41-498: Quercus coccifera , the kermes oak or holly oak , is an oak shrub or tree in section Ilex of the genus. It has many synonyms , including Quercus calliprinos . It is native to the Mediterranean region and Northern African Maghreb , south to north from Morocco to France and west to east from Portugal to Cyprus and Turkey , crossing Spain , Italy , Libya , Balkans, and Greece , including Crete . The Kermes oak
82-481: A cup covered in dense, elongated, reflexed scales. It blooms from March to May in weather still wet. It is easily propagated by seed , an acorn that lies dormant until germinated by wet weather. This might occur anywhere from late summer to late autumn or early winter (October, November or December) of the following year. The acorns are very bitter, varying greatly in size and shape from one specimen to another and tasting bad. Acorns can germinate even before falling from
123-513: A feature it shares with other oak species in similar habitats in other parts of the world, such as the chaparral communities from various parts of the Americas. The word chaparro comes from the Basque txapar meaning "little thicket". Quercus coccifera is placed in section Ilex . Quercus calliprinos Webb, syn. Quercus coccifera subsp. calliprinos (Webb) Holmboe, the 'Palestinian oak' of
164-621: A narrow belt along the Himalayas to cover mainland and island Southeast Asia as far as Sumatra , Java , Borneo , and Palawan . Finally, oaks of multiple sections ( Cyclobalanopsis , Ilex , Cerris , Quercus and related genera like Lithocarpus and Castanopsis ) extend across east Asia including China, Korea, and Japan. Potential records of Quercus have been reported from Late Cretaceous deposits in North America and East Asia. These are not considered definitive, as macrofossils older than
205-485: A range of flavours, colours, and aromas. The spongy bark of the cork oak is used to make traditional wine bottle corks. Almost a third of oak species are threatened with extinction due to climate change, invasive pests , and habitat loss . In culture, the oak tree is a symbol of strength and serves as the national tree of many countries. In Indo-European and related religions, the oak is associated with thunder gods . Individual oak trees of cultural significance include
246-535: A shrub, withstanding overgrazing . It is indifferent to soil chemistry , living on calcareous, pebbly, stony and poor soils. A lover of warm weather, it starts to fail from 1,000 m (3,300 ft) above sea level. It is capable of supporting the continental Mediterranean climate with more extreme temperatures and low rainfall, replacing Quercus ilex (holm oak) in drier areas where it excels in drought resistance. It also grows on sea cliffs and in windy areas where other species of Quercus or Pinus cannot resist
287-508: A single oak tree produces both staminate ('male') flowers in the form of catkins , and small pistillate ('female') flowers, meaning that the trees are monoecious . The fruit is a nut called an acorn , borne in a cup-like structure known as a cupule ; each acorn usually contains one seed and takes 6–18 months to mature, depending on the species. The acorns and leaves contain tannic acid , which helps to guard against fungi and insects. There are some 500 extant species of oaks. Trees in
328-486: A small tree , reaching 1–6 metres ( 3 + 1 ⁄ 2 – 19 + 1 ⁄ 2 ft) tall (with 10 m or 33 ft specimens recorded in Kouf , Libya) and 50 centimetres (20 inches) in trunk diameter. It is evergreen , with spiny-serrated coriaceous leaves 1.5–4 cm long and 1–3 cm broad. The acorns are 2–3 cm long and 1.5–2 cm in diameter when mature about 18 months after pollination. They are held in
369-546: A small area of the west of Cuba ; in Mesoamerica it occurs mainly above 1000 metres. The genus crossed the isthmus of Panama when the northern and southern continents came together and is present as one species, Q. humboldtii , above 1000 metres in Colombia. The oaks of North America are of many sections ( Protobalanus , Lobatae , Ponticae , Quercus , and Virentes ) along with related genera such as Notholithocarpus . In
410-475: A species is a binomial or binomen, and comprises two Latin words, the first denoting the genus and the second denoting the species. The scientific name of a subspecies is formed slightly differently in the different nomenclature codes. In zoology, under the International Code of Zoological Nomenclature (ICZN), the scientific name of a subspecies is termed a trinomen , and comprises three words, namely
451-426: A species. Botanists and mycologists have the choice of ranks lower than subspecies, such as variety (varietas) or form (forma), to recognize smaller differences between populations. In biological terms, rather than in relation to nomenclature, a polytypic species has two or more genetically and phenotypically divergent subspecies, races , or more generally speaking, populations that differ from each other so that
