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71-493: The order Peramelemorphia / p ɛ r ə m ɛ l ɪ ˈ m ɔːr f i ə / includes the bandicoots and bilbies . All members of the order are endemic to Australia - New Guinea and most have the characteristic bandicoot shape: a plump, arch-backed body with a long, delicately tapering snout, very large upright ears, relatively long, thin legs, and a thin tail. Their size varies from about 140 grams up to 4 kilograms, but most species are about one kilogram. The position of

142-456: A cohors (plural cohortes ). Some of the plant families still retain the names of Linnaean "natural orders" or even the names of pre-Linnaean natural groups recognized by Linnaeus as orders in his natural classification (e.g. Palmae or Labiatae ). Such names are known as descriptive family names. In the field of zoology , the Linnaean orders were used more consistently. That is,

213-466: A phylogenetic reconstruction of the members of Peramelemorphia with quite strong support. The most basal split separates Thylacomyidae ( Macrotis ) from all other bandicoots. Probably the next to diverge was the recently extinct Chaeropodidae ( Chaeropus ). The remaining taxa comprise the Peramelidae , which divides into subfamilies Peramelinae ( Isoodon and Perameles ) and a clade in which

284-635: A type section , Global Boundary Stratotype Section and Point (GSSP), for the upper Pleistocene/Holocene boundary ( i.e. the upper boundary). The proposed section is the North Greenland Ice Core Project ice core 75° 06' N 42° 18' W. The lower boundary of the Pleistocene Series is formally defined magnetostratigraphically as the base of the Matuyama (C2r) chronozone , isotopic stage 103. Above this point there are notable extinctions of

355-505: A capital letter. For some groups of organisms, their orders may follow consistent naming schemes . Orders of plants , fungi , and algae use the suffix -ales (e.g. Dictyotales ). Orders of birds and fishes use the Latin suffix -iformes meaning 'having the form of' (e.g. Passeriformes ), but orders of mammals and invertebrates are not so consistent (e.g. Artiodactyla , Actiniaria , Primates ). For some clades covered by

426-585: A change from low-amplitude glacial cycles with a dominant periodicity of 41,000 years to asymmetric high-amplitude cycles dominated by a periodicity of 100,000 years. However, a 2020 study concluded that ice age terminations might have been influenced by obliquity since the Mid-Pleistocene Transition, which caused stronger summers in the Northern Hemisphere . Glaciation in the Pleistocene

497-574: A deviation from today's annual mean temperature, taken as zero. This sort of graph is based on another isotope ratio versus time. Ratios are converted to a percentage difference from the ratio found in standard mean ocean water (SMOW). The graph in either form appears as a waveform with overtones . One half of a period is a Marine isotopic stage (MIS). It indicates a glacial (below zero) or an interglacial (above zero). Overtones are stadials or interstadials. According to this evidence, Earth experienced 102 MIS stages beginning at about 2.588 Ma BP in

568-566: A distinct rank of biological classification having its own distinctive name (and not just called a higher genus ( genus summum )) was first introduced by the German botanist Augustus Quirinus Rivinus in his classification of plants that appeared in a series of treatises in the 1690s. Carl Linnaeus was the first to apply it consistently to the division of all three kingdoms of nature (then minerals , plants , and animals ) in his Systema Naturae (1735, 1st. Ed.). For plants, Linnaeus' orders in

639-421: A few regions had been studied and the names were relatively few. Today the geologists of different nations are taking more of an interest in Pleistocene glaciology. As a consequence, the number of names is expanding rapidly and will continue to expand. Many of the advances and stadials remain unnamed. Also, the terrestrial evidence for some of them has been erased or obscured by larger ones, but evidence remains from

710-480: A fossil bandicoot was reported from two new species from Riversleigh (Travouillon et al. 2014). Named Madju variae and Madju encorensis , they are closely related to modern bandicoots, but do not fall in any modern family, as did Galadi and Bulungu . Instead they are classified as Perameloid, with all known Peremelemorphian, to the exclusion of yaralids. Madju variae is also unusual in preserving an ontogenetic series (age series from pouch young to adult),

