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147-399: A high water mark is not necessarily an actual physical mark, but it is possible for water rising to a high point to leave a lasting physical impression such as floodwater staining. A landscape marking left by the high water mark of ordinary tidal action may be called a strandline and is typically composed of debris left by high tide. The area at the top of a beach where debris is deposited

294-525: A last universal common ancestor that lived about 3.5 billion years ago . Geologists have developed a geologic time scale that divides the history of the Earth into major divisions, starting with four eons ( Hadean , Archean , Proterozoic , and Phanerozoic ), the first three of which are collectively known as the Precambrian , which lasted approximately 4 billion years. Each eon can be divided into eras, with

441-440: A lipid bilayer , including cholesterols that sit between phospholipids to maintain their fluidity at various temperatures. Cell membranes are semipermeable , allowing small molecules such as oxygen, carbon dioxide, and water to pass through while restricting the movement of larger molecules and charged particles such as ions . Cell membranes also contain membrane proteins , including integral membrane proteins that go across

588-443: A nucleus , and prokaryotic cells, which do not. Prokaryotes are single-celled organisms such as bacteria , whereas eukaryotes can be single-celled or multicellular. In multicellular organisms , every cell in the organism's body is derived ultimately from a single cell in a fertilized egg . Every cell is enclosed within a cell membrane that separates its cytoplasm from the extracellular space . A cell membrane consists of

735-504: A number of shapes , ranging from spheres to rods and spirals . Bacteria were among the first life forms to appear on Earth, and are present in most of its habitats . Bacteria inhabit soil, water, acidic hot springs , radioactive waste , and the deep biosphere of the Earth's crust . Bacteria also live in symbiotic and parasitic relationships with plants and animals. Most bacteria have not been characterised, and only about 27 percent of

882-423: A beach depends on whether the waves are constructive or destructive, and whether the material is sand or shingle. Waves are constructive if the period between their wave crests is long enough for the breaking water to recede and the sediment to settle before the succeeding wave arrives and breaks. Fine sediment transported from lower down the beach profile will compact if the receding water percolates or soaks into

1029-409: A beach tends to indicate the energy of the waves and wind in the locality. Constructive waves move material up the beach while destructive waves move the material down the beach. During seasons when destructive waves are prevalent, the shallows will carry an increased load of sediment and organic matter in suspension. On sandy beaches, the turbulent backwash of destructive waves removes material forming

1176-471: A cell's size, shape, membrane potential , metabolic activity , and responsiveness to signals, which are largely due to highly controlled modifications in gene expression and epigenetics . With a few exceptions, cellular differentiation almost never involves a change in the DNA sequence itself. Thus, different cells can have very different physical characteristics despite having the same genome . Morphogenesis, or

1323-408: A chemical (e.g., nitrous acid , benzopyrene ) or radiation (e.g., x-ray , gamma ray , ultraviolet radiation , particles emitted by unstable isotopes). Mutations can lead to phenotypic effects such as loss-of-function, gain-of-function , and conditional mutations. Some mutations are beneficial, as they are a source of genetic variation for evolution. Others are harmful if they were to result in

1470-1227: A few archaea have very different shapes, such as the flat and square cells of Haloquadratum walsbyi . Despite this morphological similarity to bacteria, archaea possess genes and several metabolic pathways that are more closely related to those of eukaryotes, notably for the enzymes involved in transcription and translation . Other aspects of archaeal biochemistry are unique, such as their reliance on ether lipids in their cell membranes , including archaeols . Archaea use more energy sources than eukaryotes: these range from organic compounds , such as sugars, to ammonia , metal ions or even hydrogen gas . Salt-tolerant archaea (the Haloarchaea ) use sunlight as an energy source, and other species of archaea fix carbon , but unlike plants and cyanobacteria , no known species of archaea does both. Archaea reproduce asexually by binary fission , fragmentation , or budding ; unlike bacteria, no known species of Archaea form endospores . The first observed archaea were extremophiles , living in extreme environments, such as hot springs and salt lakes with no other organisms. Improved molecular detection tools led to

1617-463: A filter for runoff from the coastal plain. If the runoff is naturally dispersed along the beach, water borne silt and organic matter will be retained on the land and will feed the flora in the coastal area. Runoff that is dispersed along the beach will tend to percolate through the beach and may emerge from the beach at low tide. The retention of the freshwater may also help to maintain underground water reserves and will resist salt water incursion. If

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1764-403: A final electron acceptor, which is usually the oxidized form of NADP , which is reduced to NADPH, a process that takes place in a protein complex called photosystem I (PSI). The transport of electrons is coupled to the movement of protons (or hydrogen) from the stroma to the thylakoid membrane, which forms a pH gradient across the membrane as hydrogen becomes more concentrated in the lumen than in

1911-401: A gently sloping beach. On pebble and shingle beaches the swash is dissipated more quickly because the large particle size allows greater percolation , thereby reducing the power of the backwash, and the beach remains steep. Compacted fine sediments will form a smooth beach surface that resists wind and water erosion. During hot calm seasons, a crust may form on the surface of ocean beaches as

2058-492: A lesser root ball. Erosion of beaches can expose less resilient soils and rocks to wind and wave action leading to undermining of coastal headlands eventually resulting in catastrophic collapse of large quantities of overburden into the shallows. This material may be distributed along the beach front leading to a change in the habitat as sea grasses and corals in the shallows may be buried or deprived of light and nutrients. Coastal areas settled by man inevitably become subject to

2205-759: A loss of function of genes needed for survival. Gene expression is the molecular process by which a genotype encoded in DNA gives rise to an observable phenotype in the proteins of an organism's body. This process is summarized by the central dogma of molecular biology , which was formulated by Francis Crick in 1958. According to the Central Dogma, genetic information flows from DNA to RNA to protein. There are two gene expression processes: transcription (DNA to RNA) and translation (RNA to protein). The regulation of gene expression by environmental factors and during different stages of development can occur at each step of

2352-417: A more successful evolutionary theory based on natural selection ; similar reasoning and evidence led Alfred Russel Wallace to independently reach the same conclusions. The basis for modern genetics began with the work of Gregor Mendel in 1865. This outlined the principles of biological inheritance. However, the significance of his work was not realized until the early 20th century when evolution became

2499-465: A necessary and permanent feature of beach maintenance. During beach nourishment activities, care must be taken to place new sediments so that the new sediments compact and stabilize before aggressive wave or wind action can erode them. Material that is concentrated too far down the beach may form a temporary groyne that will encourage scouring behind it. Sediments that are too fine or too light may be eroded before they have compacted or been integrated into

2646-457: A new cycle. In contrast to mitosis, meiosis results in four haploid daughter cells by undergoing one round of DNA replication followed by two divisions. Homologous chromosomes are separated in the first division ( meiosis I ), and sister chromatids are separated in the second division ( meiosis II ). Both of these cell division cycles are used in the process of sexual reproduction at some point in their life cycle. Both are believed to be present in

