210-464: (Williston, 1897) Prognathodon is an extinct genus of marine lizard belonging to the mosasaur family. It is classified as part of the Mosasaurinae subfamily, alongside genera like Mosasaurus and Clidastes . Prognathodon has been recovered from deposits ranging in age from the Campanian to the Maastrichtian in the Middle East , Europe , New Zealand , and North America . Prognathodon means "forejaw tooth", which originates from
420-522: A 2001 election ; he carried out his plan to unilaterally withdraw from the Gaza Strip and spearheaded the construction of the West Bank barrier , ending the intifada. Between 2000 and 2008, 1,063 Israelis, 5,517 Palestinians and 64 foreign citizens were killed. In 2006, a Hezbollah artillery assault on Israel's northern border communities and a cross-border abduction of two Israeli soldiers precipitated
630-438: A food chain who lose their prey. "Species coextinction is a manifestation of one of the interconnectednesses of organisms in complex ecosystems ... While coextinction may not be the most important cause of species extinctions, it is certainly an insidious one." Coextinction is especially common when a keystone species goes extinct. Models suggest that coextinction is the most common form of biodiversity loss . There may be
840-657: A nautilus to the Royal Society that was more than two feet in diameter, and morphologically distinct from any known living species. Hooke theorized that this was simply because the species lived in the deep ocean and no one had discovered them yet. While he contended that it was possible a species could be "lost", he thought this highly unlikely. Similarly, in 1695, Sir Thomas Molyneux published an account of enormous antlers found in Ireland that did not belong to any extant taxa in that area. Molyneux reasoned that they came from
1050-415: A species or a population is the variety of genetic information in its living members. A large gene pool (extensive genetic diversity ) is associated with robust populations that can survive bouts of intense selection . Meanwhile, low genetic diversity (see inbreeding and population bottlenecks ) reduces the range of adaptions possible. Replacing native with alien genes narrows genetic diversity within
1260-436: A viable population for species preservation and possible future reintroduction to the wild, through use of carefully planned breeding programs . The extinction of one species' wild population can have knock-on effects, causing further extinctions. These are also called "chains of extinction". This is especially common with extinction of keystone species . A 2018 study indicated that the sixth mass extinction started in
1470-620: A Jewish " national home " in Palestine. Weizmann's interpretation of the declaration was that negotiations on the future of the country were to happen directly between Britain and the Jews, excluding Arabs. Jewish-Arab relations in Palestine deteriorated dramatically in the following years. In 1918 the Jewish Legion , primarily Zionist volunteers, assisted in the British conquest of Palestine . In 1920
1680-474: A Lazarus species when extant individuals were described in 2019. Attenborough's long-beaked echidna ( Zaglossus attenboroughi ) is an example of a Lazarus species from Papua New Guinea that had last been sighted in 1962 and believed to be possibly extinct, until it was recorded again in November 2023. Some species currently thought to be extinct have had continued speculation that they may still exist, and in
1890-850: A Palestinian uprising against Israeli rule, broke out in 1987, with waves of uncoordinated demonstrations and violence in the occupied West Bank and Gaza. Over the following six years, the intifada became more organized and included economic and cultural measures aimed at disrupting the Israeli occupation. Over 1,000 people were killed. During the 1991 Gulf War , the PLO supported Saddam Hussein and Iraqi missile attacks against Israel . Despite public outrage, Israel heeded American calls to refrain from hitting back. In 1992, Yitzhak Rabin became prime minister following an election in which his party called for compromise with Israel's neighbours. The following year, Shimon Peres on behalf of Israel and Mahmoud Abbas for
2100-495: A cascade of coextinction across the trophic levels . Such effects are most severe in mutualistic and parasitic relationships. An example of coextinction is the Haast's eagle and the moa : the Haast's eagle was a predator that became extinct because its food source became extinct. The moa were several species of flightless birds that were a food source for the Haast's eagle. Extinction as
2310-429: A common ancestor with modern horses. Pseudoextinction is much easier to demonstrate for larger taxonomic groups. A Lazarus taxon or Lazarus species refers to instances where a species or taxon was thought to be extinct, but was later rediscovered. It can also refer to instances where large gaps in the fossil record of a taxon result in fossils reappearing much later, although the taxon may have ultimately become extinct at
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#17330934449912520-730: A comprehensive phylogenetic analysis of the Mosasauridae by Simões et al. (2017) is shown below, collapsed to only display the Mosasaurinae . The cladogram shows Prognathodon as paraphyletic, as concluded by most recent researchers, and places it as a sister group to the derived mosasaurines in the Mosasaurini . Dallasaurus turneri Clidastes liodontus Clidastes moorevillensis Clidastes propython Globidens dakotensis Globidens alabamaensis Prognathodon rapax Prognathodon overtoni Extinct Extinction
2730-616: A de facto independent emirate in the Galilee. Ottoman attempts to subdue the sheikh failed. After Zahir's death the Ottomans regained control of the area. In 1799, governor Jazzar Pasha repelled an assault on Acre by Napoleon 's troops, prompting the French to abandon the Syrian campaign. In 1834, a revolt by Palestinian Arab peasants against Egyptian conscription and taxation policies under Muhammad Ali
2940-575: A distinct lizard suborder and dividing the group into two families, the Mosasauridae and the "Plioplatecarpidae". In this early taxonomy, the Mosasauridae contained the genera Clidastes , Mosasaurus , Platecarpus , Halisaurus and Tylosaurus and the Plioplatecarpidae was monotypic, only containing Plioplatecarpus . In 1890, following further mosasaur discoveries (including that of Prognathodon ), Dollo revised his taxonomy, dividing
3150-399: A fact that was accepted by most scientists. The primary debate focused on whether this turnover caused by extinction was gradual or abrupt in nature. Cuvier understood extinction to be the result of cataclysmic events that wipe out huge numbers of species, as opposed to the gradual decline of a species over time. His catastrophic view of the nature of extinction garnered him many opponents in
3360-708: A higher risk of extinction and die out faster than less sexually dimorphic species, the least sexually dimorphic species surviving for millions of years while the most sexually dimorphic species die out within mere thousands of years. Earlier studies based on counting the number of currently living species in modern taxa have shown a higher number of species in more sexually dimorphic taxa which have been interpreted as higher survival in taxa with more sexual selection, but such studies of modern species only measure indirect effects of extinction and are subject to error sources such as dying and doomed taxa speciating more due to splitting of habitat ranges into more small isolated groups during
3570-405: A large range, a lack of individuals of both sexes (in sexually reproducing species), or other reasons. Pinpointing the extinction (or pseudoextinction ) of a species requires a clear definition of that species . If it is to be declared extinct, the species in question must be uniquely distinguishable from any ancestor or daughter species, and from any other closely related species. Extinction of
3780-592: A later point. The coelacanth , a fish related to lungfish and tetrapods , is an example of a Lazarus taxon that was known only from the fossil record and was considered to have been extinct since the end of the Cretaceous Period . In 1938, however, a living specimen was found off the Chalumna River (now Tyolomnqa) on the east coast of South Africa. Calliostoma bullatum , a species of deepwater sea snail originally described from fossils in 1844 proved to be
3990-818: A massacre of Israeli athletes at the 1972 Summer Olympics in Munich. The Israeli government responded with an assassination campaign against the organizers of the massacre, a bombing and a raid on the PLO headquarters in Lebanon . On 6 October 1973, the Egyptian and Syrian armies launched a surprise attack against Israeli forces in the Sinai Peninsula and Golan Heights, opening the Yom Kippur War . The war ended on 25 October with Israel repelling Egyptian and Syrian forces but suffering great losses. An internal inquiry exonerated
4200-587: A mathematical model that falls in all positions. By contrast, conservation biology uses the extinction vortex model to classify extinctions by cause. When concerns about human extinction have been raised, for example in Sir Martin Rees ' 2003 book Our Final Hour , those concerns lie with the effects of climate change or technological disaster. Human-driven extinction started as humans migrated out of Africa more than 60,000 years ago. Currently, environmental groups and some governments are concerned with
4410-537: A movement that sought to establish a Jewish state in Palestine, thus offering a solution to the Jewish question of the European states. Antisemitism, pogroms and official policies, in tsarist Russia led to the emigration of three million Jews in the years between 1882 and 1914, only 1% of which went to Palestine. Those who went to Palestine were driven primarily by ideas of self-determination and Jewish identity, rather than as
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#17330934449914620-456: A natural part of the evolutionary process. Only recently have extinctions been recorded and scientists have become alarmed at the current high rate of extinctions . Most species that become extinct are never scientifically documented. Some scientists estimate that up to half of presently existing plant and animal species may become extinct by 2100. A 2018 report indicated that the phylogenetic diversity of 300 mammalian species erased during
4830-441: A new mega-predator or by transporting animals and plants from one part of the world to another. Such introductions have been occurring for thousands of years, sometimes intentionally (e.g. livestock released by sailors on islands as a future source of food) and sometimes accidentally (e.g. rats escaping from boats). In most cases, the introductions are unsuccessful, but when an invasive alien species does become established,
5040-568: A part of Coele-Syria . Over the ensuing centuries, the Hellenization of the region led to cultural tensions that came to a head during the reign of Antiochus IV , giving rise to the Maccabean Revolt of 167 BCE. The civil unrest weakened Seleucid rule, and in the late 2nd century the semi-autonomous Hasmonean Kingdom of Judea arose, eventually attaining full independence and expanding into neighboring regions. The Roman Republic invaded
5250-610: A population a higher chance in the short term of surviving an adverse change in conditions. Effects that cause or reward a loss in genetic diversity can increase the chances of extinction of a species. Population bottlenecks can dramatically reduce genetic diversity by severely limiting the number of reproducing individuals and make inbreeding more frequent. Extinction sometimes results for species evolved to specific ecologies that are subjected to genetic pollution —i.e., uncontrolled hybridization , introgression and genetic swamping that lead to homogenization or out-competition from