781-528: A late Oligocene site from the Northern Territory in Australia. This species is thought to be even more primitive than Yarala burchfieldi . The second fossil skull of any fossil peramelemorphian was also recovered from Miocene sites of Riversleigh. In fact, more than one skull of this new species was found (and several lower and upper jaws), and was significantly different from any other bandicoot to erect

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852-536: A new genus, Galadi . The species was named Galadi speciosus by Travouillon and colleagues. It was short-snouted unlike modern bandicoots suggesting that it was more carnivorous than its omnivorous modern relatives. Its relationship to other bandicoots is unclear, but it was likely to be less primitive than Yarala but more primitive than living bandicoots. An additional three species of Galadi were later described in 2013 and named Galadi grandis , Galadi amplus and Galadi adversus . Gurovich et al. (2013) described

923-477: A new species Perameles , Perameles sobbei , from the Darling Downs (Pleistocene in age), south-eastern Queensland. This species was represented by a lower jaw and a few isolated lower molars. Additional material were later described in 2005 from the same site, including upper molars. A second species of Yarala , Yarala kida , was described in 2006 by Schwartz. This species was recovered from Kangaroo Well,

994-597: A new species of mouse-sized bandicoot from Riversleigh and from Kutjamarpu, Southern Australia. The species, named Bulungu palara , is represented by a skull and several lower and upper jaws. Two other species in this genus were also described from the Etadunna Formation in South Australia, Bulungu muirheadae which was the oldest fossil bandicoot recovered as of 2013 (about 24 million years old), and Bulungu campbelli . The oldest modern bandicoot (peramelid) and

1065-457: A primitive carnivore, developed the syndactylous hind foot as a specialisation for climbing, and the diprotodonts then split off and evolved the two-tooth jaw that gives them their name. Recent molecular level investigations do not so far appear to have resolved the puzzle, but do strongly suggest that whatever the relationship of the bandicoot group to the other marsupial orders may be, it is a distant one. Recent molecular analyses have resulted in

1136-568: A team of Russian scientists in collaboration with Princeton University announced that they had brought two female nematodes frozen in permafrost , from around 42,000 years ago, back to life. The two nematodes, at the time, were the oldest confirmed living animals on the planet. The evolution of anatomically modern humans took place during the Pleistocene. At the beginning of the Pleistocene Paranthropus species were still present, as well as early human ancestors, but during

1207-451: Is a combination of Ancient Greek πλεῖστος ( pleîstos ) 'most' and καινός ( kainós ; Latinized as cænus ) 'new'. At the end of the preceding Pliocene, the previously isolated North and South American continents were joined by the Isthmus of Panama , causing a faunal interchange between the two regions and changing ocean circulation patterns, with the onset of glaciation in

1278-419: Is determined by a taxonomist , as is whether a particular order should be recognized at all. Often there is no exact agreement, with different taxonomists each taking a different position. There are no hard rules that a taxonomist needs to follow in describing or recognizing an order. Some taxa are accepted almost universally, while others are recognized only rarely. The name of an order is usually written with

1349-765: Is estimated that, at maximum glacial extent, 30% of the Earth's surface was covered by ice. In addition, a zone of permafrost stretched southward from the edge of the glacial sheet, a few hundred kilometres in North America , and several hundred in Eurasia . The mean annual temperature at the edge of the ice was −6 °C (21 °F); at the edge of the permafrost, 0 °C (32 °F). Each glacial advance tied up huge volumes of water in continental ice sheets 1,500 to 3,000 metres (4,900–9,800 ft) thick, resulting in temporary sea-level drops of 100 metres (300 ft) or more over

1420-454: Is no systematic correspondence between pluvials to glacials, however. Moreover, regional pluvials do not correspond to each other globally. For example, some have used the term "Riss pluvial" in Egyptian contexts. Any coincidence is an accident of regional factors. Only a few of the names for pluvials in restricted regions have been stratigraphically defined. The sum of transient factors acting at

