116-574: Robert Francis Scharff (9 July 1858 – 13 September 1934) was an English zoologist , known for his lifetime of work in Ireland and contributions to the understanding of Irish flora and fauna. He was acting director of the National Museum of Ireland from 1916 to 1922. Scharff was born in Leeds on 9 July 1858 to German parents. He studied at University College London for a B.Sc., and furthered those studies at
232-517: A will in which he asked to be buried next to his wife. Aristotle left his works to Theophrastus, his successor as the head of the Lyceum, who in turn passed them down to Neleus of Scepsis in Asia Minor. There, the papers remained hidden for protection until they were purchased by the collector Apellicon . In the meantime, many copies of Aristotle's major works had already begun to circulate and be used in
348-541: A "survival of the fittest" origin of living things and their organs, and ridiculed the idea that accidents could lead to orderly results. To put his views into modern terms, he nowhere says that different species can have a common ancestor , or that one kind can change into another , or that kinds can become extinct . Aristotle did not do experiments in the modern sense. He used the ancient Greek term pepeiramenoi to mean observations, or at most investigative procedures like dissection. In Generation of Animals , he finds
464-543: A building known as the Lyceum (named after the sacred grove of Apollo Lykeios ), in which he established his own school. The building included a gymnasium and a colonnade ( peripatos ), from which the school acquired the name Peripatetic . Aristotle conducted courses and research at the school for the next twelve years. He often lectured small groups of distinguished students and, along with some of them, such as Theophrastus , Eudemus , and Aristoxenus , Aristotle built
580-581: A daughter. Scharff died in Worthing on 13 September 1934. Scharff was appointed as an Assistant in the Natural History Division of the National Museum of Ireland in 1887, becoming Keeper in 1890. In the years after 1916 and the resignation of George Noble Plunkett as Director of the National Museum of Ireland Scharff became the acting Director until his retirement in 1922. Scharff was one of
696-438: A fertilized hen's egg of a suitable stage and opens it to see the embryo's heart beating inside. Instead, he practiced a different style of science: systematically gathering data, discovering patterns common to whole groups of animals, and inferring possible causal explanations from these. This style is common in modern biology when large amounts of data become available in a new field, such as genomics . It does not result in
812-569: A few salient points. Aristotle was born in 384 BC in Stagira , Chalcidice , about 55 km (34 miles) east of modern-day Thessaloniki . He was the son of Nicomachus , the personal physician of King Amyntas of Macedon , and Phaestis, a woman with origins from Chalcis , Euboea . Nicomachus was said to have belonged to the medical guild of Asclepiadae and was likely responsible for Aristotle's early interest in biology and medicine. Ancient tradition held that Aristotle's family descended from
928-497: A fluid such as air. In this system, heavy bodies in steady fall indeed travel faster than light ones (whether friction is ignored, or not ), and they do fall more slowly in a denser medium. Newton's "forced" motion corresponds to Aristotle's "violent" motion with its external agent, but Aristotle's assumption that the agent's effect stops immediately it stops acting (e.g., the ball leaves the thrower's hand) has awkward consequences: he has to suppose that surrounding fluid helps to push
1044-406: A form of an apple. In this distinction, there is a particular apple and a universal form of an apple. Moreover, one can place an apple next to a book, so that one can speak of both the book and apple as being next to each other. Plato argued that there are some universal forms that are not a part of particular things. For example, it is possible that there is no particular good in existence, but "good"
1160-601: A laboratory setting. Ethologists have been particularly concerned with the evolution of behavior and the understanding of behavior in terms of the theory of natural selection . In one sense, the first modern ethologist was Charles Darwin , whose book, The Expression of the Emotions in Man and Animals , influenced many future ethologists. A subfield of ethology is behavioral ecology which attempts to answer Nikolaas Tinbergen 's four questions with regard to animal behavior: what are
1276-550: A large library which included manuscripts, maps, and museum objects. While in Athens, his wife Pythias died and Aristotle became involved with Herpyllis of Stagira. They had a son whom Aristotle named after his father, Nicomachus . This period in Athens, between 335 and 323 BC, is when Aristotle is believed to have composed many of his philosophical works. He wrote many dialogues, of which only fragments have survived. Those works that have survived are in treatise form and were not, for
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#17330937266141392-409: A library in the Lyceum, which helped him to produce many of his hundreds of books on papyrus scrolls . Though Aristotle wrote many treatises and dialogues for publication, only around a third of his original output has survived , none of it intended for publication. Aristotle provided a complex synthesis of the various philosophies existing prior to him. His teachings and methods of inquiry have had
1508-597: A linear scale, and noted various exceptions, such as that sharks had a placenta like the tetrapods. To a modern biologist, the explanation, not available to Aristotle, is convergent evolution . Philosophers of science have generally concluded that Aristotle was not interested in taxonomy, but zoologists who studied this question in the early 21st century think otherwise. He believed that purposive final causes guided all natural processes; this teleological view justified his observed data as an expression of formal design. Aristotle's psychology , given in his treatise On
1624-413: A new footing, by explaining the processes by which it can occur, and providing observational evidence that it had done so. Darwin's theory was rapidly accepted by the scientific community and soon became a central axiom of the rapidly developing science of biology. The basis for modern genetics began with the work of Gregor Mendel on peas in 1865, although the significance of his work was not realized at