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#1733092956719492-417: A subspecies. A common criterion for recognizing two distinct populations as subspecies rather than full species is the ability of them to interbreed even if some male offspring may be sterile. In the wild, subspecies do not interbreed due to geographic isolation or sexual selection . The differences between subspecies are usually less distinct than the differences between species. The scientific name of
533-438: Is a rank below species , used for populations that live in different areas and vary in size, shape, or other physical characteristics ( morphology ), but that can successfully interbreed. Not all species have subspecies, but for those that do there must be at least two. Subspecies is abbreviated as subsp. or ssp. and the singular and plural forms are the same ("the subspecies is" or "the subspecies are"). In zoology , under
574-464: Is a subspecies or a full species, the species name may be written in parentheses. Thus Larus (argentatus) smithsonianus means the American herring gull ; the notation within the parentheses means that some consider it a subspecies of a larger herring gull species and therefore call it Larus argentatus smithsonianus , while others consider it a full species and therefore call it Larus smithsonianus (and
615-966: Is as a habitat and food source in these areas (they have edible acorns, although with a very bitter taste) for nesting birds, foxes, rodents and wild boars. It forms thickets, thorny and dense, some recorded as tall as 5 m. It is sometimes accompanied by other plant species of the same size and climber plants such as asparagus or Mediterranean smilax . Q. coccifera is associated with several asparagus species, Crataegus monogyna , Mediterranean dwarf palm , ephedra , myrtle , several species of Junipers ( Juniperus, sabinas ...), Pistacia terebinthus , mastic , wild Olea europea , sarsaparilla , Rhamnus atlantica , Rhamnus lycioides , Rhamnus oleoides , Rhamnus catharticus etc. The communities receiving several characteristic names. Kermes oaks have become scarce due to their replacement in wet zones by larger species such as Holm oak. It has also suffered from extensive culling for use as charcoal. It
656-452: Is from Old English ac (seen in placenames such as Acton , from ac + tun , "oak village" ), which in turn is from Proto-Germanic *aiks , "oak". Oaks are hardwood ( dicotyledonous ) trees, deciduous or evergreen, with spirally arranged leaves , often with lobate margins ; some have serrated leaves or entire leaves with smooth margins. Many deciduous species are marcescent , not dropping dead leaves until spring. In spring,
697-987: Is located throughout the region around the Mediterranean Sea , especially in central southern and eastern halves. It is similarly found on islands in the Mediterranean, from the Balearic Islands to Cyprus . It is common in Crete and can survive heavy sheep and goat grazing for long periods as a ground cover a few centimeters high. The same is true in Mallorca , Ibiza and the Iberian Peninsula . The species grows on dry, sunny slopes. It supports both subhumid climates with dry summers or semi-desert climates with rainfall between 400 and 600 millimetres (16 and 24 in). A very hardy species, it grows well in all types of soils as
738-560: Is the only food and shelter for wildlife in some areas, such as the Ebro valley and other dry areas where chaparral replaces oaks due to low rainfall. It can survive heavy sheep and goat grazing for a long time as a ground carpet a few cm high, and will grow higher as a bush or a tree according to how much the grazing pressure is slackened. It is included as an endangered species in the Red Book of Bulgaria . In Cyprus village of Kalopanagiotis ,
779-571: Is widely disputed, with many authors, particularly in the eastern Mediterranean, considering it distinct at least at subspecies rank, if not as a species. Quercus coccifera is an important Mediterranean bush or dwarf vegetation, where the biome it dominates often bears its name ( maquis , coscojar , garrigue , carrascal , chaparral , etc.). Q. coccifera forms monospecific communities or communities integrated with Pinus , mediterranean buckthorns , Myrtus , Arecaceae , junipers , Pistacia , Rosmarinus , Thymus , etc. It
820-559: The International Code of Zoological Nomenclature , the subspecies is the only taxonomic rank below that of species that can receive a name. In botany and mycology , under the International Code of Nomenclature for algae, fungi, and plants , other infraspecific ranks , such as variety , may be named. In bacteriology and virology , under standard bacterial nomenclature and virus nomenclature , there are recommendations but not strict requirements for recognizing other important infraspecific ranks. A taxonomist decides whether to recognize
