127-503: A lyrebird is either of two species of ground-dwelling Australian birds that compose the genus Menura , and the family Menuridae . They are most notable for their impressive ability to mimic natural and artificial sounds from their environment, and the striking beauty of the male bird's huge tail when it is fanned out in courtship display. Lyrebirds have unique plumes of neutral-coloured tailfeathers and are among Australia's best-known native birds. The classification of lyrebirds
254-423: A mutation–selection balance . It is predicted that a viral quasispecies at a low but evolutionarily neutral and highly connected (that is, flat) region in the fitness landscape will outcompete a quasispecies located at a higher but narrower fitness peak in which the surrounding mutants are unfit, "the quasispecies effect" or the "survival of the flattest". There is no suggestion that a viral quasispecies resembles
381-406: A self-governing Crown colony with responsible government . Brisbane was selected as the capital city. On 10 December 1859, a proclamation was read by George Bowen , the first Governor of Queensland , formally establishing Queensland as a separate colony from New South Wales. On 22 May 1860 the first Queensland election was held and Robert Herbert , Bowen's private secretary, was appointed as
508-400: A "classical" method of determining species, such as with Linnaeus, early in evolutionary theory. However, different phenotypes are not necessarily different species (e.g. a four-winged Drosophila born to a two-winged mother is not a different species). Species named in this manner are called morphospecies . In the 1970s, Robert R. Sokal , Theodore J. Crovello and Peter Sneath proposed
635-424: A 'smallest clade' idea" (a phylogenetic species concept). Mishler and Wilkins and others concur with this approach, even though this would raise difficulties in biological nomenclature. Wilkins cited the ichthyologist Charles Tate Regan 's early 20th century remark that "a species is whatever a suitably qualified biologist chooses to call a species". Wilkins noted that the philosopher Philip Kitcher called this
762-405: A S shape that together resemble the shape of a lyre . Lyrebirds were thought to be Galliformes like the broadly similar looking partridge , junglefowl , and pheasants familiar to Europeans, reflected in the early names given to the superb lyrebird, including native pheasant. They were also called peacock-wrens and Australian birds-of-paradise. The idea that they were related to the pheasants
889-428: A connected series of neighbouring populations, each of which can sexually interbreed with adjacent related populations, but for which there exist at least two "end" populations in the series, which are too distantly related to interbreed, though there is a potential gene flow between each "linked" population. Such non-breeding, though genetically connected, "end" populations may co-exist in the same region thus closing
1016-432: A different species from its ancestors. Viruses have enormous populations, are doubtfully living since they consist of little more than a string of DNA or RNA in a protein coat, and mutate rapidly. All of these factors make conventional species concepts largely inapplicable. A viral quasispecies is a group of genotypes related by similar mutations, competing within a highly mutagenic environment, and hence governed by
1143-584: A force of settlers and native police at Battle Mountain near modern Cloncurry . The subsequent battle of Battle Mountain ended in disaster for the Kalkadoon, who suffered heavy losses. Fighting continued in North Queensland , however, with First Nations raiders attacking sheep and cattle while Native Police mounted heavy retaliatory massacres. Tens of thousands of South Sea Islanders were kidnapped from islands nearby to Australia and sold as slaves to work on
1270-508: A genetic boundary suitable for defining a species concept is present. DNA barcoding has been proposed as a way to distinguish species suitable even for non-specialists to use. One of the barcodes is a region of mitochondrial DNA within the gene for cytochrome c oxidase . A database, Barcode of Life Data System , contains DNA barcode sequences from over 190,000 species. However, scientists such as Rob DeSalle have expressed concern that classical taxonomy and DNA barcoding, which they consider
1397-557: A great rush to take up the surrounding land in the Darling Downs , Logan and Brisbane Valley and South Burnett onwards from 1840, in many cases leading to widespread fighting and heavy loss of life. The conflict later spread north to the Wide Bay and Burnett River and Hervey Bay region, and at one stage the settlement of Maryborough was virtually under siege. The largest reasonably well-documented massacres in southeast Queensland were
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#17330848902191524-465: A lineage should be divided into multiple chronospecies , or when populations have diverged to have enough distinct character states to be described as cladistic species. Species and higher taxa were seen from the time of Aristotle until the 18th century as categories that could be arranged in a hierarchy, the great chain of being . In the 19th century, biologists grasped that species could evolve given sufficient time. Charles Darwin 's 1859 book On
1651-541: A lyre, and that the tail would be held in a similar way to that of a peacock during courtship display , and so he arranged the feathers in this way. Later, John Gould (who had also never seen a live lyrebird), painted the lyrebird from the British Museum specimen. The male lyrebird's tail is not held as in John Gould's painting. Instead, the male lyrebird's tail is fanned over the lyrebird during courtship display, with
1778-500: A lyrebird chick had been a pet, although they acknowledged compelling evidence on both sides of the argument. Until the 2019–2020 Australian bushfire season , superb lyrebirds were not considered threatened in the short to medium term. Concern has since grown as early analyses have shown the extent of destruction of the lyrebird's preferred wet-forest habitats, which in less intense previous bushfire seasons have been spared, in large part due to their moisture content. Albert's lyrebird has
1905-492: A misnomer, need to be reconciled, as they delimit species differently. Genetic introgression mediated by endosymbionts and other vectors can further make barcodes ineffective in the identification of species. A phylogenetic or cladistic species is "the smallest aggregation of populations (sexual) or lineages (asexual) diagnosable by a unique combination of character states in comparable individuals (semaphoronts)". The empirical basis – observed character states – provides
2032-449: A particular species, including which genus (and higher taxa) it is placed in, is a hypothesis about the evolutionary relationships and distinguishability of that group of organisms. As further information comes to hand, the hypothesis may be corroborated or refuted. Sometimes, especially in the past when communication was more difficult, taxonomists working in isolation have given two distinct names to individual organisms later identified as
