56-548: Around 544, see text Ixora is a genus of flowering plants in the family Rubiaceae . It is the only genus in the tribe Ixoreae . It consists of tropical evergreen trees and shrubs and holds around 544 species. Though native to the tropical and subtropical areas throughout the world, its centre of diversity is in Tropical Asia. Ixora also grows commonly in subtropical climates in the United States, such as Florida where it
112-1818: A molecular phylogeny of plants placed the flowering plants in their evolutionary context: Bryophytes [REDACTED] Lycophytes [REDACTED] Ferns [REDACTED] [REDACTED] [REDACTED] The main groups of living angiosperms are: Amborellales [REDACTED] 1 sp. New Caledonia shrub Nymphaeales [REDACTED] c. 80 spp. water lilies & allies Austrobaileyales [REDACTED] c. 100 spp. woody plants Magnoliids [REDACTED] c. 10,000 spp. 3-part flowers, 1-pore pollen, usu. branch-veined leaves Chloranthales [REDACTED] 77 spp. Woody, apetalous Monocots [REDACTED] c. 70,000 spp. 3-part flowers, 1 cotyledon , 1-pore pollen, usu. parallel-veined leaves Ceratophyllales [REDACTED] c. 6 spp. aquatic plants Eudicots [REDACTED] c. 175,000 spp. 4- or 5-part flowers, 3-pore pollen, usu. branch-veined leaves Amborellales Melikyan, Bobrov & Zaytzeva 1999 Nymphaeales Salisbury ex von Berchtold & Presl 1820 Austrobaileyales Takhtajan ex Reveal 1992 Chloranthales Mart. 1835 Canellales Cronquist 1957 Piperales von Berchtold & Presl 1820 Magnoliales de Jussieu ex von Berchtold & Presl 1820 Laurales de Jussieu ex von Berchtold & Presl 1820 Acorales Link 1835 Alismatales Brown ex von Berchtold & Presl 1820 Petrosaviales Takhtajan 1997 Dioscoreales Brown 1835 Pandanales Brown ex von Berchtold & Presl 1820 Liliales Perleb 1826 Asparagales Link 1829 Arecales Bromhead 1840 Poales Small 1903 Zingiberales Grisebach 1854 Commelinales de Mirbel ex von Berchtold & Presl 1820 Rice Rice
168-416: A sustainable way. Many varieties of rice have been bred to improve crop quality and productivity. Biotechnology has created Green Revolution rice able to produce high yields when supplied with nitrogen fertiliser and managed intensively. Other products are rice able to express human proteins for medicinal use; flood-tolerant or deepwater rice ; and drought-tolerant and salt-tolerant varieties. Rice
224-442: A woody stem ), grasses and grass-like plants, a vast majority of broad-leaved trees , shrubs and vines , and most aquatic plants . Angiosperms are distinguished from the other major seed plant clade, the gymnosperms , by having flowers , xylem consisting of vessel elements instead of tracheids , endosperm within their seeds, and fruits that completely envelop the seeds. The ancestors of flowering plants diverged from
280-403: A branched inflorescence which arises from the last internode on the stem. There can be up to 350 spikelets in a panicle, each containing male and female flower parts ( anthers and ovule ). A fertilised ovule develops into the edible grain or caryopsis . Rice is a cereal belonging to the family Poaceae . As a tropical crop, it can be grown during the two distinct seasons (dry and wet) of
336-495: A fungal study, it was found that the species Pseudopestalotiopsis ixorae and Pseudopestalotiopsis taiwanensis caused leaf spots on species of Ixora , which is a popular garden plant in Taiwan. Flowering plant Basal angiosperms Core angiosperms Flowering plants are plants that bear flowers and fruits , and form the clade Angiospermae ( / ˌ æ n dʒ i ə ˈ s p ər m iː / ). The term 'angiosperm'
392-411: A role in various religions and traditions, such as in weddings . The rice plant can grow to over 1 m (3 ft) tall; if in deep water, it can reach a length of 5 m (16 ft). A single plant may have several leafy stems or tillers . The upright stem is jointed with nodes along its length; a long slender leaf arises from each node. The self-fertile flowers are produced in a panicle ,
448-506: A single domestication event for O. sativa . Both indica and japonica forms of Asian rice sprang from a single domestication event in China from the wild rice Oryza rufipogon . Despite this evidence, it appears that indica rice arose when japonica arrived in India about 4,500 years ago and hybridised with another rice, whether an undomesticated proto- indica or wild O. nivara . Rice
504-610: A total of 64 angiosperm orders and 416 families. The diversity of flowering plants is not evenly distributed. Nearly all species belong to the eudicot (75%), monocot (23%), and magnoliid (2%) clades. The remaining five clades contain a little over 250 species in total; i.e. less than 0.1% of flowering plant diversity, divided among nine families. The 25 most species-rich of 443 families, containing over 166,000 species between them in their APG circumscriptions, are: The botanical term "angiosperm", from Greek words angeíon ( ἀγγεῖον 'bottle, vessel') and spérma ( σπέρμα 'seed'),
