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30-484: See text The Saccopharyngiformes are a derived lineage of unusual eels within the order Anguilliformes , and includes families Cyematidae, Monognathidae, Eurypharyngidae, Saccopharyngidae, and the proposed family Neocyematidae. Most of the fish in this group are deep-dwelling and rarely seen, typically known from only a handful of specimens. Species include recognizable fish such as pelican eels (also commonly known as gulper eels) and bobtail eels . Some can live deep in

60-666: A cheap dish in the United Kingdom . During the 1990s, their numbers collapsed across Europe. They became a delicacy, and the UK's most expensive species. Eels, particularly the moray eel , are popular among marine aquarists . Eel blood is toxic to humans and other mammals, but both cooking and the digestive process destroy the toxic protein. High consumption of eels is seen in European countries leading to those eel species being considered endangered. In 2010, Greenpeace International added

90-590: A number of unusual characteristics, such as remarkably deep bodies in the Cyematidae, long lower jaws in the Eurypharyngidae, and unique pigmented swellings at the ends of the gut in Saccopharyngidae and Eurypharyngidae. The three established families of the suborder Saccopharyngoidei (Eupharyngidae, Monognathidae, Saccopharyngidae) all exhibit sexual dimorphism . Until recently, the order "Saccopharyngiformes"

120-441: A sucker disk that allow them to adhere to the substrate or climb structures, such as waterfalls. In priapiumfish , males have modified their pelvic structures into a spiny copulatory device that grasps the female during mating. In actinopterygian steady state swimming, the pelvic fins are actively controlled and used to provide powered corrective forces. Careful timing of the pelvic fin movement during whole-body movements allows

150-556: Is a monophyletic group that originated among the deep-sea eels. The earliest fossil eels are known from the Late Cretaceous ( Cenomanian ) of Lebanon . These early eels retain primitive traits such as pelvic fins and thus do not appear to be closely related to any extant taxa. Body fossils of modern eels do not appear until the Eocene , although otoliths assignable to extant eel families and even some genera have been recovered from

180-685: Is a traditional Māori food in New Zealand . In Italian cuisine , eels from the Valli di Comacchio , a swampy zone along the Adriatic coast, are especially prized, along with freshwater eels of Bolsena Lake and pond eels from Cabras, Sardinia . In northern Germany , the Netherlands , the Czech Republic , Poland , Denmark , and Sweden , smoked eel is considered a delicacy . Elvers, often fried, were once

210-594: Is an adjective form of an older word, ελλυ, meaning "snake", which is directly comparable to Hittite ellu-essar- "snake pit". This myth likely came to Greece via Anatolia. In the Hittite version of the myth, the dragon is called Illuyanka : the illuy- part is cognate to the word illa , and the -anka part is cognate to angu , a word for "snake". Since the words for "snake" (and similarly shaped animals) are often subject to taboo in many Indo-European (and non-Indo-European) languages, no unambiguous Proto-Indo-European form of

240-531: Is used as a metaphor for Nazi atrocities, and the sight of eels being killed by a fisherman triggers the madness of the protagonist's mother. Sinister implications of eels fishing are also referenced in Jo Nesbø 's Cockroaches , the second book of the Harry Hole detective series. The book's background includes a Norwegian village where eels in the nearby sea are rumored to feed on the corpses of drowned humans, making

270-648: The Campanian and Maastrichtian , indicating some level of diversification among the extant groups prior to the Cretaceous-Paleogene extinction , which is also supported by phylogenetic divergence estimates. One of these otolith taxa, the mud-dwelling Pythonichthys arkansasensis , appears to have thrived in the aftermath of the K-Pg extinction, based on its abundance. Taxonomy based on Eschmeyer's Catalog of Fishes : Order Anguilliformes In some classifications,

300-476: The European eel , Japanese eel , and American eel to its seafood red list. Japan consumes more than 70% of the global eel catch. The English name "eel" descends from Old English ǣl , Common Germanic *ēlaz . Also from the common Germanic are West Frisian iel , Dutch aal , German Aal , and Icelandic áll . Katz (1998) identifies a number of Indo-European cognates, among them

330-767: The Northeast . Freshwater eels, known as Kusia in Assamese , are eaten with curry, often with herbs. The European eel and other freshwater eels are mostly eaten in Europe and the United States , and is considered critically endangered. A traditional east London food is jellied eels , although the demand has significantly declined since World War II. The Spanish cuisine delicacy angulas consists of elver (young eels) sautéed in olive oil with garlic ; elvers usually reach prices of up to 1000 euro per kg. New Zealand longfin eel