1491-638: The Alpine ice sheet on the Alps . Scattered domes stretched across Siberia and the Arctic shelf. The northern seas were ice-covered. South of the ice sheets large lakes accumulated because outlets were blocked and the cooler air slowed evaporation. When the Laurentide Ice Sheet retreated, north-central North America was completely covered by Lake Agassiz . Over a hundred basins, now dry or nearly so, were overflowing in

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1562-644: The Echymiperinae ( Echymipera and Microperoryctes ) form a sister group to Peroryctinae ( Peroryctes ): Macrotis Chaeropus Isoodon Perameles Echymipera Microperoryctes Peroryctes Macrotis Rhynchomeles Isoodon Perameles Peroryctes Echymipera Microperoryctes Macrotis Isoodon Perameles Peroryctes Echymipera [incl. Rhynchomeles ] Microperoryctes Many specimens of modern peramelemorphian (e.g. Perameles spp. and Isoodon spp.) have been recovered in

1633-520: The International Code of Zoological Nomenclature , several additional classifications are sometimes used, although not all of these are officially recognized. In their 1997 classification of mammals , McKenna and Bell used two extra levels between superorder and order: grandorder and mirorder . Michael Novacek (1986) inserted them at the same position. Michael Benton (2005) inserted them between superorder and magnorder instead. This position

1704-525: The International Union of Geological Sciences , the cutoff of the Pleistocene and the preceding Pliocene was regarded as being 1.806 million years Before Present (BP). Publications from earlier years may use either definition of the period. The end of the Pleistocene corresponds with the end of the last glacial period and also with the end of the Paleolithic age used in archaeology . The name

1775-635: The Ruwenzori Range in east and central Africa were larger. Glaciers existed in the mountains of Ethiopia and to the west in the Atlas Mountains . In the northern hemisphere, many glaciers fused into one. The Cordilleran Ice Sheet covered the North American northwest; the east was covered by the Laurentide . The Fenno-Scandian ice sheet rested on northern Europe , including much of Great Britain;

1846-769: The Systema Naturae and the Species Plantarum were strictly artificial, introduced to subdivide the artificial classes into more comprehensible smaller groups. When the word ordo was first consistently used for natural units of plants, in 19th-century works such as the Prodromus Systematis Naturalis Regni Vegetabilis of Augustin Pyramus de Candolle and the Genera Plantarum of Bentham & Hooker, it indicated taxa that are now given

1917-437: The calcareous nannofossils : Discoaster pentaradiatus and Discoaster surculus . The Pleistocene covers the recent period of repeated glaciations. The name Plio-Pleistocene has, in the past, been used to mean the last ice age. Formerly, the boundary between the two epochs was drawn at the time when the foraminiferal species Hyalinea baltica first appeared in the marine section at La Castella, Calabria, Italy. However,

1988-477: The last glacial period ended about 10,000 years ago. Over 11 major glacial events have been identified, as well as many minor glacial events. A major glacial event is a general glacial excursion, termed a "glacial." Glacials are separated by "interglacials". During a glacial, the glacier experiences minor advances and retreats. The minor excursion is a "stadial"; times between stadials are "interstadials". These events are defined differently in different regions of

2059-588: The woolly rhinoceros , various giraffids , such as the Sivatherium ; ground sloths , Irish elk , cave lions , cave bears , Gomphotheres , American lions , dire wolves , and short-faced bears , began late in the Pleistocene and continued into the Holocene. Neanderthals also became extinct during this period. At the end of the last ice age, cold-blooded animals, smaller mammals like wood mice , migratory birds, and swifter animals like whitetail deer had replaced

2130-640: The "Tarantian"). In addition to these international subdivisions, various regional subdivisions are often used. In 2009 the International Union of Geological Sciences (IUGS) confirmed a change in time period for the Pleistocene, changing the start date from 1.806 to 2.588 million years BP, and accepted the base of the Gelasian as the base of the Pleistocene, namely the base of the Monte San Nicola GSSP . The start date has now been rounded down to 2.580 million years BP. The IUGS has yet to approve

