contact with non-Greek peoples in this period, especially in the Near East, inspired developments in art and architecture, the adoption of coinage, and the development of the Greek alphabet. Athens developed its democratic system over

11776-569: The government. In Athens, the population was divided into four social classes based on wealth. People could change classes if they made more money. In Sparta, all male citizens were called homoioi , meaning "peers". However, Spartan kings, who served as the city-state's dual military and religious leaders, came from two families. Women in Ancient Greece appear to have primarily performed domestic tasks, managed households, and borne and reared children. Slaves had no power or status. Slaves had

11904-584: The increasing Athenian power funded by the Delian League, Sparta offered aid to reluctant members of the League to rebel against Athenian domination. These tensions were exacerbated in 462 BC when Athens sent a force to aid Sparta in overcoming a helot revolt, but this aid was rejected by the Spartans. In the 450s, Athens took control of Boeotia, and won victories over Aegina and Corinth. However, Athens failed to win

12032-418: The local temperature to time their spring migration departure. Notably, departure responses to temperature varied between individuals but were individually repeatable (when tracked over multiple years). This suggests that individual use of temperature is a cue that allows for population-level adaptation to climate change . In other words, in a warming world, many migratory birds are predicted to depart earlier in

12160-514: The loss of Messenia's fertile land and the helot workforce it provided. The rising power of Thebes led Sparta and Athens to join forces; in 362 they were defeated by Thebes at the Battle of Mantinea . In the aftermath of Mantinea, none of the major Greek states were able to dominate. Though Thebes had won the battle, their general Epaminondas was killed, and they spent the following decades embroiled in wars with their neighbours; Athens, meanwhile, saw its second naval alliance, formed in 377, collapse in

12288-508: The loss. The typical image of migration is of northern land birds, such as swallows (Hirundinidae) and birds of prey, making long flights to the tropics. However, many Holarctic wildfowl and finch (Fringillidae) species winters in the North Temperate Zone , in regions with milder winters than their summer breeding grounds. For example, the pink-footed goose migrates from Iceland to Britain and neighbouring countries, whilst

12416-543: The mid-350s. The power vacuum in Greece after the Battle of Mantinea was filled by Macedon, under Philip II . In 338 BC, he defeated a Greek alliance at the Battle of Chaeronea , and subsequently formed the League of Corinth . Philip planned to lead the League to invade Persia, but was murdered in 336 BC. His son Alexander the Great was left to fulfil his father's ambitions. After campaigns against Macedon's western and northern enemies, and those Greek states that had broken from

12544-427: The morning and may feed for a few days before resuming their migration. These birds are referred to as passage migrants in the regions where they occur for a short period between the origin and destination. Nocturnal migrants minimize depredation, avoid overheating, and can feed during the day. One cost of nocturnal migration is the loss of sleep. Migrants may be able to alter their quality of sleep to compensate for

12672-502: The most direct line between breeding and wintering grounds. Rather, it could follow a hooked or arched line, with detours around geographical barriers or towards suitable stopover habitat. For most land birds, such barriers could consist of large water bodies or high mountain ranges, a lack of stopover or feeding sites, or a lack of thermal columns (important for broad-winged birds). Conversely, in water-birds , large areas of land without wetlands offering suitable feeding sites may present

12800-458: The nineteenth century, but 18 between the years 1937 and 2000. Red crossbills Loxia curvirostra too are irruptive, with widespread invasions across England noted in 1251, 1593, 1757, and 1791. Bird migration is primarily, but not entirely, a Northern Hemisphere phenomenon. This is because continental landmasses of the northern hemisphere are almost entirely temperate and subject to winter food shortages driving bird populations south (including

12928-456: The northern hemisphere. Species that have no long-distance migratory relatives, such as the waxwings Bombycilla , are effectively moving in response to winter weather and the loss of their usual winter food, rather than enhanced breeding opportunities. In the tropics there is little variation in the length of day throughout the year, and it is always warm enough for a food supply, but altitudinal migration occurs in some tropical birds. There

13056-553: The original wetland area remains. Migrating birds have continued to use the area, and resting in the rice fields, consumed considerable quantities of the crop. In 1937, the United States Fish and Wildlife Service , with the aid of the Civilian Conservation Corps , began the process of creating a refuge within dry, alkaline lands between the towns of Willows and Maxwell . This was the original Sacramento NWR. From

13184-524: The period of Christianization during the later 4th to early 6th centuries AD, consummated by the closure of the Academy of Athens by Justinian I in 529. The historical period of ancient Greece is unique in world history as the first period attested directly in comprehensive, narrative historiography , while earlier ancient history or protohistory is known from much more fragmentary documents such as annals, king lists, and pragmatic epigraphy . Herodotus