5460-425: A posterior bite force of 7,630–10,974 N (778–1,119 kg f ; 1,715–2,467 lb f ). The quadrates of Prognathodon , similarly to the genus Globidens , have fused suprastapedial and infrastapedial processes, which is possibly an adaptation to counteract the strong forces experienced by the bone during biting. Strong jaw musculature combined with a relatively short and tall dentary would have resulted in
5670-637: A proclamation allowing the exiled Judean population to return to Judah. The construction of the Second Temple was completed c. 520 BCE . The Achaemenids ruled the region as the province of Yehud Medinata . In 332 BCE, Alexander the Great conquered the region as part of his campaign against the Achaemenid Empire . After his death, the area was controlled by the Ptolemaic and Seleucid empires as
5880-407: A race of animals to become extinct. A series of fossils were discovered in the late 17th century that appeared unlike any living species. As a result, the scientific community embarked on a voyage of creative rationalization, seeking to understand what had happened to these species within a framework that did not account for total extinction. In October 1686, Robert Hooke presented an impression of
6090-502: A reduction in agricultural productivity. Furthermore, increased erosion contributes to poorer water quality by elevating the levels of sediment and pollutants in rivers and streams. Habitat degradation through toxicity can kill off a species very rapidly, by killing all living members through contamination or sterilizing them. It can also occur over longer periods at lower toxicity levels by affecting life span, reproductive capacity, or competitiveness. Habitat degradation can also take
6300-652: A regional power, with a capital at Samaria ; during the Omride dynasty , it controlled Samaria , Galilee , the upper Jordan Valley , the plain of Sharon and large parts of Transjordan . The Kingdom of Israel was conquered around 720 BCE by the Neo-Assyrian Empire . The Kingdom of Judah, under Davidic rule with its capital in Jerusalem , later became a client state of first the Neo-Assyrian Empire and then
6510-517: A response to pogroms or economic insecurity. The Second Aliyah (1904–1914) began after the Kishinev pogrom ; some 40,000 Jews settled in Palestine, although nearly half left eventually. Both the first and second waves of migrants were mainly Orthodox Jews . The Second Aliyah included Zionist socialist groups who established the kibbutz movement based on the idea of establishing a separate Jewish economy based exclusively on Jewish labor. Those of
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6720-486: A result of climate change has been confirmed by fossil studies. Particularly, the extinction of amphibians during the Carboniferous Rainforest Collapse , 305 million years ago. A 2003 review across 14 biodiversity research centers predicted that, because of climate change, 15–37% of land species would be "committed to extinction" by 2050. The ecologically rich areas that would potentially suffer
6930-662: A secret alliance with the UK and France and overran the Sinai Peninsula in the Suez Crisis but was pressured to withdraw by the UN in return for guarantees of Israeli shipping rights. The war resulted in significant reduction of Israeli border infiltration. In the early 1960s, Israel captured Nazi war criminal Adolf Eichmann in Argentina and brought him to Israel for trial . Eichmann remains
7140-535: A series of coordinated attacks on Israel, leading to the start of the Israel–Hamas war . On that day, approximately 1,300 Israelis, predominantly civilians, were killed in communities near the Gaza Strip border and during a music festival . Over 200 hostages were kidnapped and taken to the Gaza Strip. After clearing militants from its territory, Israel launched one of the most destructive bombing campaigns in modern history and invaded Gaza on 27 October with
7350-479: A species (or replacement by a daughter species) plays a key role in the punctuated equilibrium hypothesis of Stephen Jay Gould and Niles Eldredge . In ecology , extinction is sometimes used informally to refer to local extinction , in which a species ceases to exist in the chosen area of study, despite still existing elsewhere. Local extinctions may be made good by the reintroduction of individuals of that species taken from other locations; wolf reintroduction
7560-439: A species may come suddenly when an otherwise healthy species is wiped out completely, as when toxic pollution renders its entire habitat unliveable; or may occur gradually over thousands or millions of years, such as when a species gradually loses out in competition for food to better adapted competitors. Extinction may occur a long time after the events that set it in motion, a phenomenon known as extinction debt . Assessing
7770-404: A species or group of species. "Just as each species is unique", write Beverly and Stephen C. Stearns , "so is each extinction ... the causes for each are varied—some subtle and complex, others obvious and simple". Most simply, any species that cannot survive and reproduce in its environment and cannot move to a new environment where it can do so, dies out and becomes extinct. Extinction of
7980-518: A subsequent report, IPBES listed unsustainable fishing, hunting and logging as being some of the primary drivers of the global extinction crisis. In June 2019, one million species of plants and animals were at risk of extinction. At least 571 plant species have been lost since 1750, but likely many more. The main cause of the extinctions is the destruction of natural habitats by human activities, such as cutting down forests and converting land into fields for farming. A dagger symbol (†) placed next to
8190-405: A very powerful bite. The skull of the type specimen of Prognathodon saturator is nearly complete, only lacking the anterior portion of the premaxilla and the dentaries. Though most of the anterior marginal teeth are missing, the inclination of the preserved roots suggest that P. saturator had procumbent teeth, a trait also seen in P. solvayi . The dorsal margin of the dentary is concave, whilst
8400-630: A year of fighting, a ceasefire was declared and temporary borders, known as the Green Line , were established. Jordan annexed what became known as the West Bank , including East Jerusalem , and Egypt occupied the Gaza Strip . Over 700,000 Palestinians fled or were expelled by Zionist militias and the Israeli military —what would become known in Arabic as the nakba ('catastrophe'). The events also led to
8610-517: Is also evidence to suggest that this event was preceded by another mass extinction, known as Olson's Extinction . The Cretaceous–Paleogene extinction event (K–Pg) occurred 66 million years ago, at the end of the Cretaceous period; it is best known for having wiped out non-avian dinosaurs , among many other species. According to a 1998 survey of 400 biologists conducted by New York 's American Museum of Natural History , nearly 70% believed that
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8820-444: Is an example of this. Species that are not globally extinct are termed extant . Those species that are extant, yet are threatened with extinction, are referred to as threatened or endangered species . Currently, an important aspect of extinction is human attempts to preserve critically endangered species. These are reflected by the creation of the conservation status "extinct in the wild" (EW) . Species listed under this status by
9030-402: Is controversial. Some species have been assigned to other genera, such as Dollosaurus and Brachysaurana , but this has also been questioned. Prognathodon is known for its massively built jaws and teeth. Its distinct feeding adaptations have generated much interest in its ecology ever since its discovery, though direct evidence of its diet, such as gastric residues, is rare. Prognathodon
9240-501: Is debate about the earliest existence of the Kingdoms of Israel and Judah and their extent and power. While it is unclear if there was a United Kingdom of Israel , historians and archaeologists agree that the northern Kingdom of Israel existed by ca. 900 BCE and the Kingdom of Judah by ca. 850 BCE. The Kingdom of Israel was the more prosperous of the two and soon developed into
9450-585: Is difficult to demonstrate unless one has a strong chain of evidence linking a living species to members of a pre-existing species. For example, it is sometimes claimed that the extinct Hyracotherium , which was an early horse that shares a common ancestor with the modern horse , is pseudoextinct, rather than extinct, because there are several extant species of Equus , including zebra and donkey ; however, as fossil species typically leave no genetic material behind, one cannot say whether Hyracotherium evolved into more modern horse species or merely evolved from
9660-417: Is estimated as 100 to 1,000 times "background" rates (the average extinction rates in the evolutionary time scale of planet Earth), faster than at any other time in human history, while future rates are likely 10,000 times higher. However, some groups are going extinct much faster. Biologists Paul R. Ehrlich and Stuart Pimm , among others, contend that human population growth and overconsumption are
9870-485: Is nicknamed Carlo, after the ENCI worker who discovered it. Prognathodon constitutes one of the largest-bodied mosasaur genera, with the largest known skull (belonging to P. currii ) exceeding 1.4 metres (4.6 ft) in length. Despite its massive size, remains of the genus are often fragmentary and incomplete. To date, very few specimens are known with articulated skulls and none with an entire skeleton. In 1967, P. giganteus
10080-661: Is noted as having a very robust skull. The skull also shows adaptations towards a very powerful jaw musculature. The ratio between the length of the supratemporal fenestra and the total length of the skull has previously been used as an improvised measurement for mosasaur bite force, and is relatively high in Prognathodon (0.22 in P. overtoni and P. saturator ) compared to other genera, such as Mosasaurus (0.19 in M. hoffmannii ). In 1998, Kase and colleagues estimated that P. overtoni had an anterior bite force of 6,346–9,127 N (647–931 kg f ; 1,427–2,052 lb f ) and
10290-592: Is the destruction of ocean floors by bottom trawling . Diminished resources or introduction of new competitor species also often accompany habitat degradation. Global warming has allowed some species to expand their range, bringing competition to other species that previously occupied that area. Sometimes these new competitors are predators and directly affect prey species, while at other times they may merely outcompete vulnerable species for limited resources. Vital resources including water and food can also be limited during habitat degradation, leading to extinction. In
10500-537: Is the form of the sclerotic rings , present in all species of the genus. The function of the scleral ossicles is to maintain the shape of the cornea and support the sclera in the region of Brucke's muscle responsible for affecting accommodation in the lacertilian eye. In the type species, P. solvayi , the sclerotic ring is only partially preserved, consisting of five scleral ossicles in each ring. Similar sclerotic rings are seen in several other mosasaur genera, such as Mosasaurus . The latest published diagnosis for
10710-525: Is the most important determinant of genus extinction at background rates but becomes increasingly irrelevant as mass extinction arises. Limited geographic range is a cause both of small population size and of greater vulnerability to local environmental catastrophes. Extinction rates can be affected not just by population size, but by any factor that affects evolvability , including balancing selection , cryptic genetic variation , phenotypic plasticity , and robustness . A diverse or deep gene pool gives
