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Konrad Lorenz

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120-712: Konrad Zacharias Lorenz ( Austrian German pronunciation: [ˈkɔnʁaːd tsaxaˈʁiːas ˈloːʁɛnts] ; 7 November 1903 – 27 February 1989) was an Austrian zoologist , ethologist , and ornithologist . He shared the 1973 Nobel Prize in Physiology or Medicine with Nikolaas Tinbergen and Karl von Frisch . He is often regarded as one of the founders of modern ethology , the study of animal behavior. He developed an approach that began with an earlier generation, including his teacher Oskar Heinroth . Lorenz studied instinctive behavior in animals, especially in greylag geese and jackdaws . Working with geese, he investigated

240-426: A "psychohydraulic" model of the motivation of behavior, which tended towards group selectionist ideas, which were influential in the 1960s. Another of his contributions to ethology is his work on imprinting . His influence on a younger generation of ethologists ; and his popular works, were important in bringing ethology to the attention of the general public. Lorenz claimed that there was widespread contempt for

360-409: A Napoleon to sacrifice millions of lives in his attempt to unite the world under his scepter. We have been taught to regard some of the persons who have committed these and similar absurdities with respect, even as "great" men, we are wont to yield to the political wisdom of those in charge, and we are all so accustomed to these phenomena that most of us fail to realize how abjectly stupid and undesirable

480-668: A being of pure reason, devoid of instincts himself and unaware of the way in which all instincts in general and aggression in particular can miscarry, he would be at a complete loss how to explain history at all. The ever-recurrent phenomena of history do not have reasonable causes. It is a mere commonplace to say that they are caused by what common parlance so aptly terms "human nature." Unreasoning and unreasonable human nature causes two nations to compete, though no economic necessity compels them to do so; it induces two political parties or religions with amazingly similar programs of salvation to fight each other bitterly, and it impels an Alexander or

600-652: A cream/off-white coloring around their necks. They reach a length of 30 to 56 cm (12 to 22 in), with tails that are just as long as the body. On average, they weigh from 1.4 to 4 kg (3 to 9 pounds) and live up to 25 years old in their natural habitats, and up to 35 in captivity. Capuchins prefer environments that give them access to shelter and easy food, such as low-lying forests, mountain forests, and rain forests. They are particularly abundant in Argentina, Brazil, Costa Rica, Honduras, Paraguay, and Peru. They use these areas for shelter at night and food access during

720-441: A few of the ripest fruits, nip off the tip of the fruit and drink down the juice, then seemingly discard the rest of the fruit with the nut inside. When these discarded fruits have hardened and become slightly brittle, the capuchin will gather them up again and take them to a large flat boulder where they have previously gathered a few river stones from up to a mile away. They will then use these stones, some of them weighing as much as

840-601: A laboratory setting. Ethologists have been particularly concerned with the evolution of behavior and the understanding of behavior in terms of the theory of natural selection . In one sense, the first modern ethologist was Charles Darwin , whose book, The Expression of the Emotions in Man and Animals , influenced many future ethologists. A subfield of ethology is behavioral ecology which attempts to answer Nikolaas Tinbergen 's four questions with regard to animal behavior: what are

960-496: A lower-ranking female want to mate with an alpha male, the more dominant female will get rights to the male over the lower-ranking one. The capuchin is considered to be the most intelligent New World monkey and is often kept in captivity. The tufted monkey is especially noted for its long-term tool usage, one of the few examples of primate tool use other than by apes including humans. Upon seeing macaws eating palm nuts , cracking them open with their beaks, this monkey will select

1080-399: A mechanism which tends to encourage behavior rather than to attenuate it (p43). Positive feedback always involves the danger of an 'avalanche' effect... One particular kind of positive feedback occurs when individuals OF THE SAME SPECIES enter into competition among themselves... For many animal species, environmental factors keep... intraspecies selection from [leading to] disaster... But there

1200-441: A natural insect repellent . When presented with a reflection, capuchin monkeys react in a way that indicates an intermediate state between seeing the mirror as another individual and recognizing the image as self. Most animals react to seeing their reflections as if encountering another individual they do not recognize. An experiment with capuchins shows that they react to a reflection as a strange phenomenon, but not as if seeing

1320-413: A new footing, by explaining the processes by which it can occur, and providing observational evidence that it had done so. Darwin's theory was rapidly accepted by the scientific community and soon became a central axiom of the rapidly developing science of biology. The basis for modern genetics began with the work of Gregor Mendel on peas in 1865, although the significance of his work was not realized at

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1440-406: A new perspective on the fundamental basis of life. Having previously been the realm of gentlemen naturalists, over the 18th, 19th and 20th centuries, zoology became an increasingly professional scientific discipline . Explorer-naturalists such as Alexander von Humboldt investigated the interaction between organisms and their environment, and the ways this relationship depends on geography, laying

1560-786: A popular audience. His scientific work appeared mainly in journal articles, written in German; it became widely known to English-speaking scientists through its description in Tinbergen's 1951 book The Study of Instinct , though many of his papers were later published in English translation in the two volumes titled Studies in Animal and Human Behavior . Konrad & Adolf Lorenz Museum KALM https://www.kalm.at Zoology Zoology ( UK : / z u ˈ ɒ l ə dʒ i / zoo- OL -ə-jee , US : / z oʊ ˈ ɒ l ə dʒ i / zoh- OL -ə-jee )

1680-413: A price which is not worth paying. p53. In nature, these mechanisms tend towards a 'stable state' among the living beings of an ecology: A closer examination shows that these beings... not only do not damage each other, but often constitute a community of interests. It is obvious that the predator is strongly interested in the survival of that species, animal or vegetable, which constitutes its prey. ... It