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360-431: The slender giant moray . Adults range in weight from 30 g (1 oz) to well over 25 kg (55 lb). They possess no pelvic fins , and many species also lack pectoral fins . The dorsal and anal fins are fused with the caudal fin, forming a single ribbon running along much of the length of the animal. Eels swim by generating waves that travel the length of their bodies. They can swim backward by reversing

390-506: The Old English word for "hedgehog", which is igil (meaning "snake eater"), and perhaps in the egi- of Old High German egidehsa "wall lizard". According to this theory, the name Bellerophon ( Βελλεροφόντης , attested in a variant Ἐλλεροφόντης in Eustathius of Thessalonica ) is also related, translating to "the slayer of the serpent" ( ahihán ). In this theory, the ελλερο-

420-627: The Paleobiology Database: Freshwater eels ( unagi ) and marine eels ( conger eel , anago ) are commonly used in Japanese cuisine ; foods such as unadon and unajū are popular, but expensive. Eels are also very popular in Chinese cuisine , and are prepared in many different ways. Hong Kong eel prices have often reached 1000 HKD (128.86 US Dollars) per kg, and once exceeded 5000 HKD per kg. In India , eels are popularly eaten in

450-511: The direction of the wave. Most eels live in the shallow waters of the ocean and burrow into sand, mud, or amongst rocks. Most eel species are nocturnal , and thus are rarely seen. Sometimes, they are seen living together in holes or "eel pits". Some eels also live in deeper water on the continental shelves and over the slopes deep as 4,000 m (13,000 ft). Only members of the Anguilla regularly inhabit fresh water, but they, too, return to

480-469: The early larval stage to the eventual adult stage and are usually predators . The term "eel" is also used for some other eel-shaped fish, such as electric eels (genus Electrophorus ), swamp eels (order Synbranchiformes ), and deep-sea spiny eels (family Notacanthidae). However, these other clades , with the exception of deep-sea spiny eels, whose order Notacanthiformes is the sister clade to true eels, evolved their eel-like shapes independently from

510-578: The eating of these eels verge on cannibalism. Pelvic fin In actinopterygians , the pelvic fin consists of two endochondrally -derived bony girdles attached to bony radials. Dermal fin rays ( lepidotrichia ) are positioned distally from the radials. There are three pairs of muscles each on the dorsal and ventral side of the pelvic fin girdle that abduct and adduct the fin from the body. Pelvic fin structures can be extremely specialized in actinopterygians. Gobiids and lumpsuckers modify their pelvic fins into

540-442: The eel fisheries at Ballisodare were greatly improved by the hanging of loosely plaited grass ladders over barriers, enabling elvers to ascend more easily. Several sets of classifications of eels exist; some, such as FishBase which divide eels into 20 families, whereas other classification systems such as ITIS and Systema Naturae 2000 include additional eel families, which are noted below. Genomic studies indicate that there

570-906: The family Cyematidae of bobtail snipe eels is included in the Anguilliformes, but in the FishBase system that family is included in the order Saccopharyngiformes . The electric eel of South America is not a true eel but is a South American knifefish more closely related to the carps and catfishes . Phylogeny based on Johnson et al. 2012. Protanguillidae Synaphobranchidae Heterenchelyidae Myrocongridae Muraenidae [REDACTED] Chlopsidae Derichthyidae Nettastomatidae Congridae [REDACTED] Ophichthidae Muraenesocidae Moringuidae Eurypharyngidae Saccopharyngidae [REDACTED] Monognathidae Cyematidae [REDACTED] Nemichthyidae Serrivomeridae Anguillidae [REDACTED] Based on

600-502: The ocean, well into the aphotic zone , approximately 500 to 1,800 meters (1,600 to 5,900 ft) deep. Extensive research has not been conducted on them due to being indirectly observed, with some species known only from their larvae. All families except for the exceptionally rare individuals of proposed family Neoceymatidae (known only from the Atlantic Ocean ) are found in all major oceans. They have multiple internal differences from