2201-558: The Early Pleistocene Gelasian . Early Pleistocene stages were shallow and frequent. The latest were the most intense and most widely spaced. By convention, stages are numbered from the Holocene, which is MIS1. Glacials receive an even number and interglacials receive an odd number. The first major glacial was MIS2-4 at about 85–11 ka BP. The largest glacials were 2, 6, 12, and 16. The warmest interglacials were 1, 5, 9 and 11. For matching of MIS numbers to named stages, see under

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2272-409: The Earth's surface is cyclical: climate, ocean currents and other movements, wind currents, temperature, etc. The waveform response comes from the underlying cyclical motions of the planet, which eventually drag all the transients into harmony with them. The repeated glaciations of the Pleistocene were caused by the same factors. The Mid-Pleistocene Transition , approximately one million years ago, saw

2343-479: The North American west. Lake Bonneville , for example, stood where Great Salt Lake now does. In Eurasia, large lakes developed as a result of the runoff from the glaciers. Rivers were larger, had a more copious flow, and were braided . African lakes were fuller, apparently from decreased evaporation. Deserts, on the other hand, were drier and more extensive. Rainfall was lower because of the decreases in oceanic and other evaporation. It has been estimated that during

2414-641: The Northern Hemisphere occurring around 2.7 million years ago. During the Early Pleistocene (2.58–0.8 Ma), archaic humans of the genus Homo originated in Africa and spread throughout Afro-Eurasia . The end of the Early Pleistocene is marked by the Mid-Pleistocene Transition , with the cyclicity of glacial cycles changing from 41,000-year cycles to asymmetric 100,000-year cycles, making

2485-476: The Peramelemorphia within the marsupial family tree has long been puzzling and controversial. There are two morphological features in the order that appear to show a clear evolutionary link with another marsupial group: the type of foot, and the teeth. Unfortunately, these clear signposts point in opposite directions. All members of the order are polyprotodont (have several pairs of lower front teeth)—in

2556-490: The Pleistocene's overall climate could be characterised as a continuous El Niño with trade winds in the south Pacific weakening or heading east, warm air rising near Peru , warm water spreading from the west Pacific and the Indian Ocean to the east Pacific, and other El Niño markers. Pleistocene climate was marked by repeated glacial cycles in which continental glaciers pushed to the 40th parallel in some places. It

2627-782: The Pleistocene, the East Antarctic Ice Sheet thinned by at least 500 meters, and that thinning since the Last Glacial Maximum is less than 50 meters and probably started after ca 14 ka. During the 2.5 million years of the Pleistocene, numerous cold phases called glacials ( Quaternary ice age ), or significant advances of continental ice sheets, in Europe and North America, occurred at intervals of approximately 40,000 to 100,000 years. The long glacial periods were separated by more temperate and shorter interglacials which lasted about 10,000–15,000 years. The last cold episode of

2698-411: The Pleistocene, the plates upon which they sit probably having moved no more than 100 km (62 mi) relative to each other since the beginning of the period. In glacial periods, the sea level would drop by up to 120 m (390 ft) lower than today during peak glaciation, exposing large areas of the present continental shelf as dry land. According to Mark Lynas (through collected data),

2769-620: The Pleistocene. The climate was strongly variable depending on the glacial cycle, with the sea levels being up to 120 metres (390 ft) lower than present at peak glaciation, allowing the connection of Asia and North America via Beringia and the covering of most of northern North America by the Laurentide Ice Sheet . Charles Lyell introduced the term "Pleistocene" in 1839 to describe strata in Sicily that had at least 70% of their molluscan fauna still living today. This distinguished it from

2840-525: The Younger Dryas is the official start of the current Holocene Epoch . Although it is considered an epoch, the Holocene is not significantly different from previous interglacial intervals within the Pleistocene. In the ICS timescale, the Pleistocene is divided into four stages or ages , the Gelasian , Calabrian , Chibanian (previously the unofficial "Middle Pleistocene"), and Upper Pleistocene (unofficially

