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97-428: As of April 30, 2017, the town has an estimated population of 15,573 and a population density of 60 persons per km². The total area is 261.24 km (100.87 sq mi). Throughout Hokkaido, Kutchan is known for its proximity to the world famous Niseko powder snow region and for its production of quality potatoes and potato products. Among these products, Kutchan is most famous for manufacturing gosetsu-udon ,

194-408: A sexual population is the area where interbreeding is possible between any opposite-sex pair within the area and more probable than cross-breeding with individuals from other areas. In humans , interbreeding is unrestricted by racial differences, as all humans belong to the same species of Homo sapiens. In ecology, the population of a certain species in a certain area can be estimated using

291-412: A Japanese noodle made from potato flour . Jagata-kun, the skiing potato that is the cartoon mascot of Kutchan, indicates Kutchan's two most celebrated aspects. Jagata-kun roughly translates as, "little, fat potato boy". Kutchan has the distinction of being one of the snowiest towns on earth, with average annual snowfalls in excess of 10 m (33 ft). At only 176 m (577 ft) above sea level

388-460: A dramatic effect on community structure. Hunting of sea otters, for example, is thought to have led indirectly to the extinction of the Steller's sea cow ( Hydrodamalis gigas ). While the keystone species concept has been used extensively as a conservation tool, it has been criticized for being poorly defined from an operational stance. It is difficult to experimentally determine what species may hold

485-428: A dramatic impact on the ecology of individual species or whole ecosystems. For instance, the replacement of an ant species by another (invasive) ant species has been shown to affect how elephants reduce tree cover and thus the predation of lions on zebras . A trophic level (from Greek troph , τροφή, trophē, meaning "food" or "feeding") is "a group of organisms acquiring a considerable majority of its energy from

582-471: A few programs, most notably the Chinese government's one-child per family policy, have resorted to coercive measures. In the 1970s, tension grew between population control advocates and women's health activists who advanced women's reproductive rights as part of a human rights -based approach. Growing opposition to the narrow population control focus led to a significant change in population control policies in

679-412: A flattened body relative to the main populations that live in open savanna. The population that lives in an isolated rock outcrop hides in crevasses where its flattened body offers a selective advantage. Habitat shifts also occur in the developmental life history of amphibians, and in insects that transition from aquatic to terrestrial habitats. Biotope and habitat are sometimes used interchangeably, but

776-612: A higher one." Small scale patterns do not necessarily explain large scale phenomena, otherwise captured in the expression (coined by Aristotle) 'the sum is greater than the parts'. "Complexity in ecology is of at least six distinct types: spatial, temporal, structural, process, behavioral, and geometric." From these principles, ecologists have identified emergent and self-organizing phenomena that operate at different environmental scales of influence, ranging from molecular to planetary, and these require different explanations at each integrative level . Ecological complexity relates to

873-444: A keystone role in each ecosystem. Furthermore, food web theory suggests that keystone species may not be common, so it is unclear how generally the keystone species model can be applied. Complexity is understood as a large computational effort needed to piece together numerous interacting parts exceeding the iterative memory capacity of the human mind. Global patterns of biological diversity are complex. This biocomplexity stems from

970-408: A keystone species can result in a community collapse just as the removal of the keystone in an arch can result in the arch's loss of stability. Sea otters ( Enhydra lutris ) are commonly cited as an example of a keystone species because they limit the density of sea urchins that feed on kelp . If sea otters are removed from the system, the urchins graze until the kelp beds disappear, and this has

1067-737: A legacy niche that was constructed before their time. Biomes are larger units of organization that categorize regions of the Earth's ecosystems, mainly according to the structure and composition of vegetation. There are different methods to define the continental boundaries of biomes dominated by different functional types of vegetative communities that are limited in distribution by climate, precipitation, weather, and other environmental variables. Biomes include tropical rainforest , temperate broadleaf and mixed forest , temperate deciduous forest , taiga , tundra , hot desert , and polar desert . Other researchers have recently categorized other biomes, such as

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1164-439: A linear successional route, changes might occur quickly or slowly over thousands of years before specific forest successional stages are brought about by biological processes. An ecosystem's area can vary greatly, from tiny to vast. A single tree is of little consequence to the classification of a forest ecosystem, but is critically relevant to organisms living in and on it. Several generations of an aphid population can exist over