1800-709: A process involving the union of a male and female haploid gamete , each gamete formed by meiosis . Ordinarily, gametes produced by separate individuals unite by a process of fertilization to form a diploid zygote that can then develop into a genetically unique individual progeny. However, some animals are also capable, as an alternative reproductive process, to reproduce parthenogenetically. Parthenogenesis has been described in snakes and lizards (see Misplaced Pages Parthenogenesis in squamates ), in amphibians (see Misplaced Pages Parthenogenesis in amphibians ) and in numerous other species (see Misplaced Pages Parthenogenesis ). Generally, meiosis in parthanogenetically reproducing animals occurs by

1920-452: A separate sub-discipline of biology . A Review of General Psychology survey, published in 2002, ranked Lorenz the 65th most cited scholar of the 20th century in the technical psychology journals, introductory psychology textbooks, and survey responses. Lorenz's work was interrupted by the onset of World War II and in 1941 he was recruited into the German Army as a medic. In 1944, he

2040-407: A similar process to that in sexually reproducing animals, but the diploid zygote nucleus is generated by the union of two haploid genomes from the same individual rather than from different individuals. Capuchin monkey Cebus Sapajus The capuchin monkeys ( / ˈ k æ p j ʊ ( t ) ʃ ɪ n / ) are New World monkeys of the subfamily Cebinae . They are readily identified as

2160-510: A single male will dominate the group, and he will have primary rights to mate with the females of the group. However, the white-headed capuchin groups are led by both an alpha male and an alpha female. Each group will cover a large territory, since members must search for the best areas to feed. These primates are territorial animals, distinctly marking a central area of their territory with urine and defending it against intruders, though outer areas may overlap. The stabilization of group dynamics

2280-554: A strange capuchin. In the experiment, capuchins were presented with three different scenarios: In scenario 1, females appeared anxious and avoided eye-contact, while males made threatening gestures. In scenario 2, there was little reaction by either males or females. When presented with a reflection, females gazed into their own eyes and made friendly gestures, such as lip-smacking and swaying. Males made more eye contact than with strangers or familiar monkeys but reacted with signs of confusion or distress, such as squealing, curling up on

2400-531: A tanned color, ... with the lighter color around his eyes that melts into the white at the front, his cheeks ..., give him the looks that involuntarily reminds us of the appearance that historically in our country represents ignorance, laziness, and sensuality." The scientific name of the genus, Cebus comes from the Greek word kêbos , meaning a long-tailed monkey. The species-level taxonomy of this subfamily remains highly controversial, and alternative treatments than

2520-451: A wide collection of unlikely food, which may assure their survival in habitats with extreme food limitation. Capuchins living near water will also eat crabs and shellfish by cracking their shells with stones. Capuchin monkeys often live in large groups of 10 to 35 individuals within the forest, although they can easily adapt to places colonized by humans. The Capuchins have discrete hierarchies that are distinguished by age and sex. Usually,

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2640-468: A young goose instinctively bonds with the first moving stimulus it perceives, whether it be its mother, or a person. Lorenz showed that this behavior of imprinting is what allows the goose to learn to recognize members of its own species, enabling them to be the object of subsequent behavior patterns such as mating. He developed a theory of instinctive behavior that saw behavior patterns as largely innate but triggered through environmental stimuli, for example

2760-453: Is a form of scientific taxonomy . Modern biological classification has its root in the work of Carl Linnaeus , who grouped species according to shared physical characteristics. These groupings have since been revised to improve consistency with the Darwinian principle of common descent . Molecular phylogenetics , which uses nucleic acid sequence as data, has driven many recent revisions and

2880-723: Is a summary of his life's work and focuses on his famous studies of greylag geese. Lorenz was the son of Adolf Lorenz , a wealthy and distinguished surgeon, and his wife Emma (née Lecher), a physician who had been her husband's assistant. The family lived on a large estate at Altenberg, and had a city apartment in Vienna. He was educated at the Public Schottengymnasium of the Benedictine monks in Vienna . In his autobiographical essay, published in 1973 in Les Prix Nobel (winners of

3000-657: Is administered by the International Code of Zoological Nomenclature . A merging draft, BioCode, was published in 1997 in an attempt to standardize nomenclature, but has yet to be formally adopted. Vertebrate zoology is the biological discipline that consists of the study of vertebrate animals, that is animals with a backbone , such as fish , amphibians , reptiles , birds and mammals . The various taxonomically oriented disciplines i.e. mammalogy , biological anthropology , herpetology , ornithology , and ichthyology seek to identify and classify species and study

3120-480: Is an integrative field of study, uniting concepts and information from evolutionary biology , taxonomy , ecology , physical geography , geology , paleontology and climatology . The origin of this field of study is widely accredited to Alfred Russel Wallace , a British biologist who had some of his work jointly published with Charles Darwin . Molecular biology studies the common genetic and developmental mechanisms of animals and plants, attempting to answer

3240-402: Is being studied. For example, what is learned about the physiology of yeast cells can also apply to human cells. The field of animal physiology extends the tools and methods of human physiology to non-human species. Physiology studies how, for example, the nervous , immune , endocrine , respiratory , and circulatory systems function and interact. Developmental biology is the study of

3360-406: Is done on both the microscopic and molecular levels for single-celled organisms such as bacteria as well as the specialized cells in multicellular organisms such as humans . Understanding the structure and function of cells is fundamental to all of the biological sciences. The similarities and differences between cell types are particularly relevant to molecular biology. Anatomy considers

3480-443: Is likely to continue to do so. Biological classification belongs to the science of zoological systematics . Many scientists now consider the five-kingdom system outdated. Modern alternative classification systems generally start with the three-domain system : Archaea (originally Archaebacteria); Bacteria (originally Eubacteria); Eukaryota (including protists , fungi , plants , and animals ) These domains reflect whether