2911-687: The appearance of Homo sapiens about 300,000 years ago. Artifacts associated with modern human behavior are unambiguously attested starting 40,000–50,000 years ago. According to mitochondrial timing techniques, modern humans migrated from Africa after the Riss glaciation in the Middle Palaeolithic during the Eemian Stage , spreading all over the ice-free world during the late Pleistocene. A 2005 study posits that humans in this migration interbred with archaic human forms already outside of Africa by

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2982-515: The articles for those names. Both marine and continental faunas were essentially modern but with many more large land mammals such as Mammoths , Mastodons , Diprotodons , Smilodons , tigers , lions , Aurochs , short-faced bears , giant sloths , species within Gigantopithecus and others. Isolated landmasses such as Australia , Madagascar , New Zealand and islands in the Pacific saw

3053-585: The case of the Peramelemorphia, three pairs. This suggests that they have evolved within Dasyuromorphia (marsupial carnivores). On the other hand, they also have an unusual feature in their feet: the second and third toes are fused together. This condition is called syndactyly , and is characteristic of the Diprotodontia (the order of marsupial herbivores that includes kangaroos , wombats , possums , and many others). Attempts to resolve this puzzle include

3124-527: The climate variation more extreme. The Late Pleistocene witnessed the spread of modern humans outside of Africa as well as the extinction of all other human species. Humans also spread to the Australian continent and the Americas for the first time, co-incident with the extinction of most large-bodied animals in these regions. The aridification and cooling trends of the preceding Neogene were continued in

3195-606: The entire surface of the Earth. During interglacial times, such as at present, drowned coastlines were common, mitigated by isostatic or other emergent motion of some regions. The effects of glaciation were global. Antarctica was ice-bound throughout the Pleistocene as well as the preceding Pliocene. The Andes were covered in the south by the Patagonian ice cap. There were glaciers in New Zealand and Tasmania . The current decaying glaciers of Mount Kenya , Mount Kilimanjaro , and

3266-772: The evolution of large birds and even reptiles such as the Elephant bird , moa , Haast's eagle , Quinkana , Megalania and Meiolania . The severe climatic changes during the Ice Age had major impacts on the fauna and flora. With each advance of the ice, large areas of the continents became depopulated, and plants and animals retreating southwards in front of the advancing glacier faced tremendous stress. The most severe stress resulted from drastic climatic changes, reduced living space, and curtailed food supply. A major extinction event of large mammals ( megafauna ), which included mammoths , mastodons , saber-toothed cats , glyptodons ,

3337-546: The fossil record from Pleistocene and Holocene fossil localities. However, very few fossil species have been recovered to date. The first species of fossil peramelemorphian was described by R. A. Stirton in 1955. The specimen Stirton described was a partial lower jaw from the Tirari Desert in Central Australia, Pliocene in age. The lower jaw morphology suggested a relationship with bilbies (Family Thylacomyidae), and

3408-414: The glacial range, which have their own glacial history depending on latitude, terrain and climate. There is a general correspondence between glacials in different regions. Investigators often interchange the names if the glacial geology of a region is in the process of being defined. However, it is generally incorrect to apply the name of a glacial in one region to another. For most of the 20th century, only

3479-450: The historical terminology was established. Corresponding to the terms glacial and interglacial, the terms pluvial and interpluvial are in use (Latin: pluvia , rain). A pluvial is a warmer period of increased rainfall; an interpluvial is of decreased rainfall. Formerly a pluvial was thought to correspond to a glacial in regions not iced, and in some cases it does. Rainfall is cyclical also. Pluvials and interpluvials are widespread. There

3550-613: The large amounts of material moved about by glaciers. Less common are cave deposits, travertines and volcanic deposits (lavas, ashes). Pleistocene marine deposits are found primarily in shallow marine basins mostly (but with important exceptions) in areas within a few tens of kilometres of the modern shoreline. In a few geologically active areas such as the Southern California coast, Pleistocene marine deposits may be found at elevations of several hundred metres. The modern continents were essentially at their present positions during

3621-466: The lower Palaeolithic they disappeared, and the only hominin species found in fossilic records is Homo erectus for much of the Pleistocene. Acheulean lithics appear along with Homo erectus , some 1.8 million years ago, replacing the more primitive Oldowan industry used by A. garhi and by the earliest species of Homo . The Middle Paleolithic saw more varied speciation within Homo , including