2793-409: A process known as allopatric speciation . A phylogeny is an evolutionary history of a specific group of organisms or their genes. It can be represented using a phylogenetic tree , a diagram showing lines of descent among organisms or their genes. Each line drawn on the time axis of a tree represents a lineage of descendants of a particular species or population. When a lineage divides into two, it

2940-462: A rapid cycle of growth throughout the 1850s and 1860s. The growth was intensified by the practice among the Lancashire cotton mill owners of closing the factories for a week every year to service and repair machinery. These became known as wakes weeks . Each town's mills would close for a different week, allowing Blackpool to manage a steady and reliable stream of visitors over a prolonged period in

3087-408: A region of deoxyribonucleic acid (DNA) that carries genetic information that controls form or function of an organism. DNA is composed of two polynucleotide chains that coil around each other to form a double helix . It is found as linear chromosomes in eukaryotes , and circular chromosomes in prokaryotes . The set of chromosomes in a cell is collectively known as its genome . In eukaryotes, DNA

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3234-541: A role in the synthesis and packaging of proteins, respectively. Biomolecules such as proteins can be engulfed by lysosomes , another specialized organelle. Plant cells have additional organelles that distinguish them from animal cells such as a cell wall that provides support for the plant cell, chloroplasts that harvest sunlight energy to produce sugar, and vacuoles that provide storage and structural support as well as being involved in reproduction and breakdown of plant seeds. Eukaryotic cells also have cytoskeleton that

3381-436: A separate clade as some protists may be more closely related to plants, fungi, or animals than they are to other protists. Like groupings such as algae , invertebrates , or protozoans , the protist grouping is not a formal taxonomic group but is used for convenience. Most protists are unicellular; these are called microbial eukaryotes. Plants are mainly multicellular organisms , predominantly photosynthetic eukaryotes of

3528-435: A shoreline subject to constant erosion and loss of foreshore. This is often required where the flow of new sediment caused by the longshore current has been disrupted by construction of harbors, breakwaters, causeways or boat ramps, creating new current flows that scour the sand from behind these structures and deprive the beach of restorative sediments. If the causes of the erosion are not addressed, beach nourishment can become

3675-423: A single carbon atom can form four single covalent bonds such as in methane , two double covalent bonds such as in carbon dioxide (CO 2 ), or a triple covalent bond such as in carbon monoxide (CO). Moreover, carbon can form very long chains of interconnecting carbon–carbon bonds such as octane or ring-like structures such as glucose . The simplest form of an organic molecule is the hydrocarbon , which

3822-446: A specific enzyme. Enzymes are crucial to metabolism because they allow organisms to drive desirable reactions that require energy that will not occur by themselves, by coupling them to spontaneous reactions that release energy. Enzymes act as catalysts —they allow a reaction to proceed more rapidly without being consumed by it—by reducing the amount of activation energy needed to convert reactants into products . Enzymes also allow

3969-422: A tidal surge or tsunami which causes significant coastal flooding , substantial quantities of material may be eroded from the coastal plain or dunes behind the berm by receding water. This flow may alter the shape of the coastline, enlarge the mouths of rivers and create new deltas at the mouths of streams that had not been powerful enough to overcome longshore movement of sediment. The line between beach and dune

4116-399: A unified theory as the modern synthesis reconciled Darwinian evolution with classical genetics . In the 1940s and early 1950s, a series of experiments by Alfred Hershey and Martha Chase pointed to DNA as the component of chromosomes that held the trait-carrying units that had become known as genes . A focus on new kinds of model organisms such as viruses and bacteria, along with

4263-421: A varied mix of traits, and reproduction is able to increase any population, Darwin argued that in the natural world, it was nature that played the role of humans in selecting for specific traits. Darwin inferred that individuals who possessed heritable traits better adapted to their environments are more likely to survive and produce more offspring than other individuals. He further inferred that this would lead to

4410-492: A water molecule again. In pure water , the number of hydrogen ions balances (or equals) the number of hydroxyl ions, resulting in a pH that is neutral. Organic compounds are molecules that contain carbon bonded to another element such as hydrogen. With the exception of water, nearly all the molecules that make up each organism contain carbon. Carbon can form covalent bonds with up to four other atoms, enabling it to form diverse, large, and complex molecules. For example,

4557-536: Is evolution , which explains the unity and diversity of life. Energy processing is also important to life as it allows organisms to move , grow, and reproduce . Finally, all organisms are able to regulate their own internal environments . Biologists are able to study life at multiple levels of organization , from the molecular biology of a cell to the anatomy and physiology of plants and animals, and evolution of populations. Hence, there are multiple subdisciplines within biology , each defined by

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4704-883: Is a landform alongside a body of water which consists of loose particles. The particles composing a beach are typically made from rock , such as sand , gravel , shingle , pebbles , etc., or biological sources, such as mollusc shells or coralline algae . Sediments settle in different densities and structures, depending on the local wave action and weather , creating different textures, colors and gradients or layers of material. Though some beaches form on inland freshwater locations such as lakes and rivers , most beaches are in coastal areas where wave or current action deposits and reworks sediments. Erosion and changing of beach geologies happens through natural processes, like wave action and extreme weather events . Where wind conditions are correct, beaches can be backed by coastal dunes which offer protection and regeneration for

4851-444: Is a central feature of sexual reproduction in eukaryotes, and the most fundamental function of meiosis appears to be conservation of the integrity of the genome that is passed on to progeny by parents. Two aspects of sexual reproduction , meiotic recombination and outcrossing , are likely maintained respectively by the adaptive advantages of recombinational repair of genomic DNA damage and genetic complementation which masks

4998-641: Is a large family of organic compounds that are composed of hydrogen atoms bonded to a chain of carbon atoms. A hydrocarbon backbone can be substituted by other elements such as oxygen (O), hydrogen (H), phosphorus (P), and sulfur (S), which can change the chemical behavior of that compound. Groups of atoms that contain these elements (O-, H-, P-, and S-) and are bonded to a central carbon atom or skeleton are called functional groups . There are six prominent functional groups that can be found in organisms: amino group , carboxyl group , carbonyl group , hydroxyl group , phosphate group , and sulfhydryl group . In 1953,

5145-438: Is a significant source of sand particles. Some species of fish that feed on algae attached to coral outcrops and rocks can create substantial quantities of sand particles over their lifetime as they nibble during feeding, digesting the organic matter , and discarding the rock and coral particles which pass through their digestive tracts. The composition of the beach depends upon the nature and quantity of sediments upstream of

5292-480: Is a small polar molecule with a bent shape formed by the polar covalent bonds of two hydrogen (H) atoms to one oxygen (O) atom (H 2 O). Because the O–H bonds are polar, the oxygen atom has a slight negative charge and the two hydrogen atoms have a slight positive charge. This polar property of water allows it to attract other water molecules via hydrogen bonds, which makes water cohesive . Surface tension results from