3600-418: Is no force which exercises this type of healthy regulatory effect on humanity's cultural development; unfortunately for itself, humanity has learned to overcome all those environmental forces which are external to itself p44. Regarding aggression in human beings, Lorenz states: Let us imagine that an absolutely unbiased investigator on another planet, perhaps on Mars, is examining human behavior on earth, with

3720-431: Is not uncommon that the prey species derives specific benefits from its interaction with the predator species... pp31–33. Lorenz states that humanity is the one species not bound by these mechanisms, being the only one that has defined its own environment: [The pace of human ecology] is determined by the progress of man's technology (p35)... human ecology (economy) is governed by mechanisms of POSITIVE feedback, defined as

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3840-415: Is partly based on paleontology , which uses the fossil record to answer questions about the mode and tempo of evolution, and partly on the developments in areas such as population genetics and evolutionary theory. Following the development of DNA fingerprinting techniques in the late 20th century, the application of these techniques in zoology has increased the understanding of animal populations. In

3960-406: Is quantitative, and recently a significant amount of work has been done using computer science techniques such as bioinformatics and computational biology . Molecular genetics , the study of gene structure and function, has been among the most prominent sub-fields of molecular biology since the early 2000s. Other branches of biology are informed by molecular biology, by either directly studying

4080-408: Is recognized as one of the founding fathers of the field of ethology , the study of animal behavior. He is best known for his research of the principle of attachment, or imprinting , through which in some species a bond is formed between a newborn animal and its caregiver. This principle had been discovered by Douglas Spalding in the 19th century, and Lorenz's mentor Oskar Heinroth had also worked on

4200-698: Is reflected in zoology by the presence of animals in Egyptian hieroglyphics. Although the concept of zoology as a single coherent field arose much later, the zoological sciences emerged from natural history reaching back to the biological works of Aristotle and Galen in the ancient Greco-Roman world . In the fourth century BC, Aristotle looked at animals as living organisms, studying their structure, development and vital phenomena. He divided them into two groups: animals with blood, equivalent to our concept of vertebrates , and animals without blood, invertebrates . He spent two years on Lesbos , observing and describing

4320-592: Is served through mutual grooming, and communication occurs between the monkeys through various calls. Their vocal communications have various meanings such as creating contact with one another, warning about a predator, and forming new groups. The social experience of the capuchins directly influences the development of attention in society. They create new social behaviors within multiple groups that signify different types of interactions. These include; tests of friendship, displays against enemies, infant and sexual intimacy. This creates social rituals that are designed to test

4440-559: Is the genus, and sapiens the specific epithet, both of them combined make up the species name. When writing the scientific name of an organism, it is proper to capitalize the first letter in the genus and put all of the specific epithet in lowercase. Additionally, the entire term may be italicized or underlined. The dominant classification system is called the Linnaean taxonomy . It includes ranks and binomial nomenclature . The classification, taxonomy , and nomenclature of zoological organisms

4560-448: Is the scientific study of animals . Its studies include the structure , embryology , classification , habits , and distribution of all animals, both living and extinct , and how they interact with their ecosystems . Zoology is one of the primary branches of biology . The term is derived from Ancient Greek ζῷον , zōion ('animal'), and λόγος , logos ('knowledge', 'study'). Although humans have always been interested in

4680-614: Is the subfield of biology that studies the evolutionary processes (natural selection, common descent, speciation) that produced the diversity of life on Earth. Evolutionary research is concerned with the origin and descent of species , as well as their change over time, and includes scientists from many taxonomically oriented disciplines. For example, it generally involves scientists who have special training in particular organisms such as mammalogy , ornithology , herpetology , or entomology , but use those organisms as systems to answer general questions about evolution. Evolutionary biology

4800-792: The Konrad Lorenz Institute for Evolution and Cognition Research (KLI) was housed in Lorenz' family mansion at Altenberg before moving to Klosterneuburg in 2013; the Konrad Lorenz Forschungsstelle (KLF) at his former field station in Grünau; and the Konrad Lorenz Institute of Ethology , an external research facility of the University of Veterinary Medicine Vienna . Lorenz predicted the relationship between market economics and

4920-850: The Nazi Party in 1938 and accepted a university chair under the Nazi regime . In his application for party membership he wrote, "I'm able to say that my whole scientific work is devoted to the ideas of the National Socialists ." His publications during that time led in later years to allegations that his scientific work had been contaminated by Nazi sympathies. His published writing during the Nazi period included support for Nazi ideas of " racial hygiene " couched in pseudoscientific metaphors. In his autobiography, Lorenz wrote: The same individual geese on which we conducted these experiments, first aroused my interest in

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5040-451: The Nazi eugenics policies against this were therefore scientifically justified. In 1940 he became a professor of psychology at the University of Königsberg . He was drafted into the Wehrmacht in 1941. He sought to be a motorcycle mechanic, but instead he was assigned as a military psychologist, conducting racial studies on humans in occupied Poznań under Rudolf Hippius . The objective

5160-498: The University of Vienna . He graduated as Doctor of Medicine (MD) in 1928 and became an assistant professor at the Institute of Anatomy until 1935. He finished his zoological studies in 1933 and received his second doctorate (PhD). While still a student, Lorenz began developing what would become a large menagerie , ranging from domestic to exotic animals. In his popular book King Solomon's Ring , Lorenz recounts that while studying at

5280-545: The anatomy of different groups. It is closely related to evolutionary biology and phylogeny (the evolution of species). Physiology studies the mechanical, physical, and biochemical processes of living organisms by attempting to understand how all of the structures function as a whole. The theme of "structure to function" is central to biology. Physiological studies have traditionally been divided into plant physiology and animal physiology , but some principles of physiology are universal, no matter what particular organism