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#173309344499110920-457: Is the termination of a taxon by the death of its last member . A taxon may become functionally extinct before the death of its last member if it loses the capacity to reproduce and recover. Because a species' potential range may be very large, determining this moment is difficult, and is usually done retrospectively. This difficulty leads to phenomena such as Lazarus taxa , where a species presumed extinct abruptly "reappears" (typically in
11130-509: The jizya tax. Non-Muslim Ottoman subjects faced geographic and lifestyle restrictions, though these were not always enforced. The millet system organized non-Muslims into autonomous communities on the basis of religion. The concept of the "return" remained a symbol within religious Jewish belief which emphasized that their return should be determined by Divine Providence rather than human action. Leading Zionist historian Shlomo Avineri describes this connection: "Jews did not relate to
11340-544: The Four Holy Cities —Jerusalem, Tiberias, Hebron , and Safed —and in 1697, Rabbi Yehuda Hachasid led a group of 1,500 Jews to Jerusalem. A 1660 Druze revolt against the Ottomans destroyed Safed and Tiberias . In the second half of the 18th century, Eastern European Jews who were opponents of Hasidism , known as the Perushim , settled in Palestine. In the late 18th century, local Arab Sheikh Zahir al-Umar created
11550-879: The Gaza Strip in the ongoing Israel–Hamas war . Observers, including the UN Special Committee to investigate Israeli practices and United Nations Special Rapporteur Francesca Albanese , have cited statements by senior Israeli officials that may indicate an " intent to destroy " (in whole or in part) Gaza's population, a necessary condition for the legal threshold of genocide to be met. A majority of 758 mostly US-based Middle East scholars, polled in 2024 by Brookings , believe Israel's actions in Gaza were intended to make it uninhabitable for Palestinians, and 75% of them say Israel's actions in Gaza constitute either genocide or "major war crimes akin to genocide". On 29 December 2023, South Africa instituted proceedings against Israel at
11760-665: The Golan Heights in moves largely unrecognized internationally. After the 1973 Yom Kippur War , Israel signed peace treaties with Egypt —returning the Sinai in 1982—and Jordan . In the 2020s, it normalized relations with more Arab countries. However, efforts to resolve the Israeli–Palestinian conflict after the interim Oslo Accords have not succeeded, and the country has engaged in several wars and clashes with Palestinian militant groups . Israel's practices in its occupation of
11970-599: The Hashemites , in honor of Abdullah II of Jordan . This fossil is about 1.8 m in length, and is remarkable in that it preserves the outline of the mosasaur's tail fins, revealing that Prognathodon had a bilobed tail fluke resembling a downturned shark's tail. While this type of fluke had been suggested for mosasaurs based on skeletal anatomy (as in Platecarpus ), this is the first indisputable evidence that derived mosasaurs had bilobed tail flukes. On September 19, 2012, it
12180-474: The International Union for Conservation of Nature (IUCN) are not known to have any living specimens in the wild and are maintained only in zoos or other artificial environments. Some of these species are functionally extinct, as they are no longer part of their natural habitat and it is unlikely the species will ever be restored to the wild. When possible, modern zoological institutions try to maintain
12390-710: The Iran–Iraq War , the Israeli air force destroyed Iraq's sole nuclear reactor, then under construction, in order to impede the Iraqi nuclear weapons program. Following a series of PLO attacks in 1982, Israel invaded Lebanon to destroy the PLO bases. In the first six days, Israel destroyed the military forces of the PLO in Lebanon and decisively defeated the Syrians. An Israeli government inquiry (the Kahan Commission ) held Begin and several Israeli generals indirectly responsible for
12600-631: The Labor Party . Later that year, Egyptian President Anwar El Sadat made a trip to Israel and spoke before the Knesset in what was the first recognition of Israel by an Arab head of state. Sadat and Begin signed the Camp David Accords (1978) and the Egypt–Israel peace treaty (1979). In return, Israel withdrew from the Sinai Peninsula and agreed to enter negotiations over autonomy for Palestinians in
12810-758: The Late Bronze Age (1550–1200 BCE), large parts of Canaan formed vassal states of the New Kingdom of Egypt . As a result of the Late Bronze Age collapse , Canaan fell into chaos, and Egyptian control over the region collapsed. A people named Israel appear for the first time in the Merneptah Stele, an ancient Egyptian inscription which dates to about 1200 BCE. Ancestors of the Israelites are thought to have included ancient Semitic-speaking peoples native to this area. Modern archaeological accounts suggest that
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#173309344499113020-477: The Late Pleistocene could take up to 5 to 7 million years to restore mammal diversity to what it was before the human era. Extinction of a parent species where daughter species or subspecies are still extant is called pseudoextinction or phyletic extinction. Effectively, the old taxon vanishes, transformed ( anagenesis ) into a successor, or split into more than one ( cladogenesis ). Pseudoextinction
13230-552: The Latin pro - ("earlier" or "prior"), Greek gnathos ("jaw") and odṓn ("tooth"). Twelve nominal species of Prognathodon are recognised, from North America , northern and western Africa , the Middle East , western Europe and New Zealand . Due to the sometimes clear differences between them and the incomplete nature of many of the specimens, the systematics of the genus and which species should properly be considered Prognathodon
13440-713: The League of Nations granted Britain the Mandate for Palestine under terms which included the Balfour Declaration with its promise to the Jews and with similar provisions regarding the Arab Palestinians. The population of the area was predominantly Arab and Muslim, with Jews accounting for about 11% and Arab Christians about 9.5% of the population. The Third (1919–1923) and Fourth Aliyahs (1924–1929) brought an additional 100,000 Jews to Palestine. The rise of Nazism , and
13650-695: The Neo-Babylonian Empire . It is estimated that the region's population was around 400,000 in the Iron Age II . In 587/6 BCE, following a revolt in Judah , King Nebuchadnezzar II besieged and destroyed Jerusalem and Solomon's Temple , dissolved the kingdom and exiled much of the Judean elite to Babylon . After capturing Babylon in 539 BCE, Cyrus the Great , founder of the Achaemenid Empire , issued
13860-725: The Royal Navy and the refugees placed in detention camps in Atlit and Cyprus . On 22 July 1946, Irgun bombed the British administrative headquarters for Palestine, killing 91. The attack was a response to Operation Agatha (a series of raids, including one on the Jewish Agency , by the British) and was the deadliest directed at the British during the Mandate era. The Jewish insurgency continued throughout 1946 and 1947 despite concerted efforts by
14070-615: The Sabra and Shatila massacre and held Defense Minister Ariel Sharon as bearing "personal responsibility". Sharon was forced to resign. In 1985, Israel responded to a Palestinian terrorist attack in Cyprus by bombing the PLO headquarters in Tunisia. Israel withdrew from most of Lebanon in 1986 but maintained a borderland buffer zone in southern Lebanon until 2000, from where Israeli forces engaged in conflict with Hezbollah . The First Intifada ,
14280-456: The Sasanian conquest of Jerusalem and the short-lived Jewish revolt against Heraclius in 614 CE, the Byzantine Empire reconsolidated control of the area in 628. In 634–641 CE, the Rashidun Caliphate conquered the Levant . Over the next six centuries, control of the region transferred between the Umayyad , Abbasid , Fatimid caliphates, and subsequently the Seljuk and Ayyubid dynasties. The population drastically decreased during
14490-557: The Ubeidiya prehistoric site . The Skhul and Qafzeh hominins , dating back 120,000 years, are some of the earliest traces of anatomically modern humans outside of Africa. The Natufian culture , which may have been linked to Proto-Afroasiatic language , emerged by the 10th millennium BCE, followed by the Ghassulian culture by around 4,500 BCE. Early references to "Canaanites" and " Canaan " appear in Near Eastern and Egyptian texts ( c. 2000 BCE); these populations were structured as politically independent city-states . During
14700-413: The destruction of Jerusalem and the Second Temple and a sizable portion of the population being killed or displaced. A second uprising known as the Bar Kokhba revolt (132–136 CE) initially allowed the Jews to form an independent state, but the Romans brutally crushed the rebellion, devastating and depopulating Judea's countryside. Jerusalem was rebuilt as a Roman colony ( Aelia Capitolina ), and
14910-619: The fossil record ) after a period of apparent absence. More than 99% of all species that ever lived on Earth , amounting to over five billion species, are estimated to have died out. It is estimated that there are currently around 8.7 million species of eukaryotes globally, and possibly many times more if microorganisms , such as bacteria , are included. Notable extinct animal species include non-avian dinosaurs , saber-toothed cats , dodos , mammoths , ground sloths , thylacines , trilobites , golden toads , and passenger pigeons . Through evolution , species arise through
15120-561: The government headed by the prime minister and elects the figurehead president . Israel is the only country to have a revived official language, Hebrew . Its culture comprises Jewish and Jewish diaspora elements alongside Arab influences. Israel has one of the largest economies in the Middle East and among the highest GDP per capita and standards of living in Asia . One of the most technologically advanced and developed countries in
15330-474: The occupied West Bank , increasing friction with the Palestinians there. The 1980 Jerusalem Law was believed by some to reaffirm Israel's 1967 annexation of Jerusalem by government decree and reignited international controversy over the status of the city . No Israeli legislation has defined the territory of Israel, and no act specifically included East Jerusalem therein. In 1981 Israel effectively annexed
15540-529: The pterygoid has seven to eight teeth. The medial wing from the coronoid contacts the angular, the anterior process on the coronoid abrupts over the surangular and makes contact with the posterior process of the dentary or ends with the surangular without contacting the dentary. The retroarticular process is rectangular in outline, medially inflected or laterally lacing. The marginal teeth are stout, bicarinate and smooth or striated. Zygosphenes and zygantra are absent, incipient or large and functional. Prognathodon
15750-457: The quadrate and tympanic membrane of Plioplatecarpus houzeaui . Prognathodon giganteus , named by Dollo in 1904, is one of species with the most brief descriptions, apparently only intended to provide a name for the skeleton of the mosasaur for exhibition in the museum hall. The first comprehensive study of the Prognathodon specimens from Belgium (including the type specimen) was done by Theagarten Lingham-Soliar and Dirk Nolf in 1989 and
15960-410: The slender-billed curlew ( Numenius tenuirostris ), not seen since 2007. As long as species have been evolving, species have been going extinct. It is estimated that over 99.9% of all species that ever lived are extinct. The average lifespan of a species is 1–10 million years, although this varies widely between taxa. A variety of causes can contribute directly or indirectly to the extinction of
16170-484: The strata of the Paris basin. They saw alternating saltwater and freshwater deposits, as well as patterns of the appearance and disappearance of fossils throughout the record. From these patterns, Cuvier inferred historic cycles of catastrophic flooding, extinction, and repopulation of the earth with new species. Cuvier's fossil evidence showed that very different life forms existed in the past than those that exist today,