2159-400: A perfect model of life, it is still a useful tool to scientists and conservationists for studying life on Earth, regardless of the theoretical difficulties. If species were fixed and clearly distinct from one another, there would be no problem, but evolutionary processes cause species to change. This obliges taxonomists to decide, for example, when enough change has occurred to declare that
2286-630: A population of half a million people. Since then, Queensland has remained a federated state within Australia, and its population has significantly grown. In 1905 women voted in state elections for the first time. The state's first university, the University of Queensland , was established in Brisbane in 1909. In 1911, the first alternative treatments for polio were pioneered in Queensland and remain in use across
2413-472: A process known as blackbirding or press-ganging, and their employment conditions constituted an allegedly exploitative form of indentured labour. Italian immigrants entered the sugar cane industry from the 1890s. During the 1890s, the six Australian colonies, including Queensland, held a series of referendums which culminated in the Federation of Australia on 1 January 1901. During this time, Queensland had
2540-408: A reclassification of its status from "common" to "threatened". Beyond this new threat are the long-term vulnerabilities to predation by cats and foxes , as well as human population pressure on its habitat. The lyrebird is so called because the male bird has a spectacular tail, consisting of 16 highly modified feathers (two long slender lyrates at the centre of the plume, two broader medians on
2667-400: A short way of saying that something applies to many species within a genus, but not to all. If scientists mean that something applies to all species within a genus, they use the genus name without the specific name or epithet. The names of genera and species are usually printed in italics . However, abbreviations such as "sp." should not be italicised. When a species' identity is not clear,
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#17330848902192794-562: A single family, but evidence that they are also related to the bowerbirds remains controversial. Lyrebirds are ancient Australian animals: the Australian Museum has fossils of lyrebirds dating back to about 15 million years ago. The prehistoric Menura tyawanoides has been described from Early Miocene fossils found at the famous Riversleigh site. Two species of lyrebird are extant: [REDACTED] [REDACTED] The lyrebirds are large passerine birds, amongst
2921-404: A specialist may use "cf." before the epithet to indicate that confirmation is required. The abbreviations "nr." (near) or "aff." (affine) may be used when the identity is unclear but when the species appears to be similar to the species mentioned after. With the rise of online databases, codes have been devised to provide identifiers for species that are already defined, including: The naming of
3048-523: A species as groups of actually or potentially interbreeding natural populations, which are reproductively isolated from other such groups. It has been argued that this definition is a natural consequence of the effect of sexual reproduction on the dynamics of natural selection. Mayr's use of the adjective "potentially" has been a point of debate; some interpretations exclude unusual or artificial matings that occur only in captivity, or that involve animals capable of mating but that do not normally do so in
3175-400: A species as determined by a taxonomist. A typological species is a group of organisms in which individuals conform to certain fixed properties (a type), so that even pre-literate people often recognise the same taxon as do modern taxonomists. The clusters of variations or phenotypes within specimens (such as longer or shorter tails) would differentiate the species. This method was used as
3302-488: A species. All species definitions assume that an organism acquires its genes from one or two parents very like the "daughter" organism, but that is not what happens in HGT. There is strong evidence of HGT between very dissimilar groups of prokaryotes , and at least occasionally between dissimilar groups of eukaryotes , including some crustaceans and echinoderms . The evolutionary biologist James Mallet concludes that there
3429-685: A species. Generally the term includes the unknown element of a distinct act of creation. Many authors have argued that a simple textbook definition, following Mayr's concept, works well for most multi-celled organisms , but breaks down in several situations: Species identification is made difficult by discordance between molecular and morphological investigations; these can be categorised as two types: (i) one morphology, multiple lineages (e.g. morphological convergence , cryptic species ) and (ii) one lineage, multiple morphologies (e.g. phenotypic plasticity , multiple life-cycle stages). In addition, horizontal gene transfer (HGT) makes it difficult to define
3556-498: A superb lyrebird mimicking sounds of an electronic shooting game, workmen and chainsaws was added to the National Film and Sound Archive 's Sounds of Australia registry in 2013. Both species of lyrebird produced elaborate lyrebird-specific vocalisations including 'whistle songs'. Males also sing songs specifically associated with their song and dance displays. One researcher, Sydney Curtis, has recorded flute-like lyrebird calls in
3683-519: A taxonomic decision at the discretion of cognizant specialists, is not governed by the Codes of Zoological or Botanical Nomenclature, in contrast to the PhyloCode , and contrary to what is done in several other fields, in which the definitions of technical terms, like geochronological units and geopolitical entities, are explicitly delimited. The nomenclatural codes that guide the naming of species, including
3810-506: A traditional biological species. The International Committee on Taxonomy of Viruses has since 1962 developed a universal taxonomic scheme for viruses; this has stabilised viral taxonomy. Most modern textbooks make use of Ernst Mayr 's 1942 definition, known as the Biological Species Concept as a basis for further discussion on the definition of species. It is also called a reproductive or isolation concept. This defines
3937-447: A variation on the morphological species concept, a phenetic species, defined as a set of organisms with a similar phenotype to each other, but a different phenotype from other sets of organisms. It differs from the morphological species concept in including a numerical measure of distance or similarity to cluster entities based on multivariate comparisons of a reasonably large number of phenotypic traits. A mate-recognition species
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4064-515: A variety of reasons. Viruses are a special case, driven by a balance of mutation and selection , and can be treated as quasispecies . Biologists and taxonomists have made many attempts to define species, beginning from morphology and moving towards genetics . Early taxonomists such as Linnaeus had no option but to describe what they saw: this was later formalised as the typological or morphological species concept. Ernst Mayr emphasised reproductive isolation, but this, like other species concepts,