5400-400: The hawk/goose effect . He argued that animals have an inner drive to carry out instinctive behaviors, and that if they do not encounter the right stimulus they will eventually engage in the behavior with an inappropriate stimulus. Lorenz's approach to ethology derived from a skepticism towards the studies of animal behavior done in laboratory settings. He considered that in order to understand

5520-595: The pet trade , and humans hunting for bushmeat. According to the IUCN Red List of Threatened Species , nearly all species are decreasing in population, with many facing threats of extinction. Since capuchins have a high reproductive rate and can adapt to different living environments, they can survive forest loss more than some other species; however, habitat fragmentation is still a threat. Predators include jaguars, cougars, jaguarundis , coyotes, tayras , snakes, crocodiles, birds of prey, and humans. The main predator of

5640-550: The proximate causes of the behavior, the developmental history of the organism, the survival value and phylogeny of the behavior? Another area of study is animal cognition , which uses laboratory experiments and carefully controlled field studies to investigate an animal's intelligence and learning. Biogeography studies the spatial distribution of organisms on the Earth , focusing on topics like dispersal and migration , plate tectonics , climate change , and cladistics . It

5760-399: The taxonomist . Originally it was thought that species were immutable, but with the arrival of Darwin's theory of evolution, the field of cladistics came into being, studying the relationships between the different groups or clades . Systematics is the study of the diversification of living forms, the evolutionary history of a group is known as its phylogeny , and the relationship between

5880-510: The tufted capuchin is the harpy eagle , which has been seen bringing several capuchin back to its nest. Easily recognized as the " organ grinder " or " greyhound jockey " monkeys, capuchins are sometimes kept as exotic pets . Sometimes they plunder fields and crops and are seen as troublesome by nearby human populations. In some regions, they have become rare due to the destruction of their habitat. Capuchins have been used as service animals, and were once referred to as "nature's butlers" by

6000-535: The " organ grinder " monkey, and have been used in many movies and television shows. The range of capuchin monkeys includes some tropical forests in Central America and South America as far south as northern Argentina . In Central America, where they are called white-faced monkeys ("carablanca"), they usually occupy the wet lowland forests on the Caribbean coast of Costa Rica and Panama and deciduous dry forest on

6120-542: The 16th century. During the Renaissance and early modern period, zoological thought was revolutionized in Europe by a renewed interest in empiricism and the discovery of many novel organisms. Prominent in this movement were Andreas Vesalius and William Harvey , who used experimentation and careful observation in physiology , and naturalists such as Carl Linnaeus , Jean-Baptiste Lamarck , and Buffon who began to classify

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6240-470: The 1980s, developmental biology re-entered evolutionary biology from its initial exclusion from the modern synthesis through the study of evolutionary developmental biology . Related fields often considered part of evolutionary biology are phylogenetics , systematics , and taxonomy . Ethology is the scientific and objective study of animal behavior under natural conditions, as opposed to behaviorism , which focuses on behavioral response studies in

6360-589: The AARP. Helping Hands, a nonprofit organization, trained capuchin monkeys to assist quadriplegics as monkey helpers in a manner similar to mobility assistance dogs . In 2010, the U.S. federal government revised its definition of service animal under the Americans with Disabilities Act (ADA). Non-human primates are no longer recognized as service animals under the ADA. The American Veterinary Medical Association does not support

6480-684: The Max Planck Institute in 1973 but continued to research and publish from Altenberg and Grünau im Almtal in Austria. He died on 27 February 1989 in Altenberg. Lorenz married his childhood friend, Margarethe Gebhardt, a gynaecologist, daughter of a market gardener who lived near the Lorenz family; they had a son and two daughters. He lived at the Lorenz family estate, which included a "fantastical neo-baroque mansion", previously owned by his father. Lorenz

6600-590: The Mirror . The Max Planck Society established the Lorenz Institute for Behavioral Physiology in Buldern , Germany, in 1950. In his memoirs, Lorenz described the chronology of his war years differently from what historians have been able to document after his death. He himself claimed that he was captured in 1942, where in reality he was only sent to the front and captured in 1944, leaving out entirely his involvement with

6720-507: The Nobel prize, summarized Lorenz's major contribution to ethology as making behavior a topic of biological inquiry, considering behavior a part of an animal's evolutionary equipment. Tinbergen and Lorenz contributed to making Ethology a recognized sub-discipline within Biology and founded the first specialized journal of the field " Ethology " (originally "Zeitschift für Tierpsychologie") Lorenz joined

6840-605: The Pacific coast. The word "capuchin" derives from the Order of Friars Minor Capuchin , who wear brown robes with large hoods. When Portuguese explorers reached the Americas in the 15th century, they found small monkeys whose coloring resembled these friars, especially when in their robes with hoods down, and named them capuchins. When the scientists described a specimen (thought to be a golden-bellied capuchin ) they noted that: "his muzzle of

6960-616: The Poznań project. In 1958, Lorenz transferred to the Max Planck Institute for Behavioral Physiology in Seewiesen. He shared the 1973 Nobel Prize in Physiology or Medicine "for discoveries in individual and social behavior patterns" with two other important early ethologists , Nikolaas Tinbergen and Karl von Frisch . In 1969, he became the first recipient of the Prix mondial Cino Del Duca . He

7080-466: The University of Vienna he kept a variety of animals at his parents' apartment, ranging from fish to a capuchin monkey named Gloria. In 1936, at an international scientific symposium on instinct, Lorenz met his great friend and colleague Nikolaas Tinbergen . Together they studied geese —wild, domestic , and hybrid . One result of these studies was that Lorenz "realized that an overpowering increase in