16380-410: The "mesorhynchous" (including Mosasaurus and Clidastes ) groups. Dollo realized that Plioplatecarpus shared characters with the "microrhynchous" group in 1894 and abandoned his previous two family-system, starting to use only one family of mosasaurs, the Mosasauridae, and placing Prognathodon as closely related to Platecarpus and Plioplatecarpus . Russell (1967) retained Prognathodon within
16590-619: The 20 biodiversity goals laid out by the Aichi Biodiversity Targets in 2010, only 6 were "partially achieved" by the deadline of 2020. The report warned that biodiversity will continue to decline if the status quo is not changed, in particular the "currently unsustainable patterns of production and consumption, population growth and technological developments". In a 2021 report published in the journal Frontiers in Conservation Science , some top scientists asserted that even if
16800-545: The 2010s, increasing regional cooperation between Israel and Arab League countries have been established, culminating in the signing of the Abraham Accords . The Israeli security situation shifted from the traditional Arab–Israeli conflict towards the Iran–Israel proxy conflict and direct confrontation with Iran during the Syrian civil war . On 7 October 2023, Palestinian militant groups from Gaza, led by Hamas, launched
17010-451: The Aichi Biodiversity Targets set for 2020 had been achieved, it would not have resulted in a significant mitigation of biodiversity loss. They added that failure of the global community to reach these targets is hardly surprising given that biodiversity loss is "nowhere close to the top of any country's priorities, trailing far behind other concerns such as employment, healthcare, economic growth, or currency stability." For much of history,
17220-461: The British military and Palestine Police Force to suppress it. British efforts to mediate with Jewish and Arab representatives also failed as the Jews were unwilling to accept any solution that did not involve a Jewish state and suggested a partition of Palestine into Jewish and Arab states, while the Arabs were adamant that a Jewish state in any part of Palestine was unacceptable and that the only solution
17430-636: The British would evacuate. As Arab militias and gangs attacked Jewish areas, they were faced mainly by the Haganah as well as the smaller Irgun and Lehi. In April 1948, the Haganah moved onto the offensive. On 14 May 1948, the day before the expiration of the British Mandate, David Ben-Gurion , the head of the Jewish Agency, declared "the establishment of a Jewish state in Eretz-Israel". The following day,
17640-653: The City of Jerusalem [...] the last to be under an International Trusteeship System". Meanwhile, the Jewish insurgency continued and peaked in July 1947, with a series of widespread guerrilla raids culminating in the Sergeants affair , in which the Irgun took two British sergeants hostage as attempted leverage against the planned execution of three Irgun operatives. After the executions were carried out,
17850-605: The Earth is currently in the early stages of a human-caused mass extinction, known as the Holocene extinction . In that survey, the same proportion of respondents agreed with the prediction that up to 20% of all living populations could become extinct within 30 years (by 2028). A 2014 special edition of Science declared there is widespread consensus on the issue of human-driven mass species extinctions. A 2020 study published in PNAS stated that
18060-595: The Golan Heights from Syria. Jerusalem's boundaries were enlarged, incorporating East Jerusalem. The 1949 Green Line became the administrative boundary between Israel and the occupied territories . Following the 1967 war and the " Three Nos " resolution of the Arab League, Israel faced attacks from the Egyptians in the Sinai Peninsula during the 1967–1970 War of Attrition , and from Palestinian groups targeting Israelis in
18270-580: The Golan Heights. The international community largely rejected these moves, with the UN Security Council declaring both the Jerusalem Law and the Golan Heights Law null and void. Several waves of Ethiopian Jews immigrated to Israel since the 1980s, while between 1990 and 1994, immigration from the post-Soviet states increased Israel's population by twelve percent. On 7 June 1981, during
18480-813: The Holocaust. During the 1950s, Israel was frequently attacked by Palestinian fedayeen , nearly always against civilians, mainly from the Egyptian-occupied Gaza Strip, leading to several Israeli reprisal operations . In 1956, the UK and France aimed at regaining control of the Suez Canal , which Egypt had nationalized. The continued blockade of the Suez Canal and Straits of Tiran to Israeli shipping, together with increasing fedayeen attacks against Israel's southern population and recent Arab threatening statements, prompted Israel to attack Egypt. Israel joined
18690-472: The Irgun killed the two British soldiers, hanged their bodies from trees, and left a booby trap at the scene which injured a British soldier. The incident caused widespread outrage in the UK. In September 1947, the British cabinet decided to evacuate Palestine as the Mandate was no longer tenable. On 29 November 1947, the General Assembly adopted Resolution 181 (II) . The plan attached to the resolution
18900-790: The Israelites and their culture branched out of the Canaanite peoples through the development of a distinct monolatristic —and later monotheistic —religion centered on Yahweh . They spoke an archaic form of Hebrew , known as Biblical Hebrew . Around the same time, the Philistines settled on the southern coastal plain . Most modern scholars agree that the Exodus narrative in the Torah and Old Testament did not take place as depicted; however, some elements of these traditions do have historical roots . There
19110-638: The Levant was fairly cosmopolitan, with religious freedoms for Christians, Muslims, and Jews . In 1561 the Ottoman sultan invited Sephardi Jews escaping the Spanish Inquisition to settle in and rebuild the city of Tiberias . Under the Ottoman Empire's millet system , Christians and Jews were considered dhimmi (meaning "protected") under Ottoman law in exchange for loyalty to the state and payment of
19320-413: The Mosasauridae into three groups. These groups were based on how developed the rostrum was on the premaxilla, the size of the suprastapedial process of the quadrate and if the haemal arches were fused to the centra of the caudal vertebrae. Prognathodon was placed alongside Platecarpus in a "microrhynchous" group. The two other groups were the "megarhynchous" (including Tylosaurus and Hainosaurus ) and
19530-519: The Mosasaurinae, for the first time ever, as a close relative of the genera Globidens and Plesiotylosaurus . The tribe Prognathodontini was synonymized with the Globidensini , another tribe coined by Russell (1967) for Globidens . Bell was also the first to note that his analysis recovered Prognathodon , previously believed to be monophyletic , as paraphyletic . The view of the relationships of
19740-570: The North American moose and that the animal had once been common on the British Isles . Rather than suggest that this indicated the possibility of species going extinct, he argued that although organisms could become locally extinct, they could never be entirely lost and would continue to exist in some unknown region of the globe. The antlers were later confirmed to be from the extinct deer Megaloceros . Hooke and Molyneux's line of thinking
19950-686: The PLO signed the Oslo Accords , which gave the Palestinian National Authority (PNA) the right to govern parts of the West Bank and the Gaza Strip. The PLO also recognized Israel's right to exist and pledged an end to terrorism. In 1994, the Israel–Jordan peace treaty was signed, making Jordan the second Arab country to normalize relations with Israel. Arab public support for the Accords
20160-460: The Palestinian territories have drawn sustained international criticism—along with accusations that it has committed war crimes and crimes against humanity against the Palestinian people—from human rights organizations and United Nations officials. The country's Basic Laws establish a unicameral parliament elected by proportional representation , the Knesset , which determines the makeup of
20370-518: The Paris basin, could be formed by a slow rise and fall of sea levels . The concept of extinction was integral to Charles Darwin 's On the Origin of Species , with less fit lineages disappearing over time. For Darwin, extinction was a constant side effect of competition . Because of the wide reach of On the Origin of Species , it was widely accepted that extinction occurred gradually and evenly (a concept now referred to as background extinction ). It
20580-817: The Plioplatecarpinae, but erected a tribe for the genus and the related Plesiotylosaurus , the Prognathodontini . He considered the mosasaurs within the Prognathodontini to "clearly be of plioplatecarpine derivation", but justified the tribe by that they can be differentiated from other plioplatecarpines by their massive jaws and robust teeth. Gorden L. Bell Jr. conducted the first major phylogenetic analysis of mosasaurs in 1997, utilizing new methodologies and incorporating further taxa described since Russell's 1967 monograph (particularly basal mosasauroids, such as Aigialosaurus ). Bell recovered Prognathodon within
20790-567: The Second Aliyah who became leaders of the Yishuv in the coming decades believed that the Jewish settler economy should not depend on Arab labor. This would be a dominant source of antagonism with the Arab population, with the new Yishuv's nationalist ideology overpowering its socialist one. Though the immigrants of the Second Aliyah largely sought to create communal Jewish agricultural settlements, Tel Aviv
21000-578: The Sinai Peninsula since 1957, and blocked Israel's access to the Red Sea. Other Arab states mobilized their forces. Israel reiterated that these actions were a casus belli and launched a pre-emptive strike ( Operation Focus ) against Egypt in June. Jordan, Syria and Iraq attacked Israel. In the Six-Day War , Israel captured and occupied the West Bank from Jordan, the Gaza Strip and Sinai Peninsula from Egypt, and
21210-637: The United States government, to force the removal of Native Americans , many of whom relied on the bison for food. Israel Israel , officially the State of Israel , is a country in the Southern Levant region of West Asia . It is bordered by Lebanon and Syria to the north, the West Bank and Jordan to the east, the Gaza Strip and Egypt to the southwest, and the Mediterranean Sea to
21420-634: The West Bank and the Gaza Strip. On 11 March 1978, a PLO guerilla raid from Lebanon led to the Coastal Road massacre . Israel responded by launching an invasion of southern Lebanon to destroy PLO bases. Most PLO fighters withdrew, but Israel was able to secure southern Lebanon until a UN force and the Lebanese army could take over. The PLO soon resumed its insurgency against Israel, and Israel carried out numerous retaliatory attacks. Meanwhile, Begin's government provided incentives for Israelis to settle in
21630-429: The accumulation of slightly deleterious mutations , then a population will go extinct. Smaller populations have fewer beneficial mutations entering the population each generation, slowing adaptation. It is also easier for slightly deleterious mutations to fix in small populations; the resulting positive feedback loop between small population size and low fitness can cause mutational meltdown . Limited geographic range
21840-456: The adult male Palestinian Arab population was killed, wounded, imprisoned or exiled. The British introduced restrictions on Jewish immigration to Palestine with the White Paper of 1939 . With countries around the world turning away Jewish refugees fleeing the Holocaust , a clandestine movement known as Aliyah Bet was organized to bring Jews to Palestine. By the end of World War II , 31% of