4191-451: A very restricted habitat and had been listed as vulnerable by the IUCN , but because the species and its habitat were carefully managed, the species was re-assessed to near threatened in 2009. The superb lyrebird had already been seriously threatened by habitat destruction in the past. Its population had since recovered, but the 2019–2020 bushfires damaged much of its habitat, which may lead to
4318-438: Is "an entity composed of organisms which maintains its identity from other such entities through time and over space, and which has its own independent evolutionary fate and historical tendencies". This differs from the biological species concept in embodying persistence over time. Wiley and Mayden stated that they see the evolutionary species concept as "identical" to Willi Hennig 's species-as-lineages concept, and asserted that
4445-713: Is a state in northeastern Australia , and is the second-largest and third-most populous of the Australian states. It is bordered by the Northern Territory , South Australia and New South Wales to the west, south-west and south, respectively. To the east, Queensland is bordered by the Coral Sea and the Pacific Ocean ; to the state's north is the Torres Strait , separating the Australian mainland from Papua New Guinea , and
4572-400: Is a group of sexually reproducing organisms that recognise one another as potential mates. Expanding on this to allow for post-mating isolation, a cohesion species is the most inclusive population of individuals having the potential for phenotypic cohesion through intrinsic cohesion mechanisms; no matter whether populations can hybridise successfully, they are still distinct cohesion species if
4699-508: Is a mixture of elements of its own song and mimicry of other species. Lyrebirds render with great fidelity the individual songs of other birds and the chatter of flocks of birds, and also mimic other animals such as possums, koalas and dingoes . Lyrebirds have been recorded mimicking human sounds such as a mill whistle, a cross-cut saw, chainsaws , car engines and car alarms , fire alarms , rifle -shots, camera shutters, dogs barking, crying babies, music, mobile phone ring tones, and even
4826-452: Is a population of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring , typically by sexual reproduction . It is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity . Other ways of defining species include their karyotype , DNA sequence, morphology , behaviour, or ecological niche . In addition, paleontologists use
4953-458: Is a set of organisms adapted to a particular set of resources, called a niche, in the environment. According to this concept, populations form the discrete phenetic clusters that we recognise as species because the ecological and evolutionary processes controlling how resources are divided up tend to produce those clusters. A genetic species as defined by Robert Baker and Robert Bradley is a set of genetically isolated interbreeding populations. This
5080-414: Is called speciation . Charles Darwin was the first to describe the role of natural selection in speciation in his 1859 book The Origin of Species . Speciation depends on a measure of reproductive isolation , a reduced gene flow. This occurs most easily in allopatric speciation, where populations are separated geographically and can diverge gradually as mutations accumulate. Reproductive isolation
5207-412: Is called the specific name or the specific epithet (in botanical nomenclature , also sometimes in zoological nomenclature ). For example, Boa constrictor is one of the species of the genus Boa , with constrictor being the species' epithet. While the definitions given above may seem adequate at first glance, when looked at more closely they represent problematic species concepts. For example,
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5334-403: Is described formally, in a publication that assigns it a unique scientific name. The description typically provides means for identifying the new species, which may not be based solely on morphology (see cryptic species ), differentiating it from other previously described and related or confusable species and provides a validly published name (in botany) or an available name (in zoology) when
5461-687: Is found in areas of rainforest in Victoria , New South Wales , and south-east Queensland . It is also found in Tasmania where it was introduced in the 19th century. Many superb lyrebirds live in the Dandenong Ranges National Park and Kinglake National Park around Melbourne , the Royal National Park and Illawarra region south of Sydney, in many other parks along the east coast of Australia, and non protected bushland. Albert's lyrebird
5588-544: Is found only in a small area of Southern Queensland rainforest . Lyrebirds are shy and difficult to approach, particularly the Albert's lyrebird, with the result that little information about its behaviour has been documented. When lyrebirds detect potential danger, they pause and scan the surroundings, sound an alarm, and either flee the area on foot, or seek cover and freeze. Firefighters sheltering in mine shafts during bushfires have been joined by lyrebirds. Lyrebirds feed on
5715-671: Is further weakened by the existence of microspecies , groups of organisms, including many plants, with very little genetic variability, usually forming species aggregates . For example, the dandelion Taraxacum officinale and the blackberry Rubus fruticosus are aggregates with many microspecies—perhaps 400 in the case of the blackberry and over 200 in the dandelion, complicated by hybridisation , apomixis and polyploidy , making gene flow between populations difficult to determine, and their taxonomy debatable. Species complexes occur in insects such as Heliconius butterflies, vertebrates such as Hypsiboas treefrogs, and fungi such as
5842-726: Is hard or even impossible to test. Later biologists have tried to refine Mayr's definition with the recognition and cohesion concepts, among others. Many of the concepts are quite similar or overlap, so they are not easy to count: the biologist R. L. Mayden recorded about 24 concepts, and the philosopher of science John Wilkins counted 26. Wilkins further grouped the species concepts into seven basic kinds of concepts: (1) agamospecies for asexual organisms (2) biospecies for reproductively isolated sexual organisms (3) ecospecies based on ecological niches (4) evolutionary species based on lineage (5) genetic species based on gene pool (6) morphospecies based on form or phenotype and (7) taxonomic species,
5969-477: Is home to the Torres Strait Islander peoples . Torres Strait Islanders are ethnically and culturally distinct from mainland Aboriginal peoples. They have a long history of interaction with both Aboriginal peoples of what is now Australia and the peoples of New Guinea . In February 1606, Dutch navigator Willem Janszoon landed near the site of what is now Weipa , on the western shore of Cape York . This
