Lac Le Jeune is a lake and provincial park in British Columbia , Canada , located approximately 37 kilometres south of Kamloops and 47 kilometres north of Merritt .
109-803: The lake is located within Lac Le Jeune Provincial Park , a 213-hectare provincial park run by the British Columbia Ministry of the Environment, and is a popular fishing spot for "fighting" Rainbow Trout , which was established in 1956. It is a summer holiday spot with 144 campgrounds and two resorts; the Lac Le Jeune Wilderness Resort and the Lac Le Jeune Resort. The lake has had several names including "Batchelor", "Chuhwels", and "Le Jeune Lake". The current name
218-561: A Linear B clay tablet found in Messenia that dates to between 1450 and 1350 BC, making Greek the world's oldest recorded living language . Among the Indo-European languages, its date of earliest written attestation is matched only by the now-extinct Anatolian languages . The Greek language is conventionally divided into the following periods: In the modern era, the Greek language entered
327-570: A basis for coinages: anthropology , photography , telephony , isomer , biomechanics , cinematography , etc. Together with Latin words , they form the foundation of international scientific and technical vocabulary ; for example, all words ending in -logy ('discourse'). There are many English words of Greek origin . Greek is an independent branch of the Indo-European language family. The ancient language most closely related to it may be ancient Macedonian , which, by most accounts,
436-399: A bed of fine gravel in a riffle above a pool. A female trout clears a redd in the gravel by turning on her side and beating her tail up and down. Female rainbow trout usually produce 2000 to 3000 4-to-5-millimetre ( 5 ⁄ 32 – 3 ⁄ 16 in) eggs per kilogram of weight. During spawning, the eggs fall into spaces between the gravel, and immediately the female begins digging at
545-747: A broad reddish stripe along the lateral line , from gills to the tail, which is most vivid in breeding males. Wild-caught and hatchery -reared forms of the species have been transplanted and introduced for food or sport in at least 45 countries and every continent except Antarctica. Introductions to locations outside their native range in the United States, Southern Europe, Australia, New Zealand, and South America have damaged native fish species. Introduced populations may affect native species by preying on them, out-competing them, transmitting contagious diseases (such as whirling disease ), or hybridizing with closely related species and subspecies. The rainbow trout
654-418: A century ago, but its range has spread and it has appeared in most of Europe, northern Asia, the U.S., South Africa and other countries. In the 1980s, M. cerebralis was found to require Tubifex tubifex (a kind of segmented worm ) to complete its life cycle . The parasite infects its hosts with its cells after piercing them with polar filaments ejected from nematocyst -like capsules. This parasite
763-623: A century of hatchery stocking. Genetic analysis shows that the South Central California Coast and Southern California populations from Malibu Creek north, including the San Gabriel River , Santa Ana River , and San Mateo Creek , are not hatchery strains. Steelhead from Topanga Creek and the Sweetwater River were partly, and those from San Juan Creek completely, of hatchery origin. Genetic analysis has also shown that
872-598: A decline or even elimination of whole cohorts of fish. The parasite M. cerebralis was first recorded in North America in 1956 in Pennsylvania, but until the 1990s, whirling disease was considered a manageable problem affecting only rainbow trout in hatcheries. It eventually became established in the natural waters of the Rocky Mountain states ( Colorado , Wyoming, Utah , Montana , Idaho , New Mexico ), where it
981-508: A faster, more convenient cursive writing style with the use of ink and quill . The Greek alphabet consists of 24 letters, each with an uppercase ( majuscule ) and lowercase ( minuscule ) form. The letter sigma has an additional lowercase form (ς) used in the final position of a word: In addition to the letters, the Greek alphabet features a number of diacritical signs : three different accent marks ( acute , grave , and circumflex ), originally denoting different shapes of pitch accent on
1090-437: A generally appealing flavor. Seafood Watch ranks farmed rainbow trout as a "Best Choice" fish for human consumption. In Montana , it is illegal to sell or market wild-caught rainbow trout, which are legally classified as game fish. The color and flavor of the flesh depend on the diet and freshness of the trout. Farmed trout and some populations of wild trout, especially anadromous steelhead, have reddish or orange flesh as
1199-573: A lesser extent, in the Western Mediterranean in and around colonies such as Massalia , Monoikos , and Mainake . It was also used as the official language of government and religion in the Christian Nubian kingdoms , for most of their history. Greek, in its modern form, is the official language of Greece, where it is spoken by almost the entire population. It is also the official language of Cyprus (nominally alongside Turkish ) and
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#17329139179761308-400: A limited but productive system of compounding and a rich inflectional system. Although its morphological categories have been fairly stable over time, morphological changes are present throughout, particularly in the nominal and verbal systems. The major change in the nominal morphology since the classical stage was the disuse of the dative case (its functions being largely taken over by
1417-453: A mixed syllable structure, permitting complex syllabic onsets but very restricted codas. It has only oral vowels and a fairly stable set of consonantal contrasts . The main phonological changes occurred during the Hellenistic and Roman period (see Koine Greek phonology for details): In all its stages, the morphology of Greek shows an extensive set of productive derivational affixes ,
1526-749: A rainbow trout species that can be completely fed on a vegan diet through genetic selection. Research from a study team led by USDA research geneticist Dr. Ken Overturf concluded that such natural genetic variation of vegan trouts does exist and believe they can produce rainbow trouts that can be completely fed on a 100% plant-based diet. Populations of many rainbow trout subspecies, including anadromous forms (steelhead) of O. m. irideus (coastal rainbow trout) and O. m. gairdneri (Columbia River redband trout), have declined in their native ranges due to over-harvest, habitat loss, disease, invasive species , pollution and hybridization with other subspecies, and some introduced populations, once healthy, have declined for
