161-411: Light , visible light , or visible radiation is electromagnetic radiation that can be perceived by the human eye . Visible light spans the visible spectrum and is usually defined as having wavelengths in the range of 400–700 nanometres (nm), corresponding to frequencies of 750–420 terahertz . The visible band sits adjacent to the infrared (with longer wavelengths and lower frequencies) and
322-435: A magnetic-dipole –type that dies out with distance from the current. In a similar manner, moving charges pushed apart in a conductor by a changing electrical potential (such as in an antenna) produce an electric-dipole –type electrical field, but this also declines with distance. These fields make up the near field. Neither of these behaviours is responsible for EM radiation. Instead, they only efficiently transfer energy to
483-422: A microwave oven . These interactions produce either electric currents or heat, or both. Like radio and microwave, infrared (IR) also is reflected by metals (and also most EMR, well into the ultraviolet range). However, unlike lower-frequency radio and microwave radiation, Infrared EMR commonly interacts with dipoles present in single molecules, which change as atoms vibrate at the ends of a single chemical bond. It
644-461: A transverse wave , where the electric field E and the magnetic field B are both perpendicular to the direction of wave propagation. The electric and magnetic parts of the field in an electromagnetic wave stand in a fixed ratio of strengths to satisfy the two Maxwell equations that specify how one is produced from the other. In dissipation-less (lossless) media, these E and B fields are also in phase, with both reaching maxima and minima at
805-611: A wave form of the electric and magnetic equations , thus uncovering the wave-like nature of electric and magnetic fields and their symmetry . Because the speed of EM waves predicted by the wave equation coincided with the measured speed of light , Maxwell concluded that light itself is an EM wave. Maxwell's equations were confirmed by Heinrich Hertz through experiments with radio waves. Maxwell's equations established that some charges and currents ( sources ) produce local electromagnetic fields near them that do not radiate. Currents directly produce magnetic fields, but such fields of
966-404: A "pulse theory" and compared the spreading of light to that of waves in water in his 1665 work Micrographia ("Observation IX"). In 1672 Hooke suggested that light's vibrations could be perpendicular to the direction of propagation. Christiaan Huygens (1629–1695) worked out a mathematical wave theory of light in 1678 and published it in his Treatise on Light in 1690. He proposed that light
1127-609: A better representation of how "bright" a light appears to be than raw intensity. They relate to raw power by a quantity called luminous efficacy and are used for purposes like determining how to best achieve sufficient illumination for various tasks in indoor and outdoor settings. The illumination measured by a photocell sensor does not necessarily correspond to what is perceived by the human eye and without filters which may be costly, photocells and charge-coupled devices (CCD) tend to respond to some infrared , ultraviolet or both. Light exerts physical pressure on objects in its path,
1288-553: A body could be so massive that light could not escape from it. In other words, it would become what is now called a black hole . Laplace withdrew his suggestion later, after a wave theory of light became firmly established as the model for light (as has been explained, neither a particle or wave theory is fully correct). A translation of Newton's essay on light appears in The large scale structure of space-time , by Stephen Hawking and George F. R. Ellis . The fact that light could be polarized
1449-575: A bulk collection of charges which are spread out over large numbers of affected atoms. In electrical conductors , such induced bulk movement of charges ( electric currents ) results in absorption of the EMR, or else separations of charges that cause generation of new EMR (effective reflection of the EMR). An example is absorption or emission of radio waves by antennas, or absorption of microwaves by water or other molecules with an electric dipole moment, as for example inside
1610-465: A central band known as the visual streak. Around the fovea extends the central retina for about 6 mm and then the peripheral retina. The farthest edge of the retina is defined by the ora serrata . The distance from one ora to the other (or macula), the most sensitive area along the horizontal meridian , is about 32 mm. In section, the retina is no more than 0.5 mm thick. It has three layers of nerve cells and two of synapses , including
1771-411: A certain minimum frequency, which depended on the particular metal, no current would flow regardless of the intensity. These observations appeared to contradict the wave theory, and for years physicists tried in vain to find an explanation. In 1905, Einstein explained this puzzle by resurrecting the particle theory of light to explain the observed effect. Because of the preponderance of evidence in favor of
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#17328729849591932-605: A component wave is said to be monochromatic . A monochromatic electromagnetic wave can be characterized by its frequency or wavelength, its peak amplitude, its phase relative to some reference phase, its direction of propagation, and its polarization. Interference is the superposition of two or more waves resulting in a new wave pattern. If the fields have components in the same direction, they constructively interfere, while opposite directions cause destructive interference. Additionally, multiple polarization signals can be combined (i.e. interfered) to form new states of polarization, which
2093-579: A considered view that the bird retina depends for nutrition and oxygen supply on a specialized organ, called the "pecten" or pecten oculi , located on the blind spot or optic disk. This organ is extremely rich in blood vessels and is thought to supply nutrition and oxygen to the bird retina by diffusion through the vitreous body. The pecten is highly rich in alkaline phosphatase activity and polarized cells in its bridge portion – both befitting its secretory role. Pecten cells are packed with dark melanin granules, which have been theorized to keep this organ warm with
2254-496: A fluorescence on a nearby plate of coated glass. In one month, he discovered X-rays' main properties. The last portion of the EM spectrum to be discovered was associated with radioactivity . Henri Becquerel found that uranium salts caused fogging of an unexposed photographic plate through a covering paper in a manner similar to X-rays, and Marie Curie discovered that only certain elements gave off these rays of energy, soon discovering
2415-466: A force of about 3.3 piconewtons on the object being illuminated; thus, one could lift a U.S. penny with laser pointers, but doing so would require about 30 billion 1-mW laser pointers. However, in nanometre -scale applications such as nanoelectromechanical systems (NEMS), the effect of light pressure is more significant and exploiting light pressure to drive NEMS mechanisms and to flip nanometre-scale physical switches in integrated circuits
2576-576: A higher energy (and hence shorter wavelength) than gamma rays and vice versa. The origin of the ray differentiates them, gamma rays tend to be natural phenomena originating from the unstable nucleus of an atom and X-rays are electrically generated (and hence man-made) unless they are as a result of bremsstrahlung X-radiation caused by the interaction of fast moving particles (such as beta particles) colliding with certain materials, usually of higher atomic numbers. EM radiation (the designation 'radiation' excludes static electric and magnetic and near fields )
2737-411: A lasting molecular change (a change in conformation) in the visual molecule retinal in the human retina, which change triggers the sensation of vision. There exist animals that are sensitive to various types of infrared, but not by means of quantum-absorption. Infrared sensing in snakes depends on a kind of natural thermal imaging , in which tiny packets of cellular water are raised in temperature by