1740-406: A new perspective on the fundamental basis of life. Having previously been the realm of gentlemen naturalists, over the 18th, 19th and 20th centuries, zoology became an increasingly professional scientific discipline . Explorer-naturalists such as Alexander von Humboldt investigated the interaction between organisms and their environment, and the ways this relationship depends on geography, laying
1856-427: A plant in the soil is potentially ( dynamei ) a plant, and if it is not prevented by something, it will become a plant. Potentially, beings can either 'act' ( poiein ) or 'be acted upon' ( paschein ), which can be either innate or learned. For example, the eyes possess the potentiality of sight (innate – being acted upon), while the capability of playing the flute can be possessed by learning (exercise – acting). Actuality
1972-498: A priori principles. Aristotle's "natural philosophy" spans a wide range of natural phenomena including those now covered by physics, biology and other natural sciences. In Aristotle's terminology, "natural philosophy" is a branch of philosophy examining the phenomena of the natural world, and includes fields that would be regarded today as physics, biology and other natural sciences. Aristotle's work encompassed virtually all facets of intellectual inquiry. Aristotle makes philosophy in
2088-709: A process involving the union of a male and female haploid gamete , each gamete formed by meiosis . Ordinarily, gametes produced by separate individuals unite by a process of fertilization to form a diploid zygote that can then develop into a genetically unique individual progeny. However, some animals are also capable, as an alternative reproductive process, to reproduce parthenogenetically. Parthenogenesis has been described in snakes and lizards (see Misplaced Pages Parthenogenesis in squamates ), in amphibians (see Misplaced Pages Parthenogenesis in amphibians ) and in numerous other species (see Misplaced Pages Parthenogenesis ). Generally, meiosis in parthanogenetically reproducing animals occurs by
2204-562: A researcher and lecturer, earning for himself the nickname "mind of the school" by his tutor Plato . In Athens, he probably experienced the Eleusinian Mysteries as he wrote when describing the sights one viewed at the Mysteries, "to experience is to learn" ( παθεĩν μαθεĩν ). Aristotle remained in Athens for nearly twenty years before leaving in 348/47 BC after Plato's death. The traditional story about his departure records that he
2320-625: A significant impact across the world, and remain a subject of contemporary philosophical discussion. Aristotle's views profoundly shaped medieval scholarship . The influence of his physical science extended from late antiquity and the Early Middle Ages into the Renaissance , and was not replaced systematically until the Enlightenment and theories such as classical mechanics were developed. He influenced Judeo-Islamic philosophies during
2436-429: A similar process to that in sexually reproducing animals, but the diploid zygote nucleus is generated by the union of two haploid genomes from the same individual rather than from different individuals. Aristotle Aristotle ( Attic Greek : Ἀριστοτέλης , romanized: Aristotélēs ; 384–322 BC) was an Ancient Greek philosopher and polymath . His writings cover a broad range of subjects spanning
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#17330937266142552-549: A thrown stone, in the Physics (254b10), and "natural motion", such as of a falling object, in On the Heavens (300a20). In violent motion, as soon as the agent stops causing it, the motion stops also: in other words, the natural state of an object is to be at rest, since Aristotle does not address friction . With this understanding, it can be observed that, as Aristotle stated, heavy objects (on
2668-453: Is a form of scientific taxonomy . Modern biological classification has its root in the work of Carl Linnaeus , who grouped species according to shared physical characteristics. These groupings have since been revised to improve consistency with the Darwinian principle of common descent . Molecular phylogenetics , which uses nucleic acid sequence as data, has driven many recent revisions and
2784-454: Is a historical accident: his works on botany have been lost, but two books on plants by his pupil Theophrastus have survived. Aristotle reports on the sea-life visible from observation on Lesbos and the catches of fishermen. He describes the catfish , electric ray , and frogfish in detail, as well as cephalopods such as the octopus and paper nautilus . His description of the hectocotyl arm of cephalopods, used in sexual reproduction,
2900-657: Is administered by the International Code of Zoological Nomenclature . A merging draft, BioCode, was published in 1997 in an attempt to standardize nomenclature, but has yet to be formally adopted. Vertebrate zoology is the biological discipline that consists of the study of vertebrate animals, that is animals with a backbone , such as fish , amphibians , reptiles , birds and mammals . The various taxonomically oriented disciplines i.e. mammalogy , biological anthropology , herpetology , ornithology , and ichthyology seek to identify and classify species and study
3016-419: Is also a final cause or end. Then Aristotle proceeds and concludes that the actuality is prior to potentiality in formula, in time and in substantiality. With this definition of the particular substance (i.e., matter and form), Aristotle tries to solve the problem of the unity of the beings, for example, "what is it that makes a man one"? Since, according to Plato there are two Ideas: animal and biped, how then
3132-480: Is an integrative field of study, uniting concepts and information from evolutionary biology , taxonomy , ecology , physical geography , geology , paleontology and climatology . The origin of this field of study is widely accredited to Alfred Russel Wallace , a British biologist who had some of his work jointly published with Charles Darwin . Molecular biology studies the common genetic and developmental mechanisms of animals and plants, attempting to answer
3248-402: Is being studied. For example, what is learned about the physiology of yeast cells can also apply to human cells. The field of animal physiology extends the tools and methods of human physiology to non-human species. Physiology studies how, for example, the nervous , immune , endocrine , respiratory , and circulatory systems function and interact. Developmental biology is the study of
3364-406: Is done on both the microscopic and molecular levels for single-celled organisms such as bacteria as well as the specialized cells in multicellular organisms such as humans . Understanding the structure and function of cells is fundamental to all of the biological sciences. The similarities and differences between cell types are particularly relevant to molecular biology. Anatomy considers