1635-641: A result of high astaxanthin levels in their diets. Astaxanthin is a powerful antioxidant that may be from a natural source or a synthetic trout feed. Rainbow trout raised to have pinker flesh from a diet high in astaxanthin are sometimes sold in the U.S. with labeling calling them "steelhead". As wild steelhead are in decline in some parts of their range, farmed rainbow trout are viewed as a preferred alternative. In Chile and Norway , rainbow trout farmed in saltwater sea cages are sold labeled as steelhead. Trout can be cooked as soon as they are cleaned, without scaling , skinning , or filleting . If cooked with
1744-682: A similar latitude. The range of coastal rainbow trout ( O. m. irideus ) extends north from the Pacific basin into tributaries of the Bering Sea in northwest Alaska, while forms of the Columbia River redband trout ( O. m. gairdneri ) extend east into the upper Mackenzie River and Peace River watersheds in British Columbia and Alberta, Canada, which eventually drain into the Beaufort Sea , part of
1853-456: A state of diglossia : the coexistence of vernacular and archaizing written forms of the language. What came to be known as the Greek language question was a polarization between two competing varieties of Modern Greek: Dimotiki , the vernacular form of Modern Greek proper, and Katharevousa , meaning 'purified', a compromise between Dimotiki and Ancient Greek developed in the early 19th century that
1962-531: Is a non-profit organization dedicated to the conservation of North American freshwater streams, rivers, and associated upland habitats for trout, salmon, other aquatic species, and people. A typical TU project is the Circle Creek Fish Passage Project, in which access to a spawning stream is being improved for steelhead and other salmonid species. The Wild Salmon Center , an international coalition of Russian, Canadian and U.S. scientists, sponsors
2071-558: Is common. Rainbow trout can also be caught on various live and dead natural baits . Many anglers consider the rainbow trout the hardest-fighting trout species, as this fish is known for leaping when hooked and putting up a powerful struggle. It is considered one of the top five sport fish in North America, and the most important game fish west of the Rocky Mountains . There are tribal commercial fisheries for steelhead in Puget Sound ,
2180-448: Is completely consumed, and fry commence feeding mainly on zooplankton . The growth rate of rainbow trout varies with area, habitat, life history, and quality and quantity of food. As fry grow, they begin to develop "parr" marks or dark vertical bars on their sides. In this juvenile stage, immature trout are often called "parr" because of the marks. These small juvenile trout are sometimes called 'fingerlings' because they are approximately
2289-462: Is damaging several sport fishing rivers. Some streams in the western U.S. lost 90 percent of their trout. Whirling disease threatens recreational fishing, which is important for the tourism industry, a key component of the economies of some U.S. western states. For example, in 2005 anglers in Montana spent approximately $ 196,000,000 in activities directly related to trout fishing in the state. Some of
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#17329139179762398-523: Is evidence North American fishes are unable to digest the tiny but hard shells of the mud snail, and that their presence may result in poor growth outcomes for rainbow trout. The mud snail was first detected in the U.S. in Idaho's Snake River in 1987. Since then, the snail has spread to the Madison River , Firehole River, and other watercourses around Yellowstone National Park, and has been discovered throughout
2507-483: Is included in the list of the top 100 globally invasive species . Other introductions into waters previously devoid of fish or with severely depleted stocks of native fish have created sport fisheries , such as the Great Lakes and Wyoming 's Firehole River . Some local populations of specific subspecies, or in the case of steelhead, distinct population segments , are listed as either threatened or endangered under
2616-548: Is isolated in Lake Crescent (Washington), is threatened by the loss of its only spawning grounds in the Lyre River to siltation and other types of habitat degradation. Myxobolus cerebralis is a myxosporean parasite of salmonids (salmon, trout, and their allies) that causes whirling disease in pen farmed salmon and trout and also in wild fish populations . It was first described in rainbow trout introduced to Germany
2725-659: Is native to Greece , Cyprus , Italy (in Calabria and Salento ), southern Albania , and other regions of the Balkans , Caucasus , the Black Sea coast, Asia Minor , and the Eastern Mediterranean . It has the longest documented history of any Indo-European language, spanning at least 3,400 years of written records. Its writing system is the Greek alphabet , which has been used for approximately 2,800 years; previously, Greek
2834-559: Is popular in Western cuisine ; both wild-caught and farmed fish are eaten. It has tender flesh and a mild, somewhat nutty flavor. Wild fish has a stronger, gamier taste than farmed fish. While the taste of wild-caught trout is often promoted as superior, rainbow trout and "steelhead" sold in American restaurants is typically farmed . Farmed rainbow trout are considered one of the safest fish to eat and are noted for high levels of vitamin B and
2943-758: Is still used internationally for the writing of Ancient Greek . In Greek, the question mark is written as the English semicolon, while the functions of the colon and semicolon are performed by a raised point (•), known as the ano teleia ( άνω τελεία ). In Greek the comma also functions as a silent letter in a handful of Greek words, principally distinguishing ό,τι ( ó,ti , 'whatever') from ότι ( óti , 'that'). Ancient Greek texts often used scriptio continua ('continuous writing'), which means that ancient authors and scribes would write word after word with no spaces or punctuation between words to differentiate or mark boundaries. Boustrophedon , or bi-directional text,
3052-611: Is to modern spoken English ". Greek is spoken today by at least 13 million people, principally in Greece and Cyprus along with a sizable Greek-speaking minority in Albania near the Greek-Albanian border. A significant percentage of Albania's population has knowledge of the Greek language due in part to the Albanian wave of immigration to Greece in the 1980s and '90s and the Greek community in