2898-519: A linear medium such as a vacuum. However, in nonlinear media, such as some crystals , interactions can occur between light and static electric and magnetic fields—these interactions include the Faraday effect and the Kerr effect . In refraction , a wave crossing from one medium to another of different density alters its speed and direction upon entering the new medium. The ratio of the refractive indices of
3059-509: A linear model, this response profile is well described by a difference of Gaussians and is the basis for edge detection algorithms. Beyond this simple difference, ganglion cells are also differentiated by chromatic sensitivity and the type of spatial summation. Cells showing linear spatial summation are termed X cells (also called parvocellular, P, or midget ganglion cells), and those showing non-linear summation are Y cells (also called magnocellular, M, or parasol retinal ganglion cells), although
3220-539: A lower energy level, it emits a photon of light at a frequency corresponding to the energy difference. Since the energy levels of electrons in atoms are discrete, each element and each molecule emits and absorbs its own characteristic frequencies. Immediate photon emission is called fluorescence , a type of photoluminescence . An example is visible light emitted from fluorescent paints, in response to ultraviolet ( blacklight ). Many other fluorescent emissions are known in spectral bands other than visible light. Delayed emission
3381-450: A medium faster than the speed of light in that medium can produce visible Cherenkov radiation . Certain chemicals produce visible radiation by chemoluminescence . In living things, this process is called bioluminescence . For example, fireflies produce light by this means and boats moving through water can disturb plankton which produce a glowing wake. Certain substances produce light when they are illuminated by more energetic radiation,
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#17328729849593542-417: A particular star. Spectroscopy is also used in the determination of the distance of a star, using the red shift . When any wire (or other conducting object such as an antenna ) conducts alternating current , electromagnetic radiation is propagated at the same frequency as the current. As a wave, light is characterized by a velocity (the speed of light ), wavelength , and frequency . As particles, light
3703-424: A phenomenon which can be deduced by Maxwell's equations , but can be more easily explained by the particle nature of light: photons strike and transfer their momentum. Light pressure is equal to the power of the light beam divided by c , the speed of light. Due to the magnitude of c , the effect of light pressure is negligible for everyday objects. For example, a one-milliwatt laser pointer exerts
3864-419: A process known as fluorescence . Some substances emit light slowly after excitation by more energetic radiation. This is known as phosphorescence . Phosphorescent materials can also be excited by bombarding them with subatomic particles. Cathodoluminescence is one example. This mechanism is used in cathode-ray tube television sets and computer monitors . Certain other mechanisms can produce light: When
4025-473: A protein, retinochrome, that recycles retinal and replicates one of the functions of the vertebrate RPE, cephalopod photoreceptors are likely not maintained as well as in vertebrates, and that as a result, the useful lifetime of photoreceptors in invertebrates is much shorter than in vertebrates. Having easily replaced stalk eyes (some lobsters) or retinae (some spiders, such as Deinopis ) rarely occurs. The cephalopod retina does not originate as an outgrowth of
4186-417: A receiver very close to the source, such as inside a transformer . The near field has strong effects its source, with any energy withdrawn by a receiver causing increased load (decreased electrical reactance ) on the source. The near field does not propagate freely into space, carrying energy away without a distance limit, but rather oscillates, returning its energy to the transmitter if it is not absorbed by
4347-467: A receiver. By contrast, the far field is composed of radiation that is free of the transmitter, in the sense that the transmitter requires the same power to send changes in the field out regardless of whether anything absorbs the signal, e.g. a radio station does not need to increase its power when more receivers use the signal. This far part of the electromagnetic field is electromagnetic radiation. The far fields propagate (radiate) without allowing
4508-410: A source. One of Newton's arguments against the wave nature of light was that waves were known to bend around obstacles, while light travelled only in straight lines. He did, however, explain the phenomenon of the diffraction of light (which had been observed by Francesco Grimaldi ) by allowing that a light particle could create a localised wave in the aether . Newton's theory could be used to predict
4669-414: A surface between one transparent material and another. It is described by Snell's Law : where θ 1 is the angle between the ray and the surface normal in the first medium, θ 2 is the angle between the ray and the surface normal in the second medium and n 1 and n 2 are the indices of refraction , n = 1 in a vacuum and n > 1 in a transparent substance . When a beam of light crosses
4830-445: A third type of radiation, which in 1903 Rutherford named gamma rays . In 1910 British physicist William Henry Bragg demonstrated that gamma rays are electromagnetic radiation, not particles, and in 1914 Rutherford and Edward Andrade measured their wavelengths, finding that they were similar to X-rays but with shorter wavelengths and higher frequency, although a 'cross-over' between X and gamma rays makes it possible to have X-rays with
4991-549: A value of 298 000 000 m/s in 1862. Albert A. Michelson conducted experiments on the speed of light from 1877 until his death in 1931. He refined Foucault's methods in 1926 using improved rotating mirrors to measure the time it took light to make a round trip from Mount Wilson to Mount San Antonio in California. The precise measurements yielded a speed of 299 796 000 m/s . The effective velocity of light in various transparent substances containing ordinary matter ,
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5152-431: A very large (ideally infinite) distance from the source. Both types of waves can have a waveform which is an arbitrary time function (so long as it is sufficiently differentiable to conform to the wave equation). As with any time function, this can be decomposed by means of Fourier analysis into its frequency spectrum , or individual sinusoidal components, each of which contains a single frequency, amplitude and phase. Such
5313-464: A wave is its rate of oscillation and is measured in hertz , the SI unit of frequency, where one hertz is equal to one oscillation per second. Light usually has multiple frequencies that sum to form the resultant wave. Different frequencies undergo different angles of refraction, a phenomenon known as dispersion . A monochromatic wave (a wave of a single frequency) consists of successive troughs and crests, and
5474-518: Is quantized and proportional to frequency according to Planck's equation E = hf , where E is the energy per photon, f is the frequency of the photon, and h is the Planck constant . Thus, higher frequency photons have more energy. For example, a 10 Hz gamma ray photon has 10 times the energy of a 10 Hz extremely low frequency radio wave photon. The effects of EMR upon chemical compounds and biological organisms depend both upon
5635-528: Is a lack of one or more of the cone subtypes that causes individuals to have deficiencies in colour vision or various kinds of colour blindness . These individuals are not blind to objects of a particular colour, but are unable to distinguish between colours that can be distinguished by people with normal vision. Humans have this trichromatic vision , while most other mammals lack cones with red sensitive pigment and therefore have poorer dichromatic colour vision. However, some animals have four spectral subtypes, e.g.
