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59-443: Rice is a cereal grain. Rice , Rice's , or Rices may also refer to: Rice Rice is a cereal grain and in its domesticated form is the staple food of over half of the world's population , particularly in Asia and Africa . Rice is the seed of the grass species Oryza sativa (Asian rice)—or, much less commonly, Oryza glaberrima (African rice). Asian rice

118-403: A branched inflorescence which arises from the last internode on the stem. There can be up to 350 spikelets in a panicle, each containing male and female flower parts ( anthers and ovule ). A fertilised ovule develops into the edible grain or caryopsis . Rice is a cereal belonging to the family Poaceae . As a tropical crop, it can be grown during the two distinct seasons (dry and wet) of

177-417: A mature embryo as the resources within the seed are limited. In flowering plants, one sperm nucleus fuses with the egg cell to produce a zygote, the other fuses with the two polar nuclei of the central cell to give rise to the polyploid (typically triploid) endosperm . This double fertilization is unique to flowering plants, although in some other groups the second sperm cell does fuse with another cell in

236-411: A role in various religions and traditions, such as in weddings . The rice plant can grow to over 1 m (3 ft) tall; if in deep water, it can reach a length of 5 m (16 ft). A single plant may have several leafy stems or tillers . The upright stem is jointed with nodes along its length; a long slender leaf arises from each node. The self-fertile flowers are produced in a panicle ,

295-506: A single domestication event for O. sativa . Both indica and japonica forms of Asian rice sprang from a single domestication event in China from the wild rice Oryza rufipogon . Despite this evidence, it appears that indica rice arose when japonica arrived in India about 4,500 years ago and hybridised with another rice, whether an undomesticated proto- indica or wild O. nivara . Rice

354-440: A wide variety of pests including insects, nematodes, rodents such as rats, snails, and birds. Major rice insect pests include armyworms, rice bugs , black bugs , cutworms, field crickets, grasshoppers, leafhoppers, mealybugs, and planthoppers. High rates of nitrogen fertiliser application may worsen aphid outbreaks. Weather conditions can contribute to pest outbreaks: rice gall midge outbreaks are worsened by high rainfall in

413-436: Is rainfed like wheat or maize. Across Asia, unmilled rice or "paddy" (Indonesian and Malay padi ), was traditionally the product of smallholder agriculture, with manual harvesting . Larger farms make use of machines such as combine harvesters to reduce the input of labour. The grain is ready to harvest when the moisture content is 20–25%. Harvesting involves reaping , stacking the cut stalks, threshing to separate

472-661: Is 69% water, 29% carbohydrates , 2% protein , and contains negligible fat (table). In a reference serving of 100 grams (3.5 oz), cooked white rice provides 130 calories of food energy , and contains moderate levels of manganese (18% DV), with no other micronutrients in significant content (all less than 10% of the Daily Value ). In 2018, the World Health Organization strongly recommended fortifying rice with iron , and conditionally recommended fortifying it with vitamin A and with folic acid . Golden rice

531-613: Is a major food staple in Asia, Latin America, and some parts of Africa, feeding over half the world's population. However, a substantial part of the crop can be lost post-harvest through inefficient transportation, storage, and milling. A quarter of the crop in Nigeria is lost after harvest. Storage losses include damage by mould fungi if the rice is not dried sufficiently. In China, losses in modern metal silos were just 0.2%, compared to 7–13% when rice

590-498: Is a variety produced through genetic engineering to synthesize beta-carotene , a precursor of vitamin A, in the endosperm of the rice grain. It is intended to be grown and eaten in parts of the world where Vitamin A deficiency is prevalent. Golden rice has been opposed by activists, such as in the Philippines . In 2016 more than 100 Nobel laureates encouraged the use of genetically modified organisms , such as golden rice, for