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3692-478: The megafauna and migrated north. Late Pleistocene bighorn sheep were more slender and had longer legs than their descendants today. Scientists believe that the change in predator fauna after the late Pleistocene extinctions resulted in a change of body shape as the species adapted for increased power rather than speed. The extinctions hardly affected Africa but were especially severe in North America where native horses and camels were wiped out. In July 2018,

3763-577: The older Pliocene Epoch , which Lyell had originally thought to be the youngest fossil rock layer. He constructed the name "Pleistocene" ('most new' or 'newest') from the Greek πλεῖστος ( pleīstos ) 'most' and καινός ( kainós ( Latinized as cænus ) 'new'). This contrasts with the immediately preceding Pliocene ("newer", from πλείων ( pleíōn , "more") and kainós ) and the immediately subsequent Holocene ("wholly new" or "entirely new", from ὅλος ( hólos , "whole") and kainós ) epoch , which extends to

3834-546: The oldest bilby (Thylacomyid) were later discovered by Travouillon et al., 2014 from Riversleigh World Heritage Area, from middle Miocene fossil deposits (around 15 million years old). The peramelid, Crash bandicoot , was named after the famous video game character and is only represented by a single upper jaw. The bilby, Liyamayi dayi named after geologist and philanthropist Robert Day, is only known from 3 teeth (2 upper molar, 1 lower molar). The first record of sexual dimorphism (difference in size between males and females) in

3905-698: The orders in the zoology part of the Systema Naturae refer to natural groups. Some of his ordinal names are still in use, e.g. Lepidoptera (moths and butterflies) and Diptera (flies, mosquitoes, midges, and gnats). In virology , the International Committee on Taxonomy of Viruses 's virus classification includes fifteen taxomomic ranks to be applied for viruses , viroids and satellite nucleic acids : realm , subrealm , kingdom , subkingdom, phylum , subphylum , class, subclass, order, suborder, family, subfamily , genus, subgenus , and species. There are currently fourteen viral orders, each ending in

3976-547: The precursor of the currently used International Code of Nomenclature for algae, fungi, and plants . In the first international Rules of botanical nomenclature from the International Botanical Congress of 1905, the word family ( familia ) was assigned to the rank indicated by the French famille , while order ( ordo ) was reserved for a higher rank, for what in the 19th century had often been named

4047-458: The present time. The Pleistocene has been dated from 2.580 million (±0.005) to 11,700 years BP with the end date expressed in radiocarbon years as 10,000 carbon-14 years BP. It covers most of the latest period of repeated glaciation , up to and including the Younger Dryas cold spell. The end of the Younger Dryas has been dated to about 9700 BCE (11,700 calendar years BP). The end of

4118-494: The rank of family (see ordo naturalis , ' natural order '). In French botanical publications, from Michel Adanson 's Familles naturelles des plantes (1763) and until the end of the 19th century, the word famille (plural: familles ) was used as a French equivalent for this Latin ordo . This equivalence was explicitly stated in the Alphonse Pyramus de Candolle 's Lois de la nomenclature botanique (1868),

4189-417: The ratio of O to O (two isotopes of oxygen ) by mass (measured by a mass spectrometer ) present in the calcite of oceanic core samples is used as a diagnostic of ancient ocean temperature change and therefore of climate change. Cold oceans are richer in O , which is included in the tests of the microorganisms ( foraminifera ) contributing the calcite. A more recent version of

4260-564: The revised definition of the Quaternary , by pushing back the start date of the Pleistocene to 2.58 Ma, results in the inclusion of all the recent repeated glaciations within the Pleistocene. Radiocarbon dating is considered to be inaccurate beyond around 50,000 years ago. Marine isotope stages (MIS) derived from Oxygen isotopes are often used for giving approximate dates. Pleistocene non-marine sediments are found primarily in fluvial deposits , lakebeds, slope and loess deposits as well as in