5796-472: Is a more subtle affair. Some experiments display both the wave and particle natures of electromagnetic waves, such as the self-interference of a single photon . When a single photon is sent through an interferometer , it passes through both paths, interfering with itself, as waves do, yet is detected by a photomultiplier or other sensitive detector only once. A quantum theory of the interaction between electromagnetic radiation and matter such as electrons
5957-404: Is a stream of photons . Each has an energy related to the frequency of the wave given by Planck's relation E = hf , where E is the energy of the photon, h is the Planck constant , 6.626 × 10 J·s, and f is the frequency of the wave. In a medium (other than vacuum), velocity factor or refractive index are considered, depending on frequency and application. Both of these are ratios of
6118-527: Is also affected by the colour spectrum of light, a process known as photomorphogenesis . The speed of light in vacuum is defined to be exactly 299 792 458 m/s (approximately 186,282 miles per second). The fixed value of the speed of light in SI units results from the fact that the metre is now defined in terms of the speed of light. All forms of electromagnetic radiation move at exactly this same speed in vacuum. Different physicists have attempted to measure
6279-417: Is also available. Changes in retinal blood circulation are seen with aging and exposure to air pollution, and may indicate cardiovascular diseases such as hypertension and atherosclerosis. Determining the equivalent width of arterioles and venules near the optic disc is also a widely used technique to identify cardiovascular risks. The retina translates an optical image into neural impulses starting with
6440-450: Is an active area of research. At larger scales, light pressure can cause asteroids to spin faster, acting on their irregular shapes as on the vanes of a windmill . The possibility of making solar sails that would accelerate spaceships in space is also under investigation. Although the motion of the Crookes radiometer was originally attributed to light pressure, this interpretation
6601-523: Is associated with those EM waves that are free to propagate themselves ("radiate") without the continuing influence of the moving charges that produced them, because they have achieved sufficient distance from those charges. Thus, EMR is sometimes referred to as the far field , while the near field refers to EM fields near the charges and current that directly produced them, specifically electromagnetic induction and electrostatic induction phenomena. In quantum mechanics , an alternate way of viewing EMR
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6762-537: Is called mesopic vision . At mesopic light levels, both the rods and cones are actively contributing pattern information. What contribution the rod information makes to pattern vision under these circumstances is unclear. The response of cones to various wavelengths of light is called their spectral sensitivity. In normal human vision, the spectral sensitivity of a cone falls into one of three subtypes, often called blue, green, and red, but more accurately known as short, medium, and long wavelength-sensitive cone subtypes. It
6923-422: Is called phosphorescence . The modern theory that explains the nature of light includes the notion of wave–particle duality. Together, wave and particle effects fully explain the emission and absorption spectra of EM radiation. The matter-composition of the medium through which the light travels determines the nature of the absorption and emission spectrum. These bands correspond to the allowed energy levels in
7084-428: Is caused by the surface roughness of the reflecting surfaces, and internal scatterance is caused by the difference of refractive index between the particles and medium inside the object. Like transparent objects, translucent objects allow light to transmit through, but translucent objects also scatter certain wavelength of light via internal scatterance. Refraction is the bending of light rays when passing through
7245-563: Is classified by wavelength into radio , microwave , infrared , visible , ultraviolet , X-rays and gamma rays . Arbitrary electromagnetic waves can be expressed by Fourier analysis in terms of sinusoidal waves ( monochromatic radiation ), which in turn can each be classified into these regions of the EMR spectrum. For certain classes of EM waves, the waveform is most usefully treated as random , and then spectral analysis must be done by slightly different mathematical techniques appropriate to random or stochastic processes . In such cases,
7406-724: Is consequently absorbed by a wide range of substances, causing them to increase in temperature as the vibrations dissipate as heat. The same process, run in reverse, causes bulk substances to radiate in the infrared spontaneously (see thermal radiation section below). Infrared radiation is divided into spectral subregions. While different subdivision schemes exist, the spectrum is commonly divided as near-infrared (0.75–1.4 μm), short-wavelength infrared (1.4–3 μm), mid-wavelength infrared (3–8 μm), long-wavelength infrared (8–15 μm) and far infrared (15–1000 μm). Retina The retina (from Latin rete 'net'; pl. retinae or retinas )
7567-729: Is described by the theory of quantum electrodynamics . Electromagnetic waves can be polarized , reflected, refracted, or diffracted , and can interfere with each other. In homogeneous, isotropic media, electromagnetic radiation is a transverse wave , meaning that its oscillations are perpendicular to the direction of energy transfer and travel. It comes from the following equations : ∇ ⋅ E = 0 ∇ ⋅ B = 0 {\displaystyle {\begin{aligned}\nabla \cdot \mathbf {E} &=0\\\nabla \cdot \mathbf {B} &=0\end{aligned}}} These equations predicate that any electromagnetic wave must be
7728-464: Is hyperpolarised. The amount of neurotransmitter released is reduced in bright light and increases as light levels fall. The actual photopigment is bleached away in bright light and only replaced as a chemical process, so in a transition from bright light to darkness the eye can take up to thirty minutes to reach full sensitivity. When thus excited by light, the photoceptor sends a proportional response synaptically to bipolar cells which in turn signal
7889-455: Is incorrect; the characteristic Crookes rotation is the result of a partial vacuum. This should not be confused with the Nichols radiometer , in which the (slight) motion caused by torque (though not enough for full rotation against friction) is directly caused by light pressure. As a consequence of light pressure, Einstein in 1909 predicted the existence of "radiation friction" which would oppose
8050-453: Is known as parallel polarization state generation . The energy in electromagnetic waves is sometimes called radiant energy . An anomaly arose in the late 19th century involving a contradiction between the wave theory of light and measurements of the electromagnetic spectra that were being emitted by thermal radiators known as black bodies . Physicists struggled with this problem unsuccessfully for many years, and it later became known as
8211-640: Is less than in vacuum. For example, the speed of light in water is about 3/4 of that in vacuum. Two independent teams of physicists were said to bring light to a "complete standstill" by passing it through a Bose–Einstein condensate of the element rubidium , one team at Harvard University and the Rowland Institute for Science in Cambridge, Massachusetts and the other at the Harvard–Smithsonian Center for Astrophysics , also in Cambridge. However,
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#17328729849598372-403: Is most enhanced. The choroid supplies about 75% of these nutrients to the retina and the retinal vasculature only 25%. When light strikes 11-cis-retinal (in the disks in the rods and cones), 11-cis-retinal changes to all-trans-retinal which then triggers changes in the opsins. Now, the outer segments do not regenerate the retinal back into the cis- form once it has been changed by light. Instead
8533-438: Is not direct. Since about 150 million receptors and only 1 million optic nerve fibres exist, convergence and thus mixing of signals must occur. Moreover, the horizontal action of the horizontal and amacrine cells can allow one area of the retina to control another (e.g. one stimulus inhibiting another). This inhibition is key to lessening the sum of messages sent to the higher regions of the brain. In some lower vertebrates (e.g.
8694-408: Is one of the fundamental constants of nature. Like all types of electromagnetic radiation, visible light propagates by massless elementary particles called photons that represents the quanta of electromagnetic field, and can be analyzed as both waves and particles . The study of light, known as optics , is an important research area in modern physics . The main source of natural light on Earth
8855-448: Is regarded as the start of modern physical optics. Pierre Gassendi (1592–1655), an atomist, proposed a particle theory of light which was published posthumously in the 1660s. Isaac Newton studied Gassendi's work at an early age and preferred his view to Descartes's theory of the plenum . He stated in his Hypothesis of Light of 1675 that light was composed of corpuscles (particles of matter) which were emitted in all directions from
9016-477: Is that it consists of photons , uncharged elementary particles with zero rest mass which are the quanta of the electromagnetic field , responsible for all electromagnetic interactions. Quantum electrodynamics is the theory of how EMR interacts with matter on an atomic level. Quantum effects provide additional sources of EMR, such as the transition of electrons to lower energy levels in an atom and black-body radiation . The energy of an individual photon