5439-918: Is a term of convenience as not all algae are closely related. Algae comprise several distinct clades such as glaucophytes , which are microscopic freshwater algae that may have resembled in form to the early unicellular ancestor of Plantae. Unlike glaucophytes, the other algal clades such as red and green algae are multicellular. Green algae comprise three major clades: chlorophytes , coleochaetophytes , and stoneworts . Fungi are eukaryotes that digest foods outside their bodies, secreting digestive enzymes that break down large food molecules before absorbing them through their cell membranes. Many fungi are also saprobes , feeding on dead organic matter, making them important decomposers in ecological systems. Animals are multicellular eukaryotes. With few exceptions, animals consume organic material , breathe oxygen , are able to move , can reproduce sexually , and grow from

5586-414: Is an example of this phenomenon. Where there are tides , this line is formed by the highest position of the tide, and moves up and down the beach on a fortnightly cycle . The debris is chiefly composed of rotting seaweed , but can also include a large amount of litter , either from ships at sea or from sewage outflows. The strandline is an important habitat for a variety of animals . In parts of

5733-403: Is catalyzed by lactate dehydrogenase in a reversible reaction. Lactate can also be used as an indirect precursor for liver glycogen. During recovery, when oxygen becomes available, NAD attaches to hydrogen from lactate to form ATP. In yeast, the waste products are ethanol and carbon dioxide. This type of fermentation is known as alcoholic or ethanol fermentation . The ATP generated in this process

5880-452: Is difficult to define in the field. Over any significant period of time, sediment is always being exchanged between them. The drift line (the high point of material deposited by waves) is one potential demarcation. This would be the point at which significant wind movement of sand could occur, since the normal waves do not wet the sand beyond this area. However, the drift line is likely to move inland under assault by storm waves. Beaches are

6027-441: Is made by substrate-level phosphorylation , which does not require oxygen. Photosynthesis is a process used by plants and other organisms to convert light energy into chemical energy that can later be released to fuel the organism's metabolic activities via cellular respiration. This chemical energy is stored in carbohydrate molecules, such as sugars, which are synthesized from carbon dioxide and water. In most cases, oxygen

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6174-535: Is made up of microtubules , intermediate filaments , and microfilaments , all of which provide support for the cell and are involved in the movement of the cell and its organelles. In terms of their structural composition, the microtubules are made up of tubulin (e.g., α-tubulin and β-tubulin ) whereas intermediate filaments are made up of fibrous proteins. Microfilaments are made up of actin molecules that interact with other strands of proteins. All cells require energy to sustain cellular processes. Metabolism

6321-526: Is mainly in the cell nucleus . In prokaryotes, the DNA is held within the nucleoid . The genetic information is held within genes, and the complete assemblage in an organism is called its genotype . DNA replication is a semiconservative process whereby each strand serves as a template for a new strand of DNA. Mutations are heritable changes in DNA. They can arise spontaneously as a result of replication errors that were not corrected by proofreading or can be induced by an environmental mutagen such as

6468-399: Is one of the key ways a cell releases chemical energy to fuel cellular activity. The overall reaction occurs in a series of biochemical steps, some of which are redox reactions. Although cellular respiration is technically a combustion reaction , it clearly does not resemble one when it occurs in a cell because of the slow, controlled release of energy from the series of reactions. Sugar in

6615-407: Is pulled to the separate poles of the bacterium as it increases the size to prepare for splitting. Growth of a new cell wall begins to separate the bacterium (triggered by FtsZ polymerization and "Z-ring" formation). The new cell wall ( septum ) fully develops, resulting in the complete split of the bacterium. The new daughter cells have tightly coiled DNA rods, ribosomes , and plasmids . Meiosis

6762-399: Is released as a waste product. Most plants, algae , and cyanobacteria perform photosynthesis, which is largely responsible for producing and maintaining the oxygen content of the Earth's atmosphere, and supplies most of the energy necessary for life on Earth. Photosynthesis has four stages: Light absorption , electron transport, ATP synthesis, and carbon fixation . Light absorption is

6909-412: Is represented as a fork or split on the phylogenetic tree. Phylogenetic trees are the basis for comparing and grouping different species. Different species that share a feature inherited from a common ancestor are described as having homologous features (or synapomorphy ). Phylogeny provides the basis of biological classification. This classification system is rank-based, with the highest rank being

7056-455: Is the importing and deposition of sand or other sediments in an effort to restore a beach that has been damaged by erosion. Beach nourishment often involves excavation of sediments from riverbeds or sand quarries. This excavated sediment may be substantially different in size and appearance to the naturally occurring beach sand. In extreme cases, beach nourishment may involve placement of large pebbles or rocks in an effort to permanently restore

7203-468: Is the process by which genes and traits are passed on from parents to offspring. It has several principles. The first is that genetic characteristics, alleles , are discrete and have alternate forms (e.g., purple vs. white or tall vs. dwarf), each inherited from one of two parents. Based on the law of dominance and uniformity , which states that some alleles are dominant while others are recessive ; an organism with at least one dominant allele will display

7350-485: Is the set of chemical reactions in an organism. The three main purposes of metabolism are: the conversion of food to energy to run cellular processes; the conversion of food/fuel to monomer building blocks; and the elimination of metabolic wastes . These enzyme-catalyzed reactions allow organisms to grow and reproduce, maintain their structures, and respond to their environments. Metabolic reactions may be categorized as catabolic —the breaking down of compounds (for example,

7497-410: Is the study of chemical processes within and relating to living organisms . Molecular biology is the branch of biology that seeks to understand the molecular basis of biological activity in and between cells, including molecular synthesis, modification, mechanisms, and interactions. Life arose from the Earth's first ocean, which formed some 3.8 billion years ago. Since then, water continues to be

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7644-493: Is to store, transmit, and express hereditary information. Cell theory states that cells are the fundamental units of life, that all living things are composed of one or more cells, and that all cells arise from preexisting cells through cell division . Most cells are very small, with diameters ranging from 1 to 100  micrometers and are therefore only visible under a light or electron microscope . There are generally two types of cells: eukaryotic cells, which contain

7791-629: The Cambrian explosion . During the Permian period, synapsids , including the ancestors of mammals , dominated the land, but most of this group became extinct in the Permian–Triassic extinction event 252 million years ago. During the recovery from this catastrophe, archosaurs became the most abundant land vertebrates; one archosaur group, the dinosaurs, dominated the Jurassic and Cretaceous periods. After

7938-495: The Cretaceous–Paleogene extinction event 66 million years ago killed off the non-avian dinosaurs, mammals increased rapidly in size and diversity . Such mass extinctions may have accelerated evolution by providing opportunities for new groups of organisms to diversify. Bacteria are a type of cell that constitute a large domain of prokaryotic microorganisms . Typically a few micrometers in length, bacteria have

8085-535: The Miller–Urey experiment showed that organic compounds could be synthesized abiotically within a closed system mimicking the conditions of early Earth , thus suggesting that complex organic molecules could have arisen spontaneously in early Earth (see abiogenesis ). Macromolecules are large molecules made up of smaller subunits or monomers . Monomers include sugars, amino acids, and nucleotides. Carbohydrates include monomers and polymers of sugars. Lipids are