4331-407: The sampling process makes use of modern glacial ice cores. Although less rich in O than seawater, the snow that fell on the glacier year by year nevertheless contained O and O in a ratio that depended on the mean annual temperature. Temperature and climate change are cyclical when plotted on a graph of temperature versus time. Temperature coordinates are given in the form of

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4402-420: The second of any fossil marsupial mammal in Australia. The study of this ontogenetic series lead researchers to think that Madju variae developed slow than modern bandicoots, much more like a bilby, and therefore the rapid development of modern bandicoots must have evolved after the middle Miocene, when Australia started to become more arid. Order (biology) What does and does not belong to each order

4473-632: The sole factor responsible for the variations in climate since they explain neither the long-term cooling trend over the Plio-Pleistocene nor the millennial variations in the Greenland Ice Cores known as Dansgaard-Oeschger events and Heinrich events. Milankovitch pacing seems to best explain glaciation events with periodicity of 100,000, 40,000, and 20,000 years. Such a pattern seems to fit the information on climate change found in oxygen isotope cores. In oxygen isotope ratio analysis, variations in

4544-421: The study of cyclical climate changes. The glacials in the following tables show historical usages, are a simplification of a much more complex cycle of variation in climate and terrain, and are generally no longer used. These names have been abandoned in favour of numeric data because many of the correlations were found to be either inexact or incorrect and more than four major glacials have been recognised since

4615-474: The suffix -virales . Pleistocene The Pleistocene ( / ˈ p l aɪ s t ə ˌ s iː n , - s t oʊ -/ PLY -stə-seen, -⁠stoh- ; referred to colloquially as the Ice Age ) is the geological epoch that lasted from c.  2.58 million to 11,700 years ago, spanning the Earth's most recent period of repeated glaciations . Before a change was finally confirmed in 2009 by

4686-426: The view that the bandicoot group evolved from the carnivores, retaining the polyprotodont dentition, and independently evolving a syndactyl hind foot; the contrary view that syndactyly is so unusual that it is unlikely to have evolved twice and therefore the bandicoot group must have evolved from a possum-like diprotodont creature, and re-evolved its extra teeth. A third view suggests that the bandicoot group evolved from

4757-543: Was a series of glacials and interglacials, stadials and interstadials, mirroring periodic climate changes. The main factor at work in climate cycling is now believed to be Milankovitch cycles . These are periodic variations in regional and planetary solar radiation reaching the Earth caused by several repeating changes in the Earth's motion. The effects of Milankovitch cycles were enhanced by various positive feedbacks related to increases in atmospheric carbon dioxide concentrations and Earth's albedo. Milankovitch cycles cannot be

4828-418: Was adopted by Systema Naturae 2000 and others. In botany , the ranks of subclass and suborder are secondary ranks pre-defined as respectively above and below the rank of order. Any number of further ranks can be used as long as they are clearly defined. The superorder rank is commonly used, with the ending -anae that was initiated by Armen Takhtajan 's publications from 1966 onwards. The order as

4899-398: Was described from Riversleigh , and was named Yarala burchfieldi by Dr Jeannette Muirhead. The species was represented by several upper and lower jaws, which were smaller than any living bandicoots and had a very primitive dentition. A skull was later recovered in 2000, the first for any fossil peramelemorphian to date. Features of the skull and dentition suggested that Yarala burchfieldi

4970-473: Was distinct from other peramelemorphians, and for this reason, a new Superfamily Yaraloidea and Family Yaralidae were erected to classify this species. In 1997, Muirhead, Dawson and Archer described a new species of Perameles , Perameles bowensis , from teeth recovered from two Pliocene fossil localities, Bow and Wellington Caves . The same species was later reported in 2000 from Chinchilla, Queensland , by Mackness and colleagues. In 2002, Price described

5041-457: Was named Ischnodon australis . It was not until 1976 that Archer and Wade described the next fossil bandicoot. A single upper molar was recovered from the Bluff Downs fossil site , Allingham Formation, in northern Queensland , also Pliocene in age. The tooth was similar to that of species of Perameles , and was therefore named Perameles allinghamensis . In 1995, the first Miocene species

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