6096-403: Is no easy way to tell whether related geographic or temporal forms belong to the same or different species. Species gaps can be verified only locally and at a point of time. One is forced to admit that Darwin's insight is correct: any local reality or integrity of species is greatly reduced over large geographic ranges and time periods. The botanist Brent Mishler argued that the species concept
6223-468: Is not valid, notably because gene flux decreases gradually rather than in discrete steps, which hampers objective delimitation of species. Indeed, complex and unstable patterns of gene flux have been observed in cichlid teleosts of the East African Great Lakes . Wilkins argued that "if we were being true to evolution and the consequent phylogenetic approach to taxa, we should replace it with
6350-586: Is similar to Mayr's Biological Species Concept, but stresses genetic rather than reproductive isolation. In the 21st century, a genetic species could be established by comparing DNA sequences. Earlier, other methods were available, such as comparing karyotypes (sets of chromosomes ) and allozymes ( enzyme variants). An evolutionarily significant unit (ESU) or "wildlife species" is a population of organisms considered distinct for purposes of conservation. In palaeontology , with only comparative anatomy (morphology) and histology from fossils as evidence,
6477-488: Is sometimes an important source of genetic variation. Viruses can transfer genes between species. Bacteria can exchange plasmids with bacteria of other species, including some apparently distantly related ones in different phylogenetic domains , making analysis of their relationships difficult, and weakening the concept of a bacterial species. Queensland Queensland ( locally / ˈ k w iː n z l æ n d / KWEENZ -land , commonly abbreviated as Qld )
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#17330848902196604-488: Is the population of superb lyrebirds in Tasmania, which have retained the calls of species not native to Tasmania in their repertoire, with some local Tasmanian endemic bird songs added. The female lyrebirds of both species are also mimics capable of complex vocalisations. Superb lyrebird females are silent during courtship; however, they regularly produce sophisticated vocal displays during foraging and nest defense. A recording of
6731-594: Is threatened by hybridisation, but this can be selected against once a pair of populations have incompatible alleles of the same gene, as described in the Bateson–Dobzhansky–Muller model . A different mechanism, phyletic speciation, involves one lineage gradually changing over time into a new and distinct form (a chronospecies ), without increasing the number of resultant species. Horizontal gene transfer between organisms of different species, either through hybridisation , antigenic shift , or reassortment ,
6858-570: The 138th meridian east , and to the southwest by northeastern South Australia . The state's southern border with New South Wales is constituted in the east by the watershed from Point Danger to the Dumaresq River , and the Dumaresq, Macintyre and Barwon rivers. The west of the southern border is defined by the 29th parallel south (including some minor historical encroachments ) until it reaches South Australia. Like much of eastern Australia,
6985-619: The Australian frontier wars of the 19th century, colonists killed tens of thousands of Aboriginal people in Queensland while consolidating their control over the territory. On 6 June 1859 (now commemorated as Queensland Day ), Queen Victoria signed the letters patent to establish the colony of Queensland, separating it from New South Wales and thereby establishing Queensland as a self-governing Crown colony with responsible government . A large part of colonial Queensland's economy relied on blackbirded South Sea Islander slavery. Queensland
7112-512: The Brisbane River . He returned in 1824 and established a penal settlement at what is now Redcliffe . The settlement, initially known as Edenglassie , was then transferred to the current location of the Brisbane city centre . Edmund Lockyer discovered outcrops of coal along the banks of the upper Brisbane River in 1825. In 1839 transportation of convicts was ceased, culminating in the closure of
7239-609: The Gayiri Aboriginal people in response. Frontier violence peaked on the northern mining frontier during the 1870s, most notably in Cook district and on the Palmer and Hodgkinson River goldfields, with heavy loss of Aboriginal lives and several well-known massacres. Raids conducted by the Kalkadoon held settlers out of Western Queensland for ten years until September 1884 when they attacked
7366-601: The Great Dividing Range runs roughly parallel with, and inland from, the coast, and areas west of the range are more arid than the humid coastal regions. The Great Barrier Reef , which is the world's largest coral reef system, runs parallel to the state's Coral Sea coast between the Torres Strait and K'gari (Fraser Island) . Queensland's coastline includes the world's three largest sand islands: K'gari (Fraser Island) , Moreton , and North Stradbroke . The state contains six World Heritage -listed preservation areas:
7493-538: The Gulf of Carpentaria to the north-west. With an area of 1,723,030 square kilometres (665,270 sq mi), Queensland is the world's sixth-largest subnational entity ; it is larger than all but 16 countries . Due to its size, Queensland's geographical features and climates are diverse, and include tropical rainforests , rivers , coral reefs , mountain ranges and white sandy beaches in its tropical and sub-tropical coastal regions, as well as deserts and savanna in
7620-523: The ICZN for animals and the ICN for plants, do not make rules for defining the boundaries of the species. Research can change the boundaries, also known as circumscription, based on new evidence. Species may then need to be distinguished by the boundary definitions used, and in such cases the names may be qualified with sensu stricto ("in the narrow sense") to denote usage in the exact meaning given by an author such as
7747-619: The Kilcoy and Whiteside poisonings, each of which was said to have taken up to 70 Aboriginal lives by use of a gift of flour laced with strychnine . At the Battle of One Tree Hill in September 1843, Multuggerah and his group of warriors ambushed one group of settlers, routing them and subsequently others in the skirmishes which followed, starting in retaliation for the Kilcoy poisoning. Central Queensland
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#17330848902197874-642: The Kingdom of Great Britain on 22 August 1770 at Possession Island , naming eastern Australia, including Queensland, New South Wales . The Aboriginal population declined significantly after a smallpox epidemic during the late 18th century and massacres by the European settlers. In 1823, John Oxley , a British explorer, sailed north from what is now Sydney to scout possible penal colony sites in Gladstone (then Port Curtis ) and Moreton Bay . At Moreton Bay, he found
8001-537: The Sunshine Coast , Townsville , Cairns , Ipswich , and Toowoomba . 24.2% of the state's population were born overseas . The state has the highest inter-state net migration in Australia. Queensland was first inhabited by Aboriginal Australians , with the Torres Strait Islands inhabited by Torres Strait Islanders . Dutch navigator Willem Janszoon , the first European to land in Australia, explored