649-581: Is based on four principles: biodiversity, host plant resistance, landscape ecology, and hierarchies in a landscape—from biological to social. Farmers' pesticide applications are often unnecessary. Pesticides may actually induce resurgence of populations of rice pests such as the brown planthopper , both by destroying beneficial insects and by enhancing the pest's reproduction. The International Rice Research Institute (IRRI) demonstrated in 1993 that an 87.5% reduction in pesticide use can lead to an overall drop in pest numbers. Farmers in China, Indonesia and

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708-504: Is from Asia. The average world yield for rice was 4.7 metric tons per hectare (2.1 short tons per acre), in 2022. Yuan Longping of China's National Hybrid Rice Research and Development Center set a world record for rice yield in 1999 at 17.1 metric tons per hectare (7.6 short tons per acre) on a demonstration plot. This employed specially developed hybrid rice and the System of Rice Intensification (SRI), an innovation in rice farming. Rice

767-442: Is in downward position and chalazal end in on the upper position hence, in amphitropous the anatropous arrangement is tilted 90 degrees and in orthotropous it is completely inverted) . The ovule appears to be a megasporangium with integuments surrounding it. Ovules are initially composed of diploid maternal tissue, which includes a megasporocyte (a cell that will undergo meiosis to produce megaspores). Megaspores remain inside

826-401: Is part of the inner structure of the ovule, forming a layer of diploid ( sporophytic ) cells immediately inside the integuments. It is structurally and functionally equivalent to the megasporangium . In immature ovules, the nucellus contains a megasporocyte (megaspore mother cell), which undergoes sporogenesis via meiosis . In the megasporocyte of Arabidopsis thaliana , meiosis depends on

885-797: Is sister to the bamboos, Bambusoideae , and the cereal subfamily Pooideae . The rice genus Oryza is one of eleven in the Oryzeae; it is sister to the Phyllorachideae . The edible rice species O. sativa and O. glaberrima are among some 300 species or subspecies in the genus. other grasses (inc. the C4 grasses , maize , sorghum ) Streptogyneae Ehrharteae Phyllorachideae Wild rices inc. Zizania other rice species and subspecies O. sativa (Asian rice) O. glaberrima (African rice) Bambusoideae (bamboos) Pooideae (grasses and cereals inc. wheat , barley ) Oryza sativa rice

944-685: Is stickier, and is used for sweet dishes, and in Italy for risotto ; and sticky short-grain rice is used in Japanese sushi as it keeps its shape when cooked. White rice when cooked contains 29% carbohydrate and 2% protein, with some manganese . Golden rice is a variety produced by genetic engineering to contain vitamin A . Production of rice is estimated to have caused over 1% of global greenhouse gas emissions in 2022. Predictions of how rice yields will be affected by climate change vary across geographies and socioeconomic contexts. In human culture, rice plays

1003-438: Is subjected to a steaming process before it is milled. This makes the grain harder, and moves some of the grain's vitamins and minerals into the white part of the rice so these are retained after milling. Rice does not contain gluten , so is suitable for people on a gluten-free diet . Rice is a good source of protein and a staple food in many parts of the world, but it is not a complete protein as it does not contain all of

1062-433: Is the chalaza where the nucellus is joined to the integuments. Nutrients from the plant travel through the phloem of the vascular system to the funiculus and outer integument and from there apoplastically and symplastically through the chalaza to the nucellus inside the ovule. In chalazogamous plants, the pollen tubes enter the ovule through the chalaza instead of the micropyle opening. The nucellus (plural: nucelli)

1121-406: Is to flood the field to a depth of 5 cm (2 in), then to let the water level drop to 15 cm (6 in) below surface level, as measured by looking into a perforated field water tube sunk into the soil, and then repeating the cycle. Deepwater rice varieties tolerate flooding to a depth of over 50 centimetres for at least a month. Upland rice is grown without flooding, in hilly or mountainous regions; it