7200-415: The aid of a telescope whose magnification is too small to enable him to discern individuals and follow their separate behavior, but large enough for him to observe occurrences such as migrations of peoples, wars, and similar great historical events. He would never gain the impression that human behavior was dictated by intelligence, still less by responsible morality. If we suppose our extraneous observer to be

7320-499: The already abundant foliage that shelters the capuchin. Like most New World monkeys, capuchins are diurnal and arboreal . Capuchins are polygamous, and the females mate throughout the year, but only go through a gestation period once every 2 years between December and April. Females bear young every two years following a 160- to 180-day gestation. The young cling to their mother's chest until they are larger, then they move to her back. Adult male capuchin rarely take part in caring for

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7440-449: The animals and plants, considering the adaptations of different organisms and the function of their parts. Four hundred years later, Roman physician Galen dissected animals to study their anatomy and the function of the different parts, because the dissection of human cadavers was prohibited at the time. This resulted in some of his conclusions being false, but for many centuries it was considered heretical to challenge any of his views, so

7560-427: The cells have nuclei or not, as well as differences in the chemical composition of the cell exteriors. Further, each kingdom is broken down recursively until each species is separately classified. The order is: Domain ; kingdom ; phylum ; class ; order ; family ; genus ; species . The scientific name of an organism is generated from its genus and species. For example, humans are listed as Homo sapiens . Homo

7680-434: The clades can be shown diagrammatically in a cladogram . Although someone who made a scientific study of animals would historically have described themselves as a zoologist, the term has come to refer to those who deal with individual animals, with others describing themselves more specifically as physiologists, ethologists, evolutionary biologists, ecologists, pharmacologists, endocrinologists or parasitologists. Although

7800-852: The complete inhumanity of the Nazis". He was sent to the Russian front in 1944 where he quickly became a prisoner of war in the Soviet Union from 1944 to 1948. In captivity in Soviet Armenia , he continued to work as a medic and "became tolerably fluent in Russian and got quite friendly with some Russians, mostly doctors." When he was repatriated, he was allowed to keep the manuscript of a book he had been writing and his pet starling . He arrived back in Altenberg (his family home, near Vienna) both "with manuscript and bird intact." The manuscript became his 1973 book Behind

7920-454: The day. The canopy of the trees allows for protection from threats above, and the capuchin monkeys' innate ability to climb trees with ease allows them to escape and hide from predators on the jungle floor. This environment is mutually beneficial for the capuchins and for the ecosystem in which they inhabit. This is because they spread their seed leftovers and fecal matter across the forest floor which helps new plants to grow, therefore adding to

8040-474: The descriptive sciences. He attributed this to the denial of perception as the source of all scientific knowledge: "a denial that has been elevated to the status of religion." He wrote that in comparative behavioral research, "it is necessary to describe various patterns of movement, record them, and above all, render them unmistakably recognizable." There are three research institutions named after Lorenz in Austria:

8160-514: The determination of the double helical structure of the DNA molecule by Francis Crick and James Watson in 1953, the realm of molecular biology opened up, leading to advances in cell biology , developmental biology and molecular genetics . The study of systematics was transformed as DNA sequencing elucidated the degrees of affinity between different organisms. Zoology is the branch of science dealing with animals . A species can be defined as

8280-643: The discovery of the structure of DNA by Francis Crick and James Watson in 1953, the realm of molecular biology opened up, leading to advances in cell biology , developmental biology and molecular genetics . The history of zoology traces the study of the animal kingdom from ancient to modern times. Prehistoric people needed to study the animals and plants in their environment to exploit them and survive. Cave paintings, engravings and sculptures in France dating back 15,000 years show bison, horses, and deer in carefully rendered detail. Similar images from other parts of

8400-456: The diversity of life and the fossil record , as well as studying the development and behavior of organisms. Antonie van Leeuwenhoek did pioneering work in microscopy and revealed the previously unknown world of microorganisms , laying the groundwork for cell theory . van Leeuwenhoek's observations were endorsed by Robert Hooke ; all living organisms were composed of one or more cells and could not generate spontaneously. Cell theory provided

8520-450: The drives of feeding as well as of copulation and a waning of more differentiated social instincts is characteristic of very many domestic animals". Lorenz began to suspect and fear "that analogous processes of deterioration may be at work with civilized humanity." This observation of bird hybrids caused Lorenz to believe that domestication resulting from urbanisation in humans might also cause dysgenic effects, and to argue in two papers that

8640-724: The family Cebidae . They are omnivores , and consume a variety of plant parts such as leaves, flower and fruit, seeds, pith, woody tissue, sugarcane, bulb, and exudates, as well as arthropods , molluscs , a variety of vertebrates , and even primates . Recent findings of old stone tools in Capuchin habitats have suggested that recently the Capuchins have switched from small nuts, such as cashews, to larger and harder nuts. Capuchins have also been observed to be particularly good at catching frogs. They are characterized as innovative and extreme foragers because of their ability to acquire sustenance from

8760-759: The fear of not being up to snuff... (pp. 45–47) In his 1973 book Behind the Mirror: A Search for a Natural History of Human Knowledge , Lorenz considers the old philosophical question of whether our senses correctly inform us about the world as it is, or provide us only with an illusion. His answer comes from evolutionary biology . Only traits that help us survive and reproduce are transmitted. If our senses gave us wrong information about our environment, we would soon be extinct. Therefore, we can be sure that our senses give us correct information, for otherwise we would not be here to be deceived. Lorenz's best-known books are King Solomon's Ring and On Aggression , both written for

8880-442: The floor, or trying to escape from the test room. The question of whether capuchin monkeys have a theory of mind—whether they can understand what another creature may know or think—has been neither proven nor disproven conclusively. If confronted with a knower-guesser scenario, where one trainer can be observed to know the location of food and another trainer merely guesses the location of food, capuchin monkeys can learn to rely on

