The Majjistral Nature and History Park is a nature reserve in Mellieha , Malta. The geographic area includes the coastal area in Golden Bay ( Maltese : Il-Mixquqa ) to Il-Prajjet and Ix- Xagħra l-Ħamra . It was declared a national park in the Malta Government Gazette by Legal Notice 251 of 2007 in September 2007. The park is named for the northwestern region or wind ( Maltese : Majjistral ).
64-589: The area includes a stretch of 6 kilometres (3.7 mi) of protected coast, designated as part of a larger Special Area of Conservation of International Importance , within the Natura 2000 network of sites as per the European Habitats Directive . It has a number of historic and archaeological sites as identified in studies commissioned by the Malta Tourism Authority . The Maltese Islands have
128-464: A carbohydrate energy source. The carbohydrate source can be combinations of discrete sugars or can be derived from other sources such as banana , pineapple , peach , or even tomato puree or coconut water . After the preparation of the agar medium, it is poured into test tubes or jars which are then autoclaved (or cooked in a pressure cooker) to sterilize the medium. After cooking, the medium begins to gel as it cools. The taxonomy of this family
192-401: A seta , knocking the pollinator off the flower. After pollination, the sepals and petals fade and wilt, but they usually remain attached to the ovary. In 2011, Bulbophyllum nocturnum was discovered to flower nocturnally. Some species, such as in the genera Phalaenopsis , Dendrobium , and Vanda , produce offshoots or plantlets formed from one of the nodes along the stem , through
256-474: A spur of the labellum ( 8 in the illustration above), or on the point of the sepals, or in the septa of the ovary, the most typical position amongst the Asparagales . In orchids that produce pollinia, pollination happens as some variant of the following sequence: when the pollinator enters into the flower, it touches a viscidium, which promptly sticks to its body, generally on the head or abdomen. While leaving
320-551: A velamen , has the function of absorbing humidity. It is made of dead cells and can have a silvery-grey, white or brown appearance. In some orchids, the velamen includes spongy and fibrous bodies near the passage cells, called tilosomes. The cells of the root epidermis grow at a right angle to the axis of the root to allow them to get a firm grasp on their support. Nutrients for epiphytic orchids mainly come from mineral dust, organic detritus, animal droppings and other substances collecting among on their supporting surfaces. The base of
384-473: A different species of bee, so as to enforce proper cross-pollination. A rare achlorophyllous saprophytic orchid growing entirely underground in Australia, Rhizanthella slateri , is never exposed to light, and depends on ants and other terrestrial insects to pollinate it. Catasetum , a genus discussed briefly by Darwin , actually launches its viscid pollinia with explosive force when an insect touches
448-624: A list of the best wildlife areas containing the habitats and species listed in the Habitats Directive; this list must then be submitted to the European Commission , after which an evaluation and selection process on European level will take place in order to become a Natura 2000 site. The Habitats Directive divides the EU territory into nine biogeographic regions , each with its own ecological coherence. Natura 2000 sites are selected according to
512-522: A rethinking of the habitats assessment might be necessary in the light of a new climate law as part of e.g. the European Green Deal . In European law literature Borgstrom and Kistenkas have discussed some possible future incompatibilities of the Natura 2000 habitats-test with novel EU green infrastructure policies. In the late 2010s the habitats assessment was at the basis of the nitrogen crisis in
576-455: A semiterrestrial or rock-hugging (" lithophyte ") orchid, show a sparkling silver and gold veining on a light green background. The cordate leaves of Psychopsiella limminghei are light brownish-green with maroon-puce markings, created by flower pigments. The attractive mottle of the leaves of lady's slippers from tropical and subtropical Asia ( Paphiopedilum ), is caused by uneven distribution of chlorophyll. Also, Phalaenopsis schilleriana
640-538: A single mass. Each time pollination succeeds, thousands of ovules can be fertilized. Pollinators are often visually attracted by the shape and colours of the labellum. However, some Bulbophyllum species attract male fruit flies ( Bactrocera and Zeugodacus spp.) solely via a floral chemical which simultaneously acts as a floral reward (e.g. methyl eugenol , raspberry ketone , or zingerone ) to perform pollination. The flowers may produce attractive odours. Although absent in most species, nectar may be produced in
704-628: A sticky disc near the top of the column. Just below the pollinia is a second, larger sticky plate called the stigma . The complex mechanisms that orchids have evolved to achieve cross-pollination were investigated by Charles Darwin and described in Fertilisation of Orchids (1862). Orchids have developed highly specialized pollination systems, thus the chances of being pollinated are often scarce, so orchid flowers usually remain receptive for very long periods, rendering unpollinated flowers long-lasting in cultivation. Most orchids deliver pollen in