1180-471: Is used as a model organism in biology. Dry rice grain is milled to remove the outer layers; depending on how much is removed, products range from brown rice to rice with germ and white rice. Some is parboiled to make it easy to cook. Rice contains no gluten ; it provides protein but not all the essential amino acids needed for good health. Rice of different types is eaten around the world. Long-grain rice tends to stay intact on cooking; medium-grain rice

1239-436: The embryo sac ) is much smaller and typically consists of only seven cells and eight nuclei. This type of megagametophyte develops from the megaspore through three rounds of mitotic divisions. The cell closest to the micropyle opening of the integuments differentiates into the egg cell, with two synergid cells by its side that are involved in the production of signals that guide the pollen tube. Three antipodal cells form on

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1298-440: The essential amino acids in sufficient amounts for good health. World trade figures are much smaller than those for production, as less than 8% of rice produced is traded internationally. China, an exporter of rice in the early 2000s, had become the world's largest importer of rice by 2013. Developing countries are the main players in the world rice trade; by 2012, India was the largest exporter of rice, with Thailand and Vietnam

1357-478: The 21st century as people in Asia and elsewhere ate less grain and more meat. An exception is Sub-Saharan Africa, where both per capita consumption of rice and population are increasing. Rice is a commonly-eaten food around the world. The varieties of rice are typically classified as short-, medium-, and long-grained. Oryza sativa indica varieties are usually long-grained; Oryza sativa japonica varieties are usually short- or medium-grained. Short-grain rice, with

1416-688: The Americas by the Spanish. In British North America by the time of the start of the American War of Independence , rice had become the fourth most valuable export commodity behind only tobacco, wheat, and fish. In 2021, world production of rice was 787 million tonnes , led by China and India with a combined 52% of the total. This placed rice fourth in the list of crops by production, after sugarcane , maize , and wheat . Other major producers were Bangladesh , Indonesia and Vietnam . 90% of world production

1475-460: The Philippines have traditionally managed weeds and pests by the polycultural practice of raising ducks and sometimes fish in their rice paddies. These produce valuable additional crops, eat small pest animals, manure the rice, and in the case of ducks also control weeds. Rice plants produce their own chemical defences to protect themselves from pest attacks. Some synthetic chemicals, such as

1534-465: The benefits these could bring. In 2022, greenhouse gas emissions from rice cultivation were estimated at 5.7 billion tonnes CO2eq, representing 1.2% of total emissions. Within the agriculture sector, rice produces almost half the greenhouse gas emissions from croplands , some 30% of agricultural methane emissions , and 11% of agricultural nitrous oxide emissions. Methane is released from rice fields subject to long-term flooding, as this inhibits

1593-506: The crop would be lost under these conditions. In the Po Valley in Italy, the arborio and carnaroli risotto rice varieties have suffered poor harvests through drought in the 21st century. The Ente Nazionale Risi  [ it ] is developing drought-resistant varieties; its nuovo prometeo variety has deep roots that enable it to tolerate drought, but is not suitable for risotto. Rice yield can be reduced by weed growth, and

1652-425: The developing megasporophyte, may be described as either tenuinucellate or crassinucellate. The former has either no cells or a single cell layer between the megasporophyte and the epidermal cells, while the latter has multiple cell layers between. Embryos may be described by a number of terms including Linear (embryos have axile placentation and are longer than broad), or rudimentary (embryos are basal in which

1711-414: The embryo within the seed through a mechanism of asexual reproduction called nucellar embryony . The haploid megaspore inside the nucellus gives rise to the female gametophyte , called the megagametophyte . In gymnosperms, the megagametophyte consists of around 2000 nuclei and forms archegonia , which produce egg cells for fertilization. In flowering plants, the megagametophyte (also referred to as

1770-425: The exception of Spanish Bomba, is usually sticky when cooked, and is suitable for puddings. Thai Jasmine rice is aromatic, and unusually for a long-grain rice has some stickiness, with a soft texture. Indian Basmati rice is very long-grained and aromatic. Italian Arborio rice , used for risotto , is of medium length, oval, and quite sticky. Japanese sushi rice is a sticky short-grain variety. Cooked white rice