3161-544: The British Overseas Territory of Akrotiri and Dhekelia (alongside English ). Because of the membership of Greece and Cyprus in the European Union, Greek is one of the organization's 24 official languages . Greek is recognized as a minority language in Albania, and used co-officially in some of its municipalities, in the districts of Gjirokastër and Sarandë . It is also an official minority language in
3270-600: The Columbia River who provided Richardson with specimens. In 1855, William P. Gibbons, the curator of Geology and Mineralogy at the California Academy of Sciences , found a population and named it Salmo iridia (Latin: rainbow), later corrected to Salmo irideus . These names faded once it was determined that Walbaum's description of type specimens was conspecific and therefore had precedence . In 1989, morphological and genetic studies indicated that trout of
3379-610: The Endangered Species Act . The steelhead is the official state fish of Washington . The scientific name of the rainbow trout is Oncorhynchus mykiss . The species was originally named by German naturalist and taxonomist Johann Julius Walbaum in 1792 based on type specimens from the Kamchatka Peninsula in Siberia. Walbaum's original species name, mykiss , was derived from the local Kamchatkan name used for
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3488-637: The European canon . Greek is also the language in which many of the foundational texts in science and philosophy were originally composed. The New Testament of the Christian Bible was also originally written in Greek. Together with the Latin texts and traditions of the Roman world , the Greek texts and Greek societies of antiquity constitute the objects of study of the discipline of Classics . During antiquity , Greek
3597-510: The Indo-Iranian languages (see Graeco-Aryan ), but little definitive evidence has been found. In addition, Albanian has also been considered somewhat related to Greek and Armenian, and it has been proposed that they all form a higher-order subgroup along with other extinct languages of the ancient Balkans; this higher-order subgroup is usually termed Palaeo-Balkan , and Greek has a central position in it. Linear B , attested as early as
3706-581: The Pacific Basin were genetically closer to Pacific salmon ( Oncorhynchus species) than to the Salmo s – brown trout ( Salmo trutta ) or Atlantic salmon ( Salmo salar ) of the Atlantic Basin . Thus, in 1989, taxonomic authorities moved the rainbow, cutthroat , and other Pacific Basin trout into the genus Oncorhynchus . Walbaum's name had precedence, so the species name Oncorhynchus mykiss became
3815-633: The United Kingdom , and throughout the European Union , especially in Germany . Historically, significant Greek-speaking communities and regions were found throughout the Eastern Mediterranean , in what are today Southern Italy , Turkey , Cyprus , Syria , Lebanon , Israel , Palestine , Egypt , and Libya ; in the area of the Black Sea , in what are today Turkey, Bulgaria , Romania , Ukraine , Russia , Georgia , Armenia , and Azerbaijan ; and, to
3924-496: The Washington coast, and in the Columbia River , but there has been controversy regarding overharvesting of native stocks . The highly desirable sporting qualities and adaptability of the rainbow trout to hatchery rearing and new habitats resulted in it being introduced to many countries around the world by or at the behest of sport fishermen. Many of these introductions have resulted in environmental and ecological problems, as
4033-694: The adipose fin is a management tool used to identify hatchery-reared fish. Rainbow trout, including steelhead forms, generally spawn in early to late spring (January to June in the Northern Hemisphere and September to November in the Southern Hemisphere) when water temperatures reach at least 6 to 7 °C (42 to 44 °F). The maximum recorded lifespan for a rainbow trout is 11 years. Freshwater resident rainbow trout usually inhabit and spawn in small to moderately large, well-oxygenated, shallow rivers with gravel bottoms. They are native to
4142-518: The alluvial or freestone streams that are typical tributaries of the Pacific basin but introduced rainbow trout have established wild, self-sustaining populations in other river types such as bedrock and spring creeks . Lake-resident rainbow trout are usually found in moderately deep, cool lakes with adequate shallows and vegetation to support the production of sufficient food sources. Lake populations generally require access to gravelly-bottomed streams to be self-sustaining. Spawning sites are usually
4251-468: The genitive ). The verbal system has lost the infinitive , the synthetically -formed future, and perfect tenses and the optative mood . Many have been replaced by periphrastic ( analytical ) forms. Pronouns show distinctions in person (1st, 2nd, and 3rd), number (singular, dual , and plural in the ancient language; singular and plural alone in later stages), and gender (masculine, feminine, and neuter), and decline for case (from six cases in
4360-675: The mitochondria are the most important organelle contributing to tissue damage because of their role in metabolism and production of reactive oxygen species . In a study done by Almaida-Pagàn et al., researchers identified changes to the heart and brain mitochondrial membrane phospholipid composition in rainbow trout due to the differing levels of oxidative stress each organ faced during a high-stress time such as rapid growth and development. Stock rainbow trout of ages 1, 2, and 4 years had their heart and brain mitochondria isolated and analyzed for fatty acid composition. The tissues showed an overall similar percentage of total phospholipids but differed in
4469-417: The syntax of Greek have remained constant: verbs agree with their subject only, the use of the surviving cases is largely intact (nominative for subjects and predicates, accusative for objects of most verbs and many prepositions, genitive for possessors), articles precede nouns, adpositions are largely prepositional, relative clauses follow the noun they modify and relative pronouns are clause-initial. However,
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4578-566: The 20th century on), especially from French and English, are typically not inflected; other modern borrowings are derived from Albanian , South Slavic ( Macedonian / Bulgarian ) and Eastern Romance languages ( Aromanian and Megleno-Romanian ). Greek words have been widely borrowed into other languages, including English. Example words include: mathematics , physics , astronomy , democracy , philosophy , athletics , theatre, rhetoric , baptism , evangelist , etc. Moreover, Greek words and word elements continue to be productive as