3480-443: Is likely to continue to do so. Biological classification belongs to the science of zoological systematics . Many scientists now consider the five-kingdom system outdated. Modern alternative classification systems generally start with the three-domain system : Archaea (originally Archaebacteria); Bacteria (originally Eubacteria); Eukaryota (including protists , fungi , plants , and animals ) These domains reflect whether
3596-541: Is man a unity? However, according to Aristotle, the potential being (matter) and the actual one (form) are one and the same. Aristotle's immanent realism means his epistemology is based on the study of things that exist or happen in the world, and rises to knowledge of the universal, whereas for Plato epistemology begins with knowledge of universal Forms (or ideas) and descends to knowledge of particular imitations of these. Aristotle uses induction from examples alongside deduction , whereas Plato relies on deduction from
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3712-545: Is one of the primary branches of biology . The term is derived from Ancient Greek ζῷον , zōion ('animal'), and λόγος , logos ('knowledge', 'study'). Although humans have always been interested in the natural history of the animals they saw around them, and used this knowledge to domesticate certain species, the formal study of zoology can be said to have originated with Aristotle . He viewed animals as living organisms, studied their structure and development, and considered their adaptations to their surroundings and
3828-415: Is partly based on paleontology , which uses the fossil record to answer questions about the mode and tempo of evolution, and partly on the developments in areas such as population genetics and evolutionary theory. Following the development of DNA fingerprinting techniques in the late 20th century, the application of these techniques in zoology has increased the understanding of animal populations. In
3944-406: Is predicated. So, according to Aristotle, the form of apple exists within each apple, rather than in the world of the forms. Concerning the nature of change ( kinesis ) and its causes, as he outlines in his Physics and On Generation and Corruption ( 319b–320a), he distinguishes coming-to-be ( genesis , also translated as 'generation') from: Coming-to-be is a change where the substrate of
4060-497: Is probably not in its original form, because it was most likely edited by students and later lecturers. The logical works of Aristotle were compiled into a set of six books called the Organon around 40 BC by Andronicus of Rhodes or others among his followers. The books are: The order of the books (or the teachings from which they are composed) is not certain, but this list was derived from analysis of Aristotle's writings. It goes from
4176-406: Is quantitative, and recently a significant amount of work has been done using computer science techniques such as bioinformatics and computational biology . Molecular genetics , the study of gene structure and function, has been among the most prominent sub-fields of molecular biology since the early 2000s. Other branches of biology are informed by molecular biology, by either directly studying
4292-698: Is reflected in zoology by the presence of animals in Egyptian hieroglyphics. Although the concept of zoology as a single coherent field arose much later, the zoological sciences emerged from natural history reaching back to the biological works of Aristotle and Galen in the ancient Greco-Roman world . In the fourth century BC, Aristotle looked at animals as living organisms, studying their structure, development and vital phenomena. He divided them into two groups: animals with blood, equivalent to our concept of vertebrates , and animals without blood, invertebrates . He spent two years on Lesbos , observing and describing
4408-480: Is still a proper universal form. Aristotle disagreed with Plato on this point, arguing that all universals are instantiated at some period of time, and that there are no universals that are unattached to existing things. In addition, Aristotle disagreed with Plato about the location of universals. Where Plato spoke of the forms as existing separately from the things that participate in them, Aristotle maintained that universals exist within each thing on which each universal
4524-413: Is the fulfilment of the end of the potentiality. Because the end ( telos ) is the principle of every change, and potentiality exists for the sake of the end, actuality, accordingly, is the end. Referring then to the previous example, it can be said that an actuality is when a plant does one of the activities that plants do. For that for the sake of which ( to hou heneka ) a thing is, is its principle, and
4640-559: Is the genus, and sapiens the specific epithet, both of them combined make up the species name. When writing the scientific name of an organism, it is proper to capitalize the first letter in the genus and put all of the specific epithet in lowercase. Additionally, the entire term may be italicized or underlined. The dominant classification system is called the Linnaean taxonomy . It includes ranks and binomial nomenclature . The classification, taxonomy , and nomenclature of zoological organisms
4756-614: Is the subfield of biology that studies the evolutionary processes (natural selection, common descent, speciation) that produced the diversity of life on Earth. Evolutionary research is concerned with the origin and descent of species , as well as their change over time, and includes scientists from many taxonomically oriented disciplines. For example, it generally involves scientists who have special training in particular organisms such as mammalogy , ornithology , herpetology , or entomology , but use those organisms as systems to answer general questions about evolution. Evolutionary biology
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4872-539: The Iliad , which reportedly became one of Alexander's most prized possessions. Scholars speculate that two of Aristotle's now lost works, On kingship and On behalf of the Colonies , were composed by the philosopher for the young prince. After Philip II's assassination in 336 BC, Aristotle returned to Athens for the second and final time a year later. As a metic , Aristotle could not own property in Athens and thus rented
4988-531: The Classical period . His father, Nicomachus , died when Aristotle was a child, and he was brought up by a guardian. At around eighteen years old, he joined Plato 's Academy in Athens and remained there until the age of thirty seven ( c. 347 BC ). Shortly after Plato died, Aristotle left Athens and, at the request of Philip II of Macedon , tutored his son Alexander the Great beginning in 343 BC. He established