9177-822: Is the Sun . Historically, another important source of light for humans has been fire , from ancient campfires to modern kerosene lamps . With the development of electric lights and power systems , electric lighting has effectively replaced firelight. Generally, electromagnetic radiation (EMR) is classified by wavelength into radio waves , microwaves , infrared , the visible spectrum that we perceive as light, ultraviolet , X-rays and gamma rays . The designation " radiation " excludes static electric , magnetic and near fields . The behavior of EMR depends on its wavelength. Higher frequencies have shorter wavelengths and lower frequencies have longer wavelengths. When EMR interacts with single atoms and molecules, its behavior depends on
9338-486: Is the emission spectrum of nebulae . Rapidly moving electrons are most sharply accelerated when they encounter a region of force, so they are responsible for producing much of the highest frequency electromagnetic radiation observed in nature. These phenomena can aid various chemical determinations for the composition of gases lit from behind (absorption spectra) and for glowing gases (emission spectra). Spectroscopy (for example) determines what chemical elements comprise
9499-432: Is the innermost, light-sensitive layer of tissue of the eye of most vertebrates and some molluscs . The optics of the eye create a focused two-dimensional image of the visual world on the retina, which then processes that image within the retina and sends nerve impulses along the optic nerve to the visual cortex to create visual perception . The retina serves a function which is in many ways analogous to that of
9660-497: The Académie des Sciences in 1817. Siméon Denis Poisson added to Fresnel's mathematical work to produce a convincing argument in favor of the wave theory, helping to overturn Newton's corpuscular theory. By the year 1821, Fresnel was able to show via mathematical methods that polarization could be explained by the wave theory of light if and only if light was entirely transverse, with no longitudinal vibration whatsoever. The weakness of
9821-648: The Planck energy or exceeding it (far too high to have ever been observed) will require new physical theories to describe. When radio waves impinge upon a conductor , they couple to the conductor, travel along it and induce an electric current on the conductor surface by moving the electrons of the conducting material in correlated bunches of charge. Electromagnetic radiation phenomena with wavelengths ranging from as long as one meter to as short as one millimeter are called microwaves; with frequencies between 300 MHz (0.3 GHz) and 300 GHz. At radio and microwave frequencies, EMR interacts with matter largely as
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#17328729849599982-473: The Planck–Einstein equation . In quantum theory (see first quantization ) the energy of the photons is thus directly proportional to the frequency of the EMR wave. Likewise, the momentum p of a photon is also proportional to its frequency and inversely proportional to its wavelength: The source of Einstein's proposal that light was composed of particles (or could act as particles in some circumstances)
10143-428: The aurora borealis offer many clues as to the nature of light. A transparent object allows light to transmit or pass through. Conversely, an opaque object does not allow light to transmit through and instead reflecting or absorbing the light it receives. Most objects do not reflect or transmit light specularly and to some degree scatters the incoming light, which is called glossiness . Surface scatterance
10304-485: The blind spot . In contrast, in the cephalopod retina, the photoreceptors are in front, with processing neurons and capillaries behind them. Because of this, cephalopods do not have a blind spot. Although the overlying neural tissue is partly transparent, and the accompanying glial cells have been shown to act as fibre-optic channels to transport photons directly to the photoreceptors, light scattering does occur. Some vertebrates, including humans, have an area of
10465-414: The brain through the fibres of the optic nerve . Neural signals from the rods and cones undergo processing by other neurons, whose output takes the form of action potentials in retinal ganglion cells whose axons form the optic nerve. In vertebrate embryonic development , the retina and the optic nerve originate as outgrowths of the developing brain, specifically the embryonic diencephalon ; thus,
10626-523: The electromagnetic (EM) field , which propagate through space and carry momentum and electromagnetic radiant energy . Classically , electromagnetic radiation consists of electromagnetic waves , which are synchronized oscillations of electric and magnetic fields . In a vacuum , electromagnetic waves travel at the speed of light , commonly denoted c . There, depending on the frequency of oscillation, different wavelengths of electromagnetic spectrum are produced. In homogeneous, isotropic media,
10787-449: The film or image sensor in a camera . The neural retina consists of several layers of neurons interconnected by synapses and is supported by an outer layer of pigmented epithelial cells. The primary light-sensing cells in the retina are the photoreceptor cells , which are of two types: rods and cones . Rods function mainly in dim light and provide monochromatic vision. Cones function in well-lit conditions and are responsible for
10948-408: The ophthalmic artery bifurcates and supplies the retina via two distinct vascular networks: the choroidal network, which supplies the choroid and the outer retina, and the retinal network, which supplies the retina's inner layer. Although the inverted retina of vertebrates appears counter-intuitive, it is necessary for the proper functioning of the retina. The photoreceptor layer must be embedded in
11109-425: The outer plexiform layer and the inner plexiform layer . In the outer neuropil layer, the rods and cones connect to the vertically running bipolar cells , and the horizontally oriented horizontal cells connect to ganglion cells. The central retina predominantly contains cones, while the peripheral retina predominantly contains rods. In total, the retina has about seven million cones and a hundred million rods. At
11270-412: The photosensitive ganglion cells ; and transmission along the optic nerve. At each synaptic stage, horizontal and amacrine cells also are laterally connected. The optic nerve is a central tract of many axons of ganglion cells connecting primarily to the lateral geniculate body , a visual relay station in the diencephalon (the rear of the forebrain). It also projects to the superior colliculus ,
11431-787: The pigeon ), control of messages is "centrifugal" – that is, one layer can control another, or higher regions of the brain can drive the retinal nerve cells, but in primates, this does not occur. Using optical coherence tomography (OCT), 18 layers can be identified in the retina. The layers and anatomical correlation are: From innermost to outermost, the layers identifiable by OCT are as follows: on OCT anatomical boundaries? references (unclear if it can be observed on OCT) b) Müller cell nuclei (obliquely orientated fibres; not present in mid-peripheral or peripheral retina) Poorly distinguishable from RPE. Previously: "cone outer segment tips line" (COST) homogenous region of variable reflectivity Retinal development begins with
11592-403: The receptive field of the cell. The receptive fields of retinal ganglion cells comprise a central, approximately circular area, where light has one effect on the firing of the cell, and an annular surround, where light has the opposite effect. In ON cells, an increment in light intensity in the centre of the receptive field causes the firing rate to increase. In OFF cells, it makes it decrease. In
11753-431: The reflection of light, but could only explain refraction by incorrectly assuming that light accelerated upon entering a denser medium because the gravitational pull was greater. Newton published the final version of his theory in his Opticks of 1704. His reputation helped the particle theory of light to hold sway during the eighteenth century. The particle theory of light led Pierre-Simon Laplace to argue that
11914-613: The refraction of light in his book Optics . In ancient India , the Hindu schools of Samkhya and Vaisheshika , from around the early centuries AD developed theories on light. According to the Samkhya school, light is one of the five fundamental "subtle" elements ( tanmatra ) out of which emerge the gross elements. The atomicity of these elements is not specifically mentioned and it appears that they were actually taken to be continuous. The Vishnu Purana refers to sunlight as "the seven rays of
12075-425: The retinal ganglion cells . The photoreceptors are also cross-linked by horizontal cells and amacrine cells , which modify the synaptic signal before it reaches the ganglion cells, the neural signals being intermixed and combined. Of the retina's nerve cells, only the retinal ganglion cells and few amacrine cells create action potentials . In the retinal ganglion cells there are two types of response, depending on
12236-431: The suprachiasmatic nucleus , and the nucleus of the optic tract . It passes through the other layers, creating the optic disc in primates. Additional structures, not directly associated with vision, are found as outgrowths of the retina in some vertebrate groups. In birds , the pecten is a vascular structure of complex shape that projects from the retina into the vitreous humour ; it supplies oxygen and nutrients to