1261-439: A macroscopic view of the system. While the notion of trophic levels provides insight into energy flow and top-down control within food webs, it is troubled by the prevalence of omnivory in real ecosystems. This has led some ecologists to "reiterate that the notion that species clearly aggregate into discrete, homogeneous trophic levels is fiction." Nonetheless, recent studies have shown that real trophic levels do exist, but "above

1358-516: A mix of herbivores and predators). Omnivores do not fit neatly into a functional category because they eat both plant and animal tissues. It has been suggested that omnivores have a greater functional influence as predators because compared to herbivores, they are relatively inefficient at grazing. Trophic levels are part of the holistic or complex systems view of ecosystems. Each trophic level contains unrelated species that are grouped together because they share common ecological functions, giving

1455-420: A new appreciation for the influence that organisms have on the ecosystem and evolutionary process. The term "niche construction" is more often used in reference to the under-appreciated feedback mechanisms of natural selection imparting forces on the abiotic niche. An example of natural selection through ecosystem engineering occurs in the nests of social insects , including ants, bees, wasps, and termites. There

1552-435: A population-level phenomenon, as with the migration routes followed by plants as they occupied northern post-glacial environments. Plant ecologists use pollen records that accumulate and stratify in wetlands to reconstruct the timing of plant migration and dispersal relative to historic and contemporary climates. These migration routes involved an expansion of the range as plant populations expanded from one area to another. There

1649-401: A region), and sites are classed either as sources or sinks. A site is a generic term that refers to places where ecologists sample populations, such as ponds or defined sampling areas in a forest. Source patches are productive sites that generate a seasonal supply of juveniles that migrate to other patch locations. Sink patches are unproductive sites that only receive migrants; the population at

1746-477: A seasonal influx of new immigrants. A dynamic metapopulation structure evolves from year to year, where some patches are sinks in dry years and are sources when conditions are more favorable. Ecologists use a mixture of computer models and field studies to explain metapopulation structure. Community ecology examines how interactions among species and their environment affect the abundance, distribution and diversity of species within communities. Community ecology

1843-456: A species describes the environment over which a species is known to occur and the type of community that is formed as a result. More specifically, "habitats can be defined as regions in environmental space that are composed of multiple dimensions, each representing a biotic or abiotic environmental variable; that is, any component or characteristic of the environment related directly (e.g. forage biomass and quality) or indirectly (e.g. elevation) to

1940-402: A specific habitat, such as a cave or a pond, and principles gleaned from small-scale studies are extrapolated to larger systems. Feeding relations require extensive investigations, e.g. into the gut contents of organisms, which can be difficult to decipher, or stable isotopes can be used to trace the flow of nutrient diets and energy through a food web. Despite these limitations, food webs remain

2037-446: A valuable tool in understanding community ecosystems. Food webs illustrate important principles of ecology : some species have many weak feeding links (e.g., omnivores ) while some are more specialized with fewer stronger feeding links (e.g., primary predators ). Such linkages explain how ecological communities remain stable over time and eventually can illustrate a "complete" web of life. The disruption of food webs may have

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2134-441: A whole functional system, such as an ecosystem , cannot be predicted or understood by a simple summation of the parts. "New properties emerge because the components interact, not because the basic nature of the components is changed." Ecological studies are necessarily holistic as opposed to reductionistic . Holism has three scientific meanings or uses that identify with ecology: 1) the mechanistic complexity of ecosystems, 2)

2231-399: A wide array of interacting levels of organization spanning micro-level (e.g., cells ) to a planetary scale (e.g., biosphere ) phenomena . Ecosystems, for example, contain abiotic resources and interacting life forms (i.e., individual organisms that aggregate into populations which aggregate into distinct ecological communities). Because ecosystems are dynamic and do not necessarily follow

2328-485: Is a central concept in the ecology of organisms and is sub-divided into the fundamental and the realized niche. The fundamental niche is the set of environmental conditions under which a species is able to persist. The realized niche is the set of environmental plus ecological conditions under which a species persists. The Hutchinsonian niche is defined more technically as a " Euclidean hyperspace whose dimensions are defined as environmental variables and whose size

2425-504: Is a function of the number of values that the environmental values may assume for which an organism has positive fitness ." Biogeographical patterns and range distributions are explained or predicted through knowledge of a species' traits and niche requirements. Species have functional traits that are uniquely adapted to the ecological niche. A trait is a measurable property, phenotype , or characteristic of an organism that may influence its survival. Genes play an important role in