560-440: A wide variety of pests including insects, nematodes, rodents such as rats, snails, and birds. Major rice insect pests include armyworms, rice bugs , black bugs , cutworms, field crickets, grasshoppers, leafhoppers, mealybugs, and planthoppers. High rates of nitrogen fertiliser application may worsen aphid outbreaks. Weather conditions can contribute to pest outbreaks: rice gall midge outbreaks are worsened by high rainfall in
616-436: Is rainfed like wheat or maize. Across Asia, unmilled rice or "paddy" (Indonesian and Malay padi ), was traditionally the product of smallholder agriculture, with manual harvesting . Larger farms make use of machines such as combine harvesters to reduce the input of labour. The grain is ready to harvest when the moisture content is 20–25%. Harvesting involves reaping , stacking the cut stalks, threshing to separate
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#1732891513353672-858: Is starting to impact plants and is likely to cause many species to become extinct by 2100. Angiosperms are terrestrial vascular plants; like the gymnosperms, they have roots , stems , leaves , and seeds . They differ from other seed plants in several ways. The largest angiosperms are Eucalyptus gum trees of Australia, and Shorea faguetiana , dipterocarp rainforest trees of Southeast Asia, both of which can reach almost 100 metres (330 ft) in height. The smallest are Wolffia duckweeds which float on freshwater, each plant less than 2 millimetres (0.08 in) across. Considering their method of obtaining energy, some 99% of flowering plants are photosynthetic autotrophs , deriving their energy from sunlight and using it to create molecules such as sugars . The remainder are parasitic , whether on fungi like
728-661: Is 69% water, 29% carbohydrates , 2% protein , and contains negligible fat (table). In a reference serving of 100 grams (3.5 oz), cooked white rice provides 130 calories of food energy , and contains moderate levels of manganese (18% DV), with no other micronutrients in significant content (all less than 10% of the Daily Value ). In 2018, the World Health Organization strongly recommended fortifying rice with iron , and conditionally recommended fortifying it with vitamin A and with folic acid . Golden rice
784-732: Is a cereal grain and in its domesticated form is the staple food of over half of the world's population , particularly in Asia and Africa . Rice is the seed of the grass species Oryza sativa (Asian rice)—or, much less commonly, Oryza glaberrima (African rice). Asian rice was domesticated in China some 13,500 to 8,200 years ago; African rice was domesticated in Africa about 3,000 years ago. Rice has become commonplace in many cultures worldwide; in 2021, 787 million tons were produced, placing it fourth after sugarcane , maize , and wheat . Only some 8% of rice
840-613: Is a major food staple in Asia, Latin America, and some parts of Africa, feeding over half the world's population. However, a substantial part of the crop can be lost post-harvest through inefficient transportation, storage, and milling. A quarter of the crop in Nigeria is lost after harvest. Storage losses include damage by mould fungi if the rice is not dried sufficiently. In China, losses in modern metal silos were just 0.2%, compared to 7–13% when rice
896-498: Is a variety produced through genetic engineering to synthesize beta-carotene , a precursor of vitamin A, in the endosperm of the rice grain. It is intended to be grown and eaten in parts of the world where Vitamin A deficiency is prevalent. Golden rice has been opposed by activists, such as in the Philippines . In 2016 more than 100 Nobel laureates encouraged the use of genetically modified organisms , such as golden rice, for
952-581: Is based on four principles: biodiversity, host plant resistance, landscape ecology, and hierarchies in a landscape—from biological to social. Farmers' pesticide applications are often unnecessary. Pesticides may actually induce resurgence of populations of rice pests such as the brown planthopper , both by destroying beneficial insects and by enhancing the pest's reproduction. The International Rice Research Institute (IRRI) demonstrated in 1993 that an 87.5% reduction in pesticide use can lead to an overall drop in pest numbers. Farmers in China, Indonesia and
1008-824: Is commonly known as West Indian jasmine . Ixora is Latinized from Sanskrit Ishwara , one of the names of the Hindu god Shiva. The genus was formally created by Linnaeus in 1753, as it was noted by Hendrik van Rheede that the flowers of what he noted as schetti (and named by Rheede as Ixora coccinea ) were offered in temples in the Malabar . Other common names include viruchi , kiskaara , kepale , rangan , kheme , ponna , chann tanea , techi , pan , siantan , jarum-jarum/jejarum , cây trang thái , jungle flame , jungle geranium , and cruz de Malta , among others. The plants possess leathery leaves, ranging from 3 to 6 inches in length, and produce large clusters of tiny flowers in