5104-634: The Early Modern period. John Philoponus (in Late antiquity ) and Galileo (in Early modern period ) are said to have shown by experiment that Aristotle's claim that a heavier object falls faster than a lighter object is incorrect. A contrary opinion is given by Carlo Rovelli , who argues that Aristotle's physics of motion is correct within its domain of validity, that of objects in the Earth 's gravitational field immersed in
5220-515: The Great Comet of 371 BC . Aristotle was one of the first people to record any geological observations. He stated that geological change was too slow to be observed in one person's lifetime. The geologist Charles Lyell noted that Aristotle described such change, including "lakes that had dried up" and "deserts that had become watered by rivers", giving as examples the growth of the Nile delta since
5336-471: The History of Animals in a graded scale of perfection, a nonreligious version of the scala naturae , with man at the top. His system had eleven grades of animal, from highest potential to lowest, expressed in their form at birth: the highest gave live birth to hot and wet creatures, the lowest laid cold, dry mineral-like eggs. Animals came above plants , and these in turn were above minerals. He grouped what
5452-451: The Milky Way was made up of "those stars which are shaded by the earth from the sun's rays," pointing out partly correctly that if "the size of the sun is greater than that of the earth and the distance of the stars from the earth many times greater than that of the sun, then... the sun shines on all the stars and the earth screens none of them." He also wrote descriptions of comets, including
5568-508: The Physics (215a25), Aristotle effectively states a quantitative law, that the speed, v, of a falling body is proportional (say, with constant c) to its weight, W, and inversely proportional to the density, ρ, of the fluid in which it is falling:; Aristotle implies that in a vacuum the speed of fall would become infinite, and concludes from this apparent absurdity that a vacuum is not possible. Opinions have varied on whether Aristotle intended to state quantitative laws. Henri Carteron held
5684-755: The Royal Zoological Society of Ireland , the Royal Dublin Society , the Conchological Society of Great Britain & Ireland , the Linnean Society and the Zoological Society of London . He collected zoological specimens extensively, many of which are housed with the National Museum of Ireland. In 1899 a species of land planarian was named in his honour, Microplana scharffi (Graff, 1899) . Following his retirement, Scharff moved back to
5800-545: The anatomy of different groups. It is closely related to evolutionary biology and phylogeny (the evolution of species). Physiology studies the mechanical, physical, and biochemical processes of living organisms by attempting to understand how all of the structures function as a whole. The theme of "structure to function" is central to biology. Physiological studies have traditionally been divided into plant physiology and animal physiology , but some principles of physiology are universal, no matter what particular organism
5916-526: The natural sciences , philosophy , linguistics , economics , politics , psychology , and the arts . As the founder of the Peripatetic school of philosophy in the Lyceum in Athens , he began the wider Aristotelian tradition that followed, which set the groundwork for the development of modern science . Little is known about Aristotle's life. He was born in the city of Stagira in northern Greece during
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#17330937266146032-503: The proximate causes of the behavior, the developmental history of the organism, the survival value and phylogeny of the behavior? Another area of study is animal cognition , which uses laboratory experiments and carefully controlled field studies to investigate an animal's intelligence and learning. Biogeography studies the spatial distribution of organisms on the Earth , focusing on topics like dispersal and migration , plate tectonics , climate change , and cladistics . It
6148-399: The taxonomist . Originally it was thought that species were immutable, but with the arrival of Darwin's theory of evolution, the field of cladistics came into being, studying the relationships between the different groups or clades . Systematics is the study of the diversification of living forms, the evolutionary history of a group is known as its phylogeny , and the relationship between
6264-513: The "extreme view" that Aristotle's concept of force was basically qualitative, but other authors reject this. Archimedes corrected Aristotle's theory that bodies move towards their natural resting places; metal boats can float if they displace enough water ; floating depends in Archimedes' scheme on the mass and volume of the object, not, as Aristotle thought, its elementary composition. Aristotle's writings on motion remained influential until
6380-489: The 16th century. During the Renaissance and early modern period, zoological thought was revolutionized in Europe by a renewed interest in empiricism and the discovery of many novel organisms. Prominent in this movement were Andreas Vesalius and William Harvey , who used experimentation and careful observation in physiology , and naturalists such as Carl Linnaeus , Jean-Baptiste Lamarck , and Buffon who began to classify
6496-470: The 1980s, developmental biology re-entered evolutionary biology from its initial exclusion from the modern synthesis through the study of evolutionary developmental biology . Related fields often considered part of evolutionary biology are phylogenetics , systematics , and taxonomy . Ethology is the scientific and objective study of animal behavior under natural conditions, as opposed to behaviorism , which focuses on behavioral response studies in
6612-550: The Lyceum of Athens, Alexandria , and later in Rome . With the Prior Analytics , Aristotle is credited with the earliest study of formal logic, and his conception of it was the dominant form of Western logic until 19th-century advances in mathematical logic . Kant stated in the Critique of Pure Reason that with Aristotle, logic reached its completion. Most of Aristotle's work
6728-621: The Middle Ages, as well as Christian theology , especially the Neoplatonism of the Early Church and the scholastic tradition of the Catholic Church . Aristotle was revered among medieval Muslim scholars as "The First Teacher", and among medieval Christians like Thomas Aquinas as simply "The Philosopher", while the poet Dante called him "the master of those who know". His works contain