8128-594: The Sunshine State for its tropical and sub-tropical climates, Great Barrier Reef , and numerous beaches, tourism is also important to the state's economy. Queensland was one of the largest regions of pre-colonial Aboriginal population in Australia. The Aboriginal occupation of Queensland is thought to predate 50,000 BC, and early migrants are believed to have arrived via boat or land bridge across Torres Strait . Through time, their descendants developed into more than 90 different language and cultural groups. During
8255-414: The Torres Strait to the north, with Boigu Island off the coast of New Guinea representing the northern extreme of its territory. The triangular Cape York Peninsula , which points toward New Guinea, is the northernmost part of the state's mainland. West of the peninsula's tip, northern Queensland is bordered by the Gulf of Carpentaria . To the west, Queensland is bordered by the Northern Territory , at
8382-616: The White Australia policy came into effect, which saw most foreign workers in Australia deported under the Pacific Island Labourers Act 1901 , which saw the Pacific Islander population of the state decrease rapidly. A public meeting was held in 1851 to consider the proposed separation of Queensland from New South Wales. On 6 June 1859, Queen Victoria signed letters patent to form the separate colony of Queensland as
8509-474: The White Australia policy in 1973 saw the beginning of a wave of immigration from around the world, and most prominently from Asia, which continues to the present. In 1981 the Great Barrier Reef off Queensland's northeast coast, one of the world's largest coral reef systems, was declared a UNESCO World Heritage Site . In 2003 Queensland adopted maroon as the state's official colour. The announcement
8636-399: The fly agaric . Natural hybridisation presents a challenge to the concept of a reproductively isolated species, as fertile hybrids permit gene flow between two populations. For example, the carrion crow Corvus corone and the hooded crow Corvus cornix appear and are classified as separate species, yet they can hybridise where their geographical ranges overlap. A ring species is
8763-498: The jaguar ( Panthera onca ) of Latin America or the leopard ( Panthera pardus ) of Africa and Asia. In contrast, the scientific names of species are chosen to be unique and universal (except for some inter-code homonyms ); they are in two parts used together : the genus as in Puma , and the specific epithet as in concolor . A species is given a taxonomic name when a type specimen
8890-415: The leaf -litter. Lyrebirds are long-lived birds that can live as long as 30 years. They have long breeding cycles and start breeding later in life than other passerine birds. Female superb lyrebirds start breeding at the age of five or six, and males at the age of six to eight. Males defend territories from other males, and those territories may contain the breeding territories of up to eight females. Within
9017-519: The semi-arid and desert climatic regions of its interior . Queensland has a population of over 5.5 million, concentrated along the east coast, particularly in South East Queensland . The capital and largest city in the state is Brisbane , Australia's third-largest city . Ten of Australia's thirty largest cities are located in Queensland, the largest outside Brisbane being the Gold Coast ,
9144-406: The "cynical species concept", and arguing that far from being cynical, it usefully leads to an empirical taxonomy for any given group, based on taxonomists' experience. Other biologists have gone further and argued that we should abandon species entirely, and refer to the "Least Inclusive Taxonomic Units" (LITUs), a view that would be coherent with current evolutionary theory. The species concept
9271-406: The 1930s: "The Keel Row" and "Mosquito's Dance". Musicologist David Rothenberg has endorsed this information. However, a "flute lyrebird" research group (including Curtis and Fenton) formed to investigate the veracity of this story found no evidence of "Mosquito Dance" and only remnants of "Keel Row" in contemporary and historical lyrebird recordings from this area. Neither were they able to prove that
9398-634: The Brisbane penal settlement. In 1842 free settlement, which had already commenced, was officially permitted. In 1847, the Port of Maryborough was opened as a wool port. While most early immigrants came from New South Wales, the first free immigrant ship to arrive in Moreton Bay from Europe was the Artemisia , in 1848. Earlier than this immigrant ship was the arrival of the Irish famine orphan girls to Queensland. Devised by
9525-592: The Feisty Colleens, never set foot on Sydney soil, and instead sailed up to Brisbane (then Moreton Bay) on 21 October 1848 on board the Ann Mary . This scheme continued until 1852. In 1857, Queensland's first lighthouse was built at Cape Moreton . The frontier wars fought between European settlers and Aboriginal tribes in Queensland were the bloodiest and most brutal in colonial Australia. Many of these conflicts are now seen as acts of genocide. The wars featured
9652-1013: The Great Barrier Reef along the Coral Sea coast, K'gari (Fraser Island) on the Wide Bay–Burnett region's coastline, the wet tropics in Far North Queensland including the Daintree Rainforest , Lamington National Park in South East Queensland , the Riversleigh fossil sites in North West Queensland , and the Gondwana Rainforests in South East Queensland. The state is divided into several unofficial regions which are commonly used to refer to large areas of
9779-643: The Native Police between 1859 and 1897. The military force of the Queensland Government in this war was the Native Police , who operated from 1849 to the 1920s. The Native Police was a body of Aboriginal and Torres Strait Islander troopers that operated under the command of white officers. The Native Police were often recruited forcefully from far-away communities. Conflict spread quickly with free settlement in 1838, with settlement rapidly expanding in
9906-622: The Origin of Species explained how species could arise by natural selection . That understanding was greatly extended in the 20th century through genetics and population ecology . Genetic variability arises from mutations and recombination , while organisms themselves are mobile, leading to geographical isolation and genetic drift with varying selection pressures . Genes can sometimes be exchanged between species by horizontal gene transfer ; new species can arise rapidly through hybridisation and polyploidy ; and species may become extinct for
10033-405: The abbreviation "sp." in the singular or "spp." (standing for species pluralis , Latin for "multiple species") in the plural in place of the specific name or epithet (e.g. Canis sp.). This commonly occurs when authors are confident that some individuals belong to a particular genus but are not sure to which exact species they belong, as is common in paleontology . Authors may also use "spp." as
