141-477: Keibu Keioiba , also known as Kabui Keioiba , is a mythical creature with the head of a tiger and the body of a human in the Meitei mythology and folklore of Manipur . According to legend, he was once a skilful priest named Kabui Salang Maiba. He used witchcraft to turn himself into a ferocious tiger. As punishment of his pride , he could not completely turn back to his original human form. Keibu Keioiba
282-599: A fossil record extending back about 300 million years to the Paleozoic in the late Carboniferous period; even many of the modern genera are recognizable from fossils 60–120 million years old. Other classes and orders, now long extinct, also occur as fossils, particularly from the late Paleozoic and Mesozoic eras. Fossil conifers included many diverse forms, the most dramatically distinct from modern conifers being some herbaceous conifers with no woody stems. Major fossil orders of conifers or conifer-like plants include
423-683: A sister group to Pinales (the 'gnepine' hypothesis) or as being more derived than Pinales but sister to the rest of the group. Most recent studies favor the 'gnepine' hypothesis. The earliest conifers appear in the fossil record during the Late Carboniferous ( Pennsylvanian ), over 300 million years ago. Conifers are thought to be most closely related to the Cordaitales , a group of extinct Carboniferous-Permian trees and clambering plants whose reproductive structures had some similarities to those of conifers. The most primitive conifers belong to
564-403: A black tip marks the end. The tiger is one of only a few striped cat species. Stripes are advantageous for camouflage in vegetation with vertical patterns of light and shade, such as trees, reeds and tall grass. This is supported by a Fourier analysis study showing that the striping patterns line up with their environment. The orange colour may also aid in concealment, as the tiger's prey
705-547: A comprehensive analysis of morphological, ecological and mitochondrial DNA (mtDNA) traits of all putative tiger subspecies. In 2017, the Cat Classification Task Force of the IUCN Cat Specialist Group revised felid taxonomy in accordance with the 2015 two-subspecies proposal and recognised only P. t. tigris and P. t. sondaica . Results of a 2018 whole-genome sequencing study of 32 samples from
846-443: A deceased individual, can be taken over in days or weeks. Male tigers are generally less tolerant of other males within their home ranges than females are of other females. Disputes are usually solved by intimidation rather than fighting. Once dominance has been established, a male may tolerate a subordinate within his range, as long as they do not come near him. The most serious disputes tend to occur between two males competing for
987-422: A female multicellular gametophyte. The female gametophytes grow to produce two or more archegonia , each of which contains an egg. Upon fertilization, the diploid egg will give rise to the embryo, and a seed is produced. The female cone then opens, releasing the seeds which grow to a young seedling . Conifer reproduction is synchronous with seasonal changes in temperate zones. Reproductive development slows to
1128-399: A female cone and are drawn into a tiny opening on the ovule called the micropyle . It is within the ovule that pollen-germination occurs. From here, a pollen tube seeks out the female gametophyte, which contains archegonia each with an egg, and if successful, fertilization occurs. The resulting zygote develops into an embryo , which along with the female gametophyte (nutritional material for
1269-401: A female in oestrus . Though tigers mostly live alone, relationships between individuals can be complex. Tigers are particularly social at kills and a male tiger will sometimes share a carcass with the females and cubs within this home range and unlike male lions, will allow them to feed on the kill before he is finished with it. However, a female is more tense when encountering another female at
1410-495: A few more weeks. They can leave the denning site after two months and around the same time they start eating meat. The mother only leaves them alone to hunt and even then she does not travel far. When she suspects an area is no longer safe, she moves her cubs to a new spot, transporting them one by one by grabbing them by the scruff of the neck with her mouth. A tigress in Sikhote-Alin Biosphere Reserve maximised
1551-665: A halt during each winter season and then resumes each spring. The male strobilus development is completed in a single year. Conifers are classified by three reproductive cycles that refer to the completion of female strobilus development from initiation to seed maturation. All three types of reproductive cycle have a long gap between pollination and fertilization . One year reproductive cycle : The genera include Abies , Picea , Cedrus , Pseudotsuga , Tsuga , Keteleeria ( Pinaceae ) and Cupressus , Thuja , Cryptomeria , Cunninghamia and Sequoia ( Cupressaceae ) . Female strobili are initiated in late summer or fall of
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#17330847361571692-507: A hollow utong (bamboo pipe) open at both ends. She tells Keibu Keioiba to fetch water from the stream with the utong. In Keibu Keioiba's absence, the brothers burn down Keibu Keioiba's house, kill the baby, and run away with their sister. On the other hand, Keibu Keioiba could not collect water with the utong; when he tried, the water leaked. A crow, observing his activities from a treetop mocked his foolishness. Keibu Keioiba returns home to find his son dead, his wife missing, and his house burnt to
1833-489: A kill as early as 11 months and reach independence as a juvenile of 18 to 24 months of age; males become independent earlier than females. Radio-collared tigers in Chitwan started leaving their natal areas at the age of 19 months. Young females are sexually mature at three to four years, whereas males are at four to five years. Generation length of the tiger is about 7–10 years. Wild Bengal tigers live 12–15 years. Data from
1974-471: A kill. During friendly encounters and bonding, tigers rub against each other's bodies. Facial expressions include the "defence threat", which involves a wrinkled face, bared teeth, pulled-back ears and widened pupils. Both males and females show a flehmen response , a characteristic curled-lip grimace, when smelling urine markings. Males also use the flehmen to detect the markings made by tigresses in oestrus. Tigers will move their ears around to display
2115-413: A maximum of energy from weak sunshine at high latitudes or under forest canopy shade. Conifers from hotter areas with high sunlight levels (e.g. Turkish pine Pinus brutia ) often have yellower-green leaves, while others (e.g. blue spruce , Picea pungens ) may develop blue or silvery leaves to reflect ultraviolet light. In the great majority of genera the leaves are evergreen , usually remaining on
2256-853: A needle-like appearance, but others, including most of the Cupressaceae and some of the Podocarpaceae , have flat, triangular scale-like leaves. Some, notably Agathis in Araucariaceae and Nageia in Podocarpaceae, have broad, flat strap-shaped leaves. Others such as Araucaria columnaris have leaves that are awl-shaped. In the majority of conifers, the leaves are arranged spirally, the exceptions being most of Cupressaceae and one genus in Podocarpaceae, where they are arranged in decussate opposite pairs or whorls of 3 (−4). In many species with spirally arranged leaves, such as Abies grandis (pictured),
2397-495: A preference for sambar deer , Manchurian wapiti , barasingha , gaur and wild boar . Abundance and body weight of prey species are assumed to be the main criteria for the tiger's prey selection, both inside and outside protected areas. It also preys opportunistically on smaller species like monkeys , peafowl and other ground-based birds, porcupines and fish. Occasional attacks on Asian elephants and Indian rhinoceroses have also been reported. More often, tigers take