4687-537: The 3,400 km (2,100 mi) river system. Neither method of control is 100% effective and are best regarded as methods to change the relative population sizes of fish species. Steelhead populations in parts of its native range have declined due to a variety of human and natural causes. While populations in Alaska and along the British Columbia coast are considered healthy, populations in Kamchatka and some populations along
4796-493: The Arctic Ocean. Since 1875, the rainbow trout has been widely introduced into suitable lacustrine and riverine environments throughout the United States and around the world. Many of these introductions have established wild, self-sustaining populations. Since 1870, rainbow trout have been artificially propagated in fish hatcheries to restock streams and introduce them into non-native waters. The first rainbow trout hatchery
4905-669: The Garmat Ali River in Iraq since 2008 ), Europe (since 1859 in England), and North America (U.S. and Canada: Thunder Bay in Ontario since 2001, British Columbia since July 2007 ), most likely inadvertently during human activity. It can reach concentrations greater than 500,000 per square metre (46,000/sq ft), endangering the food chain by outcompeting native snails and water insects for food, leading to sharp declines in native populations. There
5014-399: The Great Lakes is as high as 70 percent. As young steelhead transition from freshwater to saltwater, a process called " smoltification " occurs where the trout undergoes physiological changes to allow it to survive in seawater. There are genetic differences between freshwater and steelhead populations that may account for the smoltification in steelhead. Juvenile steelhead may remain in
5123-538: The Greek alphabet since approximately the 9th century BC. It was created by modifying the Phoenician alphabet , with the innovation of adopting certain letters to represent the vowels. The variant of the alphabet in use today is essentially the late Ionic variant, introduced for writing classical Attic in 403 BC. In classical Greek, as in classical Latin, only upper-case letters existed. The lower-case Greek letters were developed much later by medieval scribes to permit
5232-540: The Greek language was the Cypriot syllabary (also a descendant of Linear A via the intermediate Cypro-Minoan syllabary ), which is closely related to Linear B but uses somewhat different syllabic conventions to represent phoneme sequences. The Cypriot syllabary is attested in Cyprus from the 11th century BC until its gradual abandonment in the late Classical period, in favor of the standard Greek alphabet. Greek has been written in
5341-559: The Kamchatka Steelhead Project, a 20-year (1994–2014) scientific program to study and conserve the present condition of Kamchatkan steelhead ("mikizha"), a species listed in the Red Data Book of Russia . Other high-profile organizations involved in rainbow trout conservation include California Trout , which protects wild trout and other salmonids in the waters of California. The Steelhead Society of British Columbia promotes
5450-848: The Pacific Ocean in North America and Asia. The steelhead (sometimes called steelhead trout ) is an anadromous (sea-run) form of the coastal rainbow trout ( O. m. irideus ) or Columbia River redband trout ( O. m. gairdneri ) that usually returns to freshwater to spawn after living two to three years in the ocean. Freshwater forms that have been introduced into the Great Lakes and migrate into tributaries to spawn are also called steelhead. Adult freshwater stream rainbow trout average between 0.5 and 2.5 kilograms (1 and 5 lb), while lake-dwelling and anadromous forms may reach 9 kg (20 lb). Coloration varies widely based on subspecies , forms, and habitat . Adult fish are distinguished by
5559-789: The Pacific basin, from the Kamchatka Peninsula in Russia, east along the Aleutian Islands , throughout southwest Alaska, the Pacific coast of British Columbia and southeast Alaska, and south along the west coast of the U.S. to northern Mexico. It is claimed that the Mexican forms of Oncorhynchus mykiss represent the southernmost native range of any trout or salmon ( Salmonidae ), though the Formosan landlocked salmon ( O. masou formosanus ) in Asia inhabits
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#17329139179765668-504: The U.S. West Coast are in decline. The U.S. National Marine Fisheries Service has 15 identified distinct population segments , in Washington , Oregon , and California. Eleven of these populations are listed under the U.S. Endangered Species Act , ten as threatened and one as endangered. One distinct population segment on the Oregon coast is designated a U.S. Species of Concern . The Southern California distinct population segment , which
5777-627: The U.S. by the Environmental Protection Agency . Once the chemicals have dissipated, native trout are released into the river. Another method is to use electrofishing which enables the fish to be caught alive and harvested or re-located. This technique has been used in the Great Smokey Mountains National Park to rid it of rainbow trout that were introduced in the 1930s and have thrived ever since. They are hoping to re-establish native brook trout in at least some of
5886-585: The U.S., Iran, Turkey, the United Kingdom, and Lesotho . While the U.S. rainbow trout industry as a whole is viewed as ecologically responsible, trout raised elsewhere are not necessarily farmed with the same methods. About three-quarters of U.S. production comes from Idaho, particularly the Snake River area, due in part to the quality and temperature of the water available there. California and Washington also produce significant numbers of farmed trout. In
5995-511: The U.S., and nearly all domestic production is consumed within the country; very little is exported. The U.S. produces about 7 percent of the world's farmed trout. Rainbow trout, especially those raised in farms and hatcheries, are susceptible to enteric redmouth disease . A considerable amount of research has been conducted on redmouth disease, given its serious implications for rainbow trout farming. The disease does not infect humans. There have been recent interest and efforts in introducing