22050-546: The area of the former Mandate the next day, beginning the First Arab–Israeli War . Subsequent armistice agreements established Israeli control over 77 percent of the former Mandate territory. The majority of Palestinian Arabs were either expelled or fled in what is known as the Nakba , with those remaining becoming the new state's main minority. Over the following decades, Israel's population increased greatly as
22260-456: The armies of four Arab countries—Egypt, Syria, Transjordan and Iraq—entered what had been Mandatory Palestine, launching the 1948 Arab–Israeli War ; contingents from Yemen , Morocco , Saudi Arabia , and Sudan joined the war. The purpose of the invasion was to prevent the establishment of the Jewish state and to "sweep them [Jews] into the sea". The Arab League stated the invasion was to restore order and prevent further bloodshed. After
22470-408: The base. The size of P. lutugini teeth also varies depending on the position in the mouth. Anterior pterygoid teeth are noted for being relatively large and comparable to the marginal teeth in size at up to 4.6 cm in height. A relatively small number of teeth in comparison to other mosasaurs (for an example, Prognathodon saturator preserves 14 teeth in the dentary, 12 in the maxilla and 6 in
22680-554: The basis that the partition plan privileged European interests over those of the Palestinians, and indicated that they would reject any other plan of partition. On 1 December 1947, the Arab Higher Committee proclaimed a three-day strike, and riots broke out in Jerusalem . The situation spiraled into a civil war . Colonial Secretary Arthur Creech Jones announced that the British Mandate would end on 15 May 1948, at which point
22890-479: The cervical vertebrae. Vertebral hypapophyseal peduncles were found on all four vertebrae and are very short and end as small laterally compressed oval facets. These facets are posteriorly inclined and located posteriorly on the ventral surfaces of the centra. The synapophyses are large, located anteriorly on the centra, and do not extend below the ventral margin of the centrum. The condyles and cotyles are weakly dorsoventrally compressed. The cervical vertebrae are almost
23100-462: The church and the authorities. Many Jews had emigrated to flourishing diaspora communities, while locally there was both Christian immigration and local conversion. By the middle of the 5th century, there was a Christian majority. Towards the end of the 5th century, Samaritan revolts erupted, continuing until the late 6th century and resulting in a large decrease in the Samaritan population. After
23310-600: The consequences can be catastrophic. Invasive alien species can affect native species directly by eating them, competing with them, and introducing pathogens or parasites that sicken or kill them; or indirectly by destroying or degrading their habitat. Human populations may themselves act as invasive predators. According to the "overkill hypothesis", the swift extinction of the megafauna in areas such as Australia (40,000 years before present), North and South America (12,000 years before present), Madagascar , Hawaii (AD 300–1000), and New Zealand (AD 1300–1500), resulted from
23520-418: The contemporary extinction crisis "may be the most serious environmental threat to the persistence of civilization, because it is irreversible." Biologist E. O. Wilson estimated in 2002 that if current rates of human destruction of the biosphere continue, one-half of all plant and animal species of life on earth will be extinct in 100 years. More significantly, the current rate of global species extinctions
23730-468: The country received an influx of Jews who emigrated, fled or were expelled from the Muslim world . Following the 1967 Six-Day War Israel occupied the West Bank, Gaza Strip, Egyptian Sinai Peninsula and Syrian Golan Heights . Israel established and continues to expand settlements across the illegally occupied territories , contrary to international law, and has effectively annexed East Jerusalem and
23940-615: The crusaders before Muslim rule was fully restored by the Mamluk sultans of Egypt in 1291. In 1516, the Ottoman Empire conquered the region and ruled it as part of Ottoman Syria . Two violent incidents took place against Jews, the 1517 Safed attacks and the 1517 Hebron attacks , after the Turkish Ottomans ousted the Mamluks during the Ottoman–Mamluk War . Under the Ottoman Empire,
24150-471: The deliberate destruction of some species, such as dangerous viruses , and the total destruction of other problematic species has been suggested. Other species were deliberately driven to extinction, or nearly so, due to poaching or because they were "undesirable", or to push for other human agendas. One example was the near extinction of the American bison , which was nearly wiped out by mass hunts sanctioned by
24360-486: The destruction of most of Palestine's Arab culture , identity , and national aspirations . Some 156,000 Arabs remained and became Arab citizens of Israel . By United Nations General Assembly Resolution 273 , Israel was admitted as a member of the UN on 11 May 1949. In the early years of the state, the Labor Zionist movement led by Prime Minister Ben-Gurion dominated Israeli politics . Immigration to Israel during
24570-527: The diagnosis in this study remains the latest published emended diagnosis for the genus. In 1998, an intact fossil skull was found in the Maastricht limestone quarries. Shortly after, it was nicknamed "Bèr", and put on display in the Maastricht Natural History Museum . This specimen was then identified as a Prognathodon , and received the species name Prognathodon saturator . This specimen
24780-402: The diverging suspensorial rami, which forms a rectangular field medially on the parietal. The ventral process of the postorbitofrontal to jugal is indistinctly separated from the moderately well exposed dorsal surface of the postorbitofrontal and the ventroposterior process on the jugal is slightly developed to absent. The squamosal wing to the parietal is large. There is a deep groove present in
24990-538: The dorsal margin of said bone a nearly 110 degree angle between the horizontal anterior end and the subvertical posterior wing. This combination of characteristics aided in determining that P. lutugini was indeed a species of Prognathodon , since some researchers place it in a genus of its own, " Dollosaurus ". Species referred to Prognathodon are variable in terms of dentition. Robust and conical teeth with blunt, serrated carinae and smooth enamel are often seen as typical Prognathodon teeth and are routinely assigned to
25200-469: The early late Campanian (c. 74.5 Ma) Bearpaw Formation in Alberta , Canada provided the first fully articulated skeletons of the genus. Detailed studies of these and previously discovered specimens allowed several characters to be established that distinguishes Prognathodon from closely related genera like Liodon and long-snouted mosasaurines. The preserved teeth and gut contents also allowed studies into
25410-403: The endangered wild water buffalo is most threatened with extinction by genetic pollution from the abundant domestic water buffalo ). Such extinctions are not always apparent from morphological (non-genetic) observations. Some degree of gene flow is a normal evolutionary process; nevertheless, hybridization (with or without introgression) threatens rare species' existence. The gene pool of
25620-570: The entirety of the Gaza Strip and over 90% of the West Bank with Jerusalem as a shared capital. Each side blamed the other for the failure of the talks. In late 2000, after a controversial visit by Sharon to the Temple Mount , the Second Intifada began. Palestinian suicide bombings were a recurrent feature. Some commentators contend that the intifada was pre-planned by Arafat after the collapse of peace talks. Sharon became prime minister in
25830-575: The event of rediscovery would be considered Lazarus species. Examples include the thylacine , or Tasmanian tiger ( Thylacinus cynocephalus ), the last known example of which died in Hobart Zoo in Tasmania in 1936; the Japanese wolf ( Canis lupus hodophilax ), last sighted over 100 years ago; the American ivory-billed woodpecker ( Campephilus principalis ), with the last universally accepted sighting in 1944; and
26040-399: The evolution of a crescent-shaped tail fluke to aid in locomotion. The tail fluke is clearly asymmetric. The lower fin lobe follows the caudal vertebrae and would have had a streamlined cross-section in life, based on the proportions of the axial skeleton and the other soft tissues. The upper fin lobe is unsupported by the skeleton and is preserved as a small, almost wing-like, structure above
26250-508: The external surface. Teeth of P. solvayi are more slender at the anterior end, broader and triangular towards the middle of the dental ramus, decreasing in size towards the posterior end with the last two teeth being rather stunted. The tooth crowns are only moderately inflated and in contrast to the teeth of P. overtoni , the teeth increase in size consistently in an antero-posterior direction and are somewhat less posteriorly recurved. In P. overtoni and other species (such as P. giganteus ),
26460-443: The extinction crisis. According to the International Union for Conservation of Nature (IUCN), 784 extinctions have been recorded since the year 1500, the arbitrary date selected to define "recent" extinctions, up to the year 2004; with many more likely to have gone unnoticed. Several species have also been listed as extinct since 2004. If adaptation increasing population fitness is slower than environmental degradation plus
26670-475: The extinction of species caused by humanity, and they try to prevent further extinctions through a variety of conservation programs. Humans can cause extinction of a species through overharvesting , pollution , habitat destruction , introduction of invasive species (such as new predators and food competitors ), overhunting, and other influences. Explosive, unsustainable human population growth and increasing per capita consumption are essential drivers of
26880-533: The field of zoology , and biology in general, and has also become an area of concern outside the scientific community. A number of organizations, such as the Worldwide Fund for Nature , have been created with the goal of preserving species from extinction. Governments have attempted, through enacting laws, to avoid habitat destruction, agricultural over-harvesting, and pollution . While many human-caused extinctions have been accidental, humans have also engaged in
27090-457: The floor of the basioccipital for the basilar artery . The suprastapedial process is fused to the infrastapedial process on the quadrate and the tympanic ala are thick. The stapedial pit is nearly circular to elliptical in form. The posterior process on the dorsal surface of the dentary is present, incipient or absent and the dentary terminates abruptly in front of the first dentary tooth. The dentary itself contains thirteen to fourteen teeth and
27300-564: The following several centuries, dropping from an estimated 1 million during Roman and Byzantine periods to about 300,000 by the early Ottoman period , and there was a steady process of Arabization and Islamization . The end of the 11th century brought the Crusades , papally -sanctioned incursions of Christian crusaders intent on wresting Jerusalem and the Holy Land from Muslim control and establishing crusader states . The Ayyubids pushed back
27510-435: The form of a physical destruction of niche habitats. The widespread destruction of tropical rainforests and replacement with open pastureland is widely cited as an example of this; elimination of the dense forest eliminated the infrastructure needed by many species to survive. For example, a fern that depends on dense shade for protection from direct sunlight can no longer survive without forest to shelter it. Another example