10160-408: The age of sixteen. In total, approximately 15,000 South Sea Islander slaves died while working in Queensland, a figure which does not include those who died in transit or who were killed in the recruitment process. This represents a mortality rate of at least 30%, which is high considering most were only on three year contracts. It is also similar to the estimated 33% death rate of enslaved Africans in
10287-570: The amount of hybridisation is insufficient to completely mix their respective gene pools . A further development of the recognition concept is provided by the biosemiotic concept of species. In microbiology , genes can move freely even between distantly related bacteria, possibly extending to the whole bacterial domain. As a rule of thumb, microbiologists have assumed that members of Bacteria or Archaea with 16S ribosomal RNA gene sequences more similar than 97% to each other need to be checked by DNA–DNA hybridisation to decide if they belong to
10414-474: The biological species concept, "the several versions" of the phylogenetic species concept, and the idea that species are of the same kind as higher taxa are not suitable for biodiversity studies (with the intention of estimating the number of species accurately). They further suggested that the concept works for both asexual and sexually-reproducing species. A version of the concept is Kevin de Queiroz 's "General Lineage Concept of Species". An ecological species
10541-505: The biological species concept, a cladistic species does not rely on reproductive isolation – its criteria are independent of processes that are integral in other concepts. Therefore, it applies to asexual lineages. However, it does not always provide clear cut and intuitively satisfying boundaries between taxa, and may require multiple sources of evidence, such as more than one polymorphic locus, to give plausible results. An evolutionary species, suggested by George Gaylord Simpson in 1951,
10668-428: The boundaries between closely related species become unclear with hybridisation , in a species complex of hundreds of similar microspecies , and in a ring species . Also, among organisms that reproduce only asexually , the concept of a reproductive species breaks down, and each clone is potentially a microspecies. Although none of these are entirely satisfactory definitions, and while the concept of species may not be
10795-456: The closure of the state borders. With a total area of 1,729,742 square kilometres (715,309 square miles), Queensland is an expansive state with a highly diverse range of climates and geographical features. If Queensland were an independent nation, it would be the world's 16th largest. Queensland's eastern coastline borders the Coral Sea , an arm of the Pacific Ocean. The state is bordered by
10922-642: The colony's agricultural plantations through a process known as blackbirding . This trade in what were then known as Kanakas was in operation from 1863 to 1908, a period of 45 years. Some 55,000 to 62,500 were brought to Australia, most being recruited or blackbirded from islands in Melanesia , such as the New Hebrides (now Vanuatu ), the Solomon Islands and the islands around New Guinea . The majority of those taken were male and around one quarter were under
11049-433: The concept of a chronospecies can be applied. During anagenesis (evolution, not necessarily involving branching), some palaeontologists seek to identify a sequence of species, each one derived from the phyletically extinct one before through continuous, slow and more or less uniform change. In such a time sequence, some palaeontologists assess how much change is required for a morphologically distinct form to be considered
11176-416: The concept of the chronospecies since fossil reproduction cannot be examined. The most recent rigorous estimate for the total number of species of eukaryotes is between 8 and 8.7 million. About 14% of these had been described by 2011. All species (except viruses ) are given a two-part name , called a "binomial". The first part of a binomial is the genus to which the species belongs. The second part
11303-435: The evidence to support hypotheses about evolutionarily divergent lineages that have maintained their hereditary integrity through time and space. Molecular markers may be used to determine diagnostic genetic differences in the nuclear or mitochondrial DNA of various species. For example, in a study done on fungi , studying the nucleotide characters using cladistic species produced the most accurate results in recognising
11430-401: The female, and the female also fosters the chick alone. A lyrebird's song is one of the more distinctive aspects of its behavioural biology. Lyrebirds sing throughout the year, but the peak of the breeding season, from June to August, is when they sing with the most intensity. During this peak males may sing for four hours of the day, almost half the hours of daylight. The song of the lyrebird
11557-507: The first Premier of Queensland . In 1865, the first rail line in the state opened between Ipswich and Grandchester . Queensland's economy expanded rapidly in 1867 after James Nash discovered gold on the Mary River near the town of Gympie , sparking a gold rush and saving the State of Freddy-Mercury-land from near economic collapse. While still significant, they were on a much smaller scale than
11684-683: The first commercial production of oil in Queensland and Australia began at Moonie . During World War II Brisbane became central to the Allied campaign when the AMP Building (now called MacArthur Central ) was used as the South West Pacific headquarters for General Douglas MacArthur , chief of the Allied Pacific forces, until his headquarters were moved to Hollandia in August 1944. In 1942, during
11811-458: The first three years of being taken to America. The trade was legally sanctioned and regulated under Queensland law, and prominent men such as Robert Towns made massive fortunes off of exploitation of slave labour, helping to establish some of the major cities in Queensland today. Towns' agent claimed that blackbirded labourers were "savages who did not know the use of money" and therefore did not deserve cash wages. Following Federation in 1901,
11938-422: The gold rushes of Victoria and New South Wales. Immigration to Australia and Queensland, in particular, began in the 1850s to support the state economy. During the period from the 1860s until the early 20th century, many labourers, known at the time as Kanakas , were brought to Queensland from neighbouring Pacific Island nations to work in the state's sugar cane fields. Some of these people had been kidnapped under
12065-437: The ground and as individuals. A range of invertebrate prey is taken, including insects such as cockroaches , beetles (both adults and larvae), earwigs , fly larvae, and the adults and larvae of moths. Other prey taken includes centipedes , spiders , earthworms . Less commonly taken prey includes stick insects, bugs, amphipods , lizards , frogs and occasionally, seeds . They find food by scratching with their feet through
12192-470: The human voice. However, while the mimicry of human noises is widely reported, the extent to which it happens is exaggerated and the phenomenon is unusual. Parts of the lyrebird's own song can resemble human-made sound effects , which has given rise to the urban legend that they frequently imitate video game or film sounds. The superb lyrebird's mimicked calls are learned from the local environment, including from other superb lyrebirds. An instructive example