6844-548: The UK, to Worthing. He still remained an active member of Irish zoological groups until his death in 1934. Zoology Zoology ( UK : / z u ˈ ɒ l ə dʒ i / zoo- OL -ə-jee , US : / z oʊ ˈ ɒ l ə dʒ i / zoh- OL -ə-jee ) is the scientific study of animals . Its studies include the structure , embryology , classification , habits , and distribution of all animals, both living and extinct , and how they interact with their ecosystems . Zoology
6960-449: The animals and plants, considering the adaptations of different organisms and the function of their parts. Four hundred years later, Roman physician Galen dissected animals to study their anatomy and the function of the different parts, because the dissection of human cadavers was prohibited at the time. This resulted in some of his conclusions being false, but for many centuries it was considered heretical to challenge any of his views, so
7076-489: The ball along to make it continue to rise even though the hand is no longer acting on it, resulting in the Medieval theory of impetus . Aristotle suggested that the reason for anything coming about can be attributed to four different types of simultaneously active factors. His term aitia is traditionally translated as "cause", but it does not always refer to temporal sequence; it might be better translated as "explanation", but
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#17330937266147192-617: The basics, the analysis of simple terms in the Categories, the analysis of propositions and their elementary relations in On Interpretation , to the study of more complex forms, namely, syllogisms (in the Analytics ) and dialectics (in the Topics and Sophistical Refutations ). The first three treatises form the core of the logical theory stricto sensu : the grammar of the language of logic and
7308-399: The becoming is for the sake of the end; and the actuality is the end, and it is for the sake of this that the potentiality is acquired. For animals do not see in order that they may have sight, but they have sight that they may see. In summary, the matter used to make a house has potentiality to be a house and both the activity of building and the form of the final house are actualities, which
7424-574: The broad sense coextensive with reasoning, which he also would describe as "science". However, his use of the term science carries a different meaning than that covered by the term "scientific method". For Aristotle, "all science ( dianoia ) is either practical, poetical or theoretical" ( Metaphysics 1025b25). His practical science includes ethics and politics; his poetical science means the study of fine arts including poetry; his theoretical science covers physics, mathematics and metaphysics. In his On Generation and Corruption , Aristotle related each of
7540-433: The cause of earthquakes was a gas or vapor ( anathymiaseis ) that was trapped inside the earth and trying to escape, following other Greek authors Anaxagoras , Empedocles and Democritus . Aristotle also made many observations about the hydrologic cycle. For example, he made some of the earliest observations about desalination: he observed early – and correctly – that when seawater is heated, freshwater evaporates and that
7656-427: The cells have nuclei or not, as well as differences in the chemical composition of the cell exteriors. Further, each kingdom is broken down recursively until each species is separately classified. The order is: Domain ; kingdom ; phylum ; class ; order ; family ; genus ; species . The scientific name of an organism is generated from its genus and species. For example, humans are listed as Homo sapiens . Homo
7772-434: The clades can be shown diagrammatically in a cladogram . Although someone who made a scientific study of animals would historically have described themselves as a zoologist, the term has come to refer to those who deal with individual animals, with others describing themselves more specifically as physiologists, ethologists, evolutionary biologists, ecologists, pharmacologists, endocrinologists or parasitologists. Although
7888-487: The correct rules of reasoning. The Rhetoric is not conventionally included, but it states that it relies on the Topics . What is today called Aristotelian logic with its types of syllogism (methods of logical argument), Aristotle himself would have labelled "analytics". The term "logic" he reserved to mean dialectics . The word "metaphysics" appears to have been coined by the first century AD editor who assembled various small selections of Aristotle's works to create
8004-399: The data to come to the theory of evolution . Aristotle's writings can seem to modern readers close to implying evolution, but while Aristotle was aware that new mutations or hybridizations could occur, he saw these as rare accidents. For Aristotle, accidents, like heat waves in winter, must be considered distinct from natural causes. He was thus critical of Empedocles's materialist theory of
8120-787: The death. Following Alexander's death, anti-Macedonian sentiment in Athens was rekindled. In 322 BC, Demophilus and Eurymedon the Hierophant reportedly denounced Aristotle for impiety, prompting him to flee to his mother's family estate in Chalcis, Euboea , at which occasion he was said to have stated "I will not allow the Athenians to sin twice against philosophy" – a reference to Athens's trial and execution of Socrates . He died in Chalcis, Euboea of natural causes later that same year, having named his student Antipater as his chief executor and leaving
8236-514: The determination of the double helical structure of the DNA molecule by Francis Crick and James Watson in 1953, the realm of molecular biology opened up, leading to advances in cell biology , developmental biology and molecular genetics . The study of systematics was transformed as DNA sequencing elucidated the degrees of affinity between different organisms. Zoology is the branch of science dealing with animals . A species can be defined as
8352-643: The discovery of the structure of DNA by Francis Crick and James Watson in 1953, the realm of molecular biology opened up, leading to advances in cell biology , developmental biology and molecular genetics . The history of zoology traces the study of the animal kingdom from ancient to modern times. Prehistoric people needed to study the animals and plants in their environment to exploit them and survive. Cave paintings, engravings and sculptures in France dating back 15,000 years show bison, horses, and deer in carefully rendered detail. Similar images from other parts of