1064-478: Is derived from the Greek words ἀγγεῖον / angeion ('container, vessel') and σπέρμα / sperma ('seed'), meaning that the seeds are enclosed within a fruit. The group was formerly called Magnoliophyta . Angiosperms are by far the most diverse group of land plants with 64 orders , 416 families , approximately 13,000 known genera and 300,000 known species . They include all forbs (flowering plants without
1120-504: Is from Asia. The average world yield for rice was 4.7 metric tons per hectare (2.1 short tons per acre), in 2022. Yuan Longping of China's National Hybrid Rice Research and Development Center set a world record for rice yield in 1999 at 17.1 metric tons per hectare (7.6 short tons per acre) on a demonstration plot. This employed specially developed hybrid rice and the System of Rice Intensification (SRI), an innovation in rice farming. Rice
1176-797: Is sister to the bamboos, Bambusoideae , and the cereal subfamily Pooideae . The rice genus Oryza is one of eleven in the Oryzeae; it is sister to the Phyllorachideae . The edible rice species O. sativa and O. glaberrima are among some 300 species or subspecies in the genus. other grasses (inc. the C4 grasses , maize , sorghum ) Streptogyneae Ehrharteae Phyllorachideae Wild rices inc. Zizania other rice species and subspecies O. sativa (Asian rice) O. glaberrima (African rice) Bambusoideae (bamboos) Pooideae (grasses and cereals inc. wheat , barley ) Oryza sativa rice
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#17328915133531232-685: Is stickier, and is used for sweet dishes, and in Italy for risotto ; and sticky short-grain rice is used in Japanese sushi as it keeps its shape when cooked. White rice when cooked contains 29% carbohydrate and 2% protein, with some manganese . Golden rice is a variety produced by genetic engineering to contain vitamin A . Production of rice is estimated to have caused over 1% of global greenhouse gas emissions in 2022. Predictions of how rice yields will be affected by climate change vary across geographies and socioeconomic contexts. In human culture, rice plays
1288-438: Is subjected to a steaming process before it is milled. This makes the grain harder, and moves some of the grain's vitamins and minerals into the white part of the rice so these are retained after milling. Rice does not contain gluten , so is suitable for people on a gluten-free diet . Rice is a good source of protein and a staple food in many parts of the world, but it is not a complete protein as it does not contain all of
1344-406: Is to flood the field to a depth of 5 cm (2 in), then to let the water level drop to 15 cm (6 in) below surface level, as measured by looking into a perforated field water tube sunk into the soil, and then repeating the cycle. Deepwater rice varieties tolerate flooding to a depth of over 50 centimetres for at least a month. Upland rice is grown without flooding, in hilly or mountainous regions; it
1400-509: Is traded internationally. China, India, and Indonesia are the largest consumers of rice. A substantial amount of the rice produced in developing nations is lost after harvest through factors such as poor transport and storage. Rice yields can be reduced by pests including insects , rodents , and birds , as well as by weeds , and by diseases such as rice blast . Traditional rice polycultures such as rice-duck farming , and modern integrated pest management seek to control damage from pests in
1456-471: Is used as a model organism in biology. Dry rice grain is milled to remove the outer layers; depending on how much is removed, products range from brown rice to rice with germ and white rice. Some is parboiled to make it easy to cook. Rice contains no gluten ; it provides protein but not all the essential amino acids needed for good health. Rice of different types is eaten around the world. Long-grain rice tends to stay intact on cooking; medium-grain rice
1512-485: The Alismatales grow in marine environments, spreading with rhizomes that grow through the mud in sheltered coastal waters. Some specialised angiosperms are able to flourish in extremely acid or alkaline habitats. The sundews , many of which live in nutrient-poor acid bogs , are carnivorous plants , able to derive nutrients such as nitrate from the bodies of trapped insects. Other flowers such as Gentiana verna ,
1568-440: The essential amino acids in sufficient amounts for good health. World trade figures are much smaller than those for production, as less than 8% of rice produced is traded internationally. China, an exporter of rice in the early 2000s, had become the world's largest importer of rice by 2013. Developing countries are the main players in the world rice trade; by 2012, India was the largest exporter of rice, with Thailand and Vietnam