12397-502: The ultraviolet (with shorter wavelengths and higher frequencies), called collectively optical radiation . In physics , the term "light" may refer more broadly to electromagnetic radiation of any wavelength, whether visible or not. In this sense, gamma rays , X-rays , microwaves and radio waves are also light. The primary properties of light are intensity , propagation direction, frequency or wavelength spectrum , and polarization . Its speed in vacuum , 299 792 458 m/s ,
12558-418: The ultraviolet catastrophe . In 1900, Max Planck developed a new theory of black-body radiation that explained the observed spectrum. Planck's theory was based on the idea that black bodies emit light (and other electromagnetic radiation) only as discrete bundles or packets of energy. These packets were called quanta . In 1905, Albert Einstein proposed that light quanta be regarded as real particles. Later
12719-520: The N-T axis is coordinated by expression of the forkhead transcription factors FOXD1 and FOXG1 . Additional gradients are formed within the retina. This spatial distribution may aid in proper targeting of RGC axons that function to establish the retinotopic map. The retina is stratified into distinct layers, each containing specific cell types or cellular compartments that have metabolisms with different nutritional requirements. To satisfy these requirements,
12880-488: The absorption of stray light falling on the pecten. This is considered to enhance metabolic rate of the pecten, thereby exporting more nutritive molecules to meet the stringent energy requirements of the retina during long periods of exposure to light. The bifurcations and other physical characteristics of the inner retinal vascular network are known to vary among individuals, and these individual variances have been used for biometric identification and for early detection of
13041-456: The amount of energy per quantum it carries. EMR in the visible light region consists of quanta (called photons ) that are at the lower end of the energies that are capable of causing electronic excitation within molecules, which leads to changes in the bonding or chemistry of the molecule. At the lower end of the visible light spectrum, EMR becomes invisible to humans (infrared) because its photons no longer have enough individual energy to cause
13202-470: The apparent size of images. Magnifying glasses , spectacles , contact lenses , microscopes and refracting telescopes are all examples of this manipulation. There are many sources of light. A body at a given temperature emits a characteristic spectrum of black-body radiation . A simple thermal source is sunlight , the radiation emitted by the chromosphere of the Sun at around 6,000 K (5,730 °C ; 10,340 °F ). Solar radiation peaks in
13363-432: The atoms in the star's atmosphere. A similar phenomenon occurs for emission , which is seen when an emitting gas glows due to excitation of the atoms from any mechanism, including heat. As electrons descend to lower energy levels, a spectrum is emitted that represents the jumps between the energy levels of the electrons, but lines are seen because again emission happens only at particular energies after excitation. An example
13524-413: The atoms. Dark bands in the absorption spectrum are due to the atoms in an intervening medium between source and observer. The atoms absorb certain frequencies of the light between emitter and detector/eye, then emit them in all directions. A dark band appears to the detector, due to the radiation scattered out of the light beam . For instance, dark bands in the light emitted by a distant star are due to
13685-403: The average number of photons in the cube of the relevant wavelength is much smaller than 1. It is not so difficult to experimentally observe non-uniform deposition of energy when light is absorbed, however this alone is not evidence of "particulate" behavior. Rather, it reflects the quantum nature of matter . Demonstrating that the light itself is quantized, not merely its interaction with matter,
13846-599: The beam from the eye travels infinitely fast this is not a problem. In 55 BC, Lucretius , a Roman who carried on the ideas of earlier Greek atomists , wrote that "The light & heat of the sun; these are composed of minute atoms which, when they are shoved off, lose no time in shooting right across the interspace of air in the direction imparted by the shove." (from On the nature of the Universe ). Despite being similar to later particle theories, Lucretius's views were not generally accepted. Ptolemy (c. second century) wrote about
14007-452: The boundary between a vacuum and another medium, or between two different media, the wavelength of the light changes, but the frequency remains constant. If the beam of light is not orthogonal (or rather normal) to the boundary, the change in wavelength results in a change in the direction of the beam. This change of direction is known as refraction . The refractive quality of lenses is frequently used to manipulate light in order to change
14168-424: The brain, as the vertebrate one does. This difference suggests that vertebrate and cephalopod eyes are not homologous , but have evolved separately. From an evolutionary perspective, a more complex structure such as the inverted retina can generally come about as a consequence of two alternate processes - an advantageous "good" compromise between competing functional limitations, or as a historical maladaptive relic of
14329-480: The central retina adapted for high-acuity vision. This area, termed the fovea centralis , is avascular (does not have blood vessels), and has minimal neural tissue in front of the photoreceptors, thereby minimizing light scattering. The cephalopods have a non-inverted retina, which is comparable in resolving power to the eyes of many vertebrates. Squid eyes do not have an analog of the vertebrate retinal pigment epithelium (RPE). Although their photoreceptors contain
14490-411: The centre of the macula is the foveal pit where the cones are narrow and long, and arranged in a hexagonal mosaic , the most dense, in contradistinction to the much fatter cones located more peripherally in the retina. At the foveal pit, the other retinal layers are displaced, before building up along the foveal slope until the rim of the fovea, or parafovea , is reached, which is the thickest portion of
14651-408: The combined energy transfer of many photons. In contrast, high frequency ultraviolet, X-rays and gamma rays are ionizing – individual photons of such high frequency have enough energy to ionize molecules or break chemical bonds . Ionizing radiation can cause chemical reactions and damage living cells beyond simply heating, and can be a health hazard and dangerous. James Clerk Maxwell derived
14812-592: The concept of light is intended to include very-high-energy photons (gamma rays), additional generation mechanisms include: Light is measured with two main alternative sets of units: radiometry consists of measurements of light power at all wavelengths, while photometry measures light with wavelength weighted with respect to a standardized model of human brightness perception. Photometry is useful, for example, to quantify Illumination (lighting) intended for human use. The photometry units are different from most systems of physical units in that they take into account how
14973-485: The convoluted path of organ evolution and transformation. Vision is an important adaptation in higher vertebrates. A third view of the "inverted" vertebrate eye is that it combines two benefits - the maintenance of the photoreceptors mentioned above, and the reduction in light intensity necessary to avoid blinding the photoreceptors, which are based on the extremely sensitive eyes of the ancestors of modern hagfish (fish that live in very deep, dark water). A recent study on
15134-422: The correspondence between X and Y cells (in the cat retina) and P and M cells (in the primate retina) is not as simple as it once seemed. In the transfer of visual signals to the brain, the visual pathway , the retina is vertically divided in two, a temporal (nearer to the temple) half and a nasal (nearer to the nose) half. The axons from the nasal half cross the brain at the optic chiasma to join with axons from
15295-508: The diameter of Earth's orbit. However, its size was not known at that time. If Rømer had known the diameter of the Earth's orbit, he would have calculated a speed of 227 000 000 m/s . Another more accurate measurement of the speed of light was performed in Europe by Hippolyte Fizeau in 1849. Fizeau directed a beam of light at a mirror several kilometers away. A rotating cog wheel was placed in
15456-400: The distance between two adjacent crests or troughs is called the wavelength . Waves of the electromagnetic spectrum vary in size, from very long radio waves longer than a continent to very short gamma rays smaller than atom nuclei. Frequency is inversely proportional to wavelength, according to the equation: where v is the speed of the wave ( c in a vacuum or less in other media), f is
15617-524: The electromagnetic spectrum includes: radio waves , microwaves , infrared , visible light , ultraviolet , X-rays , and gamma rays . Electromagnetic waves are emitted by electrically charged particles undergoing acceleration , and these waves can subsequently interact with other charged particles, exerting force on them. EM waves carry energy, momentum , and angular momentum away from their source particle and can impart those quantities to matter with which they interact. Electromagnetic radiation
15778-447: The electromagnetic vacuum. The behavior of EM radiation and its interaction with matter depends on its frequency, and changes qualitatively as the frequency changes. Lower frequencies have longer wavelengths, and higher frequencies have shorter wavelengths, and are associated with photons of higher energy. There is no fundamental limit known to these wavelengths or energies, at either end of the spectrum, although photons with energies near