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#1732883942356768-443: A variety of activities at a stretch of 6 kilometres (3.7 mi) of protected heritage coast. These include educational nature guided walks, cleanups, tree planting , seasonal olive picking, rubble wall construction, control of invasive plant species, guided snorkeling tours, and bird watching . 35°56′46″N 14°20′20″E / 35.946°N 14.339°E / 35.946; 14.339 Natura 2000 Natura 2000
832-698: A variety of habitats that are inhabited by diverse wild native plants. Some plants in the park are also endemic (found only in the Maltese islands), sub-endemic (found also in a few other Mediterranean regions), while others are scarce, localised or rare. While strolling in the Majjistral Park it is possible to observe the local environment, represented by a wide range of plants found in the different habitats: sand dunes , boulder scree , cliffs , rocky steppes and clay slopes, garrigue , temporary freshwater rock pools and maquis shrubland . The best period to observe some of
896-530: A wall or among boulders, shrubs or trees. The painted frog is the only indigenous amphibian in the Maltese Island; it is found in freshwater habitats. During the summer drought it usually hides away in damp places. The painted frog lays hundreds of eggs which hatch into tadpoles that can only live in fresh water. Several species of birds can be seen in the park and at least ten species nest there. Six of these are resident and can be found all year round. They are
960-687: Is a network of nature protection areas in the territory of the European Union . It is made up of Special Areas of Conservation and Special Protection Areas designated under the Habitats Directive and the Birds Directive , respectively. The network includes both terrestrial and Marine Protected Areas . The Natura 2000 network covered more than 18% of the European Union's land area and more than 7% of its marine area in 2022. In May 1992,
1024-418: Is a pastel pink orchid with leaves spotted dark green and light green. The jewel orchid ( Ludisia discolor ) is grown more for its colorful leaves than its white flowers. Some orchids, such as Dendrophylax lindenii (ghost orchid), Aphyllorchis and Taeniophyllum depend on their green roots for photosynthesis and lack normally developed leaves, as do all of the heterotrophic species. Orchids of
1088-437: Is a tool to explore the network and gives access to every standard data form. As of 2017 , Natura 2000 protected 27,312 sites with a terrestrial area of 787,606 km (304,096 sq mi) (around 18 percent of land of the EU countries) and marine area of 360,350 km (139,130 sq mi), and was considered almost complete in the EU terrestrial environment. The process of designation has not always been smooth as
1152-417: Is achieved by removing the pollinia with a small instrument such as a toothpick from the pollen parent and transferring them to the seed parent. Some orchids mainly or totally rely on self-pollination , especially in colder regions where pollinators are particularly rare. The caudicles may dry up if the flower has not been visited by any pollinator, and the pollinia then fall directly on the stigma. Otherwise,
1216-551: Is found in the tropics . Orchidaceae is one of the two largest families of flowering plants, along with the Asteraceae . It contains about 28,000 currently accepted species distributed across 763 genera . The Orchidaceae family encompasses about 6–11% of all species of seed plants . The largest genera are Bulbophyllum (2,000 species), Epidendrum (1,500 species), Dendrobium (1,400 species) and Pleurothallis (1,000 species). It also includes Vanilla (the genus of
1280-542: Is in constant flux, as new studies continue to clarify the relationships between species and groups of species, allowing more taxa at several ranks to be recognized. The Orchidaceae is currently placed in the order Asparagales by the APG III system of 2009. Five subfamilies are recognised. The cladogram below was made according to the APG system of 1998. It represents the view that most botanists had held up to that time. It
1344-592: Is often called a backbulb. Backbulbs still hold nutrition for the plant, but then a pseudobulb usually takes over, exploiting the last reserves accumulated in the backbulb, which eventually dies off, too. A pseudobulb typically lives for about five years. Orchids without noticeable pseudobulbs are also said to have growths, an individual component of a sympodial plant. Like most monocots , orchids generally have simple leaves with parallel veins , although some Vanilloideae have reticulate venation . Leaves may be ovate, lanceolate, or orbiculate, and very variable in size on