1624-460: The orchids for part or all of their life-cycle, or on other plants , either wholly like the broomrapes, Orobanche , or partially like the witchweeds, Striga . In terms of their environment, flowering plants are cosmopolitan, occupying a wide range of habitats on land, in fresh water and in the sea. On land, they are the dominant plant group in every habitat except for frigid moss-lichen tundra and coniferous forest . The seagrasses in
1680-883: The "Big Five" extinction events in Earth's history, only the Cretaceous–Paleogene extinction event had occurred while angiosperms dominated plant life on the planet. Today, the Holocene extinction affects all kingdoms of complex life on Earth, and conservation measures are necessary to protect plants in their habitats in the wild ( in situ ), or failing that, ex situ in seed banks or artificial habitats like botanic gardens . Otherwise, around 40% of plant species may become extinct due to human actions such as habitat destruction , introduction of invasive species , unsustainable logging , land clearing and overharvesting of medicinal or ornamental plants . Further, climate change
1736-478: The 21st century as people in Asia and elsewhere ate less grain and more meat. An exception is Sub-Saharan Africa, where both per capita consumption of rice and population are increasing. Rice is a commonly-eaten food around the world. The varieties of rice are typically classified as short-, medium-, and long-grained. Oryza sativa indica varieties are usually long-grained; Oryza sativa japonica varieties are usually short- or medium-grained. Short-grain rice, with
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1792-688: The Americas by the Spanish. In British North America by the time of the start of the American War of Independence , rice had become the fourth most valuable export commodity behind only tobacco, wheat, and fish. In 2021, world production of rice was 787 million tonnes , led by China and India with a combined 52% of the total. This placed rice fourth in the list of crops by production, after sugarcane , maize , and wheat . Other major producers were Bangladesh , Indonesia and Vietnam . 90% of world production
1848-460: The Philippines have traditionally managed weeds and pests by the polycultural practice of raising ducks and sometimes fish in their rice paddies. These produce valuable additional crops, eat small pest animals, manure the rice, and in the case of ducks also control weeds. Rice plants produce their own chemical defences to protect themselves from pest attacks. Some synthetic chemicals, such as
1904-465: The benefits these could bring. In 2022, greenhouse gas emissions from rice cultivation were estimated at 5.7 billion tonnes CO2eq, representing 1.2% of total emissions. Within the agriculture sector, rice produces almost half the greenhouse gas emissions from croplands , some 30% of agricultural methane emissions , and 11% of agricultural nitrous oxide emissions. Methane is released from rice fields subject to long-term flooding, as this inhibits
1960-547: The common ancestor of all living gymnosperms before the end of the Carboniferous , over 300 million years ago. In the Cretaceous , angiosperms diversified explosively , becoming the dominant group of plants across the planet. Agriculture is almost entirely dependent on angiosperms, and a small number of flowering plant families supply nearly all plant-based food and livestock feed. Rice , maize and wheat provide half of
2016-506: The crop would be lost under these conditions. In the Po Valley in Italy, the arborio and carnaroli risotto rice varieties have suffered poor harvests through drought in the 21st century. The Ente Nazionale Risi [ it ] is developing drought-resistant varieties; its nuovo prometeo variety has deep roots that enable it to tolerate drought, but is not suitable for risotto. Rice yield can be reduced by weed growth, and
2072-425: The exception of Spanish Bomba, is usually sticky when cooked, and is suitable for puddings. Thai Jasmine rice is aromatic, and unusually for a long-grain rice has some stickiness, with a soft texture. Indian Basmati rice is very long-grained and aromatic. Italian Arborio rice , used for risotto , is of medium length, oval, and quite sticky. Japanese sushi rice is a sticky short-grain variety. Cooked white rice
2128-425: The field where they will grow, or seedlings can be grown in a seedbed and transplanted into the field. Direct seeding needs some 60 to 80 kg of grain per hectare, while transplanting needs less, around 40 kg per hectare, but requires far more labour. Most rice in Asia is transplanted by hand. Mechanical transplanting takes less time but requires a carefully-prepared field and seedlings raised on mats or in trays to fit