15939-525: The establishment of the eye fields mediated by the SHH and SIX3 proteins, with subsequent development of the optic vesicles regulated by the PAX6 and LHX2 proteins. The role of Pax6 in eye development was elegantly demonstrated by Walter Gehring and colleagues, who showed that ectopic expression of Pax6 can lead to eye formation on Drosophila antennae, wings, and legs. The optic vesicle gives rise to three structures:
16100-575: The evolutionary purpose for the inverted retina structure from the APS (American Physical Society) says that "The directional of glial cells helps increase the clarity of human vision. But we also noticed something rather curious: the colours that best passed through the glial cells were green to red, which the eye needs most for daytime vision. The eye usually receives too much blue—and thus has fewer blue-sensitive cones. Further computer simulations showed that green and red are concentrated five to ten times more by
16261-407: The eye, and may also aid in vision. Reptiles have a similar, but much simpler, structure. In adult humans, the entire retina is about 72% of a sphere about 22 mm in diameter. The entire retina contains about 7 million cones and 75 to 150 million rods. The optic disc, a part of the retina sometimes called "the blind spot" because it lacks photoreceptors, is located at the optic papilla , where
16422-492: The eye. Another supporter of the wave theory was Leonhard Euler . He argued in Nova theoria lucis et colorum (1746) that diffraction could more easily be explained by a wave theory. In 1816 André-Marie Ampère gave Augustin-Jean Fresnel an idea that the polarization of light can be explained by the wave theory if light were a transverse wave . Later, Fresnel independently worked out his own wave theory of light and presented it to
16583-433: The eyes and rays from a source such as the sun. In about 300 BC, Euclid wrote Optica , in which he studied the properties of light. Euclid postulated that light travelled in straight lines and he described the laws of reflection and studied them mathematically. He questioned that sight is the result of a beam from the eye, for he asks how one sees the stars immediately, if one closes one's eyes, then opens them at night. If
16744-527: The fields present in the same space due to other causes. Further, as they are vector fields, all magnetic and electric field vectors add together according to vector addition . For example, in optics two or more coherent light waves may interact and by constructive or destructive interference yield a resultant irradiance deviating from the sum of the component irradiances of the individual light waves. The electromagnetic fields of light are not affected by traveling through static electric or magnetic fields in
16905-437: The fifth century BC, Empedocles postulated that everything was composed of four elements ; fire, air, earth and water. He believed that goddess Aphrodite made the human eye out of the four elements and that she lit the fire in the eye which shone out from the eye making sight possible. If this were true, then one could see during the night just as well as during the day, so Empedocles postulated an interaction between rays from
17066-424: The force of pressure acting on the back. Hence, as the resultant of the two forces, there remains a force that counteracts the motion of the plate and that increases with the velocity of the plate. We will call this resultant 'radiation friction' in brief." Usually light momentum is aligned with its direction of motion. However, for example in evanescent waves momentum is transverse to direction of propagation. In
17227-406: The frequency and λ is the wavelength. As waves cross boundaries between different media, their speeds change but their frequencies remain constant. Electromagnetic waves in free space must be solutions of Maxwell's electromagnetic wave equation . Two main classes of solutions are known, namely plane waves and spherical waves. The plane waves may be viewed as the limiting case of spherical waves at
17388-530: The glial cells, and into their respective cones, than blue light. Instead, excess blue light gets scattered to the surrounding rods. This optimization is such that color vision during the day is enhanced, while night-time vision suffers very little". The vertebrate retina has 10 distinct layers. From closest to farthest from the vitreous body: These layers can be grouped into four main processing stages—photoreception; transmission to bipolar cells ; transmission to ganglion cells , which also contain photoreceptors,
17549-420: The human eye responds to light. The cone cells in the human eye are of three types which respond differently across the visible spectrum and the cumulative response peaks at a wavelength of around 555 nm. Therefore, two sources of light which produce the same intensity (W/m) of visible light do not necessarily appear equally bright. The photometry units are designed to take this into account and therefore are
17710-473: The individual frequency components are represented in terms of their power content, and the phase information is not preserved. Such a representation is called the power spectral density of the random process. Random electromagnetic radiation requiring this kind of analysis is, for example, encountered in the interior of stars, and in certain other very wideband forms of radiation such as the Zero point wave field of
17871-418: The infrared and only a fraction in the visible spectrum. The peak of the black-body spectrum is in the deep infrared, at about 10 micrometre wavelength, for relatively cool objects like human beings. As the temperature increases, the peak shifts to shorter wavelengths, producing first a red glow, then a white one and finally a blue-white colour as the peak moves out of the visible part of the spectrum and into
18032-401: The infrared radiation. EMR in this range causes molecular vibration and heating effects, which is how these animals detect it. Above the range of visible light, ultraviolet light becomes invisible to humans, mostly because it is absorbed by the cornea below 360 nm and the internal lens below 400 nm. Furthermore, the rods and cones located in the retina of the human eye cannot detect
18193-540: The intense radiation of radium . The radiation from pitchblende was differentiated into alpha rays ( alpha particles ) and beta rays ( beta particles ) by Ernest Rutherford through simple experimentation in 1899, but these proved to be charged particulate types of radiation. However, in 1900 the French scientist Paul Villard discovered a third neutrally charged and especially penetrating type of radiation from radium, and after he described it, Rutherford realized it must be yet
18354-800: The known speed of light. Maxwell therefore suggested that visible light (as well as invisible infrared and ultraviolet rays by inference) all consisted of propagating disturbances (or radiation) in the electromagnetic field. Radio waves were first produced deliberately by Heinrich Hertz in 1887, using electrical circuits calculated to produce oscillations at a much lower frequency than that of visible light, following recipes for producing oscillating charges and currents suggested by Maxwell's equations. Hertz also developed ways to detect these waves, and produced and characterized what were later termed radio waves and microwaves . Wilhelm Röntgen discovered and named X-rays . After experimenting with high voltages applied to an evacuated tube on 8 November 1895, he noticed
18515-411: The light-sensing cells are in the back of the retina, so that light has to pass through layers of neurons and capillaries before it reaches the photosensitive sections of the rods and cones. The ganglion cells, whose axons form the optic nerve, are at the front of the retina; therefore, the optic nerve must cross through the retina en route to the brain. No photoreceptors are in this region, giving rise to
18676-424: The luminous body, rejecting the "forms" of Ibn al-Haytham and Witelo as well as the "species" of Roger Bacon , Robert Grosseteste and Johannes Kepler . In 1637 he published a theory of the refraction of light that assumed, incorrectly, that light travelled faster in a denser medium than in a less dense medium. Descartes arrived at this conclusion by analogy with the behaviour of sound waves. Although Descartes
18837-447: The media determines the degree of refraction, and is summarized by Snell's law . Light of composite wavelengths (natural sunlight) disperses into a visible spectrum passing through a prism, because of the wavelength-dependent refractive index of the prism material ( dispersion ); that is, each component wave within the composite light is bent a different amount. EM radiation exhibits both wave properties and particle properties at
18998-412: The movement of matter. He wrote, "radiation will exert pressure on both sides of the plate. The forces of pressure exerted on the two sides are equal if the plate is at rest. However, if it is in motion, more radiation will be reflected on the surface that is ahead during the motion (front surface) than on the back surface. The backwardacting force of pressure exerted on the front surface is thus larger than
19159-407: The nearby violet light. Ritter's experiments were an early precursor to what would become photography. Ritter noted that the ultraviolet rays (which at first were called "chemical rays") were capable of causing chemical reactions. In 1862–64 James Clerk Maxwell developed equations for the electromagnetic field which suggested that waves in the field would travel with a speed that was very close to