8232-492: The Precambrian about 1.5 billion years ago and can be classified into eight major clades : alveolates , excavates , stramenopiles , plants, rhizarians , amoebozoans , fungi , and animals. Five of these clades are collectively known as protists , which are mostly microscopic eukaryotic organisms that are not plants, fungi, or animals. While it is likely that protists share a common ancestor (the last eukaryotic common ancestor ), protists by themselves do not constitute

8379-473: The United Kingdom , sandhoppers such as Talitrus saltator and the seaweed fly Coelopa frigida are abundant in the rotting seaweed, and these invertebrates provide food for shore birds such as the rock pipit , turnstone and pied wagtail , and mammals such as brown hares , foxes , voles and mice . One kind of high water mark is the ordinary high water mark or average high water mark ,

8526-638: The United States Environmental Protection Agency , and other federal agencies to determine the geographical extent of their regulatory programs. Likewise, many states use similar definitions of the OHWM for the purposes of their own regulatory programs. In 2016, the Court of Appeals of Indiana ruled that land below the OHWM (as defined by common law) along Lake Michigan is held by the state in trust for public use. Beach A beach

8673-727: The activator is the transcription factor that stimulates transcription when it binds to the sequence near or at the promoter. Negative regulation occurs when another transcription factor called a repressor binds to a DNA sequence called an operator , which is part of an operon, to prevent transcription. Repressors can be inhibited by compounds called inducers (e.g., allolactose ), thereby allowing transcription to occur. Specific genes that can be activated by inducers are called inducible genes , in contrast to constitutive genes that are almost constantly active. In contrast to both, structural genes encode proteins that are not involved in gene regulation. In addition to regulatory events involving

8820-585: The bacterial phyla have species that can be grown in the laboratory. Archaea constitute the other domain of prokaryotic cells and were initially classified as bacteria, receiving the name archaebacteria (in the Archaebacteria kingdom ), a term that has fallen out of use. Archaeal cells have unique properties separating them from the other two domains , Bacteria and Eukaryota . Archaea are further divided into multiple recognized phyla . Archaea and bacteria are generally similar in size and shape, although

8967-404: The cell . In 1838, Schleiden and Schwann began promoting the now universal ideas that (1) the basic unit of organisms is the cell and (2) that individual cells have all the characteristics of life, although they opposed the idea that (3) all cells come from the division of other cells, continuing to support spontaneous generation . However, Robert Remak and Rudolf Virchow were able to reify

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9114-403: The cell membrane of another cell or located deep inside a cell. There are generally four types of chemical signals: autocrine , paracrine , juxtacrine , and hormones . In autocrine signaling, the ligand affects the same cell that releases it. Tumor cells, for example, can reproduce uncontrollably because they release signals that initiate their own self-division. In paracrine signaling,

9261-416: The circulatory systems of animals or vascular systems of plants to reach their target cells. Once a ligand binds with a receptor, it can influence the behavior of another cell, depending on the type of receptor. For instance, neurotransmitters that bind with an inotropic receptor can alter the excitability of a target cell. Other types of receptors include protein kinase receptors (e.g., receptor for

9408-528: The domain followed by kingdom , phylum , class , order , family , genus , and species . All organisms can be classified as belonging to one of three domains : Archaea (originally Archaebacteria), bacteria (originally eubacteria), or eukarya (includes the fungi, plant, and animal kingdoms). The history of life on Earth traces how organisms have evolved from the earliest emergence of life to present day. Earth formed about 4.5 billion years ago and all life on Earth, both living and extinct, descended from

9555-481: The duplication of its DNA and some of its organelles , and the subsequent partitioning of its cytoplasm into two daughter cells in a process called cell division . In eukaryotes (i.e., animal, plant, fungal , and protist cells), there are two distinct types of cell division: mitosis and meiosis . Mitosis is part of the cell cycle, in which replicated chromosomes are separated into two new nuclei. Cell division gives rise to genetically identical cells in which

9702-403: The kingdom Plantae, which would exclude fungi and some algae . Plant cells were derived by endosymbiosis of a cyanobacterium into an early eukaryote about one billion years ago, which gave rise to chloroplasts. The first several clades that emerged following primary endosymbiosis were aquatic and most of the aquatic photosynthetic eukaryotic organisms are collectively described as algae, which

9849-423: The microscope . It was then that scholars discovered spermatozoa , bacteria, infusoria and the diversity of microscopic life. Investigations by Jan Swammerdam led to new interest in entomology and helped to develop techniques of microscopic dissection and staining . Advances in microscopy had a profound impact on biological thinking. In the early 19th century, biologists pointed to the central importance of

9996-412: The mitochondrial cristae . Oxidative phosphorylation comprises the electron transport chain, which is a series of four protein complexes that transfer electrons from one complex to another, thereby releasing energy from NADH and FADH 2 that is coupled to the pumping of protons (hydrogen ions) across the inner mitochondrial membrane ( chemiosmosis ), which generates a proton motive force . Energy from

10143-546: The phenotype of that dominant allele. During gamete formation, the alleles for each gene segregate, so that each gamete carries only one allele for each gene. Heterozygotic individuals produce gametes with an equal frequency of two alleles. Finally, the law of independent assortment , states that genes of different traits can segregate independently during the formation of gametes, i.e., genes are unlinked. An exception to this rule would include traits that are sex-linked . Test crosses can be performed to experimentally determine

10290-497: The regulation of the rate of a metabolic reaction, for example in response to changes in the cell's environment or to signals from other cells. Cellular respiration is a set of metabolic reactions and processes that take place in cells to convert chemical energy from nutrients into adenosine triphosphate (ATP), and then release waste products. The reactions involved in respiration are catabolic reactions , which break large molecules into smaller ones, releasing energy. Respiration

10437-452: The seashore is most commonly associated with the word beach , beaches are also found by lakes and alongside large rivers. Beach may refer to: The former are described in detail below; the larger geological units are discussed elsewhere under bars . There are several conspicuous parts to a beach that relate to the processes that form and shape it. The part mostly above water (depending upon tide), and more or less actively influenced by

10584-409: The "ordinary high water mark" (OHWM) as "that line on the shore established by the fluctuations of water and indicated by physical characteristics such as a clear, natural line impressed on the bank, shelving, changes in the character of soil, destruction of terrestrial vegetation, the presence of litter and debris, or other appropriate means that consider the characteristics of the surrounding areas. For

10731-482: The 1720s; it had been a fashionable spa town since a stream of acidic water was discovered running from one of the cliffs to the south of the town in the 17th century. The first rolling bathing machines were introduced by 1735. The opening of the resort in Brighton and its reception of royal patronage from King George IV , extended the seaside as a resort for health and pleasure to the much larger London market, and

10878-628: The English Seas , "the Admirals power extended even to the high water mark, and into the main streams". In the United States , the high water mark is also significant because the United States Constitution gives Congress the authority to legislate for waterways, and the high water mark is used to determine the geographic extent of that authority. Federal regulations (33 CFR 328.3(e)) define

11025-475: The Phanerozoic eon that began 539 million years ago being subdivided into Paleozoic , Mesozoic , and Cenozoic eras. These three eras together comprise eleven periods ( Cambrian , Ordovician , Silurian , Devonian , Carboniferous , Permian , Triassic , Jurassic , Cretaceous , Tertiary , and Quaternary ). The similarities among all known present-day species indicate that they have diverged through