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#17328839423561408-419: Is used as a food reserve for wintry periods, and provides for the development of the other one, from which visible growth develops. In warm and constantly humid climates, many terrestrial orchids do not need pseudobulbs. Epiphytic orchids, those that grow upon a support, have modified aerial roots that can sometimes be a few meters long. In the older parts of the roots, a modified spongy epidermis , called
1472-596: The Bern Convention on the conservation of European wildlife and natural habitats . Natura 2000 is also a key contribution to the Program of Work of Protected Areas of the Convention on Biological Diversity . As a prerequisite for joining the EU, accession states have to submit proposals for Natura 2000 sites meeting the same criteria as EU member states. Some new member states have large areas which qualify to be protected under
1536-534: The blue rock thrush , the Spanish sparrow , the zitting cisticola , the Sardinian warbler , the spectacled warbler and the corn bunting . The other breeding birds — the tawny pipit , the short-toed lark , the swallow and the yelkouan shearwater — come to the park to breed and then leave after the nesting season. In the Majjistral Park, as in many other protected areas, the natural landscape has been modified during
1600-416: The natural environment . They produce oxygen and they help to clean the air. African tamarisk : A deciduous shrub or medium-sized tree, reaching up to 5 metres (16 ft) in height. Its flowers are grouped and are white, seen in late winter to early spring. Indigenous wild trees are rare. In the park these can be found near the sea. Barbary nut iris : A small herbaceous perennial that is very common in
1664-454: The vanilla plant ), the type genus Orchis , and many commonly cultivated plants such as Phalaenopsis and Cattleya . Moreover, since the introduction of tropical species into cultivation in the 19th century, horticulturists have produced many hybrids and cultivars . Orchids are easily distinguished from other plants, as they share some very evident derived characteristics or synapomorphies . Among these are: bilateral symmetry of
1728-526: The European Commission committed to raise awareness about the network and biodiversity in general with the public. In order to raise awareness about the Natura 2000 network, 21 May has been designated “Natura 2000 Day”. This precedes “International Day for Biological Diversity” on 22 May. The initiative came from SEO/BirdLife who sought and received funding from the EU LIFE+ programme in order to improve
1792-498: The Habitats Directive), dates back to the days climate change was not an issue and the concept of ecosystem services had not been well developed. Hence, in international law literature Kistenkas raised the question the habitats assessment was originally not designed for striking a balance of all relevant ecosystem services or to involve climate change into the assessment. Although well protecting nature conservation objectives,
1856-678: The Netherlands as the EU Court (C-293/17 and C-294/17, ECLI:EU:C:2018:882) ruled on agriculture nearby Natura 2000 sites. In some other member states, such as Bulgaria , there was criticism a project had not been well communicated in advance. This led to construction in the Kaliakra zone of Bulgaria, including wind turbines and a golf course. Land owners have then been told they may not use their land in certain ways, otherwise non-compliance procedures would be used against Bulgaria. This led to protests by
1920-568: The Punic period. One of the most important features of the park is It-Torri ta’ Għajn Żnuber , also known as Għajn Żnuber Tower . It was not one of the watchtowers erected by the Knights and it is believed to have been built in the late 19th century. It was used as a signal station and lookout post during the Second World War. This tower has now been restored by Il-Majjistral Park. The park offers
1984-428: The accumulation of growth hormones at that point. These shoots are known as keiki . Epipogium aphyllum exhibits a dual reproductive strategy, engaging in both sexual and asexual seed production. The likelihood of apomixis playing a substantial role in successful reproduction appears minimal. Within certain petite orchid species groups, there is a noteworthy preparation of female gametes for fertilization preceding
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2048-499: The act of pollination. The ovary typically develops into a capsule that is dehiscent by three or six longitudinal slits, while remaining closed at both ends. The seeds are generally almost microscopic and very numerous, in some species over a million per capsule. After ripening, they blow off like dust particles or spores. Most orchid species lack endosperm in their seed and must enter symbiotic relationships with various mycorrhizal basidiomyceteous fungi that provide them
2112-460: The agreement of the European Commission become Special Areas of Conservation to be designated for species other than birds, and for habitat types (e.g. particular types of forest, grasslands, wetlands, etc.). Together, Special Protection Areas and Special Areas of Conservation form the Natura 2000 network of protected areas. The Natura 2000 network is the EU contribution to the " Emerald network " of Areas of Special Conservation Interest set up under