13312-410: The power of the rich and the poor. In the middle of the sixth century, Pisistratus established himself as a tyrant, and after his death in 527 his son Hippias inherited his position; by the end of the sixth century he had been overthrown and Cleisthenes carried out further democratising reforms. In Sparta, a political system with two kings, a council of elders , and five ephors developed over

13440-613: The reforms of Solon (early 6th century), but the poorest citizens could not address the assembly or run for office. With the establishment of the democracy, the assembly became the de jure mechanism of government; all citizens had equal privileges in the assembly. However, non-citizens, such as metics (foreigners living in Athens) or slaves , had no political rights at all. After the rise of democracy in Athens, other city-states founded democracies. However, many retained more traditional forms of government. As so often in other matters, Sparta

13568-400: The refuges, not all of them migratory. Among those most likely to be seen by visitors (depending on season) are: The Sacramento National Wildlife Refuge Complex boasts a diverse flora, even though large amounts of the historic wetland has been destroyed. Among the diverse wildflowers present is the yellow mariposa lily, Calochortus luteus . Bird migration Bird migration is

13696-420: The regions of Laconia (southeast), Messenia (southwest), Elis (west), Achaia (north), Korinthia (northeast), Argolis (east), and Arcadia (center). These names survive to the present day as regional units of modern Greece , though with somewhat different boundaries. Mainland Greece to the north, nowadays known as Central Greece , consisted of Aetolia and Acarnania in the west, Locris , Doris , and Phocis in

13824-953: The reverse pattern is seen in land birds. However most bird migration is in the range of 150 to 600 m (490–2,000 ft). Bird strike Aviation records from the United States show most collisions occur below 600 m (2,000 ft) and almost none above 1,800 m (5,900 ft). Bird migration is not limited to birds that can fly. Most species of penguin (Spheniscidae) migrate by swimming. These routes can cover over 1,000 km (550 nmi). Dusky grouse Dendragapus obscurus perform altitudinal migration mostly by walking. Emus Dromaius novaehollandiae in Australia have been observed to undertake long-distance movements on foot during droughts. During nocturnal migration ("nocmig" ), many birds give nocturnal flight calls, which are short, contact-type calls. These likely serve to maintain

13952-518: The right to have a family and own property, subject to their master's goodwill and permission, but they had no political rights. By 600 BC, chattel slavery had spread in Greece. By the 5th century BC, slaves made up one-third of the total population in some city-states. Between 40–80% of the population of Classical Athens were slaves. Slaves outside of Sparta almost never revolted because they were made up of too many nationalities and were too scattered to organize. However, unlike later Western culture ,

14080-446: The role of circannual endogenous programs in controlling bird migrations. Caged birds display a preferential flight direction that corresponds with the migratory direction they would take in nature, changing their preferential direction at roughly the same time their wild conspecifics change course. Satellite tracking of 48 individual Asian houbaras ( Chlamydotis macqueenii ) across multiple migrations showed that this species uses

14208-490: The same hemisphere, but others such as semipalmated sandpiper C. pusilla travel longer distances to the tropics in the Southern Hemisphere. For some species of waders, migration success depends on the availability of certain key food resources at stopover points along the migration route. This gives the migrants an opportunity to refuel for the next leg of the voyage. Some examples of important stopover locations are

14336-470: The social system of the species. In long-lived, social species such as white storks (Ciconia ciconia), flocks are often led by the oldest members and young storks learn the route on their first journey. In short-lived species that migrate alone, such as the Eurasian blackcap Sylvia atricapilla or the yellow-billed cuckoo Coccyzus americanus , first-year migrants follow a genetically determined route that

14464-689: The start of the next, travelling not just the length of the Atlantic Ocean and the width of the Indian Ocean, but also half way across the South Pacific to the boundary between the Ross and Amundsen Seas before returning back west along the Antarctic coast and back up the Atlantic. Many tubenosed birds breed in the southern hemisphere and migrate north in the southern winter. The most pelagic species, mainly in

14592-448: The state. City-states legally owned slaves. These public slaves had a larger measure of independence than slaves owned by families, living on their own and performing specialized tasks. In Athens, public slaves were trained to look out for counterfeit coinage , while temple slaves acted as servants of the temple's deity and Scythian slaves were employed in Athens as a police force corralling citizens to political functions. Sparta had

14720-669: The territories they controlled. The most important of these rulers in the decades after Alexander's death were Antigonus I and his son Demetrius in Macedonia and the rest of Greece, Ptolemy in Egypt, and Seleucus I in Syria and the former Persian empire; smaller Hellenistic kingdoms included the Attalids in Anatolia and the Greco-Bactrian kingdom . In the early part of the Hellenistic period,