2538-603: A private, usually vegetated spot no further than 183 m (600 ft), though they have been recorded dragging them 549 m (1,801 ft). They are strong enough to drag the carcass of a fully grown buffalo for some distance. They rest for a while before eating and can consume as much as 50 kg (110 lb) of meat in one session, but feed on a carcass for several days, leaving little for scavengers. In much of their range, tigers share habitat with leopards and dholes . They typically dominate both of them, though with dholes it depends on their pack size. Interactions between
2679-402: A protective cone called a strobilus . The cones take from four months to three years to reach maturity, and vary in size from 2 to 600 millimetres ( 1 ⁄ 8 to 23 + 5 ⁄ 8 in) long. In Pinaceae , Araucariaceae , Sciadopityaceae and most Cupressaceae , the cones are woody , and when mature the scales usually spread open allowing the seeds to fall out and be dispersed by
2820-561: A reduction in leopard population densities. Similarly, at two sites in central India the size of dhole packs was negatively correlated with tiger densities. Leopard and dhole distribution in Kui Buri correlated with both prey access and tiger scarcity. In Jigme Dorji National Park , tigers were found to inhabit the deeper parts of forests while the smaller predators were pushed closer to the fringes. The tiger generally mates all year round, particularly between November and April. A tigress
2961-472: A reduction in prey numbers, tigers continued to kill favoured prey while leopards and dholes increased their consumption of small prey. Both leopards and dholes can live successfully in tiger habitat when there is abundant food and vegetation cover. Otherwise, they appear to be less common where tigers are numerous. The recovery of the tiger population in Rajaji National Park during the 2000s led to
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#17330847361573102-727: A scattered range in the Indian subcontinent, the Indochinese Peninsula , Sumatra, northeastern China and the Russian Far East . As of 2020, India had the largest extent of global tiger habitat with 300,508 km (116,027 sq mi), followed by Russia with 195,819 km (75,606 sq mi). The tiger mainly lives in forest habitats and is highly adaptable. Records in Central Asia indicate that it primarily inhabited Tugay riverine forests and hilly and lowland forests in
3243-448: A secluded location, be it in dense vegetation, in a cave or under a rocky shelter. Litters consist of as many as seven cubs, but two or three are more typical. Newborn cubs weigh 785–1,610 g (27.7–56.8 oz) and are blind and altricial . The mother licks and cleans her cubs, suckles them and viciously defends them from any potential threat. Cubs open their eyes at the age of three to 14 days and their vision becomes clear after
3384-481: A serious environmental issue causing problems for pastoral farming and for conservation . Radiata pine was introduced to Australia in the 1870s. It is "the dominant tree species in the Australian plantation estate" – so much so that many Australians are concerned by the resulting loss of native wildlife habitat. The species is widely regarded as an environmental weed across southeastern and southwestern Australia and
3525-438: A single extant class , Pinopsida . All extant conifers are perennial woody plants with secondary growth . The great majority are trees , though a few are shrubs . Examples include cedars , Douglas-firs , cypresses , firs , junipers , kauri , larches , pines , hemlocks , redwoods , spruces , and yews . As of 2002, Pinophyta contained seven families, 60 to 65 genera, and more than 600 living species. Although
3666-406: A single white spruce tree from 1926 to 1961. Apical growth of the stem was slow from 1926 through 1936 when the tree was competing with herbs and shrubs and probably shaded by larger trees. Lateral branches began to show reduced growth and some were no longer in evidence on the 36-year-old tree. Apical growth totaling about 340 m, 370 m, 420 m, 450 m, 500 m, 600 m, and 600 m was made by the tree in
3807-447: A transient in another male's home range until he is older and strong enough to challenge the resident male. Tigers mark their home ranges by spraying urine on vegetation and rocks, clawing or scent rubbing trees and marking trails with faeces , anal gland secretions and ground scrapings. Scent markings also allow an individual to pick up information on another's identity. Unclaimed home ranges, particularly those that belonged to
3948-404: A typical felid morphology, with a muscular body, shortened legs, strong forelimbs with wide front paws, a large head and a tail that is about half the length of the rest of its body. It has five digits, including a dewclaw , on the front feet and four on the back, all of which have retractile claws that are compact and curved, and can reach 10 cm (3.9 in) long. The ears are rounded and
4089-469: A useful guide by which to assess the adequacy of particular nutrients, and the ratios among the major nutrients are helpful guides to nutritional imbalances. The softwood derived from conifers is of great economic value, providing about 45% of the world's annual lumber production. Other uses of the timber include the production of paper and plastic from chemically treated wood pulp. Some conifers also provide foods such as pine nuts and juniper berries ,
4230-440: A watering hole for prey to come by, particularly during hot summer days. It is an ambush predator and when approaching potential prey, it crouches with the head lowered and hides in foliage. It switches between creeping forward and staying still. A tiger may even doze off and can stay in the same spot for as long as a day, waiting for prey and launch an attack when the prey is close enough, usually within 30 m (98 ft). If
4371-569: A work and that Thabaton was alone. The old woman helps Keibu Keioiba to trick Thabaton into opening the door to her house and succeeded in making Thabaton. Keibu Keioba kidnaps Thabaton. Shortly after this, Thabaton's brothers return home from work and discover that Thabaton is missing. They ask the old woman about their sister, and she tells them about Keibu Keioiba. The seven brothers prepare their weapons and set out in search of their sister. Keibu Keioiba has not eaten Thabaton. Instead, he has forced Thabaton to become his wife, and they live together in
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4512-591: A year, then they overwinter. Female strobili emerge followed by pollination in the following spring. Fertilization takes place in summer of the following year, only 3–4 months after pollination. Cones mature and seeds are then shed by the end of that same year. Pollination and fertilization occur in a single growing season. Two-year reproductive cycle : The genera includes Widdringtonia , Sequoiadendron ( Cupressaceae ) and most species of Pinus . Female strobilus initials are formed in late summer or fall then overwinter. Female strobili emerge and receive pollen in
4653-415: Is colour blind and possibly perceives the tiger as green and blended in with the vegetation. The three colour variants of Bengal tigers – nearly stripeless snow-white, white and golden – are now virtually non-existent in the wild due to the reduction of wild tiger populations but continue in captive populations. The white tiger has a white background colour with sepia -brown stripes. The golden tiger
4794-446: Is a coast redwood ( Sequoia sempervirens ), with a height of 115.55 metres (although one mountain ash, Eucalyptus regnans , allegedly grew to a height of 140 metres, the tallest living angiosperms are significantly smaller at around 100 metres. ) The thickest (that is, the tree with the greatest trunk diameter ) is a Montezuma cypress ( Taxodium mucronatum ), 11.42 metres in diameter. The largest tree by three-dimensional volume
4935-418: Is a giant sequoia ( Sequoiadendron giganteum ), with a volume 1486.9 cubic metres. The smallest is the pygmy pine ( Lepidothamnus laxifolius ) of New Zealand, which is seldom taller than 30 cm when mature. The oldest non-clonal living tree is a Great Basin bristlecone pine ( Pinus longaeva ), 4,700 years old. Since most conifers are evergreens, the leaves of many conifers are long, thin and have