19320-503: The neural retina, the retinal pigmented epithelium, and the optic stalk. The neural retina contains the retinal progenitor cells (RPCs) that give rise to the seven cell types of the retina. Differentiation begins with the retinal ganglion cells and concludes with production of the Muller glia. Although each cell type differentiates from the RPCs in a sequential order, there is considerable overlap in
19481-517: The onset of disease. The mapping of vascular bifurcations is one of the basic steps in biometric identification. Results of such analyses of retinal blood vessel structure can be evaluated against the ground truth data of vascular bifurcations of retinal fundus images that are obtained from the DRIVE dataset. In addition, the classes of vessels of the DRIVE dataset have also been identified, and an automated method for accurate extraction of these bifurcations
19642-499: The opposite. At that time, the speed of light could not be measured accurately enough to decide which theory was correct. The first to make a sufficiently accurate measurement was Léon Foucault , in 1850. His result supported the wave theory, and the classical particle theory was finally abandoned (only to partly re-emerge in the twentieth century as photons in quantum theory ). Electromagnetic radiation In physics , electromagnetic radiation ( EMR ) consists of waves of
19803-445: The optic nerve are devoted to the fovea. The resolution limit of the fovea has been determined to be around 10,000 points. The information capacity is estimated at 500,000 bits per second (for more information on bits, see information theory ) without colour or around 600,000 bits per second including colour. When the retina sends neural impulses representing an image to the brain, it spatially encodes (compresses) those impulses to fit
19964-478: The optic-nerve fibres leave the eye. It appears as an oval white area of 3 mm . Temporal (in the direction of the temples) to this disc is the macula , at whose centre is the fovea , a pit that is responsible for sharp central vision, but is actually less sensitive to light because of its lack of rods. Human and non-human primates possess one fovea, as opposed to certain bird species, such as hawks, that are bifoviate, and dogs and cats, that possess no fovea, but
20125-626: The oscillations of the two fields are on average perpendicular to each other and perpendicular to the direction of energy and wave propagation, forming a transverse wave . Electromagnetic radiation is commonly referred to as "light", EM, EMR, or electromagnetic waves. The position of an electromagnetic wave within the electromagnetic spectrum can be characterized by either its frequency of oscillation or its wavelength. Electromagnetic waves of different frequency are called by different names since they have different sources and effects on matter. In order of increasing frequency and decreasing wavelength,
20286-401: The particle of light was given the name photon , to correspond with other particles being described around this time, such as the electron and proton . A photon has an energy, E , proportional to its frequency, f , by where h is the Planck constant , λ {\displaystyle \lambda } is the wavelength and c is the speed of light . This is sometimes known as
20447-507: The path of the light beam as it traveled from the source, to the mirror and then returned to its origin. Fizeau found that at a certain rate of rotation, the beam would pass through one gap in the wheel on the way out and the next gap on the way back. Knowing the distance to the mirror, the number of teeth on the wheel and the rate of rotation, Fizeau was able to calculate the speed of light as 313 000 000 m/s . Léon Foucault carried out an experiment which used rotating mirrors to obtain
20608-747: The patterned excitation of the colour-sensitive pigments of its rods and cones, the retina's photoreceptor cells . The excitation is processed by the neural system and various parts of the brain working in parallel to form a representation of the external environment in the brain. The cones respond to bright light and mediate high-resolution colour vision during daylight illumination (also called photopic vision ). The rod responses are saturated at daylight levels and do not contribute to pattern vision. However, rods do respond to dim light and mediate lower-resolution, monochromatic vision under very low levels of illumination (called scotopic vision ). The illumination in most office settings falls between these two levels and
20769-478: The perception of colour through the use of a range of opsins , as well as high-acuity vision used for tasks such as reading. A third type of light-sensing cell, the photosensitive ganglion cell , is important for entrainment of circadian rhythms and reflexive responses such as the pupillary light reflex . Light striking the retina initiates a cascade of chemical and electrical events that ultimately trigger nerve impulses that are sent to various visual centres of
20930-436: The popular description of light being "stopped" in these experiments refers only to light being stored in the excited states of atoms, then re-emitted at an arbitrary later time, as stimulated by a second laser pulse. During the time it had "stopped", it had ceased to be light. The study of light and the interaction of light and matter is termed optics . The observation and study of optical phenomena such as rainbows and
21091-428: The radiation's power and its frequency. EMR of lower energy ultraviolet or lower frequencies (i.e., near ultraviolet , visible light, infrared, microwaves, and radio waves) is non-ionizing because its photons do not individually have enough energy to ionize atoms or molecules or to break chemical bonds . The effect of non-ionizing radiation on chemical systems and living tissue is primarily simply heating, through
21252-470: The red part of the spectrum, through an increase in the temperature recorded with a thermometer . These "calorific rays" were later termed infrared. In 1801, German physicist Johann Wilhelm Ritter discovered ultraviolet in an experiment similar to Herschel's, using sunlight and a glass prism. Ritter noted that invisible rays near the violet edge of a solar spectrum dispersed by a triangular prism darkened silver chloride preparations more quickly than did
21413-458: The resting state the cell is depolarised. The photon causes the retinal bound to the receptor protein to isomerise to trans-retinal . This causes the receptor to activate multiple G-proteins . This in turn causes the Ga-subunit of the protein to activate a phosphodiesterase (PDE6), which degrades cGMP, resulting in the closing of Na+ cyclic nucleotide-gated ion channels (CNGs). Thus the cell
21574-454: The retina is considered part of the central nervous system (CNS) and is actually brain tissue. It is the only part of the CNS that can be visualized noninvasively . Like most of the brain, the retina is isolated from the vascular system by the blood–brain barrier . The retina is the part of the body with the greatest continuous energy demand. The vertebrate retina is inverted in the sense that
21735-450: The retina. The macula has a yellow pigmentation, from screening pigments, and is known as the macula lutea. The area directly surrounding the fovea has the highest density of rods converging on single bipolar cells. Since its cones have a much lesser convergence of signals, the fovea allows for the sharpest vision the eye can attain. Though the rod and cones are a mosaic of sorts, transmission from receptors, to bipolars, to ganglion cells
21896-511: The retinal is pumped out to the surrounding RPE where it is regenerated and transported back into the outer segments of the photoreceptors. This recycling function of the RPE protects the photoreceptors against photo-oxidative damage and allows the photoreceptor cells to have decades-long useful lives. The bird retina is devoid of blood vessels, perhaps to give unobscured passage of light for forming images, thus giving better resolution. It is, therefore,
22057-490: The retinal pigment epithelium (RPE), which performs at least seven vital functions, one of the most obvious being to supply oxygen and other necessary nutrients needed for the photoreceptors to function. The energy requirements of the retina are even greater than that of the brain. This is due to the additional energy needed to continuously renew the photoreceptor outer segments, of which 10% are shed daily. Energy demands are greatest during dark adaptation when its sensitivity
22218-500: The rods and cones. Light is absorbed by the retinal pigment epithelium or the choroid (both of which are opaque). The white blood cells in the capillaries in front of the photoreceptors can be perceived as tiny bright moving dots when looking into blue light. This is known as the blue field entoptic phenomenon (or Scheerer's phenomenon). Between the ganglion-cell layer and the rods and cones are two layers of neuropils , where synaptic contacts are made. The neuropil layers are
22379-412: The same points in space (see illustrations). In the far-field EM radiation which is described by the two source-free Maxwell curl operator equations, a time-change in one type of field is proportional to the curl of the other. These derivatives require that the E and B fields in EMR are in-phase (see mathematics section below). An important aspect of light's nature is its frequency . The frequency of
22540-464: The same time (see wave-particle duality ). Both wave and particle characteristics have been confirmed in many experiments. Wave characteristics are more apparent when EM radiation is measured over relatively large timescales and over large distances while particle characteristics are more evident when measuring small timescales and distances. For example, when electromagnetic radiation is absorbed by matter, particle-like properties will be more obvious when