1829-441: The expression of genes that facilitate DNA repair and homologous recombination . In gymnosperms, three of the four haploid spores produced in meiosis typically degenerate, leaving one surviving megaspore inside the nucellus. Among angiosperms, however, a wide range of variation exists in what happens next. The number (and position) of surviving megaspores, the total number of cell divisions, whether nuclear fusions occur, and

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1888-425: The field where they will grow, or seedlings can be grown in a seedbed and transplanted into the field. Direct seeding needs some 60 to 80 kg of grain per hectare, while transplanting needs less, around 40 kg per hectare, but requires far more labour. Most rice in Asia is transplanted by hand. Mechanical transplanting takes less time but requires a carefully-prepared field and seedlings raised on mats or in trays to fit

1947-472: The figure) there is no separation of the megaspores following meiosis, then the nuclei fuse to form a triploid nucleus and a haploid nucleus. The subsequent arrangement of cells is similar to the Polygonum pattern, but the ploidy of the nuclei is different. After fertilization, the nucellus may develop into the perisperm that feeds the embryo. In some plants, the diploid tissue of the nucellus can give rise to

2006-546: The final number, position and ploidy of the cells or nuclei all vary. A common pattern of embryo sac development (the Polygonum type maturation pattern) includes a single functional megaspore followed by three rounds of mitosis. In some cases, however, two megaspores survive (for example, in Allium and Endymion ). In some cases all four megaspores survive, for example in the Fritillaria type of development (illustrated by Lilium in

2065-503: The grain, and cleaning by winnowing or screening . The rice grain is dried as soon as possible to bring the moisture content down to a level that is safe from mould fungi. Traditional drying relies on the heat of the sun, with the grain spread out on mats or on pavements. The edible rice species are members of the BOP clade within the grass family, the Poaceae . The rice subfamily, Oryzoideae ,

2124-414: The great diversity and continuous evolution of pests. Resistance genes are being sought from wild species of rice, and genetic engineering techniques are being applied. Ovule In flowering plants , the ovule is located inside the portion of the flower called the gynoecium . The ovary of the gynoecium produces one or more ovules and ultimately becomes the fruit wall. Ovules are attached to

2183-575: The herbicide 2,4-D , cause the plant to increase the production of certain defensive chemicals and thereby increase the plant's resistance to some types of pests. Conversely, other chemicals, such as the insecticide imidacloprid , appear to induce changes in the gene expression of the rice that make the plant more susceptible to certain pests. Plant breeders have created rice cultivars incorporating resistance to various insect pests . Conventional plant breeding of resistant varieties has been limited by challenges such as rearing insect pests for testing, and

2242-400: The inner integument (which is integral to the formation of ovules from megasporangia) has been proposed to be by enclosure of a megasporangium by sterile branches (telomes). Elkinsia , a preovulate taxon, has a lobed structure fused to the lower third of the megasporangium, with the lobes extending upwards in a ring around the megasporangium. This might, through fusion between lobes and between

2301-459: The kind produced by the Caytoniales or Glossopteridales may have evolved into the outer integument of angiosperms. The integuments develop into the seed coat when the ovule matures after fertilization. The integuments do not enclose the nucellus completely but retain an opening at the apex referred to as the micropyle . The micropyle opening allows the pollen (a male gametophyte ) to enter

2360-410: The later years of the 20th century decreased rice yield by between 10% and 20% across 200 farms in seven Asian countries. This may have been caused by increased night-time respiration. IRRI has predicted that Asian rice yields will fall by some 20% per 1°C rise in global mean temperature. Further, rice is unable to yield grain if the flowers experience a temperature of 35 °C or more for over one hour, so

2419-417: The machine. Rice does not thrive if continuously submerged. Rice can be grown in different environments, depending upon water availability. The usual arrangement is for lowland fields to be surrounded by bunds and flooded to a depth of a few centimetres until around a week before harvest time; this requires a large amount of water. The "alternate wetting and drying" technique uses less water. One form of this