2176-541: The anther may rotate and then enter the stigma cavity of the flower (as in Holcoglossum amesianum ). The slipper orchid Paphiopedilum parishii reproduces by self-fertilization . This occurs when the anther changes from a solid to a liquid state and directly contacts the stigma surface without the aid of any pollinating agent or floral assembly. The labellum of the Cypripedioideae is poke bonnet-shaped , and has
2240-522: The centuries by human hands. Ancient populations who lived here left a lot of traces that can testify to their activities. Today they represent a considerable historical heritage that enriches the park with history and culture. Walking along the planned routes is the best way to view the archaeological remains in the park. There are many examples of human intervention in this largely natural area. These include small agricultural holdings and corbelled stone huts, various cart ruts and rock cut tombs dating back to
2304-524: The clay from erosion. Flowers mainly in late spring. Evergreen honeysuckle: A shrub that can be found in the garrigue and in maquis shrubland. Giant fennel : A robust plant with huge yet low aromatic flowers grouped in largo umbels attracting several insects such as wasps. The flowers appear in late winter and spring. Over 350 species of wild plants, some of which are endemic, sub-endemic or rare are found at Il-Majjistral Nature and History Park. The stretches of rocky ground abound with orchids including
2368-425: The conditions in each biogeographical region; thus selected sites represent species and habitat types under similar natural conditions across a suite of countries. Each Natura 2000 site has a unique identification form called a standard data form. This form is used as a legal reference when assessing the management of the species and habitats through the concept of favourable conservation status. The Natura 2000 Viewer
2432-476: The directives, and implementation has not always been simple. The Natura 2000 sites are selected by member states and the European Commission following strictly scientific criteria according to the two directives mentioned above. The Special Protection Areas are designated directly by each EU member state , while the Special Areas of Conservation follow a more elaborate process: each EU member state must compile
2496-1057: The endemic Maltese pyramidal orchid , together with several species of shrubs such as Mediterranean thyme. This shrub gives these rocky grounds of the park a vivid violet at the height of its flowering season in June. At the bottom of the cliffs there is an array of wild plant species including ferns , climber plants such as the black bryony and esparto grass covering the clay slopes. Areas of farmed or fallow land hold field gladiolus and other plants associated with these habitats. Over 1300 plants were planted including olives , sandarac gum tree , evergreen oak , carob , lentisk , wolfbane, myrtle , rockrose, and golden samphire . The park's area includes diverse and unique habitats that are important for many species of fauna including rare and endemic species. These reptiles have been regularly recorded in Il-Majjistral Nature and History Park, suggesting that they are not uncommon. Other reptiles inhabiting
2560-537: The flower ( zygomorphism ), many resupinate flowers, a nearly always highly modified petal (labellum), fused stamens and carpels , and extremely small seeds . All orchids are perennial herbs that lack any permanent woody structure. They can grow according to two patterns: Terrestrial orchids may be rhizomatous or form corms or tubers . The root caps of terrestrial orchids are smooth and white. Some sympodial terrestrial orchids, such as Orchis and Ophrys , have two subterranean tuberous roots . One
2624-536: The flower develops, it undergoes a twisting through 180°, called resupination , so that the labellum lies below the column . The labellum functions to attract insects, and in resupinate flowers, also acts as a landing stage, or sometimes a trap. The reproductive parts of an orchid flower are unique in that the stamens and style are joined to form a single structure, the column . Instead of being released singly, thousands of pollen grains are contained in one or two bundles called pollinia that are attached to
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2688-436: The flower, it pulls the pollinium out of the anther, as it is connected to the viscidium by the caudicle or stipe. The caudicle then bends and the pollinium is moved forwards and downwards. When the pollinator enters another flower of the same species, the pollinium has taken such position that it will stick to the stigma of the second flower, just below the rostellum, pollinating it. In horticulture, artificial orchid pollination
2752-409: The flowers to gather volatile chemicals they require to synthesize pheromonal attractants. Males of such species as Euglossa imperialis or Eulaema meriana have been observed to leave their territories periodically to forage for aromatic compounds, such as cineole, to synthesize pheromone for attracting and mating with females. Each type of orchid places the pollinia on a different body part of