14848-510: The territory or unify it into a new province, but compelled most of the poleis to join his own Corinthian League . Initially many Greek city-states seem to have been petty kingdoms; there was often a city official carrying some residual, ceremonial functions of the king ( basileus ), e.g., the archon basileus in Athens. However, by the Archaic period and the first historical consciousness, most had already become aristocratic oligarchies . It

14976-608: The time of Alexander I of Macedon , the Argead kings of Macedon started to expand into Upper Macedonia , lands inhabited by independent Macedonian tribes like the Lyncestae , Orestae and the Elimiotae and to the west, beyond the Axius river , into Eordaia , Bottiaea , Mygdonia , and Almopia , regions settled by Thracian tribes. To the north of Macedonia lay various non-Greek peoples such as

15104-416: The tropics. As the days shorten in autumn, the birds return to warmer regions where the available food supply varies little with the season. These advantages offset the high stress, physical exertion costs, and other risks of migration. Predation can be heightened during migration: Eleonora's falcon Falco eleonorae , which breeds on Mediterranean islands, has a very late breeding season, coordinated with

15232-425: The wetlands remain productive, and provide adequate food and resting places for the birds. Between them, the refuges provide a range of habitats: seasonal marshes , uplands, permanent ponds, and riparian areas. The refuges are provided with facilities for visitors, though these have to be limited to avoid conflict with their primary purpose of conservation. The layout of the wetlands has been planned to allow visitors

15360-459: The winters. He concludes: These experiments have since been amply confirmed by ... M. Natterer , of Vienna ... and the result clearly proves, what is in fact now admitted on all hands, that Swallows do not in any material instance differ from other birds in their nature and propensities [for life in the air]; but that they leave us when this country can no longer furnish them with a supply of their proper and natural food ... In 1822,

15488-443: The word tuli referring specifically to land-finding waders, often to the Pacific golden plover. Aristotle, however, suggested that swallows and other birds hibernated. This belief persisted as late as 1878 when Elliott Coues listed the titles of no fewer than 182 papers dealing with the hibernation of swallows. Even the "highly observant" Gilbert White , in his posthumously published 1789 The Natural History of Selborne , quoted

15616-409: The world's 10,000 bird species are long-distance migrants. Many bird populations migrate long distances along a flyway. The most common pattern involves flying north in the spring to breed in the temperate or Arctic summer and returning in the autumn to wintering grounds in warmer regions to the south. Of course, in the southern hemisphere, the directions are reversed, but there is less land area in

15744-409: The year for their summer or winter destination. In polygynous species with considerable sexual dimorphism , males tend to return earlier to the breeding sites than their females. This is termed protandry. Ancient Greece Ancient Greece ( Ancient Greek : Ἑλλάς , romanized :  Hellás ) was a northeastern Mediterranean civilization, existing from the Greek Dark Ages of

15872-529: Was a notable exception to the rest of Greece, ruled through the whole period by not one, but two hereditary monarchs. This was a form of diarchy . The Kings of Sparta belonged to the Agiads and the Eurypontids, descendants respectively of Eurysthenes and Procles . Both dynasties' founders were believed to be twin sons of Aristodemus , a Heraclid ruler. However, the powers of these kings were held in check by both

16000-460: Was a situation unlike that in most other contemporary societies, which were either tribal or kingdoms ruling over relatively large territories. Undoubtedly, the geography of Greece —divided and sub-divided by hills, mountains, and rivers—contributed to the fragmentary nature of ancient Greece. On the one hand, the ancient Greeks had no doubt that they were "one people"; they had the same religion , same basic culture, and same language. Furthermore,

16128-732: Was conquered by the Romans. The Aetolian league grew wary of Roman involvement in Greece, and sided with the Seleucids in the Roman–Seleucid War ; when the Romans were victorious, the league was effectively absorbed into the Republic. Although the Achaean league outlasted both the Aetolian league and Macedon, it was also soon defeated and absorbed by the Romans in 146 BC, bringing Greek independence to an end. The Greek peninsula came under Roman rule during

16256-417: Was in permanent settlements founded by Greeks, which formed as independent poleis. The second form was in what historians refer to as emporia ; trading posts which were occupied by both Greeks and non-Greeks and which were primarily concerned with the manufacture and sale of goods. Examples of this latter type of settlement are found at Al Mina in the east and Pithekoussai in the west. From about 750 BC

16384-586: Was the Hellenistic period (323–146 BC), during which Greek culture and power expanded into the Near and Middle East from the death of Alexander until the Roman conquest. Roman Greece is usually counted from the Roman victory over the Corinthians at the Battle of Corinth in 146 BC to the establishment of Byzantium by Constantine as the capital of the Roman Empire in 330 AD. Finally, Late Antiquity refers to

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