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2478-418: The megagametophyte to produce a second embryo. The plant stores nutrients such as starch , proteins , and oils in the endosperm as a food source for the developing embryo and seedling, serving a similar function to the yolk of animal eggs. The endosperm is also called the albumen of the seed. the zygote then develops into a megasporophyte, which in turn produces one or more megasporangia. The ovule, with

2537-462: The next sporophyte generation. In flowering plants, a second sperm nucleus fuses with other nuclei in the megagametophyte forming a typically polyploid (often triploid) endosperm tissue, which serves as nourishment for the young sporophyte. An integument is a protective layer of cells surrounding the ovule. Gymnosperms typically have one integument (unitegmic) while angiosperms typically have two integuments (bitegmic). The evolutionary origin of

2596-411: The opposite (chalazal) end of the ovule and later degenerate. The large central cell of the embryo sac contains two polar nuclei . The pollen tube releases two sperm nuclei into the ovule. In gymnosperms, fertilization occurs within the archegonia produced by the female gametophyte. While it is possible that several egg cells are present and fertilized, typically only one zygote will develop into

2655-428: The other largest exporters. As of 2016, the countries that consumed the most rice were China (29% of total), India, and Indonesia. By 2020, Bangladesh had taken third place from Indonesia. On an annual average from 2020-23, China consumed 154 million tonnes of rice, India consumed 109 million tonnes, and Bangladesh and Indonesia consumed about 36 million tonnes each. Across the world, rice consumption per capita fell in

2714-440: The ovule and divide by mitosis to produce the haploid female gametophyte or megagametophyte, which also remains inside the ovule. The remnants of the megasporangium tissue (the nucellus) surround the megagametophyte. Megagametophytes produce archegonia (lost in some groups such as flowering plants), which produce egg cells. After fertilization, the ovule contains a diploid zygote and then, after cell division begins, an embryo of

2773-403: The ovule for fertilization. In gymnosperms (e.g., conifers), the pollen is drawn into the ovule on a drop of fluid that exudes out of the micropyle, the so-called pollination drop mechanism. Subsequently, the micropyle closes. In angiosperms, only a pollen tube enters the micropyle. During germination , the seedling 's radicle emerges through the micropyle. Located opposite from the micropyle

2832-438: The placenta in the ovary through a stalk-like structure known as a funiculus (plural, funiculi). Different patterns of ovule attachment, or placentation , can be found among plant species, these include: In gymnosperms such as conifers, ovules are borne on the surface of an ovuliferous (ovule-bearing) scale, usually within an ovulate cone (also called megastrobilus ). In the early extinct seed ferns , ovules were borne on

2891-812: The rice produced in developing nations is lost after harvest through factors such as poor transport and storage. Rice yields can be reduced by pests including insects , rodents , and birds , as well as by weeds , and by diseases such as rice blast . Traditional rice polycultures such as rice-duck farming , and modern integrated pest management seek to control damage from pests in a sustainable way. Many varieties of rice have been bred to improve crop quality and productivity. Biotechnology has created Green Revolution rice able to produce high yields when supplied with nitrogen fertiliser and managed intensively. Other products are rice able to express human proteins for medicinal use; flood-tolerant or deepwater rice ; and drought-tolerant and salt-tolerant varieties. Rice

2950-1011: The soil from absorbing atmospheric oxygen, resulting in anaerobic fermentation of organic matter in the soil. Emissions can be limited by planting new varieties, not flooding continuously, and removing straw. It is possible to cut methane emissions in rice cultivation by improved water management, combining dry seeding and one drawdown, or executing a sequence of wetting and drying . This results in emission reductions of up to 90% compared to full flooding and even increased yields. Predictions of climate change's effects on rice cultivation vary. Global rice yield has been projected to decrease by around 3.2% with each 1°C increase in global average temperature while another study predicts global rice cultivation will increase initially, plateauing at about 3°C warming (2091–2100 relative to 1850–1900). The impacts of climate change on rice cultivation vary across geographic location and socioeconomic context. For example, rising temperatures and decreasing solar radiation during