5076-553: Is a large cat and a member of the genus Panthera native to Asia . It has a powerful, muscular body with a large head and paws, a long tail and orange fur with black, mostly vertical stripes. It is traditionally classified into nine recent subspecies , though some recognise only two subspecies, mainland Asian tigers and the island tigers of the Sunda Islands . Throughout the tiger's range, it inhabits mainly forests, from coniferous and temperate broadleaf and mixed forests in
5217-401: Is absent, so that ligers grow far larger than either parent species. By contrast, the male tiger does not pass on a growth-promoting gene while the lioness passes on a growth inhibiting gene; hence, tigons are around the same size as their parents. Since they often develop life-threatening birth defects and can easily become obese, breeding these hybrids is regarded as unethical. The tiger has
5358-577: Is caused by a mutation of a transmembrane aminopeptidase gene. Around 37% of the Simlipal tiger population has this feature, which has been linked to genetic isolation . The tiger historically ranged from eastern Turkey, northern Iran and Afghanistan to Central Asia and from northern Pakistan through the Indian subcontinent and Indochina to southeastern Siberia, Sumatra, Java and Bali. As of 2022, it inhabits less than 7% of its historical distribution and has
5499-488: Is emitted through an open mouth and exposed teeth. In friendlier situations, tigers prusten , a soft, low-frequency snorting sound similar to purring in smaller cats. Tiger mothers communicate with their cubs by grunting, while cubs call back with miaows . When startled, they "woof". They produce a deer-like "pok" sound for unknown reasons, but most often at kills. The tiger is a carnivore and an apex predator feeding mainly on large and medium-sized ungulates, with
5640-520: Is forced through an open mouth as it closes and can be heard 3 km (1.9 mi) away. They roar multiple times in a row and others respond in kind. Tigers also roar during mating and a mother will roar to call her cubs to her. When tense, tigers moan, a sound similar to a roar but softer and made when the mouth is at least partially closed. Moaning can be heard 400 m (1,300 ft) away. Aggressive encounters involve growling , snarling and hissing. An explosive "coughing roar" or "coughing snarl"
5781-438: Is in oestrus for three to six days at a time, separated by three to nine week intervals. A resident male mates with all the females within his home range, who signal their receptiveness by roaring and marking. Younger, transient males are also attracted, leading to a fight in which the more dominant, resident male drives the usurper off. During courtship, the male is cautious with the female as he waits for her to show signs she
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5922-599: Is known to occur in Siberian tigers. A morbillivirus infection was the likely cause of death of a tigress in the Russian Far East that was also tested positive for feline panleukopenia and feline coronavirus . Blood samples from 11 adult tigers in Nepal showed antibodies for canine parvovirus -2, feline herpesvirus , feline coronavirus, leptospirosis and Toxoplasma gondii . The tiger has been listed as Endangered on
6063-437: Is less capable of climbing trees than many other cats due to its size, but cubs under 16 months old may routinely do so. An adult was recorded climbing 10 m (33 ft) up a smooth pipal tree . Adult tigers lead largely solitary lives within home ranges or territories , the size of which mainly depends on prey abundance, geographic area and sex of the individual. Males and females defend their home ranges from those of
6204-474: Is pale golden with reddish-brown stripes. The snow-white tiger is a morph with extremely faint stripes and a pale sepia-brown ringed tail. White and golden morphs are the result of an autosomal recessive trait with a white locus and a wideband locus, respectively. The snow-white variation is caused by polygenes with both white and wideband loci. The breeding of white tigers is controversial, as they have no use for conservation. Only 0.001% of wild tigers have
6345-476: Is planned. The tiger is among the most popular of the world's charismatic megafauna . It has been kept in captivity since ancient times and has been trained to perform in circuses and other entertainment shows. The tiger featured prominently in the ancient mythology and folklore of cultures throughout its historic range and has continued to appear in culture worldwide. The Old English tigras derives from Old French tigre , from Latin tigris , which
6486-402: Is probably: first to apical growth and new needle formation, then to buds for the next year's growth, with the cambium in the older parts of the branches receiving sustenance last. In the white spruce studied by Fraser et al. (1964), the needles constituted 17.5% of the over-day weight. Undoubtedly, the proportions change with time. Wind and animal dispersals are two major mechanisms involved in
6627-452: Is ready to mate. She signals to him by positioning herself in lordosis with her tail to the side. Copulation typically lasts no more than 20 seconds, with the male biting the female by the scruff of her neck. After it is finished, the male quickly pulls away as the female may turn and slap him. Tiger pairs may stay together for up to four days and mate multiple times. Gestation lasts around or over three months. A tigress gives birth in
6768-444: Is surrounded by black. The tiger is marked with distinctive black or dark brown stripes, which are uniquely patterned in each individual. The stripes are mostly vertical, but those on the limbs and forehead are horizontal. They are more concentrated towards the backside and those on the trunk may reach under the belly. The tips of stripes are generally sharp and some may split up or split and fuse again. Tail stripes are thick bands and
6909-432: Is the whitespotted sawyer ( Monochamus scutellatus ). Adults are found in summer on newly fallen or recently felled trees chewing tiny slits in the bark in which they lay eggs. The eggs hatch in about two weeks and the tiny larvae tunnel to the wood and score its surface with their feeding channels. With the onset of cooler weather, they bore into the wood, making oval entrance holes and tunnelling deeply. Feeding continues
7050-419: Is usually thin, though the Siberian tiger develops a particularly thick winter coat. The tiger has lines of fur around the face and long whiskers, especially in males. It has an orange colouration that varies from yellowish to reddish. White fur covers the underside, from head to tail, along with the inner surface of the legs and parts of the face. On the back of the ears, it has a prominent white spot, which
7191-566: The Caucasus . In the Amur - Ussuri region of Russia and China, it inhabits Korean pine and temperate broadleaf and mixed forests ; riparian forests serve as dispersal corridors, providing food and water for both tigers and ungulates . On the Indian subcontinent, it inhabits mainly tropical and subtropical moist broadleaf forests , temperate broadleaf and mixed forests , tropical moist evergreen forests , tropical dry forests , alluvial plains and
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#17330847361577332-611: The Cephalotaxaceae may be better included within the Taxaceae, and some authors additionally recognize Phyllocladaceae as distinct from Podocarpaceae (in which it is included here). The family Taxodiaceae is here included in the family Cupressaceae, but was widely recognized in the past and can still be found in many field guides. A new classification and linear sequence based on molecular data can be found in an article by Christenhusz et al. The conifers are an ancient group, with
7473-564: The Cordaitales , Vojnovskyales , Voltziales and perhaps also the Czekanowskiales (possibly more closely related to the Ginkgophyta ). Pinaceae Araucariaceae Podocarpaceae Sciadopityaceae Cupressaceae Cephalotaxaceae Taxaceae Multiple studies also indicate that the Gnetophyta belong within the conifers despite their distinct appearances, either placing them as