27720-439: The genus Prognathodon was provided by Lingham-Soliar and Nolf (1989), and states that the premaxilla lacks a rostrum anterior to the premaxillary teeth. The prefrontal forms a large portion of the posterolateral border of the external nares and the supraorbital wing with heavy triangular ala contacts the postorbitofrontal posteriorly over the orbit medial to the external edge of the frontal. The frontal does not emarginate above
27930-527: The genus to other mosasaur genera has changed little since 1997, it is routinely recovered as within the Mosasaurinae as well as paraphyletic. Cau and Madzia (2017) noted that the inclusion of Prognathodon and Plesiotylosaurus within the Globidensini would suggest a closer relationship between the genera than the reality of the situation. Though Prognathodon and Plesiotylosaurus are routinely recovered as sister genera, Cau and Madzia (2017) did not resurrect
28140-457: The genus. The first later use of " Prognathodon " was by Dale A. Russell in a comprehensive monograph on North American mosasaurs in 1967, where the priority of Prognathodon was made apparent. Russell also revised the species assigned to Prognathodon from North America , but only briefly commented on the Belgian specimens. Though the original remains of the genus were rather comprehensive and
28350-428: The genus. Despite this, the type species P. solvayi differs from this type of teeth and instead exhibits markedly labio-lingually compressed and gently facetted marginal teeth. There is also a certain degree of variation in tooth number, P. solvayi has 12 teeth on the maxilla and 13 on the dentary whilst P. overtoni has 14 dentary teeth. Of all species, P. solvayi has the most different teeth from other members of
28560-413: The genus. The tooth crowns are generally large and quite strongly striated and the anterior teeth are more procumbent than in any other mosasaurs. The premaxillary teeth are almost horizontal and the anterior dentary teeth only slightly less so. In addition to deep striae, the fragmentary material of P. solvayi reveals that the tooth crowns may also have been somewhat prismatic with seven to eight prisms on
28770-436: The government of responsibility for failures before and during the war, but public anger forced Prime Minister Golda Meir to resign. In July 1976, an airliner was hijacked in flight from Israel to France by Palestinian guerrillas; Israeli commandos rescued 102 of 106 Israeli hostages . The 1977 Knesset elections marked a major turning point in Israeli political history as Menachem Begin 's Likud party took control from
28980-580: The habitat retreat of taxa approaching extinction. Possible causes of the higher extinction risk in species with more sexual selection shown by the comprehensive fossil studies that rule out such error sources include expensive sexually selected ornaments having negative effects on the ability to survive natural selection , as well as sexual selection removing a diversity of genes that under current ecological conditions are neutral for natural selection but some of which may be important for surviving climate change. There have been at least five mass extinctions in
29190-660: The heaviest losses include the Cape Floristic Region and the Caribbean Basin . These areas might see a doubling of present carbon dioxide levels and rising temperatures that could eliminate 56,000 plant and 3,700 animal species. Climate change has also been found to be a factor in habitat loss and desertification . Studies of fossils following species from the time they evolved to their extinction show that species with high sexual dimorphism , especially characteristics in males that are used to compete for mating, are at
29400-447: The history of life on earth, and four in the last 350 million years in which many species have disappeared in a relatively short period of geological time. A massive eruptive event that released large quantities of tephra particles into the atmosphere is considered to be one likely cause of the " Permian–Triassic extinction event " about 250 million years ago, which is estimated to have killed 90% of species then existing. There
29610-475: The human era since the Late Pleistocene would require 5 to 7 million years to recover. According to the 2019 Global Assessment Report on Biodiversity and Ecosystem Services by IPBES , the biomass of wild mammals has fallen by 82%, natural ecosystems have lost about half their area and a million species are at risk of extinction—all largely as a result of human actions. Twenty-five percent of plant and animal species are threatened with extinction. In
29820-518: The increasing persecution of Jews in 1930s Europe led to the Fifth Aliyah , with an influx of a quarter of a million Jews. This was a major cause of the Arab revolt of 1936–39 , which was suppressed by British security forces and Zionist militias. Several hundred British security personnel and Jews were killed; 5,032 Arabs were killed, 14,760 were wounded, and 12,622 were detained. An estimated ten percent of
30030-507: The inferred paleoecology of the genus. In 2009, Hani F. Kaddumi described the new species P. hudae based on a complete right dentary and partial left dentary from the Maastrichtian-age Muwaqqar Chalk Marl Formation of Harrana , Jordan. The specific name hudae honors Kaddumi's daughter Huda. Also described in 2009 by Kaddumi from the same locality was P. primus , based on an articulated upper skull, with
30240-431: The introduced ( or hybrid ) species. Endemic populations can face such extinctions when new populations are imported or selectively bred by people, or when habitat modification brings previously isolated species into contact. Extinction is likeliest for rare species coming into contact with more abundant ones; interbreeding can swamp the rarer gene pool and create hybrids, depleting the purebred gene pool (for example,
30450-418: The labial. The teeth have a slight posterior and medial recurvature and are noted for having smooth surfaces (which is different from other species of Prognathodon ) except for minor wrinkles at the tips of the tooth crowns. The roots of the teeth are quite large at about 1.5 times the size of the tooth crowns and are barrel-shaped. The largest known tooth crowns are up to 5.5 cm tall and 2.5 cm wide at
30660-512: The larger specimens in the genus. Thus, it was assumed that ERMNH HFV 197 was a juvenile specimen. That tail fin would grow to account for the increased weight of bigger individuals is rather logical, and can be observed in other groups that possess tail fins, such as extant sharks and extinct ichthyosaurs . For this reason, it is likely that adult Prognathodon , particularly of the larger species, would have had larger tail fins relative to their body size. Lindgren et al. (2013) especially noted that
30870-406: The last few caudal vertebrae. The shape of the tail fluke is similar to that of the carcharhinid sharks, though turned upside down, with a small upper lobe and large lower lobe. The specimen also preserves scale impressions, particularly around the outline of the tail fluke. Though faint, these impressions reveal scales of "rhomboidal" shape. Similarly shaped scales are present on the tail fluke of
31080-485: The late 1940s and early 1950s was aided by the Israeli Immigration Department and the non-government sponsored Mossad LeAliyah Bet ( lit. "Institute for Immigration B "). The latter engaged in clandestine operations in countries, particularly in the Middle East and Eastern Europe, where the lives of Jews were in danger and exit was difficult. Mossad LeAliyah Bet was disbanded in 1953. The immigration
31290-457: The latter were often re-designated for the former, so Jews newly arrived from Arab lands generally ended up staying longer in transit camps. During this period, food, clothes and furniture were rationed in what became known as the austerity period . The need to solve the crisis led Ben-Gurion to sign a reparations agreement with West Germany that triggered mass protests by Jews angered at the idea that Israel could accept monetary compensation for
31500-440: The main drivers of the modern extinction crisis. In January 2020, the UN's Convention on Biological Diversity drafted a plan to mitigate the contemporary extinction crisis by establishing a deadline of 2030 to protect 30% of the Earth's land and oceans and reduce pollution by 50%, with the goal of allowing for the restoration of ecosystems by 2050. The 2020 United Nations ' Global Biodiversity Outlook report stated that of
31710-421: The modern understanding of extinction as the end of a species was incompatible with the prevailing worldview. Prior to the 19th century, much of Western society adhered to the belief that the world was created by God and as such was complete and perfect. This concept reached its heyday in the 1700s with the peak popularity of a theological concept called the great chain of being , in which all life on earth, from
31920-717: The month-long Second Lebanon War . In 2007 the Israeli Air Force destroyed a nuclear reactor in Syria. In 2008, a ceasefire between Hamas and Israel collapsed, resulting in the three-week Gaza War . In what Israel described as a response to over a hundred Palestinian rocket attacks on southern Israeli cities, Israel began an operation in the Gaza Strip in 2012, lasting eight days. Israel started another operation in Gaza following an escalation of rocket attacks by Hamas in July 2014. In May 2021 another round of fighting took place in Gaza and Israel, lasting eleven days. By
32130-490: The name of a species or other taxon normally indicates its status as extinct. Examples of species and subspecies that are extinct include: A species is extinct when the last existing member dies. Extinction therefore becomes a certainty when there are no surviving individuals that can reproduce and create a new generation. A species may become functionally extinct when only a handful of individuals survive, which cannot reproduce due to poor health, age, sparse distribution over
32340-474: The name referencing it being the first near-complete mosasaur skull from the Middle East. The new species and genus Tenerasaurus hashimi , also described by Kaddumi in 2009 from Harrana from fossils likely belonging to a juvenile specimen (preserving the entire skeleton aside from the head, which may have been destroyed during quarrying) was reclassified as Prognathodon hashimi by Kaddumi, Johan Lindgren, and Michael J. Polcyn in 2013. The species name references
32550-469: The natural course of events, species become extinct for a number of reasons, including but not limited to: extinction of a necessary host, prey or pollinator, interspecific competition , inability to deal with evolving diseases and changing environmental conditions (particularly sudden changes) which can act to introduce novel predators, or to remove prey. Recently in geological time, humans have become an additional cause of extinction of some species, either as
32760-410: The newly emerging school of uniformitarianism . Jean-Baptiste Lamarck , a gradualist and colleague of Cuvier, saw the fossils of different life forms as evidence of the mutable character of species. While Lamarck did not deny the possibility of extinction, he believed that it was exceptional and rare and that most of the change in species over time was due to gradual change. Unlike Cuvier, Lamarck
32970-410: The occupied territories, globally, and in Israel. Most important among the Palestinian and Arab groups was the Palestine Liberation Organization (PLO), established in 1964, which initially committed itself to "armed struggle as the only way to liberate the homeland". In the late 1960s and early 1970s, Palestinian groups launched attacks against Israeli and Jewish targets around the world, including
33180-403: The one hand, and Syria and Lebanon on the other. Arab nationalists led by Egyptian President Gamal Abdel Nasser refused to recognize Israel and called for its destruction. By 1966 Israeli-Arab relations had deteriorated to the point of battles taking place between Israeli and Arab forces. In May 1967, Egypt massed its army near the border with Israel, expelled UN peacekeepers stationed in