6104-488: The U.S., there are hundreds of hatcheries operated by the United States Fish and Wildlife Service and various state agencies and tribal governments propagating rainbow trout for conservation and recreational sport fishing. Six of ten Canadian provinces have rainbow trout farms, with Ontario leading production. Rainbow trout are commercially farmed in many countries throughout the world. The practice began in
6213-404: The acute during the late 20th century, and it has only been retained in typography . After the writing reform of 1982, most diacritics are no longer used. Since then, Greek has been written mostly in the simplified monotonic orthography (or monotonic system), which employs only the acute accent and the diaeresis. The traditional system, now called the polytonic orthography (or polytonic system),
6322-474: The arrival of Proto-Greeks, some documented in Mycenaean texts ; they include a large number of Greek toponyms . The form and meaning of many words have changed. Loanwords (words of foreign origin) have entered the language, mainly from Latin, Venetian , and Turkish . During the older periods of Greek, loanwords into Greek acquired Greek inflections, thus leaving only a foreign root word. Modern borrowings (from
6431-433: The bottom of rivers and streams in late winter. It is not considered a significant human health risk, but it can affect stream habitats and sources of food for fish, including rainbow trout, and make recreational activities unpleasant. Even though it is native in North America, it is considered a nuisance organism or invasive species. Enteric redmouth disease is a bacterial infection of freshwater and marine fish caused by
6540-485: The case of the westslope cutthroat trout ( O. clarki lewisi ), hybridization with introduced rainbow and Yellowstone cutthroat trout ( O. clarki bouvieri ) is threatening the westslope cutthroat trout with genomic extinction . Such introductions into the ranges of redband trout ( O. m. gairdneri , newberrii , and stonei ) have severely reduced the range of pure stocks of these subspecies, making them "species of concern" in their respective ranges. Within
6649-765: The country. Prior to the Greco-Turkish War and the resulting population exchange in 1923 a very large population of Greek-speakers also existed in Turkey , though very few remain today. A small Greek-speaking community is also found in Bulgaria near the Greek-Bulgarian border. Greek is also spoken worldwide by the sizable Greek diaspora which has notable communities in the United States , Australia , Canada , South Africa , Chile , Brazil , Argentina , Russia , Ukraine ,
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#17329139179766758-441: The dative led to a rise of prepositional indirect objects (and the use of the genitive to directly mark these as well). Ancient Greek tended to be verb-final, but neutral word order in the modern language is VSO or SVO. Modern Greek inherits most of its vocabulary from Ancient Greek, which in turn is an Indo-European language, but also includes a number of borrowings from the languages of the populations that inhabited Greece before
6867-528: The earliest forms attested to four in the modern language). Nouns, articles, and adjectives show all the distinctions except for a person. Both attributive and predicative adjectives agree with the noun. The inflectional categories of the Greek verb have likewise remained largely the same over the course of the language's history but with significant changes in the number of distinctions within each category and their morphological expression. Greek verbs have synthetic inflectional forms for: Many aspects of
6976-456: The east, Pennsylvania , North Carolina and West Virginia have farming operations. Rainbow trout farming is one of the largest finfish aquaculture industries in the U.S. They are raised inland in facilities where raceways or ponds have continuously flowing water with little pollution and a low risk of escape. The U.S. industry is noted for using best management practices . Imports constitute only about 15 percent of farmed rainbows sold in
7085-407: The eggs of other rainbow trout. Rainbows also consume decomposing flesh from the carcasses of other fish. They have even been observed to have consumed hatchling snakes, such as the barred grass snake ( Natrix helvetica ). Adult steelhead in the ocean feed primarily on other fish, squid and amphipods . The native range of Oncorhynchus mykiss is in the coastal waters and tributary streams of
7194-412: The extent that one can speak of a new language emerging. Greek speakers today still tend to regard literary works of ancient Greek as part of their own rather than a foreign language. It is also often stated that the historical changes have been relatively slight compared with some other languages. According to one estimation, " Homeric Greek is probably closer to Demotic than 12-century Middle English
7303-648: The first federal fish hatchery in the National Fish Hatchery System , was established on Campbell Creek, a McCloud River tributary. The McCloud River hatchery indiscriminately mixed coastal rainbow trout eggs with the eggs of local McCloud River redband trout ( O. m. stonei ) . Eggs from the McCloud hatchery were also provided to the San Leandro hatchery, thus making the origin and genetic history of hatchery-bred rainbow trout somewhat diverse and complex. In
7412-412: The fish, mykizha . The name of the genus is from the Greek ὄγκος ( ónkos ) "lump, bend, hook" and ῥύγχος ( rhúnkhos ) " snout ", in reference to the hooked jaws of males in the mating season (the " kype "). Sir John Richardson , a Scottish naturalist, named a specimen of this species Salmo gairdneri in 1836 to honor Meredith Gairdner, a Hudson's Bay Company surgeon at Fort Vancouver on
7521-468: The impassable barriers; a 2007 genetic study of fin samples collected from steelhead at 20 different sites both above and below passage barriers in the watershed found that although 30 million hatchery trout were stocked in the river from 1911 to 1925, the steelhead remain of native and not hatchery origin. Releases of conventionally reared hatchery steelhead pose ecological risks to wild steelhead populations. Hatchery steelhead are typically larger than
7630-403: The introduced rainbow trout disrupt local ecosystems and outcompete or prey upon indigenous fishes. Other introductions to support sport angling in waters either devoid of fish or with seriously depleted native stocks have created world-class fisheries such as in the Firehole River in Yellowstone National Park and in the Great Lakes . The International Game Fish Association recognizes