9000-404: The forms of macroscopic structures such as organs and organ systems. It focuses on how organs and organ systems work together in the bodies of humans and other animals, in addition to how they work independently. Anatomy and cell biology are two studies that are closely related, and can be categorized under "structural" studies. Comparative anatomy is the study of similarities and differences in

9120-431: The foundations for biogeography , ecology and ethology . Naturalists began to reject essentialism and consider the importance of extinction and the mutability of species . These developments, as well as the results from embryology and paleontology , were synthesized in the 1859 publication of Charles Darwin 's theory of evolution by natural selection ; in this Darwin placed the theory of organic evolution on

9240-748: The gracile and robust capuchins diverged approximately 6.2 million years ago. Lynch Alfaro suspects that the divergence was triggered by the creation of the Amazon River, which separated the monkeys in the Amazon north of the Amazon River, who then evolved into the gracile capuchins. Those in the Atlantic Forest south of the river evolved into the robust capuchins. Gracile capuchins have longer limbs relative to their body size than robust capuchins, and have rounder skulls, whereas robust capuchins have jaws better adapted for opening hard nuts. Robust capuchins have crests and

9360-542: The historical mass behavior of humanity actually is Lorenz does not see human independence from natural ecological processes as necessarily bad. He states that: A completely new [ecology] which corresponds in every way to [humanity's] desires... could, theoretically, prove as durable as that which would have existed without his intervention (36). However, the principle of competition, typical of Western societies, destroys any chance of this: The competition between human beings destroys with cold and diabolic brutality... Under

9480-400: The idea of an innate releasing mechanism to explain instinctive behaviors ( fixed action patterns ). They experimented with " supernormal stimuli " such as giant eggs or dummy bird beaks which they found could release the fixed action patterns more powerfully than the natural objects for which the behaviors were adapted. Influenced by the ideas of William McDougall , Lorenz developed this into

9600-476: The interactions of molecules in their own right such as in cell biology and developmental biology , or indirectly, where molecular techniques are used to infer historical attributes of populations or species , as in fields in evolutionary biology such as population genetics and phylogenetics . There is also a long tradition of studying biomolecules "from the ground up", or molecularly, in biophysics . Animals generally reproduce by sexual reproduction ,

9720-460: The knower. This has, however, been repudiated as conclusive evidence for a theory of mind as the monkeys may have learned to discriminate knower and guess by other means. Until recently it was believed that non-human great apes did not possess a theory of mind either, although recent research indicates this may not be correct. Human children commonly develop a theory of mind around the ages 3 and 4. Capuchin monkeys are threatened by deforestation,

9840-404: The largest group of organisms in which any two individuals of the appropriate sex can produce fertile offspring; about 1.5 million species of animal have been described and it has been estimated that as many as 8 million animal species may exist. An early necessity was to identify the organisms and group them according to their characteristics, differences and relationships, and this is the field of

9960-448: The males have beards. * Rediscovered species. The oldest known crown platyrrhine and member of Cebidae, Panamacebus transitus , is estimated to have lived 21 million years ago. It is the earliest known fossil evidence of a mammal travelling between South and North America. Capuchins are black, brown, buff or whitish, but their exact color and pattern depends on the species involved. Capuchin monkeys are usually dark brown with

10080-419: The mechanisms behind this contradiction. Thus "all species... are adapted to their environment... including not only inorganic components... but all the other living beings that inhabit the locality." p31. Fundamental to Lorenz's theory of ecology is the function of negative feedback mechanisms, which, in hierarchical fashion, dampen impulses that occur beneath a certain threshold. The thresholds themselves are

10200-470: The mechanisms of animal behavior, it was necessary to observe their full range of behaviors in their natural context. Lorenz did not carry out much traditional fieldwork but observed animals near his home. His method involved empathizing with animals, often using anthropomorphization to imagine their mental states. He believed that animals were capable of experiencing many of the same emotions as humans. Tinbergen, Lorenz's friend with whom he conjointly received

10320-431: The monkeys began exhibiting behaviors considered to reflect an understanding of the concept of a medium of exchange that were previously believed to be restricted to humans (such as responding rationally to price shocks). They showed the same propensity to avoid perceived losses demonstrated by human subjects and investors. During the mosquito season, they crush millipedes and rub the result on their backs. This acts as

10440-424: The monkeys, to crack open the fruit to get to the nut inside. Young capuchins will watch this process to learn from the older, more experienced adults but it takes them 8 years to master this. The learning behavior of capuchins has been demonstrated to be directly linked to a reward rather than curiosity. In 2005, experiments were conducted on the ability of capuchins to use money. After several months of training,

10560-691: The most common featured monkeys in film and television, with notable examples including: Night at the Museum ( and its sequels ), Outbreak , Monkey Shines , Pirates of the Caribbean: The Curse of the Black Pearl ( and its sequels ), Zookeeper , George of the Jungle , and The Hangover Part II . Ross Geller ( David Schwimmer ) on the NBC sitcom Friends had a capuchin monkey named Marcel. Crystal

10680-831: The natural history of the animals they saw around them, and used this knowledge to domesticate certain species, the formal study of zoology can be said to have originated with Aristotle . He viewed animals as living organisms, studied their structure and development, and considered their adaptations to their surroundings and the function of their parts. Modern zoology has its origins during the Renaissance and early modern period, with Carl Linnaeus , Antonie van Leeuwenhoek , Robert Hooke , Charles Darwin , Gregor Mendel and many others. The study of animals has largely moved on to deal with form and function, adaptations, relationships between groups, behaviour and ecology. Zoology has increasingly been subdivided into disciplines such as classification , physiology , biochemistry and evolution . With