2816-502: The flowers varies from plant to plant. Almost white to dark red with several spots. One of many orchids that can be seen in the park and can be found flowering in late winter. Dodder : This parasitic plant appears as a mass of red filaments. It has small inconspicuous flowers. Common and can be seen growing on several plants such as on the sea squill . Esparto grass : This grass frequently grows on clay forming steppe habitats. The plant spreads by sea and by rhizomes that help to protect
2880-510: The function of trapping visiting insects. The only exit leads to the anthers that deposit pollen on the visitor. In some extremely specialized orchids, such as the Eurasian genus Ophrys , the labellum is adapted to have a colour, shape, and odour which attracts male insects via mimicry of a receptive female. Pollination happens as the insect attempts to mate with flowers. Many neotropical orchids are pollinated by male orchid bees , which visit
2944-430: The genus Corallorhiza (coralroot orchids) lack leaves altogether and instead have symbiotic or parasitic associations with fungal mycelium, though which they absorb sugars. Orchid flowers have three sepals , three petals and a three-chambered ovary . The three sepals and two of the petals are often similar to each other but one petal is usually highly modified, forming a "lip" or labellum . In most orchid genera, as
3008-538: The governments of the European Communities adopted legislation designed to protect the most seriously threatened habitats and species across Europe. The Habitats Directive complements the Birds Directive adopted in 1979, and together they make up the Natura 2000 network of protected areas. The Birds Directive requires the establishment of Special Protection Areas for birds. The Habitats Directive similarly requires Sites of Community Importance which upon
3072-455: The individual plant. Their characteristics are often diagnostic. They are normally alternate on the stem, often folded lengthwise along the centre ("plicate"), and have no stipules . Orchid leaves often have siliceous bodies called stegmata in the vascular bundle sheaths (not present in the Orchidoideae ) and are fibrous. The structure of the leaves corresponds to the specific habitat of
3136-464: The infringement procedures against member states show. While designation of sites may be near complete, the management and enforcement of protection on sites is less advanced and many sites lack management plans. Natura 2000 faced criticism from developers, farmers and politicians who fear that the conservation of habitats and species places a brake on development. In 2013, there were 251,564 km (97,129 sq mi) designated as Natura 2000 in
3200-465: The knowledge of this network. In 2013, the first Natura 2000 day took place with the aim to raise awareness of citizens about the importance of Natura 2000 network in their lives. Since then, every May 21 and the weeks before, awareness actions take place all over Europe. For example, in 2014, school children and politicians across the EU were encouraged to make a butterfly gesture and post photos on social media to raise awareness. The Natura 2000 Award
3264-496: The land-owners. Germany Latvia Natura 2000 in the global context Orchids Orchids are plants that belong to the family Orchidaceae ( / ˌ ɔːr k ɪ ˈ d eɪ s i . iː , - s i . aɪ / ), a diverse and widespread group of flowering plants with blooms that are often colourful and fragrant. Orchids are cosmopolitan plants that are found in almost every habitat on Earth except glaciers . The world's richest diversity of orchid genera and species
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#17328839423563328-401: The largest orchid in the world, Grammatophyllum speciosum (giant orchid), it can reach three meters. Some Dendrobium species have long, canelike pseudobulbs with short, rounded leaves over the whole length; some other orchids have hidden or extremely small pseudobulbs, completely included inside the leaves. With ageing the pseudobulb sheds its leaves and becomes dormant. At this stage it
3392-526: The marine environment. The network in marine areas is not considered complete and acknowledged by the Commission as a "key challenge for EU biodiversity policy in the coming years". Natura 2000 sites can vary considerably in character. They are not strictly protected in terms of how they are allowed to be used by people. Many sites are farmed, forested and some are even in urban areas. Other areas are much wilder. The European Commission developed guidelines on
3456-418: The necessary nutrients to germinate, so almost all orchid species are mycoheterotrophic during germination and reliant upon fungi to complete their lifecycles. Only a handful of orchid species have seed that can germinate without mycorrhiza , namely the species within the genus Disa with hydrochorous seeds. As the chance for a seed to meet a suitable fungus is very small, only a minute fraction of all