2184-635: The flowering plants as an unranked clade without a formal Latin name (angiosperms). A formal classification was published alongside the 2009 revision in which the flowering plants rank as the subclass Magnoliidae. From 1998, the Angiosperm Phylogeny Group (APG) has reclassified the angiosperms, with updates in the APG II system in 2003, the APG III system in 2009, and the APG IV system in 2016. In 2019,
2240-503: The grain, and cleaning by winnowing or screening . The rice grain is dried as soon as possible to bring the moisture content down to a level that is safe from mould fungi. Traditional drying relies on the heat of the sun, with the grain spread out on mats or on pavements. The edible rice species are members of the BOP clade within the grass family, the Poaceae . The rice subfamily, Oryzoideae ,
2296-575: The herbicide 2,4-D , cause the plant to increase the production of certain defensive chemicals and thereby increase the plant's resistance to some types of pests. Conversely, other chemicals, such as the insecticide imidacloprid , appear to induce changes in the gene expression of the rice that make the plant more susceptible to certain pests. Plant breeders have created rice cultivars incorporating resistance to various insect pests . Conventional plant breeding of resistant varieties has been limited by challenges such as rearing insect pests for testing, and
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2352-410: The later years of the 20th century decreased rice yield by between 10% and 20% across 200 farms in seven Asian countries. This may have been caused by increased night-time respiration. IRRI has predicted that Asian rice yields will fall by some 20% per 1°C rise in global mean temperature. Further, rice is unable to yield grain if the flowers experience a temperature of 35 °C or more for over one hour, so
2408-417: The machine. Rice does not thrive if continuously submerged. Rice can be grown in different environments, depending upon water availability. The usual arrangement is for lowland fields to be surrounded by bunds and flooded to a depth of a few centimetres until around a week before harvest time; this requires a large amount of water. The "alternate wetting and drying" technique uses less water. One form of this
2464-518: The manner of vines or lianas . The number of species of flowering plants is estimated to be in the range of 250,000 to 400,000. This compares to around 12,000 species of moss and 11,000 species of pteridophytes . The APG system seeks to determine the number of families , mostly by molecular phylogenetics . In the 2009 APG III there were 415 families. The 2016 APG IV added five new orders (Boraginales, Dilleniales, Icacinales, Metteniusales and Vahliales), along with some new families, for
2520-428: The other largest exporters. As of 2016, the countries that consumed the most rice were China (29% of total), India, and Indonesia. By 2020, Bangladesh had taken third place from Indonesia. On an annual average from 2020-23, China consumed 154 million tonnes of rice, India consumed 109 million tonnes, and Bangladesh and Indonesia consumed about 36 million tonnes each. Across the world, rice consumption per capita fell in
2576-1011: The soil from absorbing atmospheric oxygen, resulting in anaerobic fermentation of organic matter in the soil. Emissions can be limited by planting new varieties, not flooding continuously, and removing straw. It is possible to cut methane emissions in rice cultivation by improved water management, combining dry seeding and one drawdown, or executing a sequence of wetting and drying . This results in emission reductions of up to 90% compared to full flooding and even increased yields. Predictions of climate change's effects on rice cultivation vary. Global rice yield has been projected to decrease by around 3.2% with each 1°C increase in global average temperature while another study predicts global rice cultivation will increase initially, plateauing at about 3°C warming (2091–2100 relative to 1850–1900). The impacts of climate change on rice cultivation vary across geographic location and socioeconomic context. For example, rising temperatures and decreasing solar radiation during
2632-568: The spring gentian, are adapted to the alkaline conditions found on calcium -rich chalk and limestone , which give rise to often dry topographies such as limestone pavement . As for their growth habit , the flowering plants range from small, soft herbaceous plants , often living as annuals or biennials that set seed and die after one growing season, to large perennial woody trees that may live for many centuries and grow to many metres in height. Some species grow tