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94-463: The term "Proavis" was first coined, although under the form "Pro-Aves", by English osteologist and zoologist William Plane Pycraft in "The Origin of Birds", a 1906 article published in the magazine Knowledge and Scientific News . Pycraft added to his article his own drawn depiction of the hypothetical animal, a restoration entitled "One of the Pro-Aves". Pycraft's "Pro-Avis" (singular of "Pro-Aves")

188-551: A cursorial origin, not arboreal. The "Pro-Avis" would thus have been a running animal, accelerating and prolonging its jumps by flapping with feathered forelimbs. There is a model of Nopcsa's restoration, still preserved, and recently restored, in the Grant Museum of Zoology , the only remaining university zoological museum in London. When Pycraft published his book A History of Birds , in 1910, he reinserted his 1906 restoration One of

282-511: A ceremonial county, or be a district tier within a shire county , be a royal or metropolitan borough , have borough or city status , or be a unitary authority . At the community level, much of England is divided into civil parishes with their own councils ; in Greater London only one such parish, Queen's Park , exists as of 2014 after they were abolished in 1965 until legislation allowed their recreation in 2007. From 1994 until

376-469: A decoupling of the locomotor functions of the forelimbs and hindlimbs, a lack of a propatagium , a shallow tail, and a weight of about one kilogramme. Paul illustrated his analysis with a skeletal diagram, accompanied by a life illustration of a "proavis". When in 2013 Aurornis was described, at the time the most basal known member of the Avialae , the group consisting of birds and their closest relatives,

470-501: A geographical frame of reference. Of these, 38 developed gradually since the Middle Ages ; these were reformed to 51 in 1974 and to their current number in 1996. Each has a Lord Lieutenant and High Sheriff ; these posts are used to represent the British monarch locally. Some counties, such as Herefordshire , are only divided further into civil parishes. The royal county of Berkshire and

564-407: A patagium. The interremigial ligament that connects the bases all the primary and secondary feathers as it passes from the tip of the hand to the elbow is thought to represent the caudal edge of the ancestral form of this patagium. The scansoriopterygid dinosaurs Yi and Ambopteryx had rather elaborate, superficially bat-like patagia in the forelimbs, unique among dinosaurs. The exact extent

658-643: A profound and permanent effect on the English language. Subsequently, the House of Plantagenet from Anjou inherited the English throne under Henry II , adding England to the budding Angevin Empire of fiefs the family had inherited in France including Aquitaine . They reigned for three centuries, some noted monarchs being Richard I , Edward I , Edward III and Henry V . The period saw changes in trade and legislation, including

752-777: A separate sovereign state on 1 May 1707, when the Acts of Union brought into effect a political union with the Kingdom of Scotland that created the Kingdom of Great Britain . England is the origin of the English language , the English legal system (which served as the basis for the common law systems of many other countries), association football , and the Anglican branch of Christianity; its parliamentary system of government has been widely adopted by other nations. The Industrial Revolution began in 18th-century England, transforming its society into

846-446: A struggle for hegemony between Northumbria and Mercia, which in the 8th century gave way to Mercian preeminence. In the early 9th century Mercia was displaced as the foremost kingdom by Wessex. Later in that century escalating attacks by the Danes culminated in the conquest of the north and east of England, overthrowing the kingdoms of Northumbria, Mercia and East Anglia. Wessex under Alfred

940-403: Is a membranous body part that assists an animal in obtaining lift when gliding or flying . The structure is found in extant and extinct groups of flying and gliding animals including bats , theropod dinosaurs (including birds and some dromaeosaurs ), pterosaurs , gliding mammals , some flying lizards , and flying frogs . The patagium that stretches between an animal's hind limbs

1034-567: Is a non-departmental public body, established in 1995 and sponsored by the Department for Environment, Food and Rural Affairs with responsibilities relating to the protection and enhancement of the environment in England. The Secretary of State for Environment, Food and Rural Affairs is the minister responsible for environmental protection, agriculture, fisheries and rural communities in England. Propatagium The patagium ( pl. : patagia )

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1128-412: Is also applied to England in a more poetic capacity, though its original meaning is the island of Britain as a whole. The earliest known evidence of human presence in the area now known as England was that of Homo antecessor , dating to about 780,000 years ago. The oldest proto-human bones discovered in England date from 500,000 years ago. Modern humans are known to have inhabited the area during

1222-429: Is called the uropatagium (especially in bats) or the interfemoral membrane . In bats, the skin forming the surface of the wing is an extension of the skin of the abdomen that runs to the tip of each digit, uniting the forelimb with the body. The patagium of a bat has four distinct parts: In the flying pterosaurs, the patagium was composed of the membrane forming the surface of the wing, supported primarily between

1316-485: Is damp relatively frequently and is changeable. The coldest months are January and February, the latter particularly on the English coast , while July is normally the warmest month. Months with mild to warm weather are May, June, September and October. Rainfall is spread fairly evenly throughout the year. Important influences on the climate of England are its proximity to the Atlantic Ocean , its northern latitude and

1410-622: Is governed directly by the Parliament of the United Kingdom , although other countries of the United Kingdom have devolved governments. There has been debate about how to counterbalance this in England. Originally it was planned that various regions of England would be devolved, but following the proposal's rejection by the North East in a 2004 referendum , this has not been carried out. In 2024, an England-only intergovernmental body, known as

1504-640: Is high moorland in upland areas, indented by fertile valleys of the region's rivers. They contain two national parks , the Yorkshire Dales and the Peak District . In the West Country , Dartmoor and Exmoor of the Southwest Peninsula include upland moorland supported by granite. The English Lowlands are in the central and southern regions of the country, consisting of green rolling hills, including

1598-758: Is known as the lost colony after it was found abandoned on the return of the late-arriving supply ship. With the East India Company , England also competed with the Dutch and French in the East. During the Elizabethan period, England was at war with Spain. An armada sailed from Spain in 1588 as part of a wider plan to invade England and re-establish a Catholic monarchy. The plan was thwarted by bad coordination, stormy weather and successful harrying attacks by an English fleet under Lord Howard of Effingham . This failure did not end

1692-603: Is mostly found in the north and west, including Dartmoor , the Lake District , the Pennines , and the Shropshire Hills . The country's capital is London , the metropolitan area of which has a population of 14.2 million as of 2021, representing the United Kingdom's largest metropolitan area. England's population of 56.3 million comprises 84% of the population of the United Kingdom, largely concentrated around London,

1786-454: Is part of the United Kingdom, a constitutional monarchy with a parliamentary system . There has not been a government of England since 1707, when the Acts of Union 1707 , putting into effect the terms of the Treaty of Union , joined England and Scotland to form the Kingdom of Great Britain . Before the union England was ruled by its monarch and the Parliament of England . Today England

1880-465: Is similar to that of other areas in the British Isles with a wide range of vertebrate and invertebrate life in a diverse range of habitats. National nature reserves in England are designated by Natural England as key places for wildlife and natural features in England. They were established to protect the most significant areas of habitat and of geological formations. NNRs are managed on behalf of

1974-509: The 1535–1542 acts . There were internal religious conflicts during the reigns of Henry's daughters, Mary I and Elizabeth I . The former took the country back to Catholicism while the latter broke from it again, forcefully asserting the supremacy of Anglicanism . The Elizabethan era is the epoch in the Tudor age of the reign of Queen Elizabeth I ("the Virgin Queen"). Historians often depict it as

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2068-624: The Age of Discovery . Henry VIII broke from communion with the Catholic Church, over issues relating to his divorce, under the Acts of Supremacy in 1534 which proclaimed the monarch head of the Church of England . In contrast with much of European Protestantism , the roots of the split were more political than theological. He also legally incorporated his ancestral land Wales into the Kingdom of England with

2162-568: The Angeln region of what is now the German state of Schleswig-Holstein . The earliest recorded use of the term, as " Engla londe ", is in the late-ninth-century translation into Old English of Bede 's Ecclesiastical History of the English People . The term was then used to mean "the land inhabited by the English", and it included English people in what is now south-east Scotland but was then part of

2256-574: The Battle of Trafalgar confirmed the naval supremacy Britain had established during the course of the eighteenth century. The Napoleonic Wars fostered a concept of Britishness and a united national British people , shared with the English, Scots and Welsh. London became the largest and most populous metropolitan area in the world during the Victorian era , and trade within the British Empire – as well as

2350-532: The Battle of Watling Street . The author of one study of Roman Britain suggested that from 43 AD to 84 AD, the Roman invaders killed somewhere between 100,000 and 250,000 people from a population of perhaps 2,000,000. This era saw a Greco-Roman culture prevail with the introduction of Roman law , Roman architecture , aqueducts , sewers , many agricultural items and silk. In the 3rd century, Emperor Septimius Severus died at Eboracum (now York ), where Constantine

2444-648: The British Isles , but also from the Commonwealth , particularly the Indian subcontinent . Since the 1970s there has been a large move away from manufacturing and an increasing emphasis on the service industry . As part of the United Kingdom, the area joined a common market initiative called the European Economic Community which became the European Union . Since the late 20th century the administration of

2538-596: The Cotswold Hills , Chiltern Hills , North and South Downs ; where they meet the sea they form white rock exposures such as the cliffs of Dover . This also includes relatively flat plains such as the Salisbury Plain , Somerset Levels , South Coast Plain and The Fens . England has a temperate maritime climate : it is mild with temperatures not much lower than 0 °C (32 °F) in winter and not much higher than 32 °C (90 °F) in summer. The weather

2632-712: The Court of Appeal , the High Court of Justice for civil cases, and the Crown Court for criminal cases. The Supreme Court of the United Kingdom is the highest court for criminal and civil cases in England and Wales . It was created in 2009 after constitutional changes, taking over the judicial functions of the House of Lords . A decision of the Supreme Court is binding on every other court in

2726-567: The Courts of England and Wales continued, under the Treaty of Union , as a separate legal system from the one used in Scotland . The general essence of English law is that it is made by judges sitting in courts, applying their common sense and knowledge of legal precedent – stare decisis – to the facts before them. The court system is headed by the Senior Courts of England and Wales, consisting of

2820-705: The French Revolution , under George III and William Pitt the Younger . The regency of George IV is noted for its elegance and achievements in the fine arts and architecture. During the Napoleonic Wars , Napoleon planned to invade from the south-east ; however, this failed to manifest and the Napoleonic forces were defeated by the British: at sea by Horatio Nelson , and on land by Arthur Wellesley . The major victory at

2914-575: The Hungarian aristocrat and paleontologist Franz Nopcsa , who, while residing at London, had seen Pycraft's restoration, drew his own vision of the animal, a picture and article published in a 1907 issue of the Proceedings of the Zoological Society of London (an English science publication nowadays known as Journal of Zoology ). But this time, for his own "Pro-Avis" restoration, Nopcsa suggested

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3008-700: The Labour Party , 116 from the Conservative Party , 65 from the Liberal Democrats , five for Reform UK and four for the Green Party of England and Wales . The English law legal system, developed over the centuries, is the basis of common law legal systems used in most Commonwealth countries and the United States (except Louisiana ). Despite now being part of the United Kingdom, the legal system of

3102-550: The Latin word Anglii is used. The etymology of the tribal name itself is disputed by scholars; it has been suggested that it derives from the shape of the Angeln peninsula, an angular shape. How and why a term derived from the name of this tribe, rather than others such as the Saxons , came to be used for the entire country is not known, but it seems this is related to the custom of calling

3196-692: The Mayoral Council for England , was established to bring together ministers from the UK Government , the Mayor of London and the leaders of combined authorities . In the House of Commons which is the lower house of the British Parliament based at the Palace of Westminster , there are 543 members of parliament (MPs) for constituencies in England, out of the 650 total. England is represented by 347 MPs from

3290-488: The Ministry of Justice , manages most prisons, housing 81,309 prisoners in England and Wales as of September 2022 . The subdivisions of England consist of up to four levels of subnational division , controlled through a variety of types of administrative entities created for the purposes of local government . Outside the London region, England's highest tier is the 48 ceremonial counties . These are used primarily as

3384-672: The Severn is the longest river flowing through England. It empties into the Bristol Channel and is notable for its Severn Bore (a tidal bore ), which can reach 2 metres (6.6 ft) in height. However, the longest river entirely in England is the Thames, which is 215 miles (346 km) in length. There are many lakes in England ; the largest is Windermere , within the aptly named Lake District . Most of England's landscape consists of low hills and plains, with upland and mountainous terrain in

3478-836: The South East , and conurbations in the Midlands , the North West , the North East , and Yorkshire , which each developed as major industrial regions during the 19th century. The name "England" is derived from the Old English name Englaland , which means "land of the Angles ". The Angles were one of the Germanic tribes that settled in Great Britain during the Early Middle Ages . They came from

3572-691: The State Opening of Parliament by the British monarch when the doors of the House of Commons are slammed in the face of the monarch's messenger, symbolising the rights of Parliament and its independence from the monarch. With the founding of the Royal Society in 1660, science was greatly encouraged. In 1666 the Great Fire of London gutted the city of London, but it was rebuilt shortly afterward with many significant buildings designed by Sir Christopher Wren . By

3666-451: The Stuart dynasty England expanded in trade, finance and prosperity. The Royal Navy developed Europe's largest merchant fleet. After the parliaments of England and Scotland agreed, the two countries joined in political union , to create the Kingdom of Great Britain in 1707. To accommodate the union, institutions such as the law and national churches of each remained separate. Under the newly formed Kingdom of Great Britain, output from

3760-433: The Trinovantes . The Romans invaded Britain in 43 AD during the reign of Emperor Claudius , subsequently conquering much of Britain , and the area was incorporated into the Roman Empire as Britannia province . The best-known of the native tribes who attempted to resist were the Catuvellauni led by Caratacus . Later, an uprising led by Boudica , Queen of the Iceni , ended with Boudica's suicide following her defeat at

3854-419: The Upper Paleolithic period, though permanent settlements were only established within the last 6,000 years. After the last ice age only large mammals such as mammoths , bison and woolly rhinoceros remained. Roughly 11,000 years ago, when the ice sheets began to recede, humans repopulated the area; genetic research suggests they came from the northern part of the Iberian Peninsula . The sea level

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3948-470: The canal age in Britain . In 1825 the world's first permanent steam locomotive-hauled passenger railway – the Stockton and Darlington Railway – opened to the public. During the Industrial Revolution , many workers moved from England's countryside to new and expanding urban industrial areas to work in factories, for instance at Birmingham and Manchester , with the latter the world's first industrial city. England maintained relative stability throughout

4042-435: The decline of the Roman Empire , Britain was left exposed by the end of Roman rule in Britain and the withdrawal of Roman army units, to defend the frontiers in continental Europe and partake in civil wars. Celtic Christian monastic and missionary movements flourished. This period of Christianity was influenced by ancient Celtic culture in its sensibilities, polity, practices and theology. Local "congregations" were centred in

4136-434: The golden age in English history that represented the apogee of the English Renaissance and saw the flowering of great art, drama, poetry, music and literature. England during this period had a centralised, well-organised, and effective government. Competing with Spain , the first English colony in the Americas was founded in 1585 by explorer Walter Raleigh in Virginia and named Roanoke . The Roanoke colony failed and

4230-473: The jet engine led to wider air travel . Residential patterns were altered in England by private motoring, and by the creation of the National Health Service in 1948, providing publicly funded health care to all permanent residents free at the point of need. Combined, these prompted the reform of local government in England in the mid-20th century. Since the 20th century, there has been significant population movement to England, mostly from other parts of

4324-526: The "thecodont" hypothesis about the origins of birds. These "proaves" were supposed to be arboreal, with short necks, long trunks, sprawling legs, non-elongated hindlimbs, small hands and feet, small general size, and gliding on membranes. Feduccia's "protoavis" had membranes on the forelimbs only and was fashioned on the model of Megalancosaurus . The term Proavis was used again in 1999, when Joseph Garner, Graham Taylor, and Adrian Thomas proposed their "pouncing proavis model", an hypothesis entailing that

4418-426: The 32 London Boroughs and the 25 Wards of the City of London . Geographically, England includes the central and southern two-thirds of the island of Great Britain, plus such offshore islands as the Isle of Wight and the Isles of Scilly . It is bordered by two other countries of the United Kingdom: to the north by Scotland and to the west by Wales. England is closer than any other part of mainland Britain to

4512-516: The 6th century. Contemporary texts describing this period are extremely scarce, giving rise to its description as a Dark Age . Details of the Anglo-Saxon settlement of Britain are consequently subject to considerable disagreement; the emerging consensus is that it occurred on a large scale in the south and east but was less substantial to the north and west, where Celtic languages continued to be spoken even in areas under Anglo-Saxon control. Roman-dominated Christianity had, in general, been replaced in

4606-448: The English kingdom of Northumbria . The Anglo-Saxon Chronicle recorded that the Domesday Book of 1086 covered the whole of England, meaning the English kingdom, but a few years later the Chronicle stated that King Malcolm III went "out of Scotlande into Lothian in Englaland", thus using it in the more ancient sense. The earliest attested reference to the Angles occurs in the 1st-century work by Tacitus , Germania , in which

4700-416: The European continent. It is separated from France ( Hauts-de-France ) by a 21-mile (34 km) sea gap, though the two countries are connected by the Channel Tunnel near Folkestone . England also has shores on the Irish Sea , North Sea and Atlantic Ocean. The ports of London, Liverpool , and Newcastle lie on the tidal rivers Thames , Mersey and Tyne respectively. At 220 miles (350 km),

4794-466: The Germanic people in Britain Angli Saxones or English Saxons to distinguish them from continental Saxons (Eald-Seaxe) of Old Saxony in Germany. In Scottish Gaelic , the Saxon tribe gave their name to the word for England ( Sasunn ); similarly, the Welsh name for the English language is Saesneg . A romantic name for England is Loegria , related to the Welsh word for England, Lloegr , and made popular by its use in Arthurian legend . Albion

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4888-403: The Great was left as the only surviving English kingdom, and under his successors, it steadily expanded at the expense of the kingdoms of the Danelaw . This brought about the political unification of England, first accomplished under Æthelstan in 927 and definitively established after further conflicts by Eadred in 953. A fresh wave of Scandinavian attacks from the late 10th century ended with

4982-452: The Italian paleontologist Andrea Cau remarked it bore an uncanny resemblance to Paul's "proavis". England – in Europe  (green & dark grey) – in the United Kingdom  (green) England is a country that is part of the United Kingdom . It is located on the island of Great Britain , of which it covers about 62%, and more than 100 smaller adjacent islands . It has land borders with Scotland to

5076-439: The North, and the successful Norman Conquest in October 1066, accomplished by an army led by Duke William of Normandy invading at Hastings late September 1066. The Normans themselves originated from Scandinavia and had settled in Normandy in the late 9th and early 10th centuries. This conquest led to the almost total dispossession of the English elite and its replacement by a new French-speaking aristocracy, whose speech had

5170-408: The Pope. During the 14th century, the Plantagenets and the House of Valois claimed to be legitimate claimants to the House of Capet and of France; the two powers clashed in the Hundred Years' War . The Black Death epidemic hit England ; starting in 1348, it eventually killed up to half of England's inhabitants . Between 1453 and 1487, a civil war known as the War of the Roses waged between

5264-555: The Pro-Aves (in page 39). In the following years, as of 1913, Danish artist and amateur zoologist Gerhard Heilmann also used and popularised the term Proavis , this time presenting the public with more accurate and anatomically probable drawings. Heilmann favoured a scientific model in which the assumed Proavis was arboreal and of thecodontian descent, thus not a dinosaur. In 1961, Czech paleontologist Josef Augusta included "Proavis" in his book Prehistoric Birds and Reptiles . The animal illustration, as were all illustrations in

5358-685: The Royal Society and other English initiatives combined with the Scottish Enlightenment to create innovations in science and engineering, while the enormous growth in British overseas trade protected by the Royal Navy paved the way for the establishment of the British Empire . Domestically it drove the Industrial Revolution , a period of profound change in the socioeconomic and cultural conditions of England, resulting in industrialised agriculture, manufacture, engineering and mining, as well as new and pioneering road, rail and water networks to facilitate their expansion and development. The opening of Northwest England's Bridgewater Canal in 1761 ushered in

5452-432: The United Kingdom has moved towards devolved governance in Scotland, Wales and Northern Ireland. England and Wales continues to exist as a jurisdiction within the United Kingdom. Devolution has stimulated a greater emphasis on a more English-specific identity and patriotism. There is no devolved English government, but an attempt to create a similar system on a sub-regional basis was rejected by referendum . England

5546-443: The United Kingdom as part of the Allies . Two decades later, in World War II , the United Kingdom was again one of the Allies . Developments in warfare technology saw many cities damaged by air-raids during the Blitz . Following the war, the British Empire experienced rapid decolonisation , and there was a speeding-up of technological innovations; automobiles became the primary means of transport and Frank Whittle 's development of

5640-413: The area, the Beaker culture people made bronze , and later iron from iron ores. The development of iron smelting allowed the construction of better ploughs , advancing agriculture (for instance, with Celtic fields ), as well as the production of more effective weapons. During the Iron Age , Celtic culture , deriving from the Hallstatt and La Tène cultures , arrived from Central Europe. Brythonic

5734-505: The auspices of James VI and I the Authorised King James Version of the Holy Bible was published in 1611. It was the standard version of the Bible read by most Protestant Christians for four hundred years until modern revisions were produced in the 20th century. Based on conflicting political, religious and social positions, the English Civil War was fought between the supporters of Parliament and those of King Charles I , known colloquially as Roundheads and Cavaliers respectively. This

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5828-510: The body and the elongated fourth finger. The patagium of a pterosaur had three distinct parts: Flying squirrels , sugar gliders , colugos , anomalures and other mammals also have patagia that extend between the limbs; as in bats and pterosaurs, they also possess propatagia and uropatagia. Though the forelimb is not as specialised as in true flyers, the membrane tends to be an equally complex organ, composed of various muscle groups and fibers. Various species have styliform bones to support

5922-507: The book, was painted by Zdeněk Burian . During the 1970s the work of John Ostrom , breathing new life into the hypothesis that birds were dinosaurs, revived the interest in the direct ancestors of birds. Ostrom assumed these were cursorial. An illustration of a running dinosaur explicitly indicated as a "Proavis" was in 1976 published by Dieter Stefan Peters . In 1991 and 1996 "proavis" or "protoavis" models were proposed by respectively Samuel Tarsitano and Alan Feduccia , both adherents of

6016-479: The conquered territories by Anglo-Saxon paganism , but was reintroduced by missionaries from Rome led by Augustine from 597. Disputes between the Roman- and Celtic-dominated forms of Christianity ended in victory for the Roman tradition at the Council of Whitby (664), which was ostensibly about tonsures (clerical haircuts) and the date of Easter, but more significantly, about the differences in Roman and Celtic forms of authority, theology, and practice. During

6110-427: The conquest of this united kingdom by Sweyn Forkbeard in 1013 and again by his son Cnut in 1016, turning it into the centre of a short-lived North Sea Empire that also included Denmark and Norway . However, the native royal dynasty was restored with the accession of Edward the Confessor in 1042. A dispute over the succession to Edward led to an unsuccessful Norwegian Invasion in September 1066 close to York in

6204-399: The course of the fifth and sixth centuries, initially in the eastern part of the country. Their advance was contained for some decades after the Britons' victory at the Battle of Mount Badon , but subsequently resumed, overrunning the fertile lowlands of Britain and reducing the area under Brittonic control to a series of separate enclaves in the more rugged country to the west by the end of

6298-460: The early 2010s England was divided for a few purposes into regions; a 1998 referendum for the London Region created the London Assembly two years later. A failed 2004 North East England devolution referendum cancelled further regional assembly devolution with the regional structure outside London abolished. Ceremonially and administratively, the region is divided between the City of London and Greater London ; these are further divided into

6392-474: The hierarchy, which must follow its directions. The Secretary of State for Justice is the minister responsible to Parliament for the judiciary , the court system and prisons and probation in England. Crime increased between 1981 and 1995 but fell by 42% in the period 1995–2006. The prison population doubled over the same period, giving it one of the highest incarceration rates in Western Europe at 147 per 100,000. His Majesty's Prison Service , reporting to

6486-442: The hindlimbs still had largely cursorial adaptations whereas the arms had been elongated in order to climb. Feathers, originally serving the insulation of an already warm-blooded animal, would by elongation have turned the arms into wings in order to fly. More generally, the proavians would, in view of their basal theropod forebears and bird descendants, have been typified by long necks, a short trunk, long fingers with opposable digits,

6580-405: The leading edge of the inner wing. Many authors use the term to describe the fold of skin between the body (behind the shoulder) and the elbow that houses the outer segments of the latissimus dorsi caudalis and triceps scapularis muscles. Similarly the fleshy pad that houses the follicles of the remiges (primary and secondary feathers) caudal to the hand and the ulna is also often referred to as

6674-426: The membranes, either on the elbow ( colugos , anomalures , greater glider , Eomys ) or on the wrist (flying squirrels). In gliding species, such as some lizards and flying frogs , the patagium is the flat parachute -like extension of skin that catches the air, which allows gliding flight . In birds , the propatagium is the elastic fold of skin extending from the shoulder to the carpal joint , making up