33390-447: The only person executed in Israel by conviction in an Israeli civilian court. In 1963, Israel was engaged in a diplomatic standoff with the United States in relation to the Israeli nuclear programme . Since 1964 Arab countries, concerned over Israeli plans to divert waters of the Jordan River into the coastal plain , had been trying to divert the headwaters to deprive Israel of water resources, provoking tensions between Israel on
33600-413: The orbits and a median dorsal ridge is either present or absent. The foramina on the parietal are small to moderately large, located anteromedially on a small prominence and are closely embraced on either side by short tongues from the frontal or located on the frontoparietal suture. The margins of the dorsal parietal surface are parallel to one another and the cranial midline to the posterior base of
33810-540: The original description was brief, no additional studies of the type material was done for a century. The lack of a comprehensive original description of the genus and the species referred to it from Belgium is strangely not unusual for mosasaur specimens discovered in the Craie de Ciply Formation of Belgium . Large amounts of work was commonly invested in extracting and mounting the specimens, but scientific study of them remained limited with diagnoses and descriptions mainly focusing on peculiar points of their anatomy, such as
34020-399: The original population, thereby increasing the chance of extinction. Habitat degradation is currently the main anthropogenic cause of species extinctions. The main cause of habitat degradation worldwide is agriculture, with urban sprawl , logging, mining, and some fishing practices close behind. The degradation of a species' habitat may alter the fitness landscape to such an extent that
34230-427: The patriarch Jacob who, according to the Hebrew Bible , was given the name after he successfully wrestled with the Angel of the Lord . The earliest known archaeological artifact to mention the word Israel as a collective is the Merneptah Stele of ancient Egypt (dated to the late 13th century BCE). Early hominin presence in the Levant , where Israel is located, dates back at least 1.5 million years based on
34440-417: The population of Palestine was Jewish. The UK found itself facing a Jewish insurgency over immigration restrictions and continued conflict with the Arab community over limit levels. The Haganah joined Irgun and Lehi in an armed struggle against British rule. The Haganah attempted to bring tens of thousands of Jewish refugees and Holocaust survivors to Palestine by ship. Most of the ships were intercepted by
34650-399: The posterior end of the splenial and the anterior blade of the prearticular and have a tooth count of 13, with at least eight teeth possessing subdental crypts with some replacement teeth having been found in the type specimen. The subdental crypts are positioned postermodeially to the functional teeth. The coronoid is saddle-shaped and has a well-developed posterodorsal process, which gives
34860-702: The process of speciation —where new varieties of organisms arise and thrive when they are able to find and exploit an ecological niche —and species become extinct when they are no longer able to survive in changing conditions or against superior competition . The relationship between animals and their ecological niches has been firmly established. A typical species becomes extinct within 10 million years of its first appearance, although some species, called living fossils , survive with little to no morphological change for hundreds of millions of years. Mass extinctions are relatively rare events; however, isolated extinctions of species and clades are quite common, and are
35070-518: The province of Judea was renamed Syria Palaestina . Jews were expelled from the districts surrounding Jerusalem. Nevertheless, there was a continuous small Jewish presence, and Galilee became its religious center. Early Christianity displaced Roman paganism in the 4th century CE, with Constantine embracing and promoting the Christian religion and Theodosius I making it the state religion . A series of laws were passed that discriminated against Jews and Judaism, and Jews were persecuted by both
35280-407: The pterygoid) is a characteristic present in all species of Prognathodon . Post-cranial fossils of Prognathodon are considerably rarer than skull material and isolated teeth. The type specimen of P. lutugini preserves several vertebrae. Four cervical vertebrae are preserved, which have well developed anterior and posterior zygapophyses. Functional zygosphenes and zygantra were found on two of
35490-444: The region and established Mandatory Palestine in 1920. Increased Jewish immigration in the leadup to the Holocaust and British colonial policy led to intercommunal conflict between Jews and Arabs , which escalated into a civil war in 1947 after the United Nations (UN) proposed partitioning the land between them. The State of Israel declared its establishment on 14 May 1948. The armies of neighboring Arab states invaded
35700-473: The region in 63 BCE, first taking control of Syria , and then intervening in the Hasmonean civil war . The struggle between pro-Roman and pro- Parthian factions in Judea led to the installation of Herod the Great as a dynastic vassal of Rome . In 6 CE, the area was annexed as the Roman province of Judaea ; tensions with Roman rule led to a series of Jewish–Roman wars , resulting in widespread destruction. The First Jewish–Roman War (66–73 CE) resulted in
35910-410: The relative importance of genetic factors compared to environmental ones as the causes of extinction has been compared to the debate on nature and nurture . The question of whether more extinctions in the fossil record have been caused by evolution or by competition or by predation or by disease or by catastrophe is a subject of discussion; Mark Newman, the author of Modeling Extinction , argues for
36120-440: The right one is at the moment almost entirely a restoration made of gypsum , with two original teeth. The left pterygoid was considerably more complete, but lacked all processes other than the basisphenoid process. The posterior alveolar margin was noted for being very small, with the teeth rising from a thin but pronounced vertical ridge. The ventral surface of the basisphenoid process is quite smooth and foramina are visible above
36330-447: The same size as the longest dorsal vertebrae and get to a length of about 7.2 cm and a height of 4 cm. 26 dorsal vertebrae are preserved in the specimen, though most are poorly preserved. Dorsal vertebrae are a little longer than the cervical and like the cervicals, the condyles and cotyles are weakly compressed dorsoventrally. The dorsal vertebrae are at a maximum about 8 cm in length and 5 cm in height. Additionally,
36540-404: The sixth tooth on the lateral surface of the pterygoid as well as above the position between the sixth and seventh teeth on the medial surface. The squamosal bone is only represented by a few fragments, but could be noted for being laterally compressed and tall, as in other species of Prognathodon . Its posteroventral surface is concave for contact with the quadrate . The dentaries are fused with
36750-408: The skull and part of the body and tail of the approximately 13-metre long skeleton. Based on stratigraphy , the age of the specimen was estimated at 67.83 million years, making it about one-and-a-half million years older than "Bèr". From what has been uncovered, this appears to be the oldest known Mosasaurid specimen from the Maastricht area, and it belongs to the genus Prognathodon . The specimen
36960-474: The soft tissue morphology of Prognathodon . The fossil is not only largely complete and articulated, which is rare for Prognathodon specimens, but also preserves significant portions of the integument and a gentle bend on the last few caudal vertebrae. Most importantly, the fossil preserves the soft tissue outline of a tail fin. This helped in providing evidence that mosasaurs were convergent with ichthyosaurs , metriorhynchid thalattosuchians and whales in
37170-460: The species is no longer able to survive and becomes extinct. This may occur by direct effects, such as the environment becoming toxic , or indirectly, by limiting a species' ability to compete effectively for diminished resources or against new competitor species. Habitat destruction, particularly the removal of vegetation that stabilizes soil, enhances erosion and diminishes nutrient availability in terrestrial ecosystems. This degradation can lead to
37380-502: The stated objectives of destroying Hamas and freeing hostages. The fifth war of the Gaza–Israel conflict since 2008, it has been the deadliest for Palestinians in the entire Israeli–Palestinian conflict and the most significant military engagement in the region since the Yom Kippur War in 1973. Israel is accused of carrying out a genocide against the Palestinian people by experts, governments, United Nations agencies, and non-governmental organisations during its invasion of
37590-430: The sudden introduction of human beings to environments full of animals that had never seen them before and were therefore completely unadapted to their predation techniques. Coextinction refers to the loss of a species due to the extinction of another; for example, the extinction of parasitic insects following the loss of their hosts. Coextinction can also occur when a species loses its pollinator , or to predators in
37800-445: The teeth are all fairly uniform in size other than the penultimate tooth base (which is larger). Another species noted for its divergent teeth is P. lutugini , where knowledge of the teeth mainly comes from isolated specimens. Known teeth are strongly bicarinate with a weak serration on both carinae. The carinae then divide the teeth into lingual and labial surfaces, with the lingual surface being more convex and large in comparison with
38010-457: The temporal region and the braincase, which are all stout compared to other species in the genus. The type specimen of P. lutugini , whilst incomplete, preserves a significantly large portion of the skull. The specimen was originally stated to have preserved a small portion of the premaxilla , though D.V. Grigoriev (2013) noted that said bone could at the moment not be located, and it is potentially missing. Both pterygoids were preserved, though
38220-427: The territory was divided between Britain and France under the mandate system , and the British-administered area (including modern Israel) was named Mandatory Palestine . Arab opposition to British rule and Jewish immigration led to the 1920 Palestine riots and the formation of a Jewish militia known as the Haganah as an outgrowth of Hashomer, from which the Irgun and Lehi paramilitaries later split. In 1922,
38430-412: The tiniest microorganism to God, is linked in a continuous chain. The extinction of a species was impossible under this model, as it would create gaps or missing links in the chain and destroy the natural order. Thomas Jefferson was a firm supporter of the great chain of being and an opponent of extinction, famously denying the extinction of the woolly mammoth on the grounds that nature never allows
38640-448: The total extinction of the dodo and the extirpation of indigenous horses to the British Isles. He similarly argued against mass extinctions , believing that any extinction must be a gradual process. Lyell also showed that Cuvier's original interpretation of the Parisian strata was incorrect. Instead of the catastrophic floods inferred by Cuvier, Lyell demonstrated that patterns of saltwater and freshwater deposits , like those seen in
38850-412: The tribe Prognathodontini in their list of mosasaur clades and their preferred definitions, offering no comment as to why not. Within Cau and Madzia's analysis (as well as previous work such as Simões et al., 2017), Prognathodon is recovered as a sister group to the Mosasaurini , and the clade Mosasaurini + Prognathodon itself was recovered as forming a sister group to the Globidensini. The results of