861-622: The Paleogene , and possibly from before the Eocene are mostly poorly preserved without critical features for certain identification. Amongst the oldest unequivocal records of Quercus are pollen from Austria, dating to the Paleocene -Eocene boundary, around 55 million years ago. The oldest records of Quercus in North America are from Oregon , dating to the Middle Eocene, around 44 million years ago, with
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#1733092956719902-810: The Royal Oak in Britain, the Charter Oak in the United States, and the Guernica Oak in the Basque Country . The generic name Quercus is Latin for "oak", derived from Proto-Indo-European *kwerkwu- , "oak", which is also the origin of the name " fir ", another important or sacred tree in Indo-European culture. The word "cork", for the bark of the cork oak , similarly derives from Quercus . The common name "oak"
943-436: The oak apple , and a large number of pests and diseases. Oak leaves and acorns contain enough tannin to be toxic to cattle, but pigs are able to digest them safely. Oak timber is strong and hard, and has found many uses in construction and furniture-making. The bark was traditionally used for tanning leather . Wine barrels are made of oak; these are used for aging alcoholic beverages such as sherry and whisky , giving them
984-767: The Northern Hemisphere; it includes some 500 species, both deciduous and evergreen . Fossil oaks date back to the Middle Eocene . Molecular phylogeny shows that the genus is divided into Old World and New World clades , but many oak species hybridise freely, making the genus's history difficult to resolve. Ecologically, oaks are keystone species in habitats from Mediterranean semi-desert to subtropical rainforest . They live in association with many kinds of fungi including truffles . Oaks support more than 950 species of caterpillar , many kinds of gall wasp which form distinctive galls , roundish woody lumps such as
1025-687: The Old World, oaks of section Quercus extend across the whole of Europe including European Russia apart from the far north, and north Africa (north of the Sahara) from Morocco to Libya. In Mediterranean Europe, they are joined by oaks of the sections Cerris and Ilex , which extend across Turkey, the Middle East, Iran, Afghanistan and Pakistan , while section Ponticae is endemic to the western Caucasus in Turkey and Georgia . Oaks of section Cyclobalanopsis extend in
1066-449: The binomen followed by the name of the subspecies. For example, the binomen for the leopard is Panthera pardus . The trinomen Panthera pardus fusca denotes a subspecies, the Indian leopard . All components of the trinomen are written in italics. In botany , subspecies is one of many ranks below that of species, such as variety , subvariety , form , and subform. To identify the rank,
1107-466: The eastern Mediterranean, has been distinguished from the kermes oak by its larger size (more often a tree, up to 18 m) and larger acorns over 2 cm diameter. In Israel it is called the 'common oak' ( Hebrew : אלון מצוי , IPA: [a'lon ma'tsuj] ) or the ' Palestine oak'. As of February 2023, Plants of the World Online regards it as a synonym of Quercus coccifera , but this
1148-404: The genus Quercus consisted of Old World and New World clades. The entire genome of Quercus robur (the pedunculate oak) has been sequenced , revealing an array of mutations that may underlie the evolution of longevity and disease resistance in oaks. In addition, hundreds of oak species have been compared (at RAD-seq loci), allowing a detailed phylogeny to be constructed. However,
1189-625: The genus are often large and slow-growing; Q. alba can reach an age of 600 years, a diameter of 13 feet (4.0 m) and a height of 145 feet (44 m). The Granit oak in Bulgaria, a Q. robur specimen, has an estimated age of 1637 years, making it the oldest oak in Europe. The Wi'aaSal tree, a live oak in the reservation of the Pechanga Band of Indians , California, is at least 1000 years old, and might be as much as 2000 years old, which would make it
1230-420: The harsh weather conditions. Populations typically occur in desert regions without any inhabited nucleus because crops are not economically profitable and the climate becomes progressively more continental and drier and therefore end in extreme temperatures accompanied by slow-growing dwarf juniper species. It is the last species of genus Quercus to disappear when rainfall is lacking. Their ecological importance