7614-520: The IUCN Red List since 1986 and the global tiger population is thought to have continuously declined from an estimated population of 5,000–8,262 tigers in the late 1990s to 3,726–5,578 individuals estimated as of 2022. During 2001–2020, landscapes where tigers live declined from 1,025,488 km (395,943 sq mi) to 911,901 km (352,087 sq mi). Habitat destruction , habitat fragmentation and poaching for fur and body parts are
7755-719: The Mesozoic era. Modern groups of conifers emerged from the Voltziales during the Late Permian through Jurassic . Conifers underwent a major decline in the Late Cretaceous corresponding to the explosive adaptive radiation of flowering plants . All living conifers are woody plants, and most are trees, the majority having a monopodial growth form (a single, straight trunk with side branches) with strong apical dominance . Many conifers have distinctly scented resin , secreted to protect
7896-629: The Russian Far East and Northeast China to tropical and subtropical moist broadleaf forests on the Indian subcontinent and Southeast Asia . The tiger is an apex predator and preys mainly on ungulates , which it takes by ambush. It lives a mostly solitary life and occupies home ranges , defending these from individuals of the same sex. The range of a male tiger overlaps with that of multiple females with whom he mates. Females give birth to usually two or three cubs that stay with their mother for about two years. When becoming independent, they leave their mother's home range and establish their own. Since
8037-423: The classification of the tiger as of 2005, and also reflect the classification recognised by the Cat Classification Task Force in 2017. Snow leopard [REDACTED] Tiger [REDACTED] Jaguar [REDACTED] Leopard [REDACTED] Lion [REDACTED] The tiger shares the genus Panthera with the lion , leopard , jaguar and snow leopard . Results of genetic analyses indicate that
8178-460: The influence of environmental conditions, their anatomical characteristics record growth rate changes produced by these changing conditions. The microscopic structure of conifer wood consists of two types of cells : parenchyma , which have an oval or polyhedral shape with approximately identical dimensions in three directions, and strongly elongated tracheids. Tracheids make up more than 90% of timber volume. The tracheids of earlywood formed at
8319-764: The mangrove forests of the Sundarbans . In the Eastern Himalayas , it was documented in temperate forest up to an elevation of 4,200 m (13,800 ft) in Bhutan, of 3,630 m (11,910 ft) in the Mishmi Hills and of 3,139 m (10,299 ft) in Mêdog County , southeastern Tibet. In Thailand, it lives in deciduous and evergreen forests. In Sumatra, it inhabits lowland peat swamp forests and rugged montane forests . Camera trapping during 2010–2015 in
8460-524: The monsoon ; three males had 84–147 km (32–57 sq mi) large home ranges in winter, 82–98 km (32–38 sq mi) in summer and 81–118 km (31–46 sq mi) during monsoon seasons. In Sikhote-Alin Biosphere Reserve , 14 females had home ranges 248–520 km (96–201 sq mi) and five resident males of 847–1,923 km (327–742 sq mi) that overlapped with those of up to five females. When tigresses in
8601-402: The wind . In some (e.g. firs and cedars ), the cones disintegrate to release the seeds, and in others (e.g. the pines that produce pine nuts ) the nut-like seeds are dispersed by birds (mainly nutcrackers , and jays ), which break up the specially adapted softer cones. Ripe cones may remain on the plant for a varied amount of time before falling to the ground; in some fire-adapted pines,
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#17330847361578742-409: The 16 nutrient elements known to be essential to plants, 13 of which are obtained from the soil, including nitrogen , phosphorus , potassium , calcium , magnesium , and sulfur , all used in relatively large amounts. Nutrient concentrations in conifers also vary with season, age, and kind of tissue sampled, and analytical technique. The ranges of concentrations occurring in well-grown plants provide
8883-481: The International Tiger Studbook 1938–2018 indicate that captive tigers lived up to 19 years. The father does not play a role in raising the young, but he encounters and interacts with them. The resident male appears to visit the female–cub families within his home range. They socialise and even share kills. One male was recorded looking after cubs whose mother had died. By defending his home range,
9024-627: The Middle-Late Pleistocene of Japan. Results of a phylogeographic study indicate that all living tigers have a common ancestor that lived between 108,000 and 72,000 years ago. Genetic studies suggest that the tiger population contracted around 115,000 years ago due to glaciation. Modern tiger populations originated from a refugium in Indochina and spread across Asia after the Last Glacial Maximum . As they colonised northeastern China,
9165-706: The North American Forest Tree Nursery Soils Workshop at Syracuse in 1980 provided strong contrary evidence: Bob Eastman, President of the Western Maine Forest Nursery Co. stated that for 15 years he has been successful in avoiding winter “burn” to Norway spruce and white spruce in his nursery operation by fertilizing with 50–80 lb/ac (56–90 kg/ha) nitrogen in September, whereas previously winter burn had been experienced annually, often severely. Eastman also stated that
9306-459: The Pinales without Taxales as paraphyletic , and the latter order is no longer considered distinct. A more accurate subdivision would be to split the class into three orders, Pinales containing only Pinaceae, Araucariales containing Araucariaceae and Podocarpaceae, and Cupressales containing the remaining families (including Taxaceae), but there has not been any significant support for such a split, with
9447-466: The ancestors of the South China tiger intermixed with a relict tiger population. Tigers can interbreed with other Panthera cats and have done so in captivity. The liger is the offspring of a female tiger and a male lion and the tigon the offspring of a male tiger and a female lion. The lion sire passes on a growth-promoting gene, but the corresponding growth-inhibiting gene from the female tiger
9588-487: The basis of fur length and colouration, striping patterns and body size of specimens in natural history museum collections that are not necessarily representative for the entire population. It was proposed to recognise only two tiger subspecies as valid, namely P. t. tigris in mainland Asia and the smaller P. t. sondaica in the Greater Sunda Islands . This two-subspecies proposal was reaffirmed in 2015 through
9729-558: The beginning of a growing season have large radial sizes and smaller, thinner cell walls . Then, the first tracheids of the transition zone are formed, where the radial size of cells and the thickness of their cell walls changes considerably. Finally, latewood tracheids are formed, with small radial sizes and greater cell wall thickness. This is the basic pattern of the internal cell structure of conifer tree rings. Most conifers are monoecious , but some are subdioecious or dioecious ; all are wind-pollinated . Conifer seeds develop inside
9870-539: The character is about the combination of a human and a beast. Various faded colours were used by the drawing team to adapt to the story. Yamata Amasung Keibu Keioiba (Yamata-no-Orochi and Keibu Keioiba) is a Meitei-language play that interweaves the stories of the two legendary creatures, Keibu Keioiba and Yamata-no-Orochi of Japanese mythology . The role of Keibu Keioiba was firstly played by Kshetrimayum Priyobrata and then secondly played by Pangambam Tyson Meitei. Tiger The tiger ( Panthera tigris )
10011-587: The cone consists of several fused scales, while in others (e.g. Taxaceae), the cone is reduced to just one seed scale or (e.g. Cephalotaxaceae) the several scales of a cone develop into individual arils, giving the appearance of a cluster of berries. The male cones have structures called microsporangia that produce yellowish pollen through meiosis. Pollen is released and carried by the wind to female cones. Pollen grains from living pinophyte species produce pollen tubes, much like those of angiosperms. The gymnosperm male gametophytes (pollen grains) are carried by wind to