22701-472: The source, the power density of EM radiation from an isotropic source decreases with the inverse square of the distance from the source; this is called the inverse-square law . This is in contrast to dipole parts of the EM field, the near field, which varies in intensity according to an inverse cube power law, and thus does not transport a conserved amount of energy over distances but instead fades with distance, with its energy (as noted) rapidly returning to
22862-601: The spectrum of each atom. Emission can be spontaneous , as in light-emitting diodes , gas discharge lamps (such as neon lamps and neon signs , mercury-vapor lamps , etc.) and flames (light from the hot gas itself—so, for example, sodium in a gas flame emits characteristic yellow light). Emission can also be stimulated , as in a laser or a microwave maser . Deceleration of a free charged particle, such as an electron , can produce visible radiation: cyclotron radiation , synchrotron radiation and bremsstrahlung radiation are all examples of this. Particles moving through
23023-494: The speed in a medium to speed in a vacuum. Electromagnetic radiation of wavelengths other than those of visible light were discovered in the early 19th century. The discovery of infrared radiation is ascribed to astronomer William Herschel , who published his results in 1800 before the Royal Society of London . Herschel used a glass prism to refract light from the Sun and detected invisible rays that caused heating beyond
23184-435: The speed of light throughout history. Galileo attempted to measure the speed of light in the seventeenth century. An early experiment to measure the speed of light was conducted by Ole Rømer , a Danish physicist, in 1676. Using a telescope , Rømer observed the motions of Jupiter and one of its moons , Io . Noting discrepancies in the apparent period of Io's orbit, he calculated that light takes about 22 minutes to traverse
23345-404: The sun". The Indian Buddhists , such as Dignāga in the fifth century and Dharmakirti in the seventh century, developed a type of atomism that is a philosophy about reality being composed of atomic entities that are momentary flashes of light or energy. They viewed light as being an atomic entity equivalent to energy. René Descartes (1596–1650) held that light was a mechanical property of
23506-450: The temporal half of the other eye before passing into the lateral geniculate body . Although there are more than 130 million retinal receptors, there are only approximately 1.2 million fibres (axons) in the optic nerve. So, a large amount of pre-processing is performed within the retina. The fovea produces the most accurate information. Despite occupying about 0.01% of the visual field (less than 2° of visual angle ), about 10% of axons in
23667-410: The term associated with the changing static electric field of the particle and the magnetic term that results from the particle's uniform velocity are both associated with the near field, and do not comprise electromagnetic radiation. Electric and magnetic fields obey the properties of superposition . Thus, a field due to any particular particle or time-varying electric or magnetic field contributes to
23828-430: The timing of when individual cell types differentiate. The cues that determine a RPC daughter cell fate are coded by multiple transcription factor families including the bHLH and homeodomain factors. In addition to guiding cell fate determination, cues exist in the retina to determine the dorsal-ventral (D-V) and nasal-temporal (N-T) axes. The D-V axis is established by a ventral to dorsal gradient of VAX2 , whereas
23989-475: The transmitter or absorbed by a nearby receiver (such as a transformer secondary coil). In the Liénard–Wiechert potential formulation of the electric and magnetic fields due to motion of a single particle (according to Maxwell's equations), the terms associated with acceleration of the particle are those that are responsible for the part of the field that is regarded as electromagnetic radiation. By contrast,
24150-427: The transmitter to affect them. This causes them to be independent in the sense that their existence and their energy, after they have left the transmitter, is completely independent of both transmitter and receiver. Due to conservation of energy , the amount of power passing through any spherical surface drawn around the source is the same. Because such a surface has an area proportional to the square of its distance from
24311-476: The trout adds an ultraviolet subgroup to short, medium, and long subtypes that are similar to humans. Some fish are sensitive to the polarization of light as well. In the photoreceptors, exposure to light hyperpolarizes the membrane in a series of graded shifts. The outer cell segment contains a photopigment . Inside the cell the normal levels of cyclic guanosine monophosphate (cGMP) keep the Na+ channel open, and thus in
24472-486: The ultraviolet. These colours can be seen when metal is heated to "red hot" or "white hot". Blue-white thermal emission is not often seen, except in stars (the commonly seen pure-blue colour in a gas flame or a welder 's torch is in fact due to molecular emission, notably by CH radicals emitting a wavelength band around 425 nm and is not seen in stars or pure thermal radiation). Atoms emit and absorb light at characteristic energies. This produces " emission lines " in
24633-436: The unique ribbon synapse . The optic nerve carries the ganglion-cell axons to the brain, and the blood vessels that supply the retina. The ganglion cells lie innermost in the eye while the photoreceptive cells lie beyond. Because of this counter-intuitive arrangement, light must first pass through and around the ganglion cells and through the thickness of the retina, (including its capillary vessels, not shown) before reaching
24794-616: The very short (below 360 nm) ultraviolet wavelengths and are in fact damaged by ultraviolet. Many animals with eyes that do not require lenses (such as insects and shrimp) are able to detect ultraviolet, by quantum photon-absorption mechanisms, in much the same chemical way that humans detect visible light. Various sources define visible light as narrowly as 420–680 nm to as broadly as 380–800 nm. Under ideal laboratory conditions, people can see infrared up to at least 1,050 nm; children and young adults may perceive ultraviolet wavelengths down to about 310–313 nm. Plant growth
24955-425: The visible region of the electromagnetic spectrum when plotted in wavelength units, and roughly 44% of the radiation that reaches the ground is visible. Another example is incandescent light bulbs , which emit only around 10% of their energy as visible light and the remainder as infrared. A common thermal light source in history is the glowing solid particles in flames , but these also emit most of their radiation in
25116-480: The wave theory was that light waves, like sound waves, would need a medium for transmission. The existence of the hypothetical substance luminiferous aether proposed by Huygens in 1678 was cast into strong doubt in the late nineteenth century by the Michelson–Morley experiment . Newton's corpuscular theory implied that light would travel faster in a denser medium, while the wave theory of Huygens and others implied
25277-519: The wave theory, however, Einstein's ideas were met initially with great skepticism among established physicists. Eventually Einstein's explanation was accepted as new particle-like behavior of light was observed, such as the Compton effect . As a photon is absorbed by an atom , it excites the atom, elevating an electron to a higher energy level (one that is on average farther from the nucleus). When an electron in an excited molecule or atom descends to
25438-422: Was an experimental anomaly not explained by the wave theory: the photoelectric effect , in which light striking a metal surface ejected electrons from the surface, causing an electric current to flow across an applied voltage . Experimental measurements demonstrated that the energy of individual ejected electrons was proportional to the frequency , rather than the intensity , of the light. Furthermore, below
25599-575: Was emitted in all directions as a series of waves in a medium called the luminiferous aether . As waves are not affected by gravity, it was assumed that they slowed down upon entering a denser medium. The wave theory predicted that light waves could interfere with each other like sound waves (as noted around 1800 by Thomas Young ). Young showed by means of a diffraction experiment that light behaved as waves. He also proposed that different colours were caused by different wavelengths of light and explained colour vision in terms of three-coloured receptors in
25760-432: Was for the first time qualitatively explained by Newton using the particle theory. Étienne-Louis Malus in 1810 created a mathematical particle theory of polarization. Jean-Baptiste Biot in 1812 showed that this theory explained all known phenomena of light polarization. At that time the polarization was considered as the proof of the particle theory. To explain the origin of colours , Robert Hooke (1635–1703) developed
25921-404: Was incorrect about the relative speeds, he was correct in assuming that light behaved like a wave and in concluding that refraction could be explained by the speed of light in different media. Descartes is not the first to use the mechanical analogies but because he clearly asserts that light is only a mechanical property of the luminous body and the transmitting medium, Descartes's theory of light
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