2522-414: Is a larger taxonomy of movement, such as commuting, foraging, territorial behavior, stasis, and ranging. Dispersal is usually distinguished from migration because it involves the one-way permanent movement of individuals from their birth population into another population. In metapopulation terminology, migrating individuals are classed as emigrants (when they leave a region) or immigrants (when they enter

2619-461: Is an emergent homeostasis or homeorhesis in the structure of the nest that regulates, maintains and defends the physiology of the entire colony. Termite mounds, for example, maintain a constant internal temperature through the design of air-conditioning chimneys. The structure of the nests themselves is subject to the forces of natural selection. Moreover, a nest can survive over successive generations, so that progeny inherit both genetic material and

2716-404: Is another statistical approach that is often used in conservation research . Metapopulation models simplify the landscape into patches of varying levels of quality, and metapopulations are linked by the migratory behaviours of organisms. Animal migration is set apart from other kinds of movement because it involves the seasonal departure and return of individuals from a habitat. Migration is also

2813-472: Is derived from the Latin word populus (a people). In sociology and population geography , population refers to a group of human beings with some predefined feature in common, such as location, race , ethnicity , nationality , or religion . In ecology , a population is a group of organisms of the same species which inhabit the same geographical area and are capable of interbreeding . The area of

2910-438: Is greater than respiration) by photosynthesis or chemosynthesis . Heterotrophs are organisms that must feed on others for nourishment and energy (respiration exceeds production). Heterotrophs can be further sub-divided into different functional groups, including primary consumers (strict herbivores), secondary consumers ( carnivorous predators that feed exclusively on herbivores), and tertiary consumers (predators that feed on

3007-503: Is mild with temperatures rarely getting above 30 °C (86 °F) and the humidity that afflicts much of Asia at this time is only felt for a short period, usually in August. Along with short grain white rice, much of Kutchan's surrounding area is devoted to potato cultivation. Kutchan hosts the Hirafu skiing area. Hirafu is commonly mistaken as a section of Niseko town since it is a part of

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3104-600: Is not even known to the nearest million, so there is a considerable margin of error in such estimates. Researcher Carl Haub calculated that a total of over 100 billion people have probably been born in the last 2000 years. Population growth increased significantly as the Industrial Revolution gathered pace from 1700 onwards. The last 50 years have seen a yet more rapid increase in the rate of population growth due to medical advances and substantial increases in agricultural productivity, particularly beginning in

3201-611: Is often referred to as the demographic transition . Human population planning is the practice of altering the rate of growth of a human population. Historically, human population control has been implemented with the goal of limiting the rate of population growth. In the period from the 1950s to the 1980s, concerns about global population growth and its effects on poverty, environmental degradation , and political stability led to efforts to reduce population growth rates. While population control can involve measures that improve people's lives by giving them greater control of their reproduction,

3298-574: Is the Park Grass Experiment , which was initiated in 1856. Another example is the Hubbard Brook study , which has been in operation since 1960. Holism remains a critical part of the theoretical foundation in contemporary ecological studies. Holism addresses the biological organization of life that self-organizes into layers of emergent whole systems that function according to non-reducible properties. This means that higher-order patterns of

3395-413: Is the biosphere: the total sum of ecosystems on the planet. Ecological relationships regulate the flux of energy, nutrients, and climate all the way up to the planetary scale. For example, the dynamic history of the planetary atmosphere's CO 2 and O 2 composition has been affected by the biogenic flux of gases coming from respiration and photosynthesis, with levels fluctuating over time in relation to

3492-436: Is the study of observable movement or behaviour in animals. This could include investigations of motile sperm of plants, mobile phytoplankton , zooplankton swimming toward the female egg, the cultivation of fungi by weevils , the mating dance of a salamander , or social gatherings of amoeba . Adaptation is the central unifying concept in behavioural ecology. Behaviours can be recorded as traits and inherited in much

3589-522: Is the study of the interactions among a collection of species that inhabit the same geographic area. Community ecologists study the determinants of patterns and processes for two or more interacting species. Research in community ecology might measure species diversity in grasslands in relation to soil fertility. It might also include the analysis of predator-prey dynamics, competition among similar plant species, or mutualistic interactions between crabs and corals. These ecosystems, as we may call them, are of

3686-405: Is undoubtedly an asset to Kutchan town, many fear that such rapid development will alter the price margin of goods and services offered in the village, and subsequently transform the local flavor of Hirafu. The local community of both Japanese and foreign "backpacker" skiers and snowboarders, who typically operate on a smaller budget than their upscale counterparts, are already being overshadowed by