10152-410: The deciduous and subtropical pine forest of Jim Corbett National Park , northern India revealed a stable tiger population density of 12–17 individuals per 100 km (39 sq mi) in an area of 521 km (201 sq mi). In northern Myanmar, the population density in a sampled area of roughly 3,250 km (1,250 sq mi) in a mosaic of tropical broadleaf forest and grassland
10293-443: The different subspecies and populations vary greatly in size and weight, the tiger's average size may be less than the lion's, while the largest tigers are bigger than their lion counterparts. The tiger's coat usually has short hairs, reaching up to 35 mm (1.4 in), though the hairs of the northern-living Siberian tiger can reach 105 mm (4.1 in). Belly hairs tend to be longer than back hairs. The density of their fur
10434-402: The dispersal of conifer seeds. Wind-born seed dispersal involves two processes, namely; local neighborhood dispersal and long-distance dispersal. Long-distance dispersal distances range from 11.9–33.7 kilometres (7.4–20.9 mi) from the source. Birds of the crow family, Corvidae , are the primary distributor of the conifer seeds. These birds are known to cache 32,000 pine seeds and transport
10575-575: The early 20th century, tiger populations have lost at least 93% of their historic range and are locally extinct in West and Central Asia , in large areas of China and on the islands of Java and Bali . Today, the tiger's range is severely fragmented. It is listed as Endangered on the IUCN Red List of Threatened Species , as its range is thought to have declined by 53% to 68% since the late 1990s. Major threats to tigers are habitat destruction and fragmentation due to deforestation , poaching for fur and
10716-407: The eyes have a round pupil. The snout ends in a triangular, pink tip with small black dots, the number of which increase with age. The tiger's skull is robust, with a constricted front region, proportionally small, elliptical orbits , long nasal bones and a lengthened cranium with a large sagittal crest . It resembles a lion's skull, but differs from it in the concave or flattened underside of
10857-567: The female cone is termed fruit , which undergoes ripening (maturation). It was found recently that the pollen of conifers transfers the mitochondrial organelles to the embryo , a sort of meiotic drive that perhaps explains why Pinus and other conifers are so productive, and perhaps also has bearing on observed sex-ratio bias. Conifers are heterosporous , generating two different types of spores: male microspores and female megaspores . These spores develop on separate male and female sporophylls on separate male and female cones. In
10998-449: The first year spring and become conelets. The conelet goes through another winter rest and, in the spring of the second year archegonia form in the conelet. Fertilization of the archegonia occurs by early summer of the second year, so the pollination-fertilization interval exceeds a year. After fertilization, the conelet is considered an immature cone. Maturation occurs by autumn of the second year, at which time seeds are shed. In summary,
11139-466: The following spring. Female strobili emerge then pollination occurs in spring of the second year then the pollinated strobili become conelets in the same year (i.e. the second year). The female gametophytes in the conelet develop so slowly that the megaspore does not go through free-nuclear divisions until autumn of the third year. The conelet then overwinters again in the free-nuclear female gametophyte stage. Fertilization takes place by early summer of
11280-492: The following summer when larvae occasionally return to the surface of the wood and extend the feeding channels generally in a U-shaped configuration. During this time, small piles of frass extruded by the larvae accumulate under logs. Early in the spring of the second year following egg-laying, the larvae, about 30 mm long, pupate in the tunnel enlargement just below the wood surface. The resulting adults chew their way out in early summer, leaving round exit holes, so completing
11421-399: The forest. The brothers never stop searching for their sister. One day, they see Thabaton. When they get close to her, they see that she is with a man and has a baby with her. The brothers discreetly signal to Thabaton so that she knows they have arrived. When Keibu Keioiba goes away for a hunt, Thabaton meet her brothers and they plan her escape. When Keibu Keioiba returns, Thabaton gives him
11562-606: The fossil record are jaw fragments from Lantion in China that are dated to the early Pleistocene. Middle- to late-Pleistocene tiger fossils have been found throughout China, Sumatra and Java. Prehistoric subspecies include Panthera tigris trinilensis and P. t. soloensis of Java and Sumatra and P. t. acutidens of China; late Pleistocene and early Holocene fossils of tigers have also been found in Borneo and Palawan, Philippines. Fossil specimens of tigers have also been reported from
11703-399: The fourth year and seeds mature in the cones by autumn of the fourth year. The growth and form of a forest tree are the result of activity in the primary and secondary meristems , influenced by the distribution of photosynthate from its needles and the hormonal gradients controlled by the apical meristems. External factors also influence growth and form. Fraser recorded the development of
11844-722: The fragmentation of potential tiger habitat, especially in the Eastern Plains. Inbreeding depression coupled with habitat destruction, insufficient prey resources and poaching is a threat to the small and isolated tiger population in the Changbai Mountains along the China–Russia border. In China, tigers became the target of large-scale 'anti-pest' campaigns in the early 1950s, where suitable habitats were fragmented following deforestation and resettlement of people to rural areas, who hunted tigers and prey species. Though tiger hunting
11985-519: The genes for this colour morph and the overrepresentation of white tigers in captivity is the result of inbreeding . Hence, their continued breeding will risk both inbreeding depression and loss of genetic variability in captive tigers. Pseudo- melanistic tigers with thick, merged stripes have been recorded in Simlipal National Park and three Indian zoos; a population genetic analysis of Indian tiger samples revealed that this phenotype
12126-463: The genus Panthera using the scientific name Panthera tigris . Nine recent tiger subspecies have been proposed between the early 19th and early 21st centuries, namely the Bengal , Malayan , Indochinese , South China , Siberian , Caspian , Javan , Bali and Sumatran tigers . The validity of several tiger subspecies was questioned in 1999 as most putative subspecies were distinguished on
12267-474: The ground. His anger knew no bounds. He charges back to the home of Thabaton and her brothers, who have prepared their weapons. As soon as Keibu Keioiba approaches them, they attack. Finally, Keibu Keioiba dies at the hands of the Thabaton's seven brothers. Thabaton and her seven brothers live happily ever after. In the 2009 Meitei language animated film Keibu Keioiba (Tiger Head) , the character of Keibu-Kei-Oiba
12408-419: The growing embryo) and its surrounding integument, becomes a seed . Eventually, the seed may fall to the ground and, if conditions permit, grow into a new plant. In forestry , the terminology of flowering plants has commonly though inaccurately been applied to cone-bearing trees as well. The male cone and unfertilized female cone are called male flower and female flower , respectively. After fertilization,