7739-449: The language show both conservative and innovative tendencies across the entire attestation of the language from the ancient to the modern period. The division into conventional periods is, as with all such periodizations, relatively arbitrary, especially because, in all periods, Ancient Greek has enjoyed high prestige, and the literate borrowed heavily from it. Across its history, the syllabic structure of Greek has varied little: Greek shows
7848-463: The late 15th century BC, was the first script used to write Greek. It is basically a syllabary , which was finally deciphered by Michael Ventris and John Chadwick in the 1950s (its precursor, Linear A , has not been deciphered and most likely encodes a non-Greek language). The language of the Linear B texts, Mycenaean Greek , is the earliest known form of Greek. Another similar system used to write
7957-546: The late 19th century, and since the 1950s commercial production has grown dramatically. Worldwide, in 2007, 604,695 tonnes (666,562 short tons) of farmed rainbow trout were harvested with a value of about US$ 2.6 billion. The largest producer is Chile. In Chile and Norway, sea cage production of steelhead has expanded to supply export markets. Inland production of rainbow trout to supply domestic markets has increased in countries such as Italy, France, Germany, Denmark, and Spain. Other significant trout-producing countries include
8066-835: The length of the body. Adult fish have a broad reddish stripe along the lateral line, from gills to the tail, which is most pronounced in breeding males. The caudal fin is squarish and only mildly forked. Lake-dwelling and anadromous forms are usually more silvery in color with the reddish stripe almost completely gone. Juvenile rainbow trout display parr marks (dark vertical bars) typical of most salmonid juveniles. In some redband and golden trout forms, parr marks are typically retained into adulthood. Some coastal rainbow trout ( O. m. irideus ) and Columbia River redband trout ( O. m. gairdneri ) populations and cutbow hybrids may also display reddish or pink throat markings similar to cutthroat trout. In many regions, hatchery-bred trout can be distinguished from native trout via fin clips . Fin clipping
8175-448: The many other countries of the Greek diaspora . Greek roots have been widely used for centuries and continue to be widely used to coin new words in other languages; Greek and Latin are the predominant sources of international scientific vocabulary . Greek has been spoken in the Balkan peninsula since around the 3rd millennium BC, or possibly earlier. The earliest written evidence is
8284-465: The morphological changes also have their counterparts in the syntax, and there are also significant differences between the syntax of the ancient and that of the modern form of the language . Ancient Greek made great use of participial constructions and of constructions involving the infinitive, and the modern variety lacks the infinitive entirely (employing a raft of new periphrastic constructions instead) and uses participles more restrictively. The loss of
8393-561: The pathogen Yersinia ruckeri . It is primarily found in rainbow trout and other cultured salmonids . The disease is characterized by subcutaneous hemorrhaging of the mouth, fins, and eyes. It is most commonly seen in fish farms with poor water quality. Redmouth disease was first discovered in Idaho rainbow trout in the 1950s. Some fisheries are focused on removing rainbow trout in order to reestablish native trout populations. This can be done by poisoning rivers with chemicals such as antimycin or rotenone which have been declared safe in
8502-525: The range of the Kern River golden trout of Southern California, hatchery-bred rainbows introduced into the Kern River have diluted the genetic purity of the Kern River rainbow trout (O. m. gilberti) and golden trout (O. m. aguabonita) through intraspecific breeding. The Beardslee trout ( O. m. irideus var. beardsleei ) , a genetically unique lake-dwelling variety of the coastal rainbow trout that
8611-622: The regions of Apulia and Calabria in Italy. In the framework of the European Charter for Regional or Minority Languages , Greek is protected and promoted officially as a regional and minority language in Armenia, Hungary , Romania, and Ukraine. It is recognized as a minority language and protected in Turkey by the 1923 Treaty of Lausanne . The phonology , morphology , syntax , and vocabulary of
8720-471: The river for one to three years before smolting and migrating to sea. Individual steelhead populations leave the ocean and migrate into their freshwater spawning tributaries at different times of the year. Two general forms exist—"summer-run steelhead" and "winter-run steelhead". Summer-run fish leave the ocean between May and October before their reproductive organs are fully mature. They mature in freshwater while en route to spawning grounds where they spawn in
8829-437: The salmonids that M. cerebralis infects ( bull trout , cutthroat trout, and anadromous forms of rainbow trout—steelhead) are already threatened or endangered, and the parasite could worsen their population decline. The New Zealand mud snail ( Potamopyrgus antipodarum ) , once endemic to New Zealand, has spread widely and has become naturalised and an invasive species in many areas including Australia, Asia (Japan, in
8938-796: The same reasons. As a consequence, some rainbow populations, particularly anadromous forms within their native range, have been classified as endangered , threatened or species of special concern by federal or state agencies. Rainbow trout, and subspecies thereof, are currently a U.S. Environmental Protection Agency -approved indicator species for acute freshwater toxicity testing. Many non-profit organizations have formed to protect, conserve and restore native rainbow trout and steelhead populations. Generally, in partnership with various universities, state, federal and tribal agencies, and private interests, these organizations sponsor projects to restore habitat, prevent habitat loss and promote awareness of threats to native trout populations. Trout Unlimited (TU)
9047-479: The same species. Like salmon, steelhead return to their original hatching grounds to spawn . Similar to Atlantic salmon, but unlike their Pacific Oncorhynchus salmonid kin, steelhead are iteroparous (able to spawn several times, each time separated by months) and make several spawning trips between fresh and salt water, although fewer than 10 percent of native spawning adults survive from one spawning to another. The survival rate for introduced populations in