11172-696: The accumulation of favorable traits over successive generations, thereby increasing the match between the organisms and their environment. A species is a group of organisms that mate with one another and speciation is the process by which one lineage splits into two lineages as a result of having evolved independently from each other. For speciation to occur, there has to be reproductive isolation . Reproductive isolation can result from incompatibilities between genes as described by Bateson–Dobzhansky–Muller model . Reproductive isolation also tends to increase with genetic divergence . Speciation can occur when there are physical barriers that divide an ancestral species,

11319-633: The all-covering beachwear of the period was considered immodest. By the end of the century the English coastline had over 100 large resort towns, some with populations exceeding 50,000. Biological Biology is the scientific study of life . It is a natural science with a broad scope but has several unifying themes that tie it together as a single, coherent field. For instance, all organisms are made up of at least one cell that processes hereditary information encoded in genes , which can be transmitted to future generations. Another major theme

11466-435: The beach and into the sea. If the flow is constant, runoff from cleared land arriving at the beach head will tend to deposit this material into the sand changing its color, odor and fauna. The concentration of pedestrian and vehicular traffic accessing the beach for recreational purposes may cause increased erosion at the access points if measures are not taken to stabilize the beach surface above high-water mark. Recognition of

11613-452: The beach as a popular leisure resort from the mid-19th century was the first manifestation of what is now the global tourist industry. The first seaside resorts were opened in the 18th century for the aristocracy, who began to frequent the seaside as well as the then fashionable spa towns, for recreation and health. One of the earliest such seaside resorts, was Scarborough in Yorkshire during

11760-612: The beach became a centre for upper-class pleasure and frivolity. This trend was praised and artistically elevated by the new romantic ideal of the picturesque landscape; Jane Austen 's unfinished novel Sanditon is an example of that. Later, Queen Victoria 's long-standing patronage of the Isle of Wight and Ramsgate in Kent ensured that a seaside residence was considered as a highly fashionable possession for those wealthy enough to afford more than one home. The extension of this form of leisure to

11907-401: The beach head, they may erode the beach creating a lagoon or delta. Dense vegetation tends to absorb rainfall reducing the speed of runoff and releasing it over longer periods of time. Destruction by burning or clearance of the natural vegetation tends to increase the speed and erosive power of runoff from rainfall. This runoff will tend to carry more silt and organic matter from the land onto

12054-523: The beach profile is higher in summer. The gentle wave action during this season tends to transport sediment up the beach towards the berm where it is deposited and remains while the water recedes. Onshore winds carry it further inland forming and enhancing dunes. Conversely, the beach profile is lower in the storm season (winter in temperate areas) due to the increased wave energy, and the shorter periods between breaking wave crests. Higher energy waves breaking in quick succession tend to mobilise sediment from

12201-404: The beach, and the speed of flow and turbidity of water and wind. Sediments are moved by moving water and wind according to their particle size and state of compaction. Particles tend to settle and compact in still water. Once compacted, they are more resistant to erosion . Established vegetation (especially species with complex network root systems) will resist erosion by slowing the fluid flow at

12348-1686: The beach. However, these natural forces have become more extreme due to climate change , permanently altering beaches at very rapid rates. Some estimates describe as much as 50 percent of the earth's sandy beaches disappearing by 2100 due to climate-change driven sea level rise. Sandy beaches occupy about one third of global coastlines. These beaches are popular for recreation , playing important economic and cultural roles—often driving local tourism industries. To support these uses, some beaches have human-made infrastructure, such as lifeguard posts, changing rooms , showers, shacks and bars. They may also have hospitality venues (such as resorts, camps, hotels, and restaurants) nearby or housing, both for permanent and seasonal residents. Human forces have significantly changed beaches globally: direct impacts include bad construction practices on dunes and coastlines, while indirect human impacts include water pollution , plastic pollution and coastal erosion from sea level rise and climate change . Some coastal management practices are designed to preserve or restore natural beach processes, while some beaches are actively restored through practices like beach nourishment . Wild beaches, also known as undeveloped or undiscovered beaches, are not developed for tourism or recreation. Preserved beaches are important biomes with important roles in aquatic or marine biodiversity, such as for breeding grounds for sea turtles or nesting areas for seabirds or penguins . Preserved beaches and their associated dune are important for protection from extreme weather for inland ecosystems and human infrastructure. Although

12495-413: The beach. Compacted sediment is more resistant to movement by turbulent water from succeeding waves. Conversely, waves are destructive if the period between the wave crests is short. Sediment that remains in suspension when the following wave crest arrives will not be able to settle and compact and will be more susceptible to erosion by longshore currents and receding tides. The nature of sediments found on

12642-586: The body plan and the number, identity, and pattern of body parts. Among the most important toolkit genes are the Hox genes . Hox genes determine where repeating parts, such as the many vertebrae of snakes, will grow in a developing embryo or larva. Evolution is a central organizing concept in biology. It is the change in heritable characteristics of populations over successive generations . In artificial selection , animals were selectively bred for specific traits. Given that traits are inherited, populations contain

12789-450: The breaking down of glucose to pyruvate by cellular respiration ); or anabolic —the building up ( synthesis ) of compounds (such as proteins, carbohydrates, lipids, and nucleic acids). Usually, catabolism releases energy, and anabolism consumes energy. The chemical reactions of metabolism are organized into metabolic pathways , in which one chemical is transformed through a series of steps into another chemical, each step being facilitated by

12936-452: The character of underwater flora and fauna in the coastal shallows. Burning or clearance of vegetation on the land adjacent to the beach head, for farming and residential development, changes the surface wind patterns, and exposes the surface of the beach to wind erosion. Farming and residential development are also commonly associated with changes in local surface water flows. If these flows are concentrated in stormwater drains emptying onto

13083-410: The cohesive force due to the attraction between molecules at the surface of the liquid. Water is also adhesive as it is able to adhere to the surface of any polar or charged non-water molecules. Water is denser as a liquid than it is as a solid (or ice). This unique property of water allows ice to float above liquid water such as ponds, lakes, and oceans, thereby insulating the liquid below from

13230-433: The cold air above. Water has the capacity to absorb energy, giving it a higher specific heat capacity than other solvents such as ethanol . Thus, a large amount of energy is needed to break the hydrogen bonds between water molecules to convert liquid water into water vapor . As a molecule, water is not completely stable as each water molecule continuously dissociates into hydrogen and hydroxyl ions before reforming into

13377-501: The cytoplasm of a cell, there are many biomolecules such as proteins and nucleic acids . In addition to biomolecules, eukaryotic cells have specialized structures called organelles that have their own lipid bilayers or are spatially units. These organelles include the cell nucleus , which contains most of the cell's DNA, or mitochondria , which generate adenosine triphosphate (ATP) to power cellular processes. Other organelles such as endoplasmic reticulum and Golgi apparatus play