3783-552: Is used for managing wildlife stocks and setting harvest quotas. In cases where basic models are insufficient, ecologists may adopt different kinds of statistical methods, such as the Akaike information criterion , or use models that can become mathematically complex as "several competing hypotheses are simultaneously confronted with the data." The concept of metapopulations was defined in 1969 as "a population of populations which go extinct locally and recolonize". Metapopulation ecology

3880-703: Is very likely that the world's population will stop growing before the end of the 21st century. Further, there is some likelihood that population will actually decline before 2100. Population has already declined in the last decade or two in Eastern Europe, the Baltics and in the former Commonwealth of Independent States. The population pattern of less-developed regions of the world in recent years has been marked by gradually declining birth rates. These followed an earlier sharp reduction in death rates. This transition from high birth and death rates to low birth and death rates

3977-473: The Lincoln index to calculate the total population of an area based on the number of individuals observed. In genetics, a population is often defined as a set of organisms in which any pair of members can breed together. They can thus routinely exchange gametes in order to have usually fertile progeny, and such a breeding group is also known therefore as a gamodeme. This also implies that all members belong to

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4074-405: The abundance or biomass at each level. When the relative abundance or biomass of each species is sorted into its respective trophic level, they naturally sort into a 'pyramid of numbers'. Species are broadly categorized as autotrophs (or primary producers ), heterotrophs (or consumers ), and Detritivores (or decomposers ). Autotrophs are organisms that produce their own food (production

4171-436: The logistic equation by Pierre Verhulst : where N(t) is the number of individuals measured as biomass density as a function of time, t , r is the maximum per-capita rate of change commonly known as the intrinsic rate of growth, and α {\displaystyle \alpha } is the crowding coefficient, which represents the reduction in population growth rate per individual added. The formula states that

4268-547: The 1960s, made by the Green Revolution . In 2017 the United Nations Population Division projected that the world's population would reach about 9.8 billion in 2050 and 11.2 billion in 2100. In the future, the world's population is expected to peak at some point, after which it will decline due to economic reasons, health concerns, land exhaustion and environmental hazards. According to one report, it

4365-545: The Earth and atmospheric conditions within a narrow self-regulating range of tolerance. Population ecology studies the dynamics of species populations and how these populations interact with the wider environment. A population consists of individuals of the same species that live, interact, and migrate through the same niche and habitat. A primary law of population ecology is the Malthusian growth model which states, "a population will grow (or decline) exponentially as long as

4462-485: The United States Census Bureau, the world population hit 6.5 billion on 24 February 2006. The United Nations Population Fund designated 12 October 1999 as the approximate day on which world population reached 6 billion. This was about 12 years after the world population reached 5 billion in 1987, and six years after the world population reached 5.5 billion in 1993. The population of countries such as Nigeria

4559-705: The analytical tools of ecologists and evolutionists overlap as they organize, classify, and investigate life through common systematic principles, such as phylogenetics or the Linnaean system of taxonomy . The two disciplines often appear together, such as in the title of the journal Trends in Ecology and Evolution . There is no sharp boundary separating ecology from evolution, and they differ more in their areas of applied focus. Both disciplines discover and explain emergent and unique properties and processes operating across different spatial or temporal scales of organization. While

4656-471: The area and spawned businesses such as pensions, bars, and cafes. Hirafu is also frequented by a considerable number of tourists from Hong Kong , as well as tourists coming from within Japan. Hirafu's expanding tourism industry is often attributed to the steady and copious supply of fresh powder snow, which many [1] claim is among the best in the entire hemisphere, if not the world. While Hirafu's economic boom

4753-498: The boundary between ecology and evolution is not always clear, ecologists study the abiotic and biotic factors that influence evolutionary processes, and evolution can be rapid, occurring on ecological timescales as short as one generation. All organisms can exhibit behaviours. Even plants express complex behaviour, including memory and communication. Behavioural ecology is the study of an organism's behaviour in its environment and its ecological and evolutionary implications. Ethology

4850-480: The breaking up of a large sexual population (panmictic) into smaller overlapping sexual populations. This failure of panmixia leads to two important changes in overall population structure: (1) the component gamodemes vary (through gamete sampling) in their allele frequencies when compared with each other and with the theoretical panmictic original (this is known as dispersion, and its details can be estimated using expansion of an appropriate binomial equation ); and (2)