8468-456: The diversity of life and the fossil record , as well as studying the development and behavior of organisms. Antonie van Leeuwenhoek did pioneering work in microscopy and revealed the previously unknown world of microorganisms , laying the groundwork for cell theory . van Leeuwenhoek's observations were endorsed by Robert Hooke ; all living organisms were composed of one or more cells and could not generate spontaneously. Cell theory provided
8584-479: The earliest known formal study of logic, and were studied by medieval scholars such as Peter Abelard and Jean Buridan . Aristotle's influence on logic continued well into the 19th century. In addition, his ethics , although always influential, gained renewed interest with the modern advent of virtue ethics . In general, the details of Aristotle's life are not well-established. The biographies written in ancient times are often speculative and historians only agree on
8700-543: The end of his life, the two men became estranged having diverging opinions over issues, like the optimal administration of city-states, the treatment of conquered populations, such as the Persians, and philosophical questions, like the definition of braveness. A widespread speculation in antiquity suggested that Aristotle played a role in Alexander's death, but the only evidence of this is an unlikely claim made some six years after
8816-533: The following. Brood size decreases with (adult) body mass, so that an elephant has fewer young (usually just one) per brood than a mouse. Lifespan increases with gestation period , and also with body mass, so that elephants live longer than mice, have a longer period of gestation, and are heavier. As a final example, fecundity decreases with lifespan, so long-lived kinds like elephants have fewer young in total than short-lived kinds like mice. Aristotle distinguished about 500 species of animals , arranging these in
8932-403: The form of the substance is the actual house, namely 'covering for bodies and chattels' or any other differentia that let us define something as a house. The formula that gives the components is the account of the matter, and the formula that gives the differentia is the account of the form. Like his teacher Plato, Aristotle's philosophy aims at the universal . Aristotle's ontology places
9048-404: The forms of macroscopic structures such as organs and organ systems. It focuses on how organs and organ systems work together in the bodies of humans and other animals, in addition to how they work independently. Anatomy and cell biology are two studies that are closely related, and can be categorized under "structural" studies. Comparative anatomy is the study of similarities and differences in
9164-431: The foundations for biogeography , ecology and ethology . Naturalists began to reject essentialism and consider the importance of extinction and the mutability of species . These developments, as well as the results from embryology and paleontology , were synthesized in the 1859 publication of Charles Darwin 's theory of evolution by natural selection ; in this Darwin placed the theory of organic evolution on
9280-509: The four elements proposed earlier by Empedocles , earth , water , air , and fire , to two of the four sensible qualities, hot, cold, wet, and dry. In the Empedoclean scheme, all matter was made of the four elements, in differing proportions. Aristotle's scheme added the heavenly aether , the divine substance of the heavenly spheres , stars and planets. Aristotle describes two kinds of motion: "violent" or "unnatural motion", such as that of
9396-502: The function of their parts. Modern zoology has its origins during the Renaissance and early modern period, with Carl Linnaeus , Antonie van Leeuwenhoek , Robert Hooke , Charles Darwin , Gregor Mendel and many others. The study of animals has largely moved on to deal with form and function, adaptations, relationships between groups, behaviour and ecology. Zoology has increasingly been subdivided into disciplines such as classification , physiology , biochemistry and evolution . With
9512-428: The ground, say) require more force to make them move; and objects pushed with greater force move faster. This would imply the equation incorrect in modern physics. Natural motion depends on the element concerned: the aether naturally moves in a circle around the heavens, while the 4 Empedoclean elements move vertically up (like fire, as is observed) or down (like earth) towards their natural resting places. In
9628-440: The image. According to Aristotle, spontaneity and chance are causes of some things, distinguishable from other types of cause such as simple necessity. Chance as an incidental cause lies in the realm of accidental things , "from what is spontaneous". There is also more a specific kind of chance, which Aristotle names "luck", that only applies to people's moral choices. In astronomy , Aristotle refuted Democritus 's claim that
9744-476: The interactions of molecules in their own right such as in cell biology and developmental biology , or indirectly, where molecular techniques are used to infer historical attributes of populations or species , as in fields in evolutionary biology such as population genetics and phylogenetics . There is also a long tradition of studying biomolecules "from the ground up", or molecularly, in biophysics . Animals generally reproduce by sexual reproduction ,
9860-404: The largest group of organisms in which any two individuals of the appropriate sex can produce fertile offspring; about 1.5 million species of animal have been described and it has been estimated that as many as 8 million animal species may exist. An early necessity was to identify the organisms and group them according to their characteristics, differences and relationships, and this is the field of
9976-558: The legendary physician Asclepius and his son Machaon . Both of Aristotle's parents died when he was still at a young age and Proxenus of Atarneus became his guardian. Although little information about Aristotle's childhood has survived, he probably spent some time in the Macedonian capital, making his first connections with the Macedonian monarchy . At the age of seventeen or eighteen, Aristotle moved to Athens to continue his education at Plato's Academy . He became distinguished as
10092-549: The main organisers and contributors to the Clare Island Survey , serving as a chair of the organising committee. He is best known for his work on biogeography , particularly the recolonisation of Ireland after the ice age. He was a vocal proponent of the land-bridge hypothesis of animal migration. This work was informed by his experience excavating caves in Kesh . Scharff was a well-known figure in his time receiving many honours like