10800-404: The new rulers. The precedent narrow-minded catholic regime in Austria induced better and more intelligent men than I was to cherish this naive hope. Practically all my friends and teachers did so, including my own father who certainly was a kindly and humane man. None of us as much as suspected that the word "selection", when used by these rulers, meant murder. I regret those writings not so much for

10920-439: The one listed below have been suggested. In 2011, Jessica Lynch Alfaro et al. proposed that the robust capuchins (formerly the C. apella group) be placed in a separate genus, Sapajus , from the gracile capuchins (formerly the C. capucinus group) which retain the genus Cebus . Other primatologists, such as Paul Garber , have begun using this classification. According to genetic studies led by Lynch Alfaro in 2011,

11040-435: The pressure of this competitive fury we have not only forgotten what is useful to humanity as a whole, but even that which is good and advantageous to the individual. [...] One asks, which is more damaging to modern humanity: the thirst for money or consuming haste... in either case, fear plays a very important role: the fear of being overtaken by one's competitors, the fear of becoming poor, the fear of making wrong decisions or

11160-419: The principle of imprinting , the process by which some nidifugous birds (i.e. birds that leave their nest early) bond instinctively with the first moving object that they see within the first hours of hatching. Although Lorenz did not discover the topic, he became widely known for his descriptions of imprinting as an instinctive bond. In 1936, he met Tinbergen, and the two collaborated in developing ethology as

11280-483: The prizes are requested to provide such essays), Lorenz credits his career to his parents, who "were supremely tolerant of my inordinate love for animals", and to his childhood encounter with Selma Lagerlöf 's The Wonderful Adventures of Nils , which filled him with a great enthusiasm about wild geese." At the request of his father, Adolf Lorenz, he began a premedical curriculum in 1922 at Columbia University , but he returned to Vienna in 1923 to continue his studies at

11400-418: The process of domestication. They were F1 hybrids of wild Greylags and domestic geese and they showed surprising deviations from the normal social and sexual behaviour of the wild birds. I realised that an overpowering increase in the drives of feeding as well as of copulation and a waning of more differentiated social instincts is characteristic of very many domestic animals. I was frightened – as I still am – by

11520-517: The processes by which animals and plants reproduce and grow. The discipline includes the study of embryonic development , cellular differentiation , regeneration , asexual and sexual reproduction , metamorphosis , and the growth and differentiation of stem cells in the adult organism. Development of both animals and plants is further considered in the articles on evolution , population genetics , heredity , genetic variability , Mendelian inheritance , and reproduction . Evolutionary biology

11640-468: The product of the interaction of contrasting mechanisms. Thus pain and pleasure act as checks on each other: To gain a desired prey, a dog or wolf will do things that, in other contexts, they would shy away from: run through thorn bushes, jump into cold water and expose themselves to risks which would normally frighten them. All these inhibitory mechanisms... act as a counterweight to the effects of learning mechanisms... The organism cannot allow itself to pay

11760-493: The questions regarding the mechanisms of genetic inheritance and the structure of the gene . In 1953, James Watson and Francis Crick described the structure of DNA and the interactions within the molecule, and this publication jump-started research into molecular biology and increased interest in the subject. While researchers practice techniques specific to molecular biology, it is common to combine these with methods from genetics and biochemistry . Much of molecular biology

11880-633: The racist ideology of National Socialism". During the final years of his life, Lorenz supported the fledgling Austrian Green Party and in 1984 became the figurehead of the Konrad Lorenz Volksbegehren , a grass-roots movement that was formed to prevent the building of a power plant at the Danube near Hainburg an der Donau and thus the destruction of the surrounding woodland. Lorenz has been called 'The father of ethology', by Niko Tinbergen. Perhaps Lorenz's most important contribution to ethology

12000-494: The results of his own investigations. His general approach was surprisingly modern, and he wrote, "For it is [the task] of natural science not simply to accept what we are told but to inquire into the causes of natural things." An early pioneer was Conrad Gessner , whose monumental 4,500-page encyclopedia of animals, Historia animalium , was published in four volumes between 1551 and 1558. In Europe, Galen's work on anatomy remained largely unsurpassed and unchallenged up until

12120-503: The rise of the germ theory of disease , though the mechanism of inheritance remained a mystery. In the early 20th century, the rediscovery of Mendel's work led to the rapid development of genetics , and by the 1930s the combination of population genetics and natural selection in the modern synthesis created evolutionary biology . Research in cell biology is interconnected to other fields such as genetics, biochemistry , medical microbiology , immunology , and cytochemistry . With

12240-478: The strength of social bonds and a reliance on social learning. Capuchin females often direct most of their proceptive and mating behavior towards the alpha male. However, when the female reaches the end of her proceptive period, she may sometimes mate with up to six different subordinate males in one day. Strictly targeting the alpha male does not happen every time, as some females have been observed to mate with three to four different males. When an alpha female and

12360-499: The structures and mechanisms specific to those groups. The rest of the animal kingdom is dealt with by invertebrate zoology , a vast and very diverse group of animals that includes sponges , echinoderms , tunicates , worms , molluscs , arthropods and many other phyla , but single-celled organisms or protists are not usually included. Cell biology studies the structural and physiological properties of cells , including their behavior , interactions, and environment . This

12480-529: The study of anatomy stultified. During the post-classical era , Middle Eastern science and medicine was the most advanced in the world, integrating concepts from Ancient Greece, Rome, Mesopotamia and Persia as well as the ancient Indian tradition of Ayurveda , while making numerous advances and innovations. In the 13th century, Albertus Magnus produced commentaries and paraphrases of all Aristotle's works; his books on topics like botany , zoology, and minerals included information from ancient sources, but also