12549-461: The illegal trade of body parts for medicinal purposes. Tigers are also victims of human–wildlife conflict as they attack and prey on livestock in areas where natural prey is scarce. The tiger is legally protected in all range countries. National conservation measures consist of action plans, anti-poaching patrols and schemes for monitoring tiger populations. In several range countries, wildlife corridors have been established and tiger reintroduction
12690-522: The immense conifer forests of the world represent the largest terrestrial carbon sink . Conifers are of great economic value for softwood lumber and paper production. Conifer is a Latin word, a compound of conus (cone) and ferre (to bear), meaning "the one that bears (a) cone(s)". The division name Pinophyta conforms to the rules of the International Code of Nomenclature for algae, fungi, and plants (ICN), which state (Article 16.1) that
12831-535: The largest and economically most important component group of gymnosperms, but nevertheless they comprise only one of the four groups. The division Pinophyta consists of just one class, Pinopsida, which includes both living and fossil taxa. Subdivision of the living conifers into two or more orders has been proposed from time to time. The most commonly seen in the past was a split into two orders, Taxales (Taxaceae only) and Pinales (the rest), but recent research into DNA sequences suggests that this interpretation leaves
12972-417: The leaf bases are twisted to present the leaves in a very flat plane for maximum light capture. Leaf size varies from 2 mm in many scale-leaved species, up to 400 mm long in the needles of some pines (e.g. Apache pine, Pinus engelmannii ). The stomata are in lines or patches on the leaves and can be closed when it is very dry or cold. The leaves are often dark green in colour, which may help absorb
13113-442: The lower jaw and in its longer nasals. The tiger has 30 fairly robust teeth and its somewhat curved canines are the longest in the cat family at 6.4–7.6 cm (2.5–3.0 in). The tiger has a head-body length of 1.4–2.8 m (4 ft 7 in – 9 ft 2 in) with a 0.6–1.1 m (2 ft 0 in – 3 ft 7 in) tail and stands 0.8–1.1 m (2 ft 7 in – 3 ft 7 in) at
13254-489: The major threats that contributed to the decrease of tiger populations in all range countries. Protected areas in central India are highly fragmented due to linear infrastructure like roads, railway lines, transmission lines , irrigation channels and mining activities in their vicinity. In the Tanintharyi Region of southern Myanmar, deforestation coupled with mining activities and high hunting pressure threatens
13395-402: The majority of opinion preferring retention of all the families within a single order Pinales, despite their antiquity and diverse morphology . There were seven families of conifers c. 2011 , with 65–70 genera and over 600 living species ( c. 2002 ). The seven most distinct families are linked in the box above right and phylogenetic diagram left. In other interpretations,
13536-403: The male cones, microspores are produced from microsporocytes by meiosis . The microspores develop into pollen grains, which contain the male gametophytes. Large amounts of pollen are released and carried by the wind. Some pollen grains will land on a female cone for pollination. The generative cell in the pollen grain divides into two haploid sperm cells by mitosis leading to the development of
13677-808: The male protects the females and cubs from other males. When a new male takes over, dependent cubs are at risk of infanticide as the male attempts to sire his own young with the females. A seven-year long study in Chitwan National Park revealed that 12 of 56 detected cubs and juveniles were killed by new males taking over home ranges. Tigers are recorded as hosts for various parasites including tapeworms like Diphyllobothrium erinacei , Taenia pisiformis in India and nematodes like Toxocara species in India and Physaloptera preputialis , Dirofilaria ursi and Uiteinarta species in Siberia. Canine distemper
13818-523: The more vulnerable calves. They sometimes prey on livestock and dogs in close proximity to settlements. Tigers occasionally consume vegetation, fruit and minerals for dietary fibre and supplements. Tigers learn to hunt from their mothers, though the ability to hunt may be partially inborn. Depending on the size of the prey, they typically kill weekly though mothers must kill more often. Families hunt together when cubs are old enough. They search for prey using vision and hearing. A tiger will also wait at
13959-546: The morning. A three-year-long camera trap survey in Shuklaphanta National Park revealed that tigers were most active from dusk until midnight. In northeastern China, tigers were crepuscular and active at night with activity peaking at dawn and dusk; they were largely active at the same time as their prey. The tiger is a powerful swimmer and easily transverses rivers as wide as 8 km (5.0 mi); it immerses in water, particularly on hot days. In general, it
14100-538: The mother and fatal injuries. After around two months, the cubs are able to follow their mother. They still hide in vegetation when she goes hunting. Young bond through play fighting and practice stalking. A hierarchy develops in the litter, with the biggest cub, often a male, being the most dominant and the first to eat its fill at a kill. Around the age of six months, cubs are fully weaned and have more freedom to explore their environment. Between eight and ten months, they accompany their mother on hunts. A cub can make
14241-463: The names of higher taxa in plants (above the rank of family) are either formed from the name of an included family (usually the most common and/or representative), in this case Pinaceae (the pine family), or are descriptive. A descriptive name in widespread use for the conifers (at whatever rank is chosen) is Coniferae (Art 16 Ex 2). According to the ICN, it is possible to use a name formed by replacing
14382-477: The neck. Large prey may be disabled by a bite to the back of the hock , severing the tendon. Swipes from the large paws are capable of stunning or breaking the skull of a water buffalo . They kill small prey with a bite to the back of the neck or head. Estimates of the success rate for hunting tigers range from a low of 5% to a high of 50%. They are sometimes killed or injured by large or dangerous prey like gaur, buffalo and boar. Tigers typically move kills to
14523-486: The one 11 m tall white spruce, Fraser et al. (1964) speculated that if the photosynthate used in making apical growth in 1961 was manufactured the previous year, then the 4 million needles that were produced up to 1960 manufactured food for about 600,000 mm of apical growth or 730 g dry weight, over 12 million mm of wood for the 1961 annual ring, plus 1 million new needles, in addition to new tissue in branches, bark, and roots in 1960. Added to this would be
14664-424: The one-year and the two-year cycles differ mainly in the duration of the pollination-fertilization interval. Three-year reproductive cycle : Three of the conifer species are pine species ( Pinus pinea , Pinus leiophylla , Pinus torreyana ) which have pollination and fertilization events separated by a two-year interval. Female strobili initiated during late summer or autumn of a year, then overwinter until
14805-425: The overwintering storage capacity of stock thus treated was much improved (Eastman 1980). The concentrations of nutrients in plant tissues depend on many factors, including growing conditions. Interpretation of concentrations determined by analysis is easy only when a nutrient occurs in excessively low or occasionally excessively high concentration. Values are influenced by environmental factors and interactions among
14946-562: The paraphyletic assemblage of " walchian conifers ", which were small trees, and probably originated in dry upland habitats. The range of conifers expanded during the Early Permian ( Cisuralian ) to lowlands due to increasing aridity. Walchian conifers were gradually replaced by more advanced voltzialean or "transition" conifers. Conifers were largely unaffected by the Permian–Triassic extinction event , and were dominant land plants of