6768-453: The metropolitan counties have different types of status to other ceremonial counties. The second tier is made up of combined authorities and the 27 county-tier shire counties . In 1974, all ceremonial counties were two-tier; and with the metropolitan county tier phased out, the 1996 reform separated the ceremonial county and the administrative county tier. England is also divided into local government districts. The district can align to

6862-680: The mid-to-late 17th century, two political factions had emerged – the Tories and Whigs . Though the Tories initially supported Catholic king James II , some of them, along with the Whigs, during the Revolution of 1688 invited the Dutch Prince William of Orange to defeat James and become the king. Some English people, especially in the north, were Jacobites and continued to support James and his sons. Under

6956-558: The monastic community and monastic leaders were more like chieftains, as peers, rather than in the more hierarchical system of the Roman-dominated church. Roman military withdrawals left Britain open to invasion by pagan, seafaring warriors from north-western continental Europe, chiefly the Saxons, Angles , Jutes and Frisians who had long raided the coasts of the Roman province. These groups then began to settle in increasing numbers over

7050-536: The nation, many by Natural England themselves, but also by non-governmental organisations, including the members of The Wildlife Trusts partnership, the National Trust , and the Royal Society for the Protection of Birds . There are 221 NNRs in England covering 110,000 hectares (1,100 square kilometres). Often they contain rare species or nationally important populations of plants and animals. . The Environment Agency

7144-621: The north and Wales to the west, and is otherwise surrounded by the North Sea to the east, the English Channel to the south, the Celtic Sea to the south-west, and the Irish Sea to the west. Continental Europe lies to the south-east, and Ireland to the west. At the 2021 census , the population was 56,490,048. London is both the largest city and the capital . The area now called England

7238-594: The north and west of the country. The northern uplands include the Pennines , a chain of uplands dividing east and west, the Lake District mountains in Cumbria, and the Cheviot Hills , straddling the border between England and Scotland. The highest point in England, at 978 metres (3,209 ft), is Scafell Pike in the Lake District. The Shropshire Hills are near Wales while Dartmoor and Exmoor are two upland areas in

7332-592: The origin of avian flight rested in a predatory behaviour, the "proavis" jumping on its prey from trees (see pouncing proavis model in Origin of avian flight ). In his 2002 book Dinosaurs of the Air , Gregory S. Paul tried to conceptually model a "pro-avian". In his view the direct ancestors of birds cannot have been completely arboreal, because in that case they would probably have used membranes to fly. He thought they must have represented an intermediate ecological stage, in which

7426-450: The settlement period the lands ruled by the incomers seem to have been fragmented into numerous tribal territories, but by the 7th century, when substantial evidence of the situation again becomes available, these had coalesced into roughly a dozen kingdoms including Northumbria , Mercia , Wessex , East Anglia , Essex , Kent and Sussex . Over the following centuries, this process of political consolidation continued. The 7th century saw

7520-472: The signing of Magna Carta , an English legal charter used to limit the sovereign's powers by law and protect the privileges of freemen. Catholic monasticism flourished, providing philosophers, and the universities of Oxford and Cambridge were founded with royal patronage. The Principality of Wales became a Plantagenet fief during the 13th century and the Lordship of Ireland was given to the English monarchy by

7614-629: The south-west of the country. The approximate dividing line between terrain types is often indicated by the Tees–Exe line . The Pennines, known as the "backbone of England", are the oldest range of mountains in the country, originating from the end of the Paleozoic Era around 300 million years ago. Their geological composition includes, among others, sandstone and limestone , and also coal. There are karst landscapes in calcite areas such as parts of Yorkshire and Derbyshire . The Pennine landscape

7708-487: The standing of the British military and navy – was prestigious. Technologically, this era saw many innovations that proved key to the United Kingdom's power and prosperity. Political agitation at home from radicals such as the Chartists and the suffragettes enabled legislative reform and universal suffrage . Power shifts in east-central Europe led to World War I; hundreds of thousands of English soldiers died fighting for

7802-487: The threat: Spain launched two further armadas, in 1596 and 1597 , but both were driven back by storms. The political structure of the island changed in 1603, when the King of Scots , James VI , a kingdom which had been a long-time rival to English interests, inherited the throne of England as James I, thereby creating a personal union . He styled himself King of Great Britain , although this had no basis in English law. Under