12600-516: The study of animal life is ancient, its scientific incarnation is relatively modern. This mirrors the transition from natural history to biology at the start of the 19th century. Since Hunter and Cuvier , comparative anatomical study has been associated with morphography , shaping the modern areas of zoological investigation: anatomy , physiology , histology , embryology , teratology and ethology . Modern zoology first arose in German and British universities. In Britain, Thomas Henry Huxley

12720-470: The thought that analogous genetical processes of deterioration may be at work with civilized humanity. Moved by this fear, I did a very ill-advised thing soon after the Germans had invaded Austria: I wrote about the dangers of domestication and, in order to be understood, I couched my writing in the worst of nazi-terminology. I do not want to extenuate this action. I did, indeed, believe that some good might come of

12840-462: The threat of ecological catastrophe. In his 1973 book, Civilized Man's Eight Deadly Sins , Lorenz addresses the following paradox: All the advantages that man has gained from his ever-deepening understanding of the natural world that surrounds him, his technological, chemical and medical progress, all of which should seem to alleviate human suffering... tends instead to favor humanity's destruction Lorenz adopts an ecological model to attempt to grasp

12960-433: The time. Darwin gave a new direction to morphology and physiology , by uniting them in a common biological theory: the theory of organic evolution. The result was a reconstruction of the classification of animals upon a genealogical basis, fresh investigation of the development of animals, and early attempts to determine their genetic relationships. The end of the 19th century saw the fall of spontaneous generation and

13080-515: The topic, but Lorenz's description of Prägung , imprinting, in nidifugous birds such as greylag geese in his 1935 book Der Kumpan in der Umwelt des Vogels ("The Companion in the Environment of Birds") became the foundational description of the phenomenon. Here, Lorenz used Jakob von Uexküll 's concept of Umwelt to understand how the limited perception of animals filtered out certain phenomena with which they interacted instinctively. For example,

13200-486: The undeniable discredit they reflect on my person as for their effect of hampering the future recognition of the dangers of domestication. With significant post-war activity After the war, Lorenz denied having been a party member, until his membership application was made public; and he denied having known the extent of the genocide, despite his position as a psychologist in the Office of Racial Policy . This practice of denial

13320-440: The use of nonhuman primates as assistance animals because of animal welfare concerns, the potential for serious injury to people, and risks that primates may transfer dangerous diseases to humans. In 2021, Helping Hands (the organization that provided helper monkeys to disabled persons) rebranded, changing its name to Envisioning Access and replaced the use of monkeys with a focus on new assistive technologies. Capuchin monkeys are

13440-489: The world illustrated mostly the animals hunted for food and the savage animals. The Neolithic Revolution , which is characterized by the domestication of animals , continued throughout Antiquity. Ancient knowledge of wildlife is illustrated by the realistic depictions of wild and domestic animals in the Near East, Mesopotamia, and Egypt, including husbandry practices and techniques, hunting and fishing. The invention of writing

13560-423: The young. Juveniles become fully mature within four years for females and eight years for males. In captivity, individuals have reached an age of 50 years, although natural life expectancy is only 15 to 25 years. Capuchins live in groups of 6–40 members, consisting of related females, their offspring, and several males. The capuchin monkey feeds on a vast range of food types, and is more varied than other monkeys in

13680-401: Was a friend and student of renowned biologist Sir Julian Huxley (grandson of "Darwin's bulldog", Thomas Henry Huxley ). Famed psychoanalyst Ralph Greenson and Sir Peter Scott were good friends. Lorenz and Karl Popper were childhood friends; many years after they met, during the celebration of Popper's 80 years, they wrote together a book entitled Die Zukunft ist offen . He retired from

13800-502: Was a prominent figure. His ideas were centered on the morphology of animals. Many consider him the greatest comparative anatomist of the latter half of the 19th century. Similar to Hunter , his courses were composed of lectures and laboratory practical classes in contrast to the previous format of lectures only. Scientific classification in zoology , is a method by which zoologists group and categorize organisms by biological type , such as genus or species . Biological classification

13920-405: Was common practice in postwar Austria, as it allowed Nazi-involved academics to return to their posts after WWII and the postwar administration was all too happy not to ask too many questions. These rehirings included Nazi functionaries (e.g. Eberhard Kranzmayer , Richard Wolfram ), an very early NSDAP members (e.g. Otto Höfler ), who were thus able to influence entire fields. Lorenz, for instance,

14040-446: Was his idea that behavior patterns can be studied as anatomical organs. This concept forms the foundation of ethological research. However, Richard Dawkins called Lorenz a "'good of the species' man", stating that the idea of group selection was "so deeply ingrained" in Lorenz's thinking that he "evidently did not realize that his statements contravened orthodox Darwinian theory ." Together with Nikolaas Tinbergen , Lorenz developed

14160-514: Was sent to the Eastern Front where he was captured by the Soviet Red Army and spent four years as a German prisoner of war in Soviet Armenia . After the war, he regretted his membership in the Nazi Party . Lorenz wrote numerous books, some of which, such as King Solomon's Ring , On Aggression , and Man Meets Dog , became popular reading. His last work Here I Am – Where Are You?

14280-470: Was shown to have made anti-Semitic jokes on 'Jewish characteristics' in letters to his mentor Heinroth. In 2015, the University of Salzburg posthumously rescinded an honorary doctorate awarded to Lorenz in 1983, citing his party membership and his assertions in his application that he was "always a National Socialist", and that his work "stands to serve National Socialist thought". The university also accused him of using his work to spread "basic elements of

14400-470: Was to study the biological characteristics of "German-Polish half-breeds" to determine whether they 'benefited' from the same work ethics as 'pure' Germans. The degree to which Lorenz participated in the project is unknown, but the project director Hippius referred a couple of times to Lorenz as an "examining psychologist". Lorenz later described that he once saw transports of concentration camp inmates at Fort VII near Poznań , which made him "fully realize

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