3009-441: The structure and the megasporangium, have produced an integument. The origin of the second or outer integument has been an area of active contention for some time. The cupules of some extinct taxa have been suggested as the origin of the outer integument. A few angiosperms produce vascular tissue in the outer integument, the orientation of which suggests that the outer surface is morphologically abaxial. This suggests that cupules of

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3068-424: The surface of leaves. In the most recent of these taxa, a cupule (a modified branch or group of branches) surrounded the ovule (e.g. Caytonia or Glossopteris ). Ovule orientation may be anatropous , such that when inverted the micropyle faces the placenta (this is the most common ovule orientation in flowering plants), amphitropous , campylotropous , or orthotropous ( anatropous are common and micropyle

3127-677: The voyage to the rest of the Pacific. It reached Austroasiatic and Kra-Dai -speakers in Mainland Southeast Asia and southern China by 5000 years ago. Rice spread around the rest of the world through cultivation, migration and trade, eventually to the Americas as part of the Columbian exchange after 1492. The now less common Oryza glaberrima (African rice) was independently domesticated in Africa around 3,000 years ago, and introduced to

3186-785: The wet season, while thrips outbreaks are associated with drought. Rice blast , caused by the fungus Magnaporthe grisea , is the most serious disease of growing rice. It and bacterial leaf streak (caused by Xanthomonas oryzae pv. oryzae ) are perennially the two worst rice diseases worldwide; they are both among the ten most important diseases of all crop plants. Other major rice diseases include sheath blight (caused by Rhizoctonia solani ), false smut ( Ustilaginoidea virens ), and bacterial panicle blight ( Burkholderia glumae ). Viral diseases include rice bunchy stunt, rice dwarf, rice tungro , and rice yellow mottle. Crop protection scientists are developing sustainable techniques for managing rice pests. Sustainable pest management

3245-539: The year provided that sufficient water is made available. It is normally an annual, but in the tropics it can survive as a perennial , producing a ratoon crop. Like all crops, rice depends for its growth on both biotic and abiotic environmental factors. The principal biotic factors are crop variety, pests , and plant diseases . Abiotic factors include the soil type, whether lowland or upland, amount of rain or irrigation water, temperature, day length , and intensity of sunlight. Rice grains can be planted directly into

3304-460: Was domesticated in China some 13,500 to 8,200 years ago; African rice was domesticated in Africa about 3,000 years ago. Rice has become commonplace in many cultures worldwide; in 2021, 787 million tons were produced, placing it fourth after sugarcane , maize , and wheat . Only some 8% of rice is traded internationally. China, India, and Indonesia are the largest consumers of rice. A substantial amount of

3363-465: Was first domesticated in China 9,000 years ago, by people of Neolithic cultures in the Upper and Lower Yangtze , associated with Hmong-Mien -speakers and pre-Austronesians , respectively. The functional allele for nonshattering , the critical indicator of domestication in grains, as well as five other single-nucleotide polymorphisms , is identical in both indica and japonica . This implies

3422-559: Was introduced early into Sino-Tibetan cultures in northern China by around 6000 to 5600 years ago, and to the Korean peninsula and Japan by around 5500 to 3200 years ago. It was also carried into Taiwan by the Dapenkeng culture by 5500 to 4000 years ago, before spreading southwards via the Austronesian migrations to Island Southeast Asia , Madagascar , and Guam , but did not survive

3481-439: Was stored by rural households. The dry grain is milled to remove the outer layers, namely the husk and bran . These can be removed in a single step, in two steps, or as in commercial milling in a multi-step process of cleaning, dehusking, separation, polishing, grading, and weighing. Brown rice only has the inedible husk removed. Further milling removes bran and the germ to create successively whiter products. Parboiled rice

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