15087-429: The photosynthate to produce energy to sustain respiration over this period, an amount estimated to be about 10% of the total annual photosynthate production of a young healthy tree. On this basis, one needle produced food for about 0.19 mg dry weight of apical growth, 3 mm wood, one-quarter of a new needle, plus an unknown amount of branch wood, bark and roots. The order of priority of photosynthate distribution
15228-550: The plant for several (2–40) years before falling, but five genera ( Larix , Pseudolarix , Glyptostrobus , Metasequoia and Taxodium ) are deciduous , shedding their leaves in autumn. The seedlings of many conifers, including most of the Cupressaceae, and Pinus in Pinaceae, have a distinct juvenile foliage period where the leaves are different, often markedly so, from the typical adult leaves. Tree rings are records of
15369-416: The pollen tube. At fertilization, one of the sperm cells unites its haploid nucleus with the haploid nucleus of an egg cell. The female cone develops two ovules, each of which contains haploid megaspores. A megasporocyte is divided by meiosis in each ovule. Each winged pollen grain is a four celled male gametophyte . Three of the four cells break down leaving only a single surviving cell which will develop into
15510-405: The prey spots it before then, the cat does not pursue further. A tiger can sprint 56 km/h (35 mph) and leap 10 m (33 ft); it is not a long-distance runner and gives up a chase if prey outpaces it over a certain distance. The tiger attacks from behind or at the sides and tries to knock the target off balance. It latches onto prey with its forelimbs, twisting and turning during
15651-507: The removal of individual plants beyond plantations is encouraged. At least 20 species of roundheaded borers of the family Cerambycidae feed on the wood of spruce , fir , and hemlock (Rose and Lindquist 1985). Borers rarely bore tunnels in living trees, although when populations are high, adult beetles feed on tender twig bark, and may damage young living trees. One of the most common and widely distributed borer species in North America
15792-453: The same effect in 105-day-old white spruce. The general short-term effect of nitrogen fertilization on coniferous seedlings is to stimulate shoot growth more so than root growth (Armson and Carman 1961). Over a longer period, root growth is also stimulated. Many nursery managers were long reluctant to apply nitrogenous fertilizers late in the growing season, for fear of increased danger of frost damage to succulent tissues. A presentation at
15933-668: The same reserve had cubs of up to four months of age, they reduced their home ranges to stay near their young and steadily enlarged them until their offspring were 13–18 months old. The tiger is a long-ranging species and individuals disperse over distances of up to 650 km (400 mi) to reach tiger populations in other areas. Young tigresses establish their first home ranges close to their mothers' while males migrate further than their female counterparts. Four radio-collared females in Chitwan dispersed between 0 and 43.2 km (0.0 and 26.8 mi) and 10 males between 9.5 and 65.7 km (5.9 and 40.8 mi). A subadult male lives as
16074-424: The same sex and the home range of a male encompasses that of multiple females. Two females in the Sundarbans had home ranges of 10.6 and 14.1 km (4.1 and 5.4 sq mi). In Panna Tiger Reserve , the home ranges of five reintroduced females varied from 53–67 km (20–26 sq mi) in winter to 55–60 km (21–23 sq mi) in summer and to 46–94 km (18–36 sq mi) during
16215-402: The same time and place, and was suggested to be a sister species of the modern tiger when it was examined in 2014. However, as of 2023, at least two subsequent studies considered P. zdanskyi likely to be a synonym of P. palaeosinensis , noting that its proposed differences from that species fell within the range of individual variation. The earliest appearance of the modern tiger species in
16356-918: The seeds as far as 12–22 km (7.5–13.7 mi) from the source. The birds store the seeds in the soil at depths of 2–3 cm ( 3 ⁄ 4 – 1 + 1 ⁄ 4 in) under conditions which favor germination . Conifers are the dominant plants over large areas of land, most notably the taiga of the Northern Hemisphere , but also in similar cool climates in mountains further south. A number of conifers originally introduced for forestry have become invasive species in parts of New Zealand , including radiata pine ( Pinus radiata ), lodgepole pine ( P. contorta ), Douglas fir ( Pseudotsuga mensiezii ) and European larch ( Larix decidua ). In parts of South Africa , maritime pine ( Pinus pinaster ), patula pine ( P. patula ) and radiata pine have been declared invasive species. These wilding conifers are
16497-540: The seeds may be stored in closed cones for up to 60–80 years, being released only when a fire kills the parent tree. In the families Podocarpaceae , Cephalotaxaceae , Taxaceae , and one Cupressaceae genus ( Juniperus ), the scales are soft, fleshy, sweet, and brightly colored, and are eaten by fruit-eating birds, which then pass the seeds in their droppings. These fleshy scales are (except in Juniperus ) known as arils . In some of these conifers (e.g. most Podocarpaceae),
16638-449: The shoulder. The Siberian and Bengal tigers are the largest. Male Bengal tigers weigh 200–260 kg (440–570 lb), and females weigh 100–160 kg (220–350 lb); island tigers are the smallest, likely due to insular dwarfism . Male Sumatran tigers weigh 100–140 kg (220–310 lb), and females weigh 75–110 kg (165–243 lb). The tiger is popularly thought to be the largest living felid species; but since tigers of
16779-557: The six living putative subspecies—the Bengal, Malayan, Indochinese, South China, Siberian and Sumatran tiger—found them to be distinct and separate clades . These results were corroborated in 2021 and 2023. The Cat Specialist Group states that "Given the varied interpretations of data, the [subspecific] taxonomy of this species is currently under review by the IUCN SSC Cat Specialist Group." The following tables are based on
16920-429: The sole source of nitrogen guanidine compounds were less prominent. Durzan and Steward noted that their results, drawn from determinations made in late summer, did not rule out the occurrence of different interim responses at other times of the year. Ammonium nitrogen produced significantly heavier (dry weight) seedlings with a higher nitrogen content after 5 weeks than did the same amount of nitrate nitrogen. Swan found
17061-401: The struggle and tries to pull it to the ground. The tiger generally applies a bite to the throat until its victim dies of strangulation . It has an average bite force at the canine tips of 1234.3 newtons . Holding onto the throat puts the cat out of reach of horns, antlers, tusks and hooves. Tigers are adaptable killers and may use other methods, including ripping the throat or breaking
17202-517: The termination -aceae in the name of an included family, in this case preferably Pinaceae , by the appropriate termination, in the case of this division -ophyta . Alternatively, " descriptive botanical names " may also be used at any rank above family. Both are allowed. This means that if conifers are considered a division, they may be called Pinophyta or Coniferae. As a class, they may be called Pinopsida or Coniferae. As an order they may be called Pinales or Coniferae or Coniferales . Conifers are
17343-422: The three predators involve chasing, stealing kills and direct killing. Large dhole packs may kill tigers. Tigers, leopards and dholes coexist by hunting different sized prey. In Nagarhole National Park , the average weight for tiger kills was found to be 91.5 kg (202 lb), compared to 37.6 kg (83 lb) for leopards and 43.4 kg (96 lb) for dholes. In Kui Buri National Park , following
17484-498: The tiger and snow leopard are sister species whose lineages split from each other between 2.70 and 3.70 million years ago. The tiger's whole genome sequencing shows repeated sequences that parallel those in other cat genomes. The fossil species Panthera palaeosinensis of early Pleistocene northern China was described as a possible tiger ancestor when it was discovered in 1924, but modern cladistics places it as basal to modern Panthera . Panthera zdanskyi lived around