13524-455: The cytoplasm, where it is converted to waste products that may be removed from the cell. This serves the purpose of oxidizing the electron carriers so that they can perform glycolysis again and removing the excess pyruvate. Fermentation oxidizes NADH to NAD so it can be re-used in glycolysis. In the absence of oxygen, fermentation prevents the buildup of NADH in the cytoplasm and provides NAD for glycolysis. This waste product varies depending on

13671-444: The dangers of loss of beach front flora has caused many local authorities responsible for managing coastal areas to restrict beach access points by physical structures or legal sanctions, and fence off foredunes in an effort to protect the flora. These measures are often associated with the construction of structures at these access points to allow traffic to pass over or through the dunes without causing further damage. Beaches provide

13818-673: The development of biological knowledge. He explored biological causation and the diversity of life. His successor, Theophrastus , began the scientific study of plants. Scholars of the medieval Islamic world who wrote on biology included al-Jahiz (781–869), Al-Dīnawarī (828–896), who wrote on botany, and Rhazes (865–925) who wrote on anatomy and physiology . Medicine was especially well studied by Islamic scholars working in Greek philosopher traditions, while natural history drew heavily on Aristotelian thought. Biology began to quickly develop with Anton van Leeuwenhoek 's dramatic improvement of

13965-418: The development of body form, is the result of spatial differences in gene expression. A small fraction of the genes in an organism's genome called the developmental-genetic toolkit control the development of that organism. These toolkit genes are highly conserved among phyla , meaning that they are ancient and very similar in widely separated groups of animals. Differences in deployment of toolkit genes affect

14112-440: The developmental fate of a cell, which becomes more restrictive during development. Differentiation is the process by which specialized cells arise from less specialized cells such as stem cells . Stem cells are undifferentiated or partially differentiated cells that can differentiate into various types of cells and proliferate indefinitely to produce more of the same stem cell. Cellular differentiation dramatically changes

14259-411: The discovery of archaea in almost every habitat , including soil, oceans, and marshlands . Archaea are particularly numerous in the oceans, and the archaea in plankton may be one of the most abundant groups of organisms on the planet. Archaea are a major part of Earth's life . They are part of the microbiota of all organisms. In the human microbiome , they are important in the gut , mouth, and on

14406-420: The discovery of the double-helical structure of DNA by James Watson and Francis Crick in 1953, marked the transition to the era of molecular genetics . From the 1950s onwards, biology has been vastly extended in the molecular domain. The genetic code was cracked by Har Gobind Khorana , Robert W. Holley and Marshall Warren Nirenberg after DNA was understood to contain codons . The Human Genome Project

14553-616: The earliest terrestrial ecosystems , at least 2.7 billion years ago. Microorganisms are thought to have paved the way for the inception of land plants in the Ordovician period. Land plants were so successful that they are thought to have contributed to the Late Devonian extinction event . Ediacara biota appear during the Ediacaran period, while vertebrates , along with most other modern phyla originated about 525 million years ago during

14700-504: The effects of human-made structures and processes. Over long periods of time, these influences may substantially alter the shape of the coastline, and the character of the beach. Beachfront flora plays a major role in stabilizing the foredunes and preventing beach head erosion and inland movement of dunes. If flora with network root systems (creepers, grasses, and palms) are able to become established, they provide an effective coastal defense as they trap sand particles and rainwater and enrich

14847-434: The established vegetation. Foreign unwashed sediments may introduce flora or fauna that are not usually found in that locality. Brighton Beach, on the south coast of England, is a shingle beach that has been nourished with very large pebbles in an effort to withstand the erosion of the upper area of the beach. These large pebbles made the beach unwelcoming for pedestrians for a period of time until natural processes integrated

14994-473: The expression of deleterious recessive mutations . The beneficial effect of genetic complementation, derived from outcrossing (cross-fertilization) is also referred to as hybrid vigor or heterosis. Charles Darwin in his 1878 book The Effects of Cross and Self-Fertilization in the Vegetable Kingdom at the start of chapter XII noted “The first and most important of the conclusions which may be drawn from

15141-417: The feature. Where wind is the force distributing the grains inland, the deposit behind the beach becomes a dune . These geomorphic features compose what is called the beach profile . The beach profile changes seasonally due to the change in wave energy experienced during summer and winter months. In temperate areas where summer is characterised by calmer seas and longer periods between breaking wave crests,

15288-455: The form of glucose is the main nutrient used by animal and plant cells in respiration. Cellular respiration involving oxygen is called aerobic respiration, which has four stages: glycolysis , citric acid cycle (or Krebs cycle), electron transport chain , and oxidative phosphorylation . Glycolysis is a metabolic process that occurs in the cytoplasm whereby glucose is converted into two pyruvates , with two net molecules of ATP being produced at

15435-450: The heat of the sun evaporates the water leaving the salt which crystallises around the sand particles. This crust forms an additional protective layer that resists wind erosion unless disturbed by animals or dissolved by the advancing tide. Cusps and horns form where incoming waves divide, depositing sand as horns and scouring out sand to form cusps. This forms the uneven face on some sand shorelines . White sand beaches look white because

15582-449: The height of the water table. Some flora naturally occurring on the beach head requires freshwater runoff from the land. Diversion of freshwater runoff into drains may deprive these plants of their water supplies and allow sea water incursion, increasing the saltiness of the groundwater. Species that are not able to survive in salt water may die and be replaced by mangroves or other species adapted to salty environments. Beach nourishment

15729-480: The high water mark that can be expected to be produced by a body of water in non-flood conditions. The ordinary high water mark may have legal significance and is often being used to demarcate property boundaries . The ordinary high water mark has also been used for other legal demarcations. For example, a 1651 analysis of laws passed by the English Parliament notes that for persons granted the title Admiral of

15876-426: The hormone insulin ) and G protein-coupled receptors . Activation of G protein-coupled receptors can initiate second messenger cascades. The process by which a chemical or physical signal is transmitted through a cell as a series of molecular events is called signal transduction . The cell cycle is a series of events that take place in a cell that cause it to divide into two daughter cells. These events include

16023-414: The initial step of photosynthesis whereby light energy is absorbed by chlorophyll pigments attached to proteins in the thylakoid membranes . The absorbed light energy is used to remove electrons from a donor (water) to a primary electron acceptor, a quinone designated as Q. In the second stage, electrons move from the quinone primary electron acceptor through a series of electron carriers until they reach

16170-406: The last eukaryotic common ancestor. Prokaryotes (i.e., archaea and bacteria) can also undergo cell division (or binary fission ). Unlike the processes of mitosis and meiosis in eukaryotes, binary fission in prokaryotes takes place without the formation of a spindle apparatus on the cell. Before binary fission, DNA in the bacterium is tightly coiled. After it has uncoiled and duplicated, it

16317-405: The ligand diffuses to nearby cells and affects them. For example, brain cells called neurons release ligands called neurotransmitters that diffuse across a synaptic cleft to bind with a receptor on an adjacent cell such as another neuron or muscle cell . In juxtacrine signaling, there is direct contact between the signaling and responding cells. Finally, hormones are ligands that travel through