7896-774: The two branches of the royal family, the Yorkists and Lancastrians . Eventually it led to the Yorkists losing the throne entirely to a Welsh noble family the Tudors , a branch of the Lancastrians headed by Henry Tudor who invaded with Welsh and Breton mercenaries, gaining victory at the Battle of Bosworth Field where the Yorkist king Richard III was killed. During the Tudor period , England began to develop naval skills , and exploration intensified in

7990-530: The warming of the sea by the Gulf Stream . Rainfall is higher in the west, and parts of the Lake District receive more rain than anywhere else in the country. Since weather records began, the highest temperature recorded was 40.3 °C (104.5 °F) on 19 July 2022 at Coningsby , Lincolnshire , while the lowest was −26.1 °C (−15.0 °F) on 10 January 1982 in Edgmond , Shropshire. The fauna of England

8084-536: The world's first industrialised nation. England is home to the two oldest universities in the English-speaking world: the University of Oxford , founded in 1096, and the University of Cambridge , founded in 1209. Both universities are ranked among the most prestigious in the world. England's terrain chiefly consists of low hills and plains , especially in the centre and south . Upland and mountainous terrain

8178-456: Was arboreal , as suggested Professor Osborn six years before, in 1900, in an article dealing with the hypothetical common ancestors of dinosaurs and birds. Pycraft assumed that birds had developed as tree-dwelling dinosaurs, gliding on membranes between the limbs and the trunk. These membranes would gradually have been covered by increasingly more elongated scales, which eventually would have evolved into feathers. A year after Pycraft's article,

8272-563: Was an interwoven part of the wider multifaceted Wars of the Three Kingdoms , involving Scotland and Ireland . The Parliamentarians were victorious, Charles I was executed and the kingdom replaced by the Commonwealth . Leader of the Parliament forces, Oliver Cromwell declared himself Lord Protector in 1653; a period of personal rule followed. After Cromwell's death and the resignation of his son Richard as Lord Protector, Charles II

8366-494: Was established with the Bill of Rights in 1689. Among the statutes set down were that the law could only be made by Parliament and could not be suspended by the King, also that the King could not impose taxes or raise an army without the prior approval of Parliament. Also since that time, no British monarch has entered the House of Commons when it is sitting, which is annually commemorated at

8460-487: Was first inhabited by modern humans during the Upper Paleolithic . It takes its name from the Angles , a Germanic tribe who settled during the 5th and 6th centuries. England became a unified state in the 10th century and has had extensive cultural and legal impact on the wider world since the Age of Discovery , which began during the 15th century. The Kingdom of England , which included Wales after 1535, ceased to be

8554-567: Was invited to return as monarch in 1660, in a move called the Restoration . With the reopening of theatres, fine arts, literature and performing arts flourished throughout the Restoration of the "Merry Monarch" Charles II. After the Glorious Revolution of 1688, it was constitutionally established that King and Parliament should rule together, though Parliament would have the real power. This

8648-572: Was lower than the present day and Britain was connected by land bridge to Ireland and Eurasia . As the seas rose, it was separated from Ireland 10,000 years ago and from Eurasia two millennia later. The Beaker culture arrived around 2,500 BC, introducing drinking and food vessels constructed from clay, as well as vessels used as reduction pots to smelt copper ores. It was during this time that major Neolithic monuments such as Stonehenge (phase III) and Avebury were constructed. By heating together tin and copper, which were in abundance in

8742-616: Was subsequently proclaimed emperor a century later. There is debate about when Christianity was first introduced; it was no later than the 4th century, probably much earlier. According to Bede , missionaries were sent from Rome by Eleutherius at the request of the chieftain Lucius of Britain in 180 AD, to settle differences as to Eastern and Western ceremonials, which were disturbing the church. There are traditions linked to Glastonbury claiming an introduction through Joseph of Arimathea , while others claim through Lucius of Britain. By 410, during

8836-506: Was the spoken language during this time. Society was tribal; according to Ptolemy 's Geographia there were around 20 tribes in the area. Like other regions on the edge of the Empire, Britain had long enjoyed trading links with the Romans. Julius Caesar of the Roman Republic attempted to invade twice in 55 BC; although largely unsuccessful, he managed to set up a client king from

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