630-432: The pelvic fins to generate forces that dampen the forces from the entire body, therefore stabilizing the fish. For maneuvers, electromyogram data shows that pelvic fin muscles are activated after the start of the maneuver, indicating that the fins are used more for stabilization instead of generating the maneuver. In rays and skates, pelvic fins can be used for "punting," where they asynchronously or synchronously push off

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660-672: The rest of Anguilliformes. Notably, they have no symplectic bone , opercular bones, ribs, or swim bladders . Like many other eels, they lack scales and pelvic fins . Their myomeres (muscle segments) are V-shaped instead of W-shaped as in all other fishes, and their lateral lines have no pores, instead being modified to groups of elevated tubules. The jaws are quite large, lined with small teeth, and several types are notable for being able to consume fish larger than themselves. Some species in families Eurypharyngidae and Saccopharyngidae are bioluminescent . Like other eels, saccopharyngids have leptocephalus larvae. However, these larvae also have

690-426: The sea to breed. The heaviest true eel is the European conger . The maximum size of this species has been reported as reaching a length of 3 m (10 ft) and a weight of 110 kg (240 lb). Other eels are longer, but do not weigh as much, such as the slender giant moray , which reaches 4 m (13 ft). Eels begin life as flat and transparent larvae , called leptocephali . Eel larvae drift in

720-508: The sea's surface waters, feeding on marine snow , small particles that float in the water. Eel larvae then metamorphose into glass eels and become elvers before finally seeking out their juvenile and adult habitats. Some individuals of anguillid elvers remains in brackish and marine areas close to coastlines, but most of them enter freshwater where they travel upstream and are forced to climb up obstructions, such as weirs , dam walls, and natural waterfalls. Gertrude Elizabeth Blood found that

750-753: The second part of the Latin word for eels, anguilla , attested in its simplex form illa (in a glossary only), and the Greek word for "eel", egkhelys (the second part of which is attested in Hesychius as elyes ). The first compound member, anguis ("snake"), is cognate to other Indo-European words for "snake" (compare Old Irish escung "eel", Old High German unc "snake", Lithuanian angìs , Greek ophis, okhis , Vedic Sanskrit áhi , Avestan aži , Armenian auj, iž , Old Church Slavonic *ǫžь , all from Proto-Indo-European *h₁ogʷʰis ). The word also appears in

780-406: The substrate to propel the animal forwards. Unlike limb development in tetrapods, where the forelimb and hindlimb buds emerge at roughly the same timepoint, the pelvic fin bud emerges much later than the pectoral fin. While the pectoral fin bud is apparent at 36 hours post fertilization (hpf) in zebrafish , the pelvic fin bud is only clear at around 21 days post fertilization (dpf), roughly when

810-841: The town is located. The daylight passage in the spring of elvers upstream along the Thames was at one time called "eel fare". The word 'elver' is thought to be a corruption of "eel fare". A famous attraction on the French Polynesian island of Huahine (part of the Society Islands ) is the bridge across a stream hosting three- to six-foot-long eels, deemed sacred by local culture. Eel fishing in Nazi -era Danzig plays an important role in Günter Grass ' novel The Tin Drum . The cruelty of humans to eels

840-437: The true eels. As a main rule, most eels are marine. Exceptions are the catadromous genus Anguilla and the freshwater moray , which spend most of their life in freshwater, the anadromous rice-paddy eel , which spawns in freshwater, and the freshwater snake eel Stictorhinus . Eels are elongated fish, ranging in length from 5 cm (2 in) in the one-jawed eel ( Monognathus ahlstromi ) to 4 m (13 ft) in

870-545: The word for eel can be reconstructed. It may have been *ēl(l)-u- , *ēl(l)-o- , or something similar. The large lake of Almere , which existed in the early Medieval Netherlands , got its name from the eels which lived in its water (the Dutch word for eel is aal or ael , so: " ael mere " = "eel lake"). The name is preserved in the new city of Almere in Flevoland , given in 1984 in memory of this body of water on whose site

900-593: Was accepted as a separate order from Anguilliformes, based on their distinct morphological differences from the rest of the "true eels". Genetic work over the past few years has shown that it is instead a derived lineage within the Anguilliformes. The four (proposed five) families in two suborders are: Eel see text Eels are ray-finned fish belonging to the order Anguilliformes ( / æ ŋ ˈ ɡ w ɪ l ɪ f ɔːr m iː z / ), which consists of eight suborders, 20 families , 164 genera , and about 1000 species . Eels undergo considerable development from

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