16464-444: The membrane serving as membrane transporters , and peripheral proteins that loosely attach to the outer side of the cell membrane, acting as enzymes shaping the cell. Cell membranes are involved in various cellular processes such as cell adhesion , storing electrical energy , and cell signalling and serve as the attachment surface for several extracellular structures such as a cell wall , glycocalyx , and cytoskeleton . Within

16611-445: The middle and working classes began with the development of the railways in the 1840s, which offered cheap fares to fast-growing resort towns. In particular, the completion of a branch line to the small seaside town of Blackpool from Poulton led to a sustained economic and demographic boom. A sudden influx of visitors, arriving by rail, led entrepreneurs to build accommodation and create new attractions, leading to more visitors and

16758-403: The mitotic phase of an animal cell cycle—the division of the mother cell into two genetically identical daughter cells. The cell cycle is a vital process by which a single-celled fertilized egg develops into a mature organism, as well as the process by which hair, skin, blood cells , and some internal organs are renewed. After cell division, each of the daughter cells begin the interphase of

16905-437: The most abundant molecule in every organism. Water is important to life because it is an effective solvent , capable of dissolving solutes such as sodium and chloride ions or other small molecules to form an aqueous solution . Once dissolved in water, these solutes are more likely to come in contact with one another and therefore take part in chemical reactions that sustain life. In terms of its molecular structure , water

17052-612: The naturally occurring shingle into the pebble base. Even in Roman times, wealthy people spent their free time on the coast. They also built large villa complexes with bathing facilities (so-called maritime villas) in particularly beautiful locations. Excavations of Roman architecture can still be found today, for example on the Amalfi Coast near Naples and in Barcola in Trieste. The development of

17199-403: The nature of their research questions and the tools that they use. Like other scientists, biologists use the scientific method to make observations , pose questions, generate hypotheses , perform experiments, and form conclusions about the world around them. Life on Earth, which emerged more than 3.7 billion years ago, is immensely diverse. Biologists have sought to study and classify

17346-441: The observations given in this volume, is that generally cross-fertilisation is beneficial and self-fertilisation often injurious, at least with the plants on which I experimented.” Genetic variation , often produced as a byproduct of sexual reproduction, may provide long-term advantages to those sexual lineages that engage in outcrossing . Genetics is the scientific study of inheritance. Mendelian inheritance , specifically,

17493-517: The only class of macromolecules that are not made up of polymers. They include steroids , phospholipids , and fats, largely nonpolar and hydrophobic (water-repelling) substances. Proteins are the most diverse of the macromolecules. They include enzymes , transport proteins , large signaling molecules, antibodies , and structural proteins . The basic unit (or monomer) of a protein is an amino acid . Twenty amino acids are used in proteins. Nucleic acids are polymers of nucleotides . Their function

17640-404: The organism. In skeletal muscles, the waste product is lactic acid . This type of fermentation is called lactic acid fermentation . In strenuous exercise, when energy demands exceed energy supply, the respiratory chain cannot process all of the hydrogen atoms joined by NADH. During anaerobic glycolysis, NAD regenerates when pairs of hydrogen combine with pyruvate to form lactate. Lactate formation

17787-621: The permanent water forming offshore bars, lagoons or increasing the area of the beach exposed at low tide. Large and rapid movements of exposed sand can bury and smother flora in adjacent areas, aggravating the loss of habitat for fauna, and enlarging the area of instability. If there is an adequate supply of sand, and weather conditions do not allow vegetation to recover and stabilize the sediment, wind-blown sand can continue to advance, engulfing and permanently altering downwind landscapes. Sediment moved by waves or receding floodwaters can be deposited in coastal shallows, engulfing reed beds and changing

17934-428: The possibility of common descent . Serious evolutionary thinking originated with the works of Jean-Baptiste Lamarck , who presented a coherent theory of evolution. The British naturalist Charles Darwin , combining the biogeographical approach of Humboldt , the uniformitarian geology of Lyell , Malthus's writings on population growth, and his own morphological expertise and extensive natural observations, forged

18081-907: The process of evolution from their common ancestor. Biologists regard the ubiquity of the genetic code as evidence of universal common descent for all bacteria , archaea , and eukaryotes . Microbial mats of coexisting bacteria and archaea were the dominant form of life in the early Archean eon and many of the major steps in early evolution are thought to have taken place in this environment. The earliest evidence of eukaryotes dates from 1.85 billion years ago, and while they may have been present earlier, their diversification accelerated when they started using oxygen in their metabolism . Later, around 1.7 billion years ago, multicellular organisms began to appear, with differentiated cells performing specialised functions. Algae-like multicellular land plants are dated back to about 1 billion years ago, although evidence suggests that microorganisms formed

18228-539: The process such as transcription , RNA splicing , translation , and post-translational modification of a protein. Gene expression can be influenced by positive or negative regulation, depending on which of the two types of regulatory proteins called transcription factors bind to the DNA sequence close to or at a promoter. A cluster of genes that share the same promoter is called an operon , found mainly in prokaryotes and some lower eukaryotes (e.g., Caenorhabditis elegans ). In positive regulation of gene expression,

18375-517: The promoter, gene expression can also be regulated by epigenetic changes to chromatin , which is a complex of DNA and protein found in eukaryotic cells. Development is the process by which a multicellular organism (plant or animal) goes through a series of changes, starting from a single cell, and taking on various forms that are characteristic of its life cycle. There are four key processes that underlie development: Determination , differentiation , morphogenesis , and growth. Determination sets

18522-406: The proton motive force drives the enzyme ATP synthase to synthesize more ATPs by phosphorylating ADPs . The transfer of electrons terminates with molecular oxygen being the final electron acceptor . If oxygen were not present, pyruvate would not be metabolized by cellular respiration but undergoes a process of fermentation . The pyruvate is not transported into the mitochondrion but remains in

18669-643: The purposes of Section 404 of the Clean Water Act , the OHWM defines the lateral limits of federal jurisdiction over non-tidal water bodies in the absence of adjacent wetlands. For the purposes of Sections 9 and 10 of the Rivers and Harbors Act of 1899 , the OHWM defines the lateral limits of federal jurisdiction over traditional navigable waters of the US. The OHWM is used by the United States Army Corps of Engineers ,

18816-564: The quartz or eroded limestone in the sand reflects or scatters sunlight without absorbing other colors. The composition of the sand varies depending on the local minerals and geology. Some of the types of sand found in beaches around the world are: Beaches are changed in shape chiefly by the movement of water and wind. Any weather event that is associated with turbid or fast-flowing water or high winds will erode exposed beaches. Longshore currents will tend to replenish beach sediments and repair storm damage. Tidal waterways generally change

18963-415: The result of wave action by which waves or currents move sand or other loose sediments of which the beach is made as these particles are held in suspension . Alternatively, sand may be moved by saltation (a bouncing movement of large particles). Beach materials come from erosion of rocks offshore, as well as from headland erosion and slumping producing deposits of scree . A coral reef offshore

