Llewellyn Ivor Price (October 9, 1905 – June 9, 1980) was one of the first Brazilian paleontologists . His work contributed not only to the development of Brazilian but also to global paleontology . He collected Staurikosaurus in 1936, the first dinosaur discovered in Brazil.
13-616: See text Romeriidae Price , 1937 Captorhinidae is an extinct family of tetrapods , typically considered primitive reptiles , known from the late Carboniferous to the Late Permian . They had a cosmopolitan distribution across Pangea . Captorhinids are a clade of small to very large lizard-like animals that date from the Late Carboniferous through the Permian . Their skulls were much stronger than those of their relatives,
26-465: A 2018 study that captorhinids were the first amniotes to develop caudal autotomy as a defensive function. In studied specimens a split line is present in certain caudal vertebrae that is similar to those found in modern reptiles that perform caudal autonomy. This behaviour represented significant evolutionary benefit for the animals, allowing for escape and distracting predators, as well as minimizing blood loss at an injury site. Euconcordia cunninghami
39-545: A comparatively large pineal foramen (relative to skull size), which are similar to the related basal species Procaptorhinus . Rhiodenticulatus was first named by David S. Berman and Robert R. Reisz in 1986 and the type species is Rhiodenticulatus heatoni . The generic name is derived from Greek , meaning "peak with small teeth". The specific name honors the paleontologist Malcolm J. Heaton. [REDACTED] [REDACTED] [REDACTED] [REDACTED] [REDACTED] [REDACTED] This article about
52-471: Is an extinct genus of Early Permian captorhinid known from Rio Arriba County , New Mexico of the United States . Rhiodenticulatus is known from the holotype UCMP 35757, three-dimensionally preserved nearly complete skull and postcranial elements, and from the referred specimens, which come from a second individual, UCMP 40209 and UCMP 40210. All specimens were collected in
65-577: Is thought to be the basalmost known member of Captorhinidae. A phylogenic study of primitive reptile relationships by Muller & Reisz in 2006 recovered Thuringothyris as a sister taxon of the Captorhinidae. The same results were obtained in later phylogenic analyses. Captorhinidae contains a single subfamily, the Moradisaurinae . Moradisaurinae was named and assigned to the family Captorhinidae by A. D. Ricqlès and P. Taquet in 1982. Moradisaurinae
78-616: The Camp Quarry (UCMP V-2814 locality), from the Cutler Formation of New Mexico, dating to the Sakmarian stage of the early Cisuralian Series . It is small for a captorhinid, with a skull length of about 45 mm, and in most respects the skull is fairly primitive. However, it does have some unusual features; the lacrimal bone is very high dorsoventrally and so the snout has a domed appearance. The teeth are also unusual, in that
91-433: The premaxillary teeth are homodont - most captorhinids had one large premaxillary tooth and several slightly smaller ones - and the middle maxillary teeth are uneven, with one about twice as wide at the base as the others but the same height. The postcranial bones show no differences to those of other captorhinids. It is fairly basal among captorhinids, as it only has one row of teeth, a slender supratemporal bone, and
104-594: The protorothyridids , and had teeth that were better able to deal with tough plant material. The postcranial skeleton is similar to those of seymouriamorphs and diadectomorphs ; these animals were grouped together with the captorhinids in the order Cotylosauria as the first reptiles in the early 20th century, but are now usually regarded as stem - amniotes no closer to reptiles than to mammals . Captorhinids have broad, robust skulls that are generally triangular in shape when seen in dorsal view. The premaxillae are characteristically downturned. The largest captorhinid,
117-795: The early 20th century. However, more recent fossil finds have shown that Eunotosaurus was either a parareptile or a diapsid , and therefore unrelated to captorhinids. The following taxonomy follows Reisz et al. , 2011 and Sumida et al. , 2010 unless otherwise noted. The cladogram below follows the topology from a 2011 analysis by paleontologists Robert R. Reisz , Jun Liu, Jin-Ling Li and Johannes Müller. Paleothyris Thuringothyris Concordia Rhiodenticulatus Romeria Protocaptorhinus Saurorictus C. laticeps C. aguti C. magnus Captorhinikos Labidosaurus Labidosaurikos Moradisaurus Rothianiscus Gansurhinus Simões et al. (2022) recovered captorhinids as stem-amniotes instead, as
130-566: The herbivorous Moradisaurus , could reach an estimated snout-vent length of 2 meters (6.5 feet). Early, smaller forms possessed single rows of teeth, and were likely carnivorous or omnivorous, while the larger, more derived captorhinids belonging to the subfamily Moradisaurinae were herbivorous and developed multiple (up to 11) rows of teeth in the jaws alongside propalinal (back and forth) jaw motion, which created an effective apparatus for grinding and shredding plant matter. Histological and SEM analysis of captorhinid tail vertebrae concluded in
143-701: The sister group to Protorothyris archeri , while the clade including captorhinids and P. archeri was recovered as the sister group to Araeoscelidia . A cladogram from that study is shown below. Using the same data matrix, Klembara et al. (2023) found a similar result. Seymouria Limnoscelis paludis Diadectes Orobates pabsti Sauropsida Synapsida Araeoscelidia Protorothyris archeri Captorhinus aguti Labidosaurus hamatus Euconcordia cunninghami Protocaptorhinus pricei [REDACTED] [REDACTED] [REDACTED] [REDACTED] [REDACTED] [REDACTED] [REDACTED] Llewellyn Ivor Price Price
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#1732891652772156-589: Was born in Santa Maria on October 9, 1905. The son of American parents, he studied chemistry and graduated in zoology and geology in the United States. After being professor at Harvard he returned to Brazil . He died of a heart attack in Rio de Janeiro on June 9, 1980, aged 74. This Brazilian scientist article is a stub . You can help Misplaced Pages by expanding it . Rhiodenticulatus Rhiodenticulatus
169-423: Was defined as "all captorhinids more closely related to Moradisaurus than to Captorhinus ". The moradisaurines inhabited what is now China , Morocco , Niger , Russia , Texas and Oklahoma . Captorhinids were once thought to be the ancestors of turtles . The Middle Permian reptile Eunotosaurus from South Africa was seen as the " missing link " between cotylosaurs and chelonians throughout much of
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