10208-468: The modern zoologist would call vertebrates as the hotter "animals with blood", and below them the colder invertebrates as "animals without blood". Those with blood were divided into the live-bearing ( mammals ), and the egg-laying ( birds , reptiles , fish ). Those without blood were insects, crustacea (non-shelled – cephalopods, and shelled ) and the hard-shelled molluscs ( bivalves and gastropods ). He recognised that animals did not exactly fit into
10324-528: The most part, intended for widespread publication; they are generally thought to be lecture aids for his students. His most important treatises include Physics , Metaphysics , Nicomachean Ethics , Politics , On the Soul and Poetics . Aristotle studied and made significant contributions to "logic, metaphysics, mathematics, physics, biology, botany, ethics, politics, agriculture, medicine, dance, and theatre." While Alexander deeply admired Aristotle, near
10440-498: The near-by island of Lesbos . During this time, Aristotle married Pythias , Hermias's adoptive daughter and niece, and had a daughter whom they also named Pythias. In 343/42 BC, Aristotle was invited to Pella by Philip II of Macedon in order to become the tutor to his thirteen-year-old son Alexander ; a choice perhaps influenced by the relationship of Aristotle's family with the Macedonian dynasty. Aristotle taught Alexander at
10556-404: The oceans are then replenished by the cycle of rainfall and river runoff ("I have proved by experiment that salt water evaporated forms fresh and the vapor does not when it condenses condense into sea water again.") Aristotle was the first person to study biology systematically, and biology forms a large part of his writings. He spent two years observing and describing the zoology of Lesbos and
10672-437: The primary kind of knowledge; but if there is some motionless independent thing, the knowledge of this precedes it and is first philosophy, and it is universal in just this way , because it is first. And it belongs to this sort of philosophy to study being as being, both what it is and what belongs to it just by virtue of being. Aristotle examines the concepts of substance ( ousia ) and essence ( to ti ên einai , "the what it
10788-567: The private school of Mieza , in the gardens of the Nymphs , the royal estate near Pella. Alexander's education probably included a number of subjects, such as ethics and politics , as well as standard literary texts, like Euripides and Homer . It is likely that during Aristotle's time in the Macedonian court, other prominent nobles, like Ptolemy and Cassander , would have occasionally attended his lectures. Aristotle encouraged Alexander toward eastern conquest, and his own attitude towards Persia
10904-562: The prize of Emperor Nicholas II in 1897, and he holding the Swiney lectureship in geology in London in 1906 and 1908. He was an active member of many societies in Ireland and the UK. Whilst a member of the Royal Irish Academy over a forty-year period, he served as vice-president, secretary for foreign correspondence, and chairman of the flora and fauna committee. He was also an active member of
11020-517: The processes by which animals and plants reproduce and grow. The discipline includes the study of embryonic development , cellular differentiation , regeneration , asexual and sexual reproduction , metamorphosis , and the growth and differentiation of stem cells in the adult organism. Development of both animals and plants is further considered in the articles on evolution , population genetics , heredity , genetic variability , Mendelian inheritance , and reproduction . Evolutionary biology
11136-493: The questions regarding the mechanisms of genetic inheritance and the structure of the gene . In 1953, James Watson and Francis Crick described the structure of DNA and the interactions within the molecule, and this publication jump-started research into molecular biology and increased interest in the subject. While researchers practice techniques specific to molecular biology, it is common to combine these with methods from genetics and biochemistry . Much of molecular biology
11252-494: The results of his own investigations. His general approach was surprisingly modern, and he wrote, "For it is [the task] of natural science not simply to accept what we are told but to inquire into the causes of natural things." An early pioneer was Conrad Gessner , whose monumental 4,500-page encyclopedia of animals, Historia animalium , was published in four volumes between 1551 and 1558. In Europe, Galen's work on anatomy remained largely unsurpassed and unchallenged up until
11368-503: The rise of the germ theory of disease , though the mechanism of inheritance remained a mystery. In the early 20th century, the rediscovery of Mendel's work led to the rapid development of genetics , and by the 1930s the combination of population genetics and natural selection in the modern synthesis created evolutionary biology . Research in cell biology is interconnected to other fields such as genetics, biochemistry , medical microbiology , immunology , and cytochemistry . With
11484-433: The same certainty as experimental science, but it sets out testable hypotheses and constructs a narrative explanation of what is observed. In this sense, Aristotle's biology is scientific. From the data he collected and documented, Aristotle inferred quite a number of rules relating the life-history features of the live-bearing tetrapods (terrestrial placental mammals) that he studied. Among these correct predictions are
11600-499: The structures and mechanisms specific to those groups. The rest of the animal kingdom is dealt with by invertebrate zoology , a vast and very diverse group of animals that includes sponges , echinoderms , tunicates , worms , molluscs , arthropods and many other phyla , but single-celled organisms or protists are not usually included. Cell biology studies the structural and physiological properties of cells , including their behavior , interactions, and environment . This
11716-529: The study of anatomy stultified. During the post-classical era , Middle Eastern science and medicine was the most advanced in the world, integrating concepts from Ancient Greece, Rome, Mesopotamia and Persia as well as the ancient Indian tradition of Ayurveda , while making numerous advances and innovations. In the 13th century, Albertus Magnus produced commentaries and paraphrases of all Aristotle's works; his books on topics like botany , zoology, and minerals included information from ancient sources, but also