4947-657: The burgeoning influx of wealthy tourists. As the main regional center of population, Kutchan hosts a variety of cultural institutions and events. The town is home to two museums, the Kutchan Natural History Museum and the Shu Ogawara Art Museum. It also hosts the annual Mt. Niseko Kutchan Jazz Festival every July in the center of town, drawing thousands to enjoy the music of Japanese and Australian musicians. Jagata-kun ( じゃが太くん ) and Jagako-chan ( じゃが子ちゃん ) are Kutchan's mascot cartoon character. Due to

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5044-976: The closely related sciences of biogeography , evolutionary biology , genetics , ethology , and natural history . Ecology is a branch of biology , and is the study of abundance , biomass , and distribution of organisms in the context of the environment. It encompasses life processes, interactions, and adaptations ; movement of materials and energy through living communities; successional development of ecosystems; cooperation, competition, and predation within and between species ; and patterns of biodiversity and its effect on ecosystem processes. Ecology has practical applications in conservation biology , wetland management, natural resource management ( agroecology , agriculture , forestry , agroforestry , fisheries , mining , tourism ), urban planning ( urban ecology ), community health , economics , basic and applied science , and human social interaction ( human ecology ). The word ecology ( German : Ökologie )

5141-714: The diversity of life from genes to ecosystems and spans every level of biological organization. The term has several interpretations, and there are many ways to index, measure, characterize, and represent its complex organization. Biodiversity includes species diversity , ecosystem diversity , and genetic diversity and scientists are interested in the way that this diversity affects the complex ecological processes operating at and among these respective levels. Biodiversity plays an important role in ecosystem services which by definition maintain and improve human quality of life. Conservation priorities and management techniques require different approaches and considerations to address

5238-514: The dynamic resilience of ecosystems that transition to multiple shifting steady-states directed by random fluctuations of history. Long-term ecological studies provide important track records to better understand the complexity and resilience of ecosystems over longer temporal and broader spatial scales. These studies are managed by the International Long Term Ecological Network (LTER). The longest experiment in existence

5335-409: The early 1980s. Ecology Ecology (from Ancient Greek οἶκος ( oîkos )  'house' and -λογία ( -logía )  'study of') is the natural science of the relationships among living organisms and their environment . Ecology considers organisms at the individual, population , community , ecosystem , and biosphere levels. Ecology overlaps with

5432-452: The ecology and evolution of plants and animals. Ecological theory has also been used to explain self-emergent regulatory phenomena at the planetary scale: for example, the Gaia hypothesis is an example of holism applied in ecological theory. The Gaia hypothesis states that there is an emergent feedback loop generated by the metabolism of living organisms that maintains the core temperature of

5529-434: The energy is transferred through a chain of organisms by consumption. The simplified linear feeding pathways that move from a basal trophic species to a top consumer is called the food chain . Food chains in an ecological community create a complex food web. Food webs are a type of concept map that is used to illustrate and study pathways of energy and material flows. Empirical measurements are generally restricted to

5626-460: The environment experienced by all individuals in the population remains constant." Simplified population models usually starts with four variables: death, birth, immigration , and emigration . An example of an introductory population model describes a closed population, such as on an island, where immigration and emigration does not take place. Hypotheses are evaluated with reference to a null hypothesis which states that random processes create

5723-451: The equilibrium, r / α {\displaystyle r/\alpha } as K , which is known as the "carrying capacity." Population ecology builds upon these introductory models to further understand demographic processes in real study populations. Commonly used types of data include life history , fecundity , and survivorship, and these are analyzed using mathematical techniques such as matrix algebra . The information

5820-589: The flux of energy and matter through an environment. Ecosystems have biophysical feedback mechanisms that moderate processes acting on living ( biotic ) and abiotic components of the planet. Ecosystems sustain life-supporting functions and provide ecosystem services like biomass production (food, fuel, fiber, and medicine), the regulation of climate , global biogeochemical cycles , water filtration , soil formation , erosion control, flood protection, and many other natural features of scientific, historical, economic, or intrinsic value. The scope of ecology contains

5917-400: The former applies to a community's environment, whereas the latter applies to a species' environment. Definitions of the niche date back to 1917, but G. Evelyn Hutchinson made conceptual advances in 1957 by introducing a widely adopted definition: "the set of biotic and abiotic conditions in which a species is able to persist and maintain stable population sizes." The ecological niche