39060-399: The type specimen also preserved many rib fragments, though all incomplete. Two grooves run along the distal portion of the ribs from the articular head but only one groove remains behind the middle of the rib. The discovery of the exceptionally well-preserved specimen ERMNH HFV 197 from Maastrichtian deposits in Harrana , central Jordan allowed detailed examinations of unique details of
39270-443: The upper fin lobe likely would have grown to be proportionally larger in bigger individuals. Modern phylogenetic analyses continually place Prognathodon within the Mosasaurinae subfamily, despite this Prognathodon has historically been seen as a genus sharing close relations with Platecarpus and the Plioplatecarpinae . Louis Dollo was one of the earliest researchers to work on mosasaur systematics, initially placing them as
39480-401: The ventral margin of the maxilla is slightly convex. The marginal teeth are massive, smooth and rounded as opposed to most other mosasaur teeth, which are typically facetted and laterally compressed. The mandible of P. saturator is very tall and massive, even more so than in other members of the genus. This is matched by the also massive pterygoid and various other portions of the skull, such as
39690-434: The vision of the Return in a more active way than most Christians viewed the Second Coming ." The religious Judaic notion of being a nation was distinct from the modern European notion of nationalism. The Jewish population of Palestine from the Ottoman rule to the beginning of the Zionist movement, known as the Old Yishuv , comprised a minority and fluctuated in size. During the 16th century, Jewish communities struck roots in
39900-453: The well-preserved Platecarpus specimen LACM 128319. The proportions of the soft tissue structures and their relation to the skeletal elements of the specimen can be used to infer the shape and size of the fins in other species of Prognathodon and potentially in other mosasaur genera as well. In their description of the specimen, Lindgren et al. (2013) noted that the specimen is strangely small by Prognathodon standards compared to some of
40110-440: The west. The country also has a small coastline on the Red Sea at its southernmost point, and part of the Dead Sea lies along its eastern border. Israel's proclaimed capital is in Jerusalem , while Tel Aviv is the country's largest urban area and economic center . Israel is located in a region known to Jews as the Land of Israel , synonymous with the Palestine region , the Holy Land , and Canaan . In antiquity, it
40320-530: The wider scientific community of his theory. Cuvier was a well-regarded geologist, lauded for his ability to reconstruct the anatomy of an unknown species from a few fragments of bone. His primary evidence for extinction came from mammoth skulls found in the Paris basin . Cuvier recognized them as distinct from any known living species of elephant, and argued that it was highly unlikely such an enormous animal would go undiscovered. In 1812, Cuvier, along with Alexandre Brongniart and Geoffroy Saint-Hilaire , mapped
40530-689: The world, it spends proportionally more on research and development than any other and is widely believed to possess nuclear weapons . Under the British Mandate (1920–1948), the entire region was known as Palestine . Upon establishment in 1948 , the country formally adopted the name State of Israel ( Hebrew : מְדִינַת יִשְׂרָאֵל , Medīnat Yisrā'el [mediˈnat jisʁaˈʔel] ; Arabic : دَوْلَة إِسْرَائِيل , Dawlat Isrāʼīl , [dawlat ʔisraːˈʔiːl] ) after other proposed names including Land of Israel ( Eretz Israel ), Ever (from ancestor Eber ), Zion , and Judea , were considered but rejected. The name Israel
40740-427: Was a unified Palestine under Arab rule. In February 1947, the British referred the Palestine issue to the newly formed United Nations . On 15 May 1947, the UN General Assembly resolved that a Special Committee be created "to prepare ... a report on the question of Palestine". The Report of the Committee proposed a plan to replace the British Mandate with "an independent Arab State, an independent Jewish State, and
40950-462: Was announced that nine days earlier, again a skeleton of what appears to be a Mosasaur was found in the limestone quarry just outside Maastricht, the same quarry that yielded the type specimen of Mosasaurus hoffmanni . Carlo Brauer, an excavator operator at the ENCI quarry, discovered the teeth of the fossil in the shovel of his digger on Monday morning, September 10. In the days following the discovery, museum staff retrieved several large sections of
41160-431: Was damaged by the continuation of Israeli settlements and checkpoints , and the deterioration of economic conditions. Israeli public support for the Accords waned after Palestinian suicide attacks . In November 1995, Rabin was assassinated by Yigal Amir , a far-right Jew who opposed the Accords. During Benjamin Netanyahu's premiership at the end of the 1990s, Israel agreed to withdraw from Hebron , though this
41370-425: Was difficult to disprove. When parts of the world had not been thoroughly examined and charted, scientists could not rule out that animals found only in the fossil record were not simply "hiding" in unexplored regions of the Earth. Georges Cuvier is credited with establishing the modern conception of extinction in a 1796 lecture to the French Institute , though he would spend most of his career trying to convince
41580-519: Was essentially that proposed in the report of 3 September. The Jewish Agency, the recognized representative of the Jewish community, accepted the plan, which assigned 55–56% of Mandatory Palestine to the Jews. At the time, the Jews were about a third of the population and owned around 6–7% of the land. Arabs constituted the majority and owned about 20% of the land, with the remainder held by the Mandate authorities or foreign landowners. The Arab League and Arab Higher Committee of Palestine rejected it on
41790-409: Was established as the first planned Jewish town in 1909. Jewish armed militias emerged during this period, the first being Bar-Giora in 1907. Two years later, the larger Hashomer organization was founded as its replacement. Chaim Weizmann's efforts to garner British support for the Zionist movement eventually secured the Balfour Declaration of 1917, stating Britain's support for the creation of
42000-544: Was estimated to have reached 10 metres (33 ft) in length. In 2002, Christiansen and Bonde estimated the maximum length of P. currii at 11 metres (36 ft). Considering this 2002 study, Schupl and colleagues estimated that P. saturator would have reached 12 metres (39 ft) in length. Though many species (such as P. currii , P. saturator and P. giganteus ) were large with sizes approaching or potentially exceeding 10 metres (33 ft), many species were considerably smaller in size. The type species , P. solvayi ,
42210-403: Was first described by Louis Dollo in 1889 based on specimens gathered in Belgium . There is some confusion over the correct generic name for the taxon. Dollo first mentioned the taxon as " Prognathodon " in some preliminary notes and provided a provisional diagnosis, but replaced the name Prognathodon with " Prognathosaurus " and used Prognathosaurus in all of his subsequent papers mentioning
42420-429: Was home to the Canaanite civilization followed by the kingdoms of Israel and Judah . Situated at a continental crossroad , the region experienced demographic changes under the rule of various empires from the Romans to the Ottomans . European antisemitism in the late 19th century galvanized Zionism , which sought a Jewish homeland in Palestine and gained British support . After World War I , Britain occupied
42630-825: Was in accordance with the One Million Plan . Some immigrants held Zionist beliefs or came for the promise of a better life, while others moved to escape persecution or were expelled from their homes. An influx of Holocaust survivors and Jews from Arab and Muslim countries to Israel during the first three years increased the number of Jews from 700,000 to 1,400,000. By 1958, the population had risen to two million. Between 1948 and 1970, approximately 1,150,000 Jewish refugees relocated to Israel. Some immigrants arrived as refugees and were housed in temporary camps known as ma'abarot ; by 1952, over 200,000 people were living in these tent cities. Jews of European background were often treated more favorably than Jews from Middle Eastern and North African countries—housing units reserved for
42840-406: Was never ratified or implemented, and he signed the Wye River Memorandum , giving greater control to the PNA. Ehud Barak , elected prime minister in 1999, withdrew forces from southern Lebanon and conducted negotiations with PNA Chairman Yasser Arafat and U.S. President Bill Clinton at the 2000 Camp David Summit . Barak offered a plan for the establishment of a Palestinian state , including
43050-430: Was not until 1982, when David Raup and Jack Sepkoski published their seminal paper on mass extinctions, that Cuvier was vindicated and catastrophic extinction was accepted as an important mechanism . The current understanding of extinction is a synthesis of the cataclysmic extinction events proposed by Cuvier, and the background extinction events proposed by Lyell and Darwin. Extinction is an important research topic in
43260-571: Was skeptical that catastrophic events of a scale large enough to cause total extinction were possible. In his geological history of the earth titled Hydrogeologie, Lamarck instead argued that the surface of the earth was shaped by gradual erosion and deposition by water, and that species changed over time in response to the changing environment. Charles Lyell , a noted geologist and founder of uniformitarianism , believed that past processes should be understood using present day processes. Like Lamarck, Lyell acknowledged that extinction could occur, noting
43470-564: Was suggested by David Ben-Gurion and passed by a vote of 6–3. In the early weeks after establishment, the government chose the term Israeli to denote a citizen of the Israeli state. The names Land of Israel and Children of Israel have historically been used to refer to the biblical Kingdom of Israel and the entire Jewish people respectively. The name Israel (Hebrew: Yīsrāʾēl ; Septuagint Ancient Greek : Ἰσραήλ , Israēl , " El (God) persists/rules", though after Hosea 12:4 often interpreted as "struggle with God") refers to
43680-486: Was suppressed; Muhammad Ali's army retreated and Ottoman rule was restored with British support in 1840. The Tanzimat reforms were implemented across the Ottoman Empire. The first wave of modern Jewish migration to Ottoman-ruled Palestine , known as the First Aliyah , began in 1881, as Jews fled pogroms in Eastern Europe. The 1882 May Laws increased economic discrimination against Jews, and restricted where they could live. In response, political Zionism took form,
43890-460: Was the first reasonably complete mosasaur specimen recovered from the Maastricht area since 1957; the skeleton is on display at the Natural History Museum at Maastricht, and is from an animal that was probably 12 metres long. A very large specimen found in Israel was for some time informally named " Oronosaurus ", but eventually described as a new species of Prognathodon , P. currii . Two specimens of Prognathodon overtoni described in 2011 from
44100-472: Was the smallest, reaching around 5–6 metres (16–20 ft) in length. P. hudae is estimated to have reached 6.4 metres (21 ft) in length. The largest P. overtoni specimen is TMP 2018.042.0005, a nearly completely preserved individual measuring over 7–8 metres (23–26 ft) in length. P. sectorius would have approached the same size, about 8 metres (26 ft) in length. Other than its size and robust nature, another feature that defines Prognathodon
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