17625-411: The tiger and the river is doubted. In 1758, Carl Linnaeus described the tiger in his work Systema Naturae and gave it the scientific name Felis tigris , as the genus Felis was being used for all cats at the time. His scientific description was based on descriptions by earlier naturalists such as Conrad Gessner and Ulisse Aldrovandi . In 1929, Reginald Innes Pocock placed the species in
17766-564: The tiger population in far eastern Russia, where logging roads facilitate access for poachers and people harvesting forest products that are important for prey species to survive in winter. Conifer Conifers are a group of cone-bearing seed plants , a subset of gymnosperms . Scientifically, they make up the division Pinophyta ( / p ɪ ˈ n ɒ f ɪ t ə , ˈ p aɪ n oʊ f aɪ t ə / ), also known as Coniferophyta ( / ˌ k ɒ n ɪ f ə ˈ r ɒ f ɪ t ə , - oʊ f aɪ t ə / ) or Coniferae . The division contains
17907-584: The tiger population. In Thailand, nine of 15 protected areas hosting tigers are isolated and fragmented, offering a low probability for dispersal between them; four of these have not harboured tigers since about 2013. In Peninsular Malaysia, 8,315.7 km (3,210.7 sq mi) of tiger habitat was cleared during 1988–2012, most of it for industrial plantations . Large-scale land acquisitions of about 23,000 km (8,900 sq mi) for commercial agriculture and timber extraction in Cambodia contributed to
18048-786: The time spent with her cubs by reducing her home range, killing larger prey and returning to her den more rapidly than without cubs; when the cubs started to eat meat, she took them to kill sites, thereby optimising their protection and access to food. In the same reserve, one of 21 cubs died in over eight years of monitoring and mortality did not differ between male and female juveniles. Tiger monitoring over six years in Ranthambore Tiger Reserve indicated an average annual survival rate of around 85 percent for 74 male and female cubs; survival rate increased to 97 percent for both males and female juveniles of one to two years of age. Causes of cub mortality include predators, floods, fires, death of
18189-570: The total number of species is relatively small, conifers are ecologically important. They are the dominant plants over large areas of land, most notably the taiga of the Northern Hemisphere , but also in similar cool climates in mountains further south. Boreal conifers have many wintertime adaptations. The narrow conical shape of northern conifers, and their downward-drooping limbs, help them shed snow. Many of them seasonally alter their biochemistry to make them more resistant to freezing. While tropical rainforests have more biodiversity and turnover,
18330-469: The tree against insect infestation and fungal infection of wounds. Fossilized resin hardens into amber , which has been commercially exploited historically (for example, in New Zealand's 19th-century kauri gum industry). The size of mature conifers varies from less than one metre to over 100 metres in height. The world's tallest, thickest, largest, and oldest living trees are all conifers. The tallest
18471-416: The usual 2-year life cycle. Conifers – notably Abies (fir), Cedrus , Chamaecyparis lawsoniana (Lawson's cypress), Cupressus (cypress), juniper , Picea (spruce), Pinus (pine), Taxus (yew), Thuja (cedar) – have been the subject of selection for ornamental purposes. Plants with unusual growth habits, sizes, and colours are propagated and planted in parks and gardens throughout
18612-463: The white spots, particularly during aggressive encounters and between mothers and cubs. They also use their tails to signal their mood. To show cordiality, the tail sticks up and sways slowly, while an apprehensive tiger lowers its tail or wags it side-to-side. When calm, the tail hangs low. Tigers are normally silent but can produce numerous vocalisations. They roar to signal their presence to other individuals over long distances. This vocalisation
18753-468: The world. Conifers can absorb nitrogen in either the ammonium (NH 4 ) or nitrate (NO 3 ) form, but the forms are not physiologically equivalent. Form of nitrogen affected both the total amount and relative composition of the soluble nitrogen in white spruce tissues (Durzan and Steward). Ammonium nitrogen was shown to foster arginine and amides and lead to a large increase of free guanidine compounds, whereas in leaves nourished by nitrate as
18894-403: The years 1955 through 1961, respectively. The total number of needles of all ages present on the 36-year-old tree in 1961 was 5.25 million weighing 14.25 kg. In 1961, needles as old as 13 years remained on the tree. The ash weight of needles increased progressively with age from about 4% in first-year needles in 1961 to about 8% in needles 10 years old. In discussing the data obtained from
19035-465: Was a borrowing from Classical Greek [τίγρις] Error: {{Transliteration}}: transliteration text not Latin script (pos 1) ( help ) 'tigris'. Since ancient times, the word tigris has been suggested to originate from the Armenian or Persian word for 'arrow', which may also be the origin of the name for the river Tigris . However, today, the names are thought to be homonyms , and the connection between
19176-435: Was a human in the daytime and a tiger at night. During night, he prowled around looking for food. One night, he found an older woman's house and planned to devour her. The old woman said that her wrinkled skin would not be tasty and suggested that he eat a beautiful and young woman named Thabaton instead. Thabaton was the only sister in a family with seven brothers. The old woman told Keibu Keioiba that her brothers had gone for
19317-507: Was considered the last important site for the tiger in Laos, but it has not been recorded there at least since 2013; this population likely fell victim to indiscriminate snaring. Anti-poaching units in Sumatra's Kerinci Seblat landscape removed 362 tiger snare traps and seized 91 tiger skins during 2005–2016; annual poaching rates increased with rising skin prices. Poaching is also the main threat to
19458-420: Was estimated to be 0.21–0.44 tigers per 100 km (39 sq mi) as of 2009. Population density in mixed deciduous and semi-evergreen forests of Thailand's Huai Kha Khaeng Wildlife Sanctuary was estimated at 2.01 tigers per 100 km (39 sq mi); during the 1970s and 1980s, logging and poaching had occurred in the adjacent Mae Wong and Khlong Lan National Parks , where population density
19599-839: Was much lower, estimated at only 0.359 tigers per 100 km (39 sq mi) as of 2016. Population density in dipterocarp and montane forests in northern Malaysia was estimated at 1.47–2.43 adult tigers per 100 km (39 sq mi) in Royal Belum State Park , but 0.3–0.92 adult tigers per 100 km (39 sq mi) in the unprotected selectively logged Temengor Forest Reserve. Camera trap data show that tigers in Chitwan National Park avoided locations frequented by people and were more active at night than during day. In Sundarbans National Park , six radio-collared tigers were most active from dawn to early morning and reached their zenith around 7:00 o'clock in
19740-569: Was prohibited in 1977, the population continued to decline and is considered extinct in South China since 2001. Tiger populations in India have been targeted by poachers since the 1990s and were extirpated in two tiger reserves in 2005 and 2009. Between March 2017 and January 2020, 630 activities of hunters using snares , drift nets, hunting platforms and hunting dogs were discovered in a reserve forest of about 1,000 km (390 sq mi) in southern Myanmar. Nam Et-Phou Louey National Park
19881-400: Was uniquely designed to a humanoid tiger, which had to speak dialogues in human voices. The facial structure of the character was twisted out of the normal shape of a real tiger to a little bit extend. It was done so as to bring adjustment to the human voice and the speaking actions during digitalization. Artist Bhumenjoy faced great challenges in making the visual character of Keibu Kei-Oiba, as
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