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6014-544: The former relates only to the physical modifications of the habitat whereas the latter also considers the evolutionary implications of physical changes to the environment and the feedback this causes on the process of natural selection. Ecosystem engineers are defined as: "organisms that directly or indirectly modulate the availability of resources to other species, by causing physical state changes in biotic or abiotic materials. In so doing they modify, maintain and create habitats." The ecosystem engineering concept has stimulated

6111-518: The full ecological scope of biodiversity. Natural capital that supports populations is critical for maintaining ecosystem services and species migration (e.g., riverine fish runs and avian insect control) has been implicated as one mechanism by which those service losses are experienced. An understanding of biodiversity has practical applications for species and ecosystem-level conservation planners as they make management recommendations to consulting firms, governments, and industry. The habitat of

6208-581: The greater Niseko skiing region. However, to the financial benefit of Kutchan, Hirafu lies within the confines of Kutchan town. Hirafu's largest ski resort, Niseko Grand Hirafu , draws the highest number of both domestic and foreign skiers and snowboarders than any other resort in the Niseko region. It is linked to the ski areas of Annupuri, Higashiyama, and Hanazono which together may be skied on one lift pass. In recent years, an explosively high number of Australians have frequented Hirafu, many of whom have settled in

6305-468: The heavy snowfalls in the region, Kutchan hosts several alpine resorts frequented by domestic and international skiers and snowboarders. In the summer, Kutchan is best known for its primary agricultural product: potatoes. The town's two main industries (ski resorts and potato farming) are both reflected in Jagata-kun, a skiing potato. Population Population is the term typically used to refer to

6402-408: The herbivore trophic level, food webs are better characterized as a tangled web of omnivores." A keystone species is a species that is connected to a disproportionately large number of other species in the food-web . Keystone species have lower levels of biomass in the trophic pyramid relative to the importance of their role. The many connections that a keystone species holds means that it maintains

6499-406: The human and oceanic microbiomes . To a microbe , the human body is a habitat and a landscape. Microbiomes were discovered largely through advances in molecular genetics , which have revealed a hidden richness of microbial diversity on the planet. The oceanic microbiome plays a significant role in the ecological biogeochemistry of the planet's oceans. The largest scale of ecological organization

6596-404: The interplay among ecological processes that operate and influence patterns at different scales that grade into each other, such as transitional areas or ecotones spanning landscapes. Complexity stems from the interplay among levels of biological organization as energy, and matter is integrated into larger units that superimpose onto the smaller parts. "What were wholes on one level become parts on

6693-412: The interplay of development and environmental expression of traits. Resident species evolve traits that are fitted to the selection pressures of their local environment. This tends to afford them a competitive advantage and discourages similarly adapted species from having an overlapping geographic range. The competitive exclusion principle states that two species cannot coexist indefinitely by living off

6790-402: The latter focuses on materials and energy fluxes. System behaviors must first be arrayed into different levels of the organization. Behaviors corresponding to higher levels occur at slow rates. Conversely, lower organizational levels exhibit rapid rates. For example, individual tree leaves respond rapidly to momentary changes in light intensity, CO 2 concentration, and the like. The growth of

6887-463: The level of homozygosity rises in the entire collection of gamodemes. The overall rise in homozygosity is quantified by the inbreeding coefficient (f or φ). All homozygotes are increased in frequency – both the deleterious and the desirable. The mean phenotype of the gamodemes collection is lower than that of the panmictic original – which is known as inbreeding depression. It is most important to note, however, that some dispersion lines will be superior to

6984-450: The life sciences. Natural selection , life history , development , adaptation , populations , and inheritance are examples of concepts that thread equally into ecological and evolutionary theory. Morphological, behavioural, and genetic traits, for example, can be mapped onto evolutionary trees to study the historical development of a species in relation to their functions and roles in different ecological circumstances. In this framework,

7081-424: The lifespan of a single leaf. Each of those aphids, in turn, supports diverse bacterial communities. The nature of connections in ecological communities cannot be explained by knowing the details of each species in isolation, because the emergent pattern is neither revealed nor predicted until the ecosystem is studied as an integrated whole. Some ecological principles, however, do exhibit collective properties where

7178-432: The lower adjacent level (according to ecological pyramids ) nearer the abiotic source." Links in food webs primarily connect feeding relations or trophism among species. Biodiversity within ecosystems can be organized into trophic pyramids, in which the vertical dimension represents feeding relations that become further removed from the base of the food chain up toward top predators, and the horizontal dimension represents