12319-437: The largest in the order. They are ground living birds with strong legs and feet and short rounded wings. They are poor fliers and rarely fly except for periods of downhill gliding. The superb lyrebird is the larger of the two species. Lyrebirds measure 31 to 39 inches in length, including their tail. Males tend to be slightly larger than females. Females weigh around 2 pounds, and males weigh around 2.4 pounds. The superb lyrebird
12446-424: The last ice age , Queensland's landscape became more arid and largely desolate, making food and other supplies scarce. The people developed the world's first seed-grinding technology. The end of the glacial period brought about a warming climate, making the land more hospitable. It brought high rainfall along the eastern coast, stimulating the growth of the state's tropical rainforests. The Torres Strait Islands
12573-490: The later decades of the 20th century, the humid subtropical climate —regulated by the availability of air conditioning—saw Queensland become a popular destination for migrants from interstate. Since that time, Queensland has continuously seen high levels of migration from the other states and territories of Australia. In 1966, Lyndon B. Johnson became the first U.S. president to visit Queensland. During his visit, he met with Australia prime minister Harold Holt . The end of
12700-426: The male lyrebird has several. The strength, volume, and location of the nest built by the female lyrebird is dependent on the rainfall and predation during the nest building period. It is important for the nest to be water resistant and hidden in secluded areas so predators cannot attack. Once the nest is made in the preferred location, the female lyrebird lays a single egg . The egg is incubated over 50 days solely by
12827-408: The male territories, the males create or use display platforms; for the superb lyrebird, this is a mound of bare soil; for the Albert's lyrebird, it is a pile of twigs on the forest floor. Male lyrebirds call mostly during winter, when they construct and maintain an open arena-mound in dense bush , on which they sing and dance in an elaborate courtship display performed for potential mates, of which
12954-452: The most frequent massacres of First Nations people, the three deadliest massacres on white settlers, the most disreputable frontier police force, and the highest number of white victims to frontier violence on record in any Australian colony. Across at least 644 collisions at least 66,680 were killed — with Aboriginal fatalities alone comprising no less than 65,180. Of these deaths, around 24,000 Aboriginal men, women and children were killed by
13081-478: The numerous fungi species of all the concepts studied. Versions of the phylogenetic species concept that emphasise monophyly or diagnosability may lead to splitting of existing species, for example in Bovidae , by recognising old subspecies as species, despite the fact that there are no reproductive barriers, and populations may intergrade morphologically. Others have called this approach taxonomic inflation , diluting
13208-399: The only Australian state with a unicameral parliament . In 1935 cane toads were deliberately introduced to Queensland from Hawaii in an unsuccessful attempt to reduce the number of French's cane and greyback cane beetles that were destroying the roots of sugar cane plants, which are integral to Queensland's economy. The toads have remained an environmental pest since that time. In 1962,
13335-469: The outside edges and twelve filamentaries arrayed between them), which was originally thought to resemble a lyre . This happened when a superb lyrebird specimen (which had been taken from Australia to England during the early 19th century) was prepared for display at the British Museum by a taxidermist who had never seen a live lyrebird. The taxidermist mistakenly thought that the tail would resemble
13462-585: The paper is accepted for publication. The type material is usually held in a permanent repository, often the research collection of a major museum or university, that allows independent verification and the means to compare specimens. Describers of new species are asked to choose names that, in the words of the International Code of Zoological Nomenclature , are "appropriate, compact, euphonious, memorable, and do not cause offence". Books and articles sometimes intentionally do not identify species fully, using
13589-674: The person who named the species, while the antonym sensu lato ("in the broad sense") denotes a wider usage, for instance including other subspecies. Other abbreviations such as "auct." ("author"), and qualifiers such as "non" ("not") may be used to further clarify the sense in which the specified authors delineated or described the species. Species are subject to change, whether by evolving into new species, exchanging genes with other species, merging with other species or by becoming extinct. The evolutionary process by which biological populations of sexually-reproducing organisms evolve to become distinct or reproductively isolated as species
13716-487: The result of misclassification leading to questions on whether there really are any ring species. The commonly used names for kinds of organisms are often ambiguous: "cat" could mean the domestic cat, Felis catus , or the cat family, Felidae . Another problem with common names is that they often vary from place to place, so that puma, cougar, catamount, panther, painter and mountain lion all mean Puma concolor in various parts of America, while "panther" may also mean
13843-573: The ring. Ring species thus present a difficulty for any species concept that relies on reproductive isolation. However, ring species are at best rare. Proposed examples include the herring gull – lesser black-backed gull complex around the North pole, the Ensatina eschscholtzii group of 19 populations of salamanders in America, and the greenish warbler in Asia, but many so-called ring species have turned out to be
13970-508: The same species. This concept was narrowed in 2006 to a similarity of 98.7%. The average nucleotide identity (ANI) method quantifies genetic distance between entire genomes , using regions of about 10,000 base pairs . With enough data from genomes of one genus, algorithms can be used to categorize species, as for Pseudomonas avellanae in 2013, and for all sequenced bacteria and archaea since 2020. Observed ANI values among sequences appear to have an "ANI gap" at 85–95%, suggesting that
14097-529: The same species. When two species names are discovered to apply to the same species, the older species name is given priority and usually retained, and the newer name considered as a junior synonym, a process called synonymy . Dividing a taxon into multiple, often new, taxa is called splitting . Taxonomists are often referred to as "lumpers" or "splitters" by their colleagues, depending on their personal approach to recognising differences or commonalities between organisms. The circumscription of taxa, considered