3520-450: The park include the Moorish gecko , often seen basking in the sun on walls, ocellated skink , and Western whip snake , the most likely of the snakes to be encountered by day. Judging by the burrows and their droppings, wild rabbits are numerous. They are shy and rarely show up during the day. If lucky, one can also see a top predator – the weasel , hurriedly crossing a path and disappearing in
3584-646: The park. It has attractive bluish flowers that open in the early afternoon and close in the evening. Frequently met with along paths in the park. Blue stonecrop : A small often reddish succulent annual with small flowers. These plants often grow in patches and in shallow rock pools and depressions in the garrigue. Common and flowers in spring. Borage : An annual plant with blue flowers in winter and spring and bristly leaves. Common and often found in disturbed ground. Branched asphodel : An attractive rather robust plant with white flowers that can be seen in winter and spring. Clustered sulla : Flowers in spring colouring
3648-658: The plant. Species that typically bask in sunlight, or grow on sites which can be occasionally very dry, have thick, leathery leaves and the laminae are covered by a waxy cuticle to retain their necessary water supply. Shade-loving species, on the other hand, have long, thin leaves. The leaves of most orchids are perennial, that is, they live for several years, while others, especially those with plicate leaves as in Catasetum , shed them annually and develop new leaves together with new pseudobulbs. The leaves of some orchids are considered ornamental. The leaves of Macodes sanderiana ,
3712-540: The relation between Natura 2000 and wild areas which are thought to make up around 13% of the network. This was in response to a report by Members of the European Parliament in 2009 which called for further protection of Europe's wilderness. The Natura 2000 network is not well known among European Union citizens; only 11% of citizens knew what it was in 2013. As part of the EU Biodiversity Strategy ,
3776-459: The rocky ground with its purplish flowers. It is an annual plant and often grows with stems spreading over the ground. The seeds are found in segmented spiny pods, each segment with one seed each. Common pyramidal orchid : The park's symbol. One of the commonest orchids mainly in garrigue and rocky steppe. Its flowers are grouped in cone-shaped to cylindrical inflorescence, pale or dark pink/purple. Flowers in spring. Conical orchid : The colour of
3840-417: The seeds released grow into adult plants. In cultivation, germination typically takes weeks. Horticultural techniques have been devised for germinating orchid seeds on an artificial nutrient medium, eliminating the requirement of the fungus for germination and greatly aiding the propagation of ornamental orchids. The usual medium for the sowing of orchids in artificial conditions is agar gel combined with
3904-531: The species found in the park is the time of flowering — which for the majority of them extends from late winter to spring. Plants, in particular the endemic ones, are an intrinsic part of Malta's heritage and their preservation demands striking a constant balance between human presence and nature protection. Wild plants are important in providing habitat and food for wildlife, protecting the soil from erosion, and are of great value to human life. They also provide aesthetic value. Wild plants are an important component of
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#17328839423563968-419: The stem of sympodial epiphytes, or in some species essentially the entire stem, may be thickened to form a pseudobulb that contains nutrients and water for drier periods. The pseudobulb typically has a smooth surface with lengthwise grooves, and can have different shapes, often conical or oblong. Its size is very variable; in some small species of Bulbophyllum , it is no longer than two millimeters, while in
4032-726: Was launched by the European Commission in 2013, with the first prizes awarded on Natura 2000 Day in 2014. The annual Award aims to raise awareness about the Natura 2000 network, showcase excellence and encourage networking between people working on Natura 2000 sites. It awards prizes to those working on the management of or to raise awareness about Natura 2000 in five award categories: Conservation; Socio-Economic Benefits; Communication; Reconciling Interests and Perceptions; Cross-Border Cooperation and Networking. In its first year, winners were selected from Bulgaria, Romania, Czech Republic, Belgium and Spain. The Natura 2000 legislation, notably with its habitats-test (or: habitats assessment, Article 6 of
4096-594: Was supported by morphological studies , but never received strong support in molecular phylogenetic studies. Apostasioideae : 2 genera and 16 species, south-eastern Asia Cypripedioideae : 5 genera and 130 species, from the temperate regions of the world, as well as tropical America and tropical Asia Vanilloideae : 15 genera and 180 species, humid tropical and subtropical regions, eastern North America Epidendroideae : more than 500 genera and more or less 20,000 species, cosmopolitan Orchidoideae : 208 genera and 3,630 species, cosmopolitan In 2015,
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