7275-535: The most various kinds and sizes. They form one category of the multitudinous physical systems of the universe, which range from the universe as a whole down to the atom. Ecosystems may be habitats within biomes that form an integrated whole and a dynamically responsive system having both physical and biological complexes. Ecosystem ecology is the science of determining the fluxes of materials (e.g. carbon, phosphorus) between different pools (e.g., tree biomass, soil organic material). Ecosystem ecologists attempt to determine

7372-456: The nature of the unique physical environments that shapes the biodiversity within each. A more recent addition to ecosystem ecology are technoecosystems , which are affected by or primarily the result of human activity. A food web is the archetypal ecological network . Plants capture solar energy and use it to synthesize simple sugars during photosynthesis . As plants grow, they accumulate nutrients and are eaten by grazing herbivores , and

7469-573: The niche is called the ecotope , which is defined as the full range of environmental and biological variables affecting an entire species. Organisms are subject to environmental pressures, but they also modify their habitats. The regulatory feedback between organisms and their environment can affect conditions from local (e.g., a beaver pond ) to global scales, over time and even after death, such as decaying logs or silica skeleton deposits from marine organisms. The process and concept of ecosystem engineering are related to niche construction , but

7566-495: The number of nitrogen fixers , can lead to disproportionate, perhaps irreversible, changes in the system properties." Biodiversity refers to the variety of life and its processes. It includes the variety of living organisms, the genetic differences among them, the communities and ecosystems in which they occur, and the ecological and evolutionary processes that keep them functioning, yet ever-changing and adapting. Biodiversity (an abbreviation of "biological diversity") describes

7663-452: The number of people in a single area. Governments conduct a census to quantify the size of a resident population within a given jurisdiction. The term is also applied to non-human animals , microorganisms , and plants , and has specific uses within such fields as ecology and genetics . The word population is derived from the Late Latin populatio (a people, a multitude), which itself

7760-399: The observed data. In these island models, the rate of population change is described by: where N is the total number of individuals in the population, b and d are the per capita rates of birth and death respectively, and r is the per capita rate of population change. Using these modeling techniques, Malthus' population principle of growth was later transformed into a model known as

7857-406: The organization and structure of entire communities. The loss of a keystone species results in a range of dramatic cascading effects (termed trophic cascades ) that alters trophic dynamics, other food web connections, and can cause the extinction of other species. The term keystone species was coined by Robert Paine in 1969 and is a reference to the keystone architectural feature as the removal of

7954-418: The panmictic original, while some will be about the same, and some will be inferior. The probabilities of each can be estimated from those binomial equations. In plant and animal breeding , procedures have been developed which deliberately utilize the effects of dispersion (such as line breeding, pure-line breeding, backcrossing). Dispersion-assisted selection leads to the greatest genetic advance (ΔG=change in

8051-756: The phenotypic mean), and is much more powerful than selection acting without attendant dispersion. This is so for both allogamous (random fertilization) and autogamous (self-fertilization) gamodemes. According to the UN, the world's population surpassed 8 billion on 15 November 2022, an increase of 1 billion since 12 March 2012. According to a separate estimate by the United Nations, Earth's population exceeded seven billion in October 2011. According to UNFPA , growth to such an extent offers unprecedented challenges and opportunities to all of humanity. According to papers published by

8148-413: The practical description of patterns in quantitative reductionist terms where correlations may be identified but nothing is understood about the causal relations without reference to the whole system, which leads to 3) a metaphysical hierarchy whereby the causal relations of larger systems are understood without reference to the smaller parts. Scientific holism differs from mysticism that has appropriated

8245-468: The published work of George Perkins Marsh ("Man and Nature"). Within an ecosystem, organisms are linked to the physical and biological components of their environment to which they are adapted. Ecosystems are complex adaptive systems where the interaction of life processes form self-organizing patterns across different scales of time and space. Ecosystems are broadly categorized as terrestrial , freshwater , atmospheric, or marine . Differences stem from

8342-502: The rate of change in population size ( d N ( t ) / d t {\displaystyle \mathrm {d} N(t)/\mathrm {d} t} ) will grow to approach equilibrium, where ( d N ( t ) / d t = 0 {\displaystyle \mathrm {d} N(t)/\mathrm {d} t=0} ), when the rates of increase and crowding are balanced, r / α {\displaystyle r/\alpha } . A common, analogous model fixes