1271-1166: The high signal of introgressive hybridization (the transfer of genetic material by repeated backcrossing with hybrid offspring) in the genus has made it difficult to resolve an unambiguous, unitary history of oaks. The phylogeny from Hipp et al. 2019 is: CTB lineage [REDACTED] Cyclobalanoides Glauca [REDACTED] Acuta [REDACTED] Semiserrata East Asian Cerris [REDACTED] West Eurasian Cerris [REDACTED] Early-diverging Ilex [REDACTED] East Asian Ilex [REDACTED] Himalaya-Mediterranean [REDACTED] Himalayan subalpine [REDACTED] Agrifoliae [REDACTED] Palustres [REDACTED] Coccineae (Rubrae) [REDACTED] Phellos (Laurifoliae) [REDACTED] Texas red oaks [REDACTED] Erythromexicana [REDACTED] [REDACTED] [REDACTED] [REDACTED] Dumosae [REDACTED] Prinoids [REDACTED] Albae [REDACTED] Roburoids [REDACTED] Stellatae [REDACTED] Texas white oaks [REDACTED] Leucomexicana [REDACTED] Subspecies In biological classification , subspecies ( pl. : subspecies)
Quercus coccifera - Misplaced Pages Continue
1312-459: The largest number of oak species, with approximately 160 species in Mexico, of which 109 are endemic, and about 90 in the United States. The second greatest area of oak diversity is China, with approximately 100 species. In the Americas, Quercus is widespread from Vancouver and Nova Scotia in the south of Canada, south to Mexico and across the whole of the eastern United States. It is present in
1353-547: The oldest oak in the US. Among the smallest oaks is Q. acuta , the Japanese evergreen oak. It forms a bush or small tree to a height of some 30 feet (9.1 m). The genus Quercus is native to the Northern Hemisphere and includes deciduous and evergreen species extending from cool temperate to tropical latitudes in the Americas , Asia, Europe, and North Africa. North America has
1394-855: The oldest records in Asia from the Middle Eocene of Japan; both forms have affinities to the Cyclobalanopsis group. Quercus forms part, or rather two parts, of the Quercoideae subfamily of the Fagaceae , the beech family. Modern molecular phylogenetics suggests the following relationships: Fagus (beeches) Trigonobalanus (3 evergreen species) Lithocarpus (stone oaks) Chrysolepis (chinquapins) Quercus pro parte Notholithocarpus (tan oak) Quercus pro parte Castanopsis (also called chinquapins) Castanea (chestnuts) Molecular techniques for phylogenetic analysis show that
1435-492: The plant, but Q. coccifera is also multiplied by root suckers and layering. Quercus coccifera was first described by Carl Linnaeus in 1753. It is called " chêne des garrigues " (garrigue oak) in French. The term " garrigue " comes from Catalan or Occitan " garric " (meaning "twisted") the name for Q. coccifera in those languages. The common Spanish name of Q. coccifera is chaparro , which refers to its small size,
1476-467: The rank of variety are taken to be names of subspecies (see International Code of Nomenclature of Prokaryotes ). As in botany, subspecies is conventionally abbreviated as "subsp.", and is used in the scientific name: Bacillus subtilis subsp. spizizenii . In zoological nomenclature , when a species is split into subspecies, the originally described population is retained as the "nominotypical subspecies" or "nominate subspecies", which repeats
1517-418: The same name as the species. For example, Motacilla alba alba (often abbreviated M. a. alba ) is the nominotypical subspecies of the white wagtail ( Motacilla alba ). The subspecies name that repeats the species name is referred to in botanical nomenclature as the subspecies " autonym ", and the subspecific taxon as the "autonymous subspecies". When zoologists disagree over whether a certain population
1558-556: The subspecific name must be preceded by "subspecies" (which can be abbreviated to "subsp." or "ssp."), as in Schoenoplectus californicus subsp. tatora . In bacteriology , the only rank below species that is regulated explicitly by the code of nomenclature is subspecies , but infrasubspecific taxa are extremely important in bacteriology; Appendix 10 of the code lays out some recommendations that are intended to encourage uniformity in describing such taxa. Names published before 1992 in
1599-400: The tallest tree of this species has grown to a height of 17 meters. The tree is approximately 700 years old. Oak See also List of Quercus species . An oak is a hardwood tree or shrub in the genus Quercus of the beech family . They have spirally arranged leaves, often with lobed edges, and a nut called an acorn , borne within a cup. The genus is widely distributed in
1640-425: The user of the notation is not taking a position). A subspecies is a taxonomic rank below species – the only such rank recognized in the zoological code, and one of three main ranks below species in the botanical code. When geographically separate populations of a species exhibit recognizable phenotypic differences, biologists may identify these as separate subspecies; a subspecies is a recognized local variant of
1681-453: Was historically important as the food plant of Kermes scale insects, from which a red dye called crimson was obtained. The etymology of the specific name coccifera is related to the production of red cochineal (crimson) dye and derived from Latin coccum which was from Greek κόκκος , the kermes insect. The Latin -fera means 'bearer'. Quercus coccifera is usually a shrub less than 2 metres ( 6 + 1 ⁄ 2 feet) high, rarely