9156-829: The scientific name of the rainbow trout. The previous species names irideus and gairdneri were adopted as subspecies names for the coastal rainbow and Columbia River redband trout, respectively. Anadromous forms of the coastal rainbow trout ( O. m. irideus ) or Columbia River redband trout ( O. m. gairdneri ) are commonly known as steelhead . Subspecies of Oncorhynchus mykiss are listed below as described by fisheries biologist Robert J. Behnke (2002). Resident freshwater rainbow trout adults average between 0.5 and 2.5 kg (1 and 5 lb) in riverine environments, while lake-dwelling, and anadromous forms may reach 9 kg (20 lb). Coloration varies widely between regions and subspecies. Adult freshwater forms are generally blue-green or olive green with heavy black spotting over
9265-526: The size of a human finger. In streams where rainbow trout are stocked for sport fishing, but no natural reproduction occurs, some of the stocked trout may survive and grow or "carryover" for several seasons before they are caught or perish. The oceangoing (anadromous) form, including those returning for spawning, are known as steelhead in Canada and the U.S. In Tasmania they are commercially propagated in sea cages and are known as ocean trout , although they are
9374-485: The skin on, the meat tends to hold together better. While trout sold commercially in Europe is often prepared and served this way, most trout sold commercially in the United States have the fish heads removed and have been fully or partially deboned and filleted. Medium to heavy-bodied white wines , such as chardonnay , sauvignon blanc , or pinot gris , are typical wine pairings for trout. In Mainland China since 2018, it
9483-909: The spring. Summer-run fish generally spawn in longer, more inland rivers such as the Columbia River. Winter-run fish are ready to spawn when they leave the ocean, typically between November and April, and spawn shortly after returning to fresh water. Winter-run fish generally spawn in shorter, coastal rivers typically found along the Olympic Peninsula and British Columbia coastline, and summer-run fish are found in some shorter, coastal streams. Once steelhead enter riverine systems and reach suitable spawning grounds, they spawn just like resident freshwater rainbow trout. During periods of rapid growth and aging, trout display high levels of metabolic activity. High metabolic activity has been correlated with increased levels of oxidative stress and decreased machinery repair in rainbow trout. During high oxidative stress,
9592-531: The steelhead in the streams of the Santa Clara County and Monterey Bay basins are not of hatchery origin, including the Coyote Creek , Guadalupe River , Pajaro River , Permanente Creek , Stevens Creek , San Francisquito Creek , San Lorenzo River , and San Tomas Aquino Creek basins. Natural waterfalls and two major dams have isolated Russian River steelhead from freshwater rainbow trout forms above
9701-437: The stressed vowel; the so-called breathing marks ( rough and smooth breathing ), originally used to signal presence or absence of word-initial /h/; and the diaeresis , used to mark the full syllabic value of a vowel that would otherwise be read as part of a diphthong. These marks were introduced during the course of the Hellenistic period. Actual usage of the grave in handwriting saw a rapid decline in favor of uniform usage of
9810-853: The trout's development into an adult, a time of intense growth, the mitochondrial membrane composition and fluidity changes, which can cause defects in the electron transport chain . These defects combined with altered machinery repair and reactive oxygen species may cause more detrimental effects on the mitochondria in the fish as it matures. Rainbow trout are predators with a varied diet and will eat nearly anything they can capture. They are not as piscivorous or aggressive as brown trout or chars . Rainbow trout, including juvenile steelhead in fresh water, routinely feed on larval , pupal , and adult forms of aquatic insects (typically caddisflies , stoneflies , mayflies and aquatic diptera ). They also eat fish eggs and adult forms of terrestrial insects (typically ants, beetles, grasshoppers, and crickets) that fall into
9919-422: The types and proportions of phospholipids. With age, the heart showed more unsaturated phospholipids, which are more susceptible to peroxidation, and thus, damage. The brain mitochondria of rainbow trout show decreased levels of docosahexaenoic acid and a lower peroxidation index, suggesting a lower susceptibility to damage by oxidative stress and a different reaction to growth compared to heart mitochondria. During
10028-409: The upstream edge of the nest, covering the eggs with the displaced gravel. As eggs are released by the female, a male moves alongside and deposits milt (sperm) over the eggs to fertilize them. The eggs usually hatch in about four to seven weeks although the time of hatching varies greatly with region and habitat. Newly hatched trout are called sac fry or alevin . In approximately two weeks, the yolk sac
10137-459: The water, as well as algae. Other prey includes small fish up to one-third of their length, crayfish , shrimp, and other crustaceans . As rainbow trout grow, the proportion of fish consumed increases in most populations. Some lake-dwelling forms may become planktonic feeders. In rivers and streams populated with other salmonid species, rainbow trout eat varied fish eggs, including those of salmon, brown and cutthroat trout, mountain whitefish , and
10246-570: The wellbeing of wild salmonids in British Columbia. In 1997, a group of approximately 40 ichthyologists, biologists and naturalists from several U.S. and Mexican institutions formed a collaborative group—Truchas Mexicanas—to study the diversity of Mexican native trout , most of which are considered subspecies of O. mykiss . Rainbow trout, primarily hatchery-raised fish of the coastal rainbow trout subspecies ( O. m. irideus ) introduced into waters inhabited with cutthroat trout, will breed with cutthroats and produce fertile hybrids called cutbows. In
10355-502: The western U.S. The exact means of transmission is unknown, but it is likely that it was introduced in water transferred with live game fish and has been spread by ship ballast or contaminated recreational equipment such as wading gear. Didymosphenia geminata , commonly known as didymo or rock snot, is a species of diatom that produces nuisance growths in freshwater rivers and streams with consistently cold water temperatures. In New Zealand, invasive didymo can form large mats on