8439-424: The same limiting resource ; one will always out-compete the other. When similarly adapted species overlap geographically, closer inspection reveals subtle ecological differences in their habitat or dietary requirements. Some models and empirical studies, however, suggest that disturbances can stabilize the co-evolution and shared niche occupancy of similar species inhabiting species-rich communities. The habitat plus

8536-829: The same species. If the gamodeme is very large (theoretically, approaching infinity), and all gene alleles are uniformly distributed by the gametes within it, the gamodeme is said to be panmictic. Under this state, allele ( gamete ) frequencies can be converted to genotype ( zygote ) frequencies by expanding an appropriate quadratic equation , as shown by Sir Ronald Fisher in his establishment of quantitative genetics. This seldom occurs in nature: localization of gamete exchange – through dispersal limitations, preferential mating, cataclysm, or other cause – may lead to small actual gamodemes which exchange gametes reasonably uniformly within themselves but are virtually separated from their neighboring gamodemes. However, there may be low frequencies of exchange with these neighbors. This may be viewed as

8633-431: The same term. An example of metaphysical holism is identified in the trend of increased exterior thickness in shells of different species. The reason for a thickness increase can be understood through reference to principles of natural selection via predation without the need to reference or understand the biomolecular properties of the exterior shells. Ecology and evolutionary biology are considered sister disciplines of

8730-399: The site will disappear unless rescued by an adjacent source patch or environmental conditions become more favorable. Metapopulation models examine patch dynamics over time to answer potential questions about spatial and demographic ecology. The ecology of metapopulations is a dynamic process of extinction and colonization. Small patches of lower quality (i.e., sinks) are maintained or rescued by

8827-524: The study of ecology into a conceptually manageable framework, the biological world is organized into a nested hierarchy , ranging in scale from genes , to cells , to tissues , to organs , to organisms , to species , to populations , to guilds , to communities , to ecosystems , to biomes , and up to the level of the biosphere . This framework forms a panarchy and exhibits non-linear behaviors; this means that "effect and cause are disproportionate, so that small changes to critical variables, such as

8924-432: The sum of the components explain the properties of the whole, such as birth rates of a population being equal to the sum of individual births over a designated time frame. The main subdisciplines of ecology, population (or community ) ecology and ecosystem ecology , exhibit a difference not only in scale but also in two contrasting paradigms in the field. The former focuses on organisms' distribution and abundance, while

9021-426: The town receives greater amounts of snow than the world's leading ski areas such as Whistler, Aspen or Chamonix. Wintery weather lasts from early November through to early April with the snowiest month being January which averages 2.9 m (9.5 ft). January is also the coldest month averaging −5.7 °C (21.7 °F). Kutchan has 4 very distinct seasons with reasonably short spring and autumn periods. The summer

9118-640: The tree responds more slowly and integrates these short-term changes. The scale of ecological dynamics can operate like a closed system, such as aphids migrating on a single tree, while at the same time remaining open about broader scale influences, such as atmosphere or climate. Hence, ecologists classify ecosystems hierarchically by analyzing data collected from finer scale units, such as vegetation associations , climate, and soil types , and integrate this information to identify emergent patterns of uniform organization and processes that operate on local to regional, landscape , and chronological scales. To structure

9215-415: The underlying causes of these fluxes. Research in ecosystem ecology might measure primary production (g C/m^2) in a wetland in relation to decomposition and consumption rates (g C/m^2/y). This requires an understanding of the community connections between plants (i.e., primary producers) and the decomposers (e.g., fungi and bacteria). The underlying concept of an ecosystem can be traced back to 1864 in

9312-439: The use of a location by the animal." For example, a habitat might be an aquatic or terrestrial environment that can be further categorized as a montane or alpine ecosystem. Habitat shifts provide important evidence of competition in nature where one population changes relative to the habitats that most other individuals of the species occupy. For example, one population of a species of tropical lizard ( Tropidurus hispidus ) has

9409-589: Was coined in 1866 by the German scientist Ernst Haeckel . The science of ecology as we know it today began with a group of American botanists in the 1890s. Evolutionary concepts relating to adaptation and natural selection are cornerstones of modern ecological theory . Ecosystems are dynamically interacting systems of organisms, the communities they make up, and the non-living ( abiotic ) components of their environment. Ecosystem processes, such as primary production , nutrient cycling , and niche construction , regulate

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