19110-449: The same time. Each pyruvate is then oxidized into acetyl-CoA by the pyruvate dehydrogenase complex , which also generates NADH and carbon dioxide. Acetyl-CoA enters the citric acid cycle, which takes places inside the mitochondrial matrix. At the end of the cycle, the total yield from 1 glucose (or 2 pyruvates) is 6 NADH, 2 FADH 2 , and 2 ATP molecules. Finally, the next stage is oxidative phosphorylation, which in eukaryotes, occurs in

19257-676: The second and third stages, respectively, provide the energy and electrons to drive the synthesis of glucose by fixing atmospheric carbon dioxide into existing organic carbon compounds, such as ribulose bisphosphate (RuBP) in a sequence of light-independent (or dark) reactions called the Calvin cycle . Cell signaling (or communication) is the ability of cells to receive, process, and transmit signals with its environment and with itself. Signals can be non-chemical such as light, electrical impulses , and heat, or chemical signals (or ligands ) that interact with receptors , which can be found embedded in

19404-412: The shallows, keeping it in suspension where it is prone to be carried along the beach by longshore currents, or carried out to sea to form longshore bars, especially if the longshore current meets an outflow from a river or flooding stream. The removal of sediment from the beach berm and dune thus decreases the beach profile. If storms coincide with unusually high tides, or with a freak wave event such as

19551-487: The shape of their adjacent beaches by small degrees with every tidal cycle. Over time these changes can become substantial leading to significant changes in the size and location of the beach. Changes in the shape of the beach may undermine the roots of large trees and other flora. Many beach adapted species (such as coconut palms) have a fine root system and large root ball which tends to withstand wave and wind action and tends to stabilize beaches better than other trees with

19698-562: The shape, profile and location of a beach within hours. Destruction of flora on the berm by the use of herbicides, excessive pedestrian or vehicle traffic, or disruption to freshwater flows may lead to erosion of the berm and dunes. While the destruction of flora may be a gradual process that is imperceptible to regular beach users, it often becomes immediately apparent after storms associated with high winds and freak wave events that can rapidly move large volumes of exposed and unstable sand, depositing them further inland, or carrying them out into

19845-553: The skin. Their morphological, metabolic, and geographical diversity permits them to play multiple ecological roles: carbon fixation; nitrogen cycling; organic compound turnover; and maintaining microbial symbiotic and syntrophic communities, for example. Eukaryotes are hypothesized to have split from archaea, which was followed by their endosymbioses with bacteria (or symbiogenesis ) that gave rise to mitochondria and chloroplasts, both of which are now part of modern-day eukaryotic cells. The major lineages of eukaryotes diversified in

19992-476: The stroma. This is analogous to the proton-motive force generated across the inner mitochondrial membrane in aerobic respiration. During the third stage of photosynthesis, the movement of protons down their concentration gradients from the thylakoid lumen to the stroma through the ATP synthase is coupled to the synthesis of ATP by that same ATP synthase. The NADPH and ATPs generated by the light-dependent reactions in

20139-566: The summer. A prominent feature of the resort was the promenade and the pleasure piers , where an eclectic variety of performances vied for the people's attention. In 1863, the North Pier in Blackpool was completed, rapidly becoming a centre of attraction for upper class visitors. Central Pier was completed in 1868, with a theatre and a large open-air dance floor. Many of the popular beach resorts were equipped with bathing machines , because even

20286-425: The surface flow of the runoff is diverted and concentrated by drains that create constant flows over the beach above the sea or river level, the beach will be eroded and ultimately form an inlet unless longshore flows deposit sediments to repair the breach. Once eroded, an inlet may allow tidal inflows of salt water to pollute areas inland from the beach and may also affect the quality of underground water supplies and

20433-519: The surface layer of the dunes, allowing other plant species to become established. They also protect the berm from erosion by high winds, freak waves and subsiding floodwaters. Over long periods of time, well-stabilized foreshore areas will tend to accrete, while unstabilized foreshores will tend to erode, leading to substantial changes in the shape of the coastline. These changes usually occur over periods of many years. Freak wave events such as tsunami, tidal waves, and storm surges may substantially alter

20580-657: The surface layer. When affected by moving water or wind, particles that are eroded and held in suspension will increase the erosive power of the fluid that holds them by increasing the average density, viscosity, and volume of the moving fluid. Coastlines facing very energetic wind and wave systems will tend to hold only large rocks as smaller particles will be held in suspension in the turbid water column and carried to calmer areas by longshore currents and tides. Coastlines that are protected from waves and winds will tend to allow finer sediments such as clay and mud to precipitate creating mud flats and mangrove forests. The shape of

20727-471: The third tenet, and by the 1860s most biologists accepted all three tenets which consolidated into cell theory . Meanwhile, taxonomy and classification became the focus of natural historians. Carl Linnaeus published a basic taxonomy for the natural world in 1735, and in the 1750s introduced scientific names for all his species. Georges-Louis Leclerc, Comte de Buffon , treated species as artificial categories and living forms as malleable—even suggesting

20874-439: The total number of chromosomes is maintained. In general, mitosis (division of the nucleus) is preceded by the S stage of interphase (during which the DNA is replicated) and is often followed by telophase and cytokinesis ; which divides the cytoplasm , organelles and cell membrane of one cell into two new cells containing roughly equal shares of these cellular components. The different stages of mitosis all together define

21021-431: The underlying genotype of an organism with a dominant phenotype. A Punnett square can be used to predict the results of a test cross. The chromosome theory of inheritance , which states that genes are found on chromosomes, was supported by Thomas Morgans 's experiments with fruit flies , which established the sex linkage between eye color and sex in these insects. A gene is a unit of heredity that corresponds to

21168-656: The various forms of life, from prokaryotic organisms such as archaea and bacteria to eukaryotic organisms such as protists , fungi, plants, and animals. These various organisms contribute to the biodiversity of an ecosystem , where they play specialized roles in the cycling of nutrients and energy through their biophysical environment . The earliest of roots of science, which included medicine, can be traced to ancient Egypt and Mesopotamia in around 3000 to 1200 BCE . Their contributions shaped ancient Greek natural philosophy . Ancient Greek philosophers such as Aristotle (384–322 BCE) contributed extensively to

21315-408: The waves at some point in the tide, is termed the beach berm. The berm is the deposit of material comprising the active shoreline. The berm has a crest (top) and a face —the latter being the slope leading down towards the water from the crest. At the very bottom of the face, there may be a trough , and further seaward one or more long shore bars: slightly raised, underwater embankments formed where

21462-434: The waves first start to break. The sand deposit may extend well inland from the berm crest , where there may be evidence of one or more older crests (the storm beach ) resulting from very large storm waves and beyond the influence of the normal waves. At some point the influence of the waves (even storm waves) on the material comprising the beach stops, and if the particles are small enough (sand size or smaller), winds shape

21609-434: Was launched in 1990 to map the human genome . All organisms are made up of chemical elements ; oxygen , carbon , hydrogen , and nitrogen account for most (96%) of the mass of all organisms, with calcium , phosphorus , sulfur , sodium , chlorine , and magnesium constituting essentially all the remainder. Different elements can combine to form compounds such as water, which is fundamental to life. Biochemistry

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