10464-520: The wild forms and can displace wild-form juveniles from optimal habitats. The dominance of hatchery steelhead for optimal microhabitats within streams may reduce wild steelhead survival as a result of reduced foraging opportunities and increased rates of predation. Rainbow trout and steelhead are highly regarded game fish among anglers . Rainbow trout are a popular target for fly fishers , and several lure fishing methods are used. The use of lures presented via spinning, casting, or trolling techniques
10573-489: The world record for rainbow trout as a fish caught on Saskatchewan 's Lake Diefenbaker by Sean Konrad on September 5, 2009, which weighed 48 lb (22 kg). The record is controversial because the fish was a genetically modified triploid and was part of a large number of triploid rainbow trout which escaped from an aquaculture facility. Rainbow trout is sometimes used as a biological indicator for water quality in water purification facilities. Rainbow trout
10682-568: Was a distinct dialect of Greek itself. Aside from the Macedonian question, current consensus regards Phrygian as the closest relative of Greek, since they share a number of phonological, morphological and lexical isoglosses , with some being exclusive between them. Scholars have proposed a Graeco-Phrygian subgroup out of which Greek and Phrygian originated. Among living languages, some Indo-Europeanists suggest that Greek may be most closely related to Armenian (see Graeco-Armenian ) or
10791-444: Was adopted in 1956, and commemorates Father Jean-Marie-Raphaël Le Jeune , a French Catholic priest who spent much of his life in the region. This British Columbia protected areas related article is a stub . You can help Misplaced Pages by expanding it . Rainbow Trout This is an accepted version of this page The rainbow trout ( Oncorhynchus mykiss ) is a species of trout native to cold-water tributaries of
10900-546: Was also used in Ancient Greek. Greek has occasionally been written in the Latin script , especially in areas under Venetian rule or by Greek Catholics . The term Frankolevantinika / Φραγκολεβαντίνικα applies when the Latin script is used to write Greek in the cultural ambit of Catholicism (because Frankos / Φράγκος is an older Greek term for West-European dating to when most of (Roman Catholic Christian) West Europe
11009-541: Was by far the most widely spoken lingua franca in the Mediterranean world . It eventually became the official language of the Byzantine Empire and developed into Medieval Greek . In its modern form , Greek is the official language of Greece and Cyprus and one of the 24 official languages of the European Union . It is spoken by at least 13.5 million people today in Greece, Cyprus, Italy, Albania, Turkey , and
11118-489: Was established on San Leandro Creek , a tributary of San Francisco Bay , in 1870, and trout production began in 1871. The hatchery was stocked with the locally native rainbow trout, and likely steelhead of the coastal rainbow trout subspecies ( O. m. irideus ) . The fish raised in this hatchery were shipped to hatcheries out of state for the first time in 1875, to Caledonia, New York , and then in 1876 to Northville, Michigan . In 1877, another California rainbow trout hatchery,
11227-615: Was listed as endangered in 2011, has been affected by habitat loss due to dams, confinement of streams in concrete channels, water pollution , groundwater pumping, urban heat island effects, and other byproducts of urbanization. Steelhead in the Kamchatka Peninsula are threatened by over-harvest, particularly from poaching and potential development, and are listed in the Red Data Book of Russia that documents rare and endangered species. Several studies have shown that almost all California coastal steelhead are of native origin, despite over
11336-526: Was originally a mild pathogen of brown trout in central Europe and other salmonids in northeast Asia, and the spread of the rainbow trout has greatly increased its impact. Having no innate immunity to M. cerebralis , rainbow trout are particularly susceptible, and can release so many spores that even more resistant species in the same area, such as Salmo trutta , can become overloaded with parasites and incur mortalities of 80 to 90 percent. Where M. cerebralis has become well-established, it has caused
11445-480: Was recorded in writing systems such as Linear B and the Cypriot syllabary . The alphabet arose from the Phoenician script and was in turn the basis of the Latin , Cyrillic , Coptic , Gothic , and many other writing systems. The Greek language holds a very important place in the history of the Western world. Beginning with the epics of Homer , ancient Greek literature includes many works of lasting importance in
11554-415: Was ruled by an industrial association that rainbow trout can be labelled and sold as salmon . Greek language Greek ( Modern Greek : Ελληνικά , romanized : Elliniká , [eliniˈka] ; Ancient Greek : Ἑλληνική , romanized : Hellēnikḗ ) is an Indo-European language, constituting an independent Hellenic branch within the Indo-European language family. It
11663-566: Was under the control of the Frankish Empire ). Frankochiotika / Φραγκοχιώτικα (meaning 'Catholic Chiot') alludes to the significant presence of Catholic missionaries based on the island of Chios . Additionally, the term Greeklish is often used when the Greek language is written in a Latin script in online communications. The Latin script is nowadays used by the Greek-speaking communities of Southern Italy . The Yevanic dialect
11772-637: Was used for literary and official purposes in the newly formed Greek state. In 1976, Dimotiki was declared the official language of Greece, after having incorporated features of Katharevousa and thus giving birth to Standard Modern Greek , used today for all official purposes and in education . The historical unity and continuing identity between the various stages of the Greek language are often emphasized. Although Greek has undergone morphological and phonological changes comparable to those seen in other languages, never since classical antiquity has its cultural, literary, and orthographic tradition been interrupted to
11881-653: Was written by Romaniote and Constantinopolitan Karaite Jews using the Hebrew Alphabet . In a tradition, that in modern time, has come to be known as Greek Aljamiado , some Greek Muslims from Crete wrote their Cretan Greek in the Arabic alphabet . The same happened among Epirote Muslims in Ioannina . This also happened among Arabic-speaking Byzantine rite Christians in the Levant ( Lebanon , Palestine, and Syria ). This usage
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