11832-565: The study of animal life is ancient, its scientific incarnation is relatively modern. This mirrors the transition from natural history to biology at the start of the 19th century. Since Hunter and Cuvier , comparative anatomical study has been associated with morphography , shaping the modern areas of zoological investigation: anatomy , physiology , histology , embryology , teratology and ethology . Modern zoology first arose in German and British universities. In Britain, Thomas Henry Huxley
11948-549: The surrounding seas, including in particular the Pyrrha lagoon in the centre of Lesbos. His data in History of Animals , Generation of Animals , Movement of Animals , and Parts of Animals are assembled from his own observations, statements given by people with specialized knowledge, such as beekeepers and fishermen, and less accurate accounts provided by travellers from overseas. His apparent emphasis on animals rather than plants
12064-402: The thing that has undergone the change has itself changed. In that particular change he introduces the concept of potentiality ( dynamis ) and actuality ( entelecheia ) in association with the matter and the form. Referring to potentiality, this is what a thing is capable of doing or being acted upon if the conditions are right and it is not prevented by something else. For example, the seed of
12180-470: The time of Homer , and "the upheaving of one of the Aeolian islands , previous to a volcanic eruption ."' Meteorologica lends its name to the modern study of meteorology, but its modern usage diverges from the content of Aristotle's ancient treatise on meteors . The ancient Greeks did use the term for a range of atmospheric phenomena, but also for earthquakes and volcanic eruptions. Aristotle proposed that
12296-433: The time. Darwin gave a new direction to morphology and physiology , by uniting them in a common biological theory: the theory of organic evolution. The result was a reconstruction of the classification of animals upon a genealogical basis, fresh investigation of the development of animals, and early attempts to determine their genetic relationships. The end of the 19th century saw the fall of spontaneous generation and
12412-473: The traditional rendering will be employed here. Aristotle describes experiments in optics using a camera obscura in Problems , book 15. The apparatus consisted of a dark chamber with a small aperture that let light in. With it, he saw that whatever shape he made the hole, the sun's image always remained circular. He also noted that increasing the distance between the aperture and the image surface magnified
12528-401: The treatise we know by the name Metaphysics . Aristotle called it "first philosophy", and distinguished it from mathematics and natural science (physics) as the contemplative ( theoretikē ) philosophy which is "theological" and studies the divine. He wrote in his Metaphysics (1026a16): If there were no other independent things besides the composite natural ones, the study of nature would be
12644-492: The universal ( katholou ) in particulars ( kath' hekaston ), things in the world, whereas for Plato the universal is a separately existing form which actual things imitate. For Aristotle, "form" is still what phenomena are based on, but is "instantiated" in a particular substance. Plato argued that all things have a universal form , which could be either a property or a relation to other things. When one looks at an apple, for example, one sees an apple, and one can also analyse
12760-505: The universities of Edinburgh with an MA graduating in 1885, and Heidelberg where he completed a Ph.D. From there he went on to study at St. Andrews Biological Station in New Brunswick and Stazione Zoologica in Naples. Scharff was twice married, his first wife Alice Hutton died during the 1918 flu pandemic , they had two sons together. In 1920 he married Jane Stephens with whom he had
12876-489: The world illustrated mostly the animals hunted for food and the savage animals. The Neolithic Revolution , which is characterized by the domestication of animals , continued throughout Antiquity. Ancient knowledge of wildlife is illustrated by the realistic depictions of wild and domestic animals in the Near East, Mesopotamia, and Egypt, including husbandry practices and techniques, hunting and fishing. The invention of writing
12992-502: Was a prominent figure. His ideas were centered on the morphology of animals. Many consider him the greatest comparative anatomist of the latter half of the 19th century. Similar to Hunter , his courses were composed of lectures and laboratory practical classes in contrast to the previous format of lectures only. Scientific classification in zoology , is a method by which zoologists group and categorize organisms by biological type , such as genus or species . Biological classification
13108-670: Was disappointed with the Academy's direction after control passed to Plato's nephew Speusippus , although it is possible that the anti-Macedonian sentiments in Athens could have also influenced his decision. Aristotle left with Xenocrates to Assos in Asia Minor , where he was invited by his former fellow student Hermias of Atarneus ; he stayed there for a few years and left around the time of Hermias' death. While at Assos, Aristotle and his colleague Theophrastus did extensive research in botany and marine biology , which they later continued at
13224-474: Was strongly ethnocentric . In one famous example, he counsels Alexander to be "a leader to the Greeks and a despot to the barbarians". Alexander's education under the guardianship of Aristotle likely lasted for only a few years, as at around the age of sixteen he returned to Pella and was appointed regent of Macedon by his father Philip. During this time, Aristotle is said to have gifted Alexander an annotated copy of
13340-421: Was to be") in his Metaphysics (Book VII), and he concludes that a particular substance is a combination of both matter and form, a philosophical theory called hylomorphism . In Book VIII, he distinguishes the matter of the substance as the substratum , or the stuff of which it is composed. For example, the matter of a house is the bricks, stones, timbers, etc., or whatever constitutes the potential house, while
13456-588: Was widely disbelieved until the 19th century. He gives accurate descriptions of the four-chambered fore-stomachs of ruminants , and of the ovoviviparous embryological development of the hound shark . He notes that an animal's structure is well matched to function so birds like the heron (which live in marshes with soft mud and live by catching fish) have a long neck, long legs, and a sharp spear-like beak, whereas ducks that swim have short legs and webbed feet. Darwin , too, noted these sorts of differences between similar kinds of animal, but unlike Aristotle used
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