14224-456: The species concept and making taxonomy unstable. Yet others defend this approach, considering "taxonomic inflation" pejorative and labelling the opposing view as "taxonomic conservatism"; claiming it is politically expedient to split species and recognise smaller populations at the species level, because this means they can more easily be included as endangered in the IUCN red list and can attract conservation legislation and funding. Unlike
14351-511: The species was described and named Menura novaehollandiae by John Latham in 1801, and this is the accepted name by virtue of nomenclatural priority . The genus name Menura refers to the pattern of repeated transparent crescents (or "lunules") on the superb lyrebird's outer tail-feathers, from the Ancient Greek words μήνη mēnē "moon" and ουρά ourá "tail". Lyrebirds are named because their outer tail feathers are broad and curved in
14478-636: The tail completely covering his head and back—as can be seen in the image in the " breeding " section of this page, and also the image of the 10-cent coin, where the superb lyrebird's tail (in courtship display) is portrayed accurately. The lyrebird has been featured as a symbol and emblem many times, especially in New South Wales and Victoria (where the superb lyrebird has its natural habitat), and in Queensland (where Albert's lyrebird has its natural habitat). Species A species ( pl. : species)
14605-545: The then British Secretary of State for the Colonies, The Earl Grey Scheme established a special emigration scheme which was designed to resettle destitute girls from the workhouses of Ireland during the Great Famine. The first ship, the "Earl Grey", departed Ireland for a 124-day sail to Sydney. After controversy developed upon their arrival in Australia, a small group of 37 young orphans, sometimes referred to as The Belfast Girls or
14732-400: The tunes into his repertoire, and retained them after release into the park. Neville Fenton forwarded a tape of his recording to Norman Robinson . Because a lyrebird is able to carry two tunes at the same time, Robinson filtered out one of the tunes and put it on the phonograph for the purposes of analysis. One witness suggested that the song represents a modified version of two popular tunes in
14859-560: The vicinity of the New England National Park . Similarly, in 1969, a park ranger, Neville Fenton, recorded a lyrebird song which resembled flute sounds in the New England National Park, near Dorrigo in northern coastal New South Wales. After much detective work by Fenton, it was discovered that in the 1930s, a flute player living on a farm adjoining the park used to play tunes near his pet lyrebird. The lyrebird adopted
14986-428: The war, Brisbane was the site of a violent clash between visiting US military personnel and Australian servicemen and civilians, which resulted in one death and hundreds of injuries. This incident became known colloquially as the Battle of Brisbane . The end of World War II saw a wave of immigration from across Europe, with many more immigrants coming from southern and eastern Europe than in previous decades. In
15113-534: The west coast of the Cape York Peninsula in 1606. In 1770, James Cook claimed the east coast of Australia for the Kingdom of Great Britain . In 1788, Arthur Phillip founded the colony of New South Wales, which included all of what is now Queensland. Queensland was explored in subsequent decades, and the Moreton Bay Penal Settlement was established at Brisbane in 1824 by John Oxley . During
15240-537: The wild. It is difficult to define a species in a way that applies to all organisms. The debate about species concepts is called the species problem. The problem was recognised even in 1859, when Darwin wrote in On the Origin of Species : I was much struck how entirely vague and arbitrary is the distinction between species and varieties. He went on to write: No one definition has satisfied all naturalists; yet every naturalist knows vaguely what he means when he speaks of
15367-582: The world today. World War I had a major impact on Queensland . Over 58,000 Queenslanders fought in World War I and over 10,000 of them died. Australia's first major airline, Qantas (originally standing for "Queensland and Northern Territory Aerial Services"), was founded in Winton in 1920 to serve outback Queensland. In 1922 Queensland abolished the Queensland Legislative Council , becoming
15494-435: Was abandoned when the first chicks, which are altricial , were described. They were not classed with the passerines until a paper was published in 1840, twelve years after they were assigned a discrete family, Menuridae. Within that family they compose a single genus, Menura . It is generally accepted that the lyrebird family is most closely related to the scrub -birds ( Atrichornithidae ) and some authorities combine both in
15621-580: Was among the six colonies which became the founding states of Australia with Federation on 1 January 1901. Since the Bjelke-Petersen era of the late 20th century, Queensland has received a high level of internal migration from the other states and territories of Australia and remains a popular destination for interstate migration. Queensland has the third-largest economy among Australian states, with strengths in mining, agriculture, transportation, international education , insurance, and banking. Nicknamed
15748-570: Was made as a result of an informal tradition to use maroon to represent the state in association with sporting events. After three decades of record population growth, Queensland was impacted by major floods between late 2010 and early 2011 , causing extensive damage and disruption across the state. In 2020 Queensland was impacted by the COVID-19 pandemic . Despite a low number and abrupt decline in cases from April 2020 onward, social distancing requirements were implemented from March 2020 including
15875-552: Was particularly hard hit during the 1860s and 1870s, several contemporary writers mention the Skull Hole, Bladensburg, or Mistake Creek massacre on Bladensburg Station near Winton , which in 1901 was said to have taken up to 200 Aboriginal lives. First Nations warriors killed 19 settlers during the Cullin-La-Ringo massacre on 17 October 1861. In the weeks afterwards, police, native police and civilians killed up to 370 members of
16002-545: Was the first recorded landing of a European in Australia , and it also marked the first reported contact between Europeans and the Aboriginal people of Australia . The region was also explored by French and Spanish explorers (commanded by Louis Antoine de Bougainville and Luís Vaez de Torres , respectively) before the arrival of Lieutenant James Cook in 1770. Cook claimed the east coast under instruction from King George III of
16129-486: Was the subject of much debate after the first specimens reached European scientists after 1798. Based on specimens sent from New South Wales to England, Major-General Thomas Davies illustrated and described this species as the superb lyrebird , which he called Menura superba , in an 1800 presentation to the Linnean Society of London , but this work was not published until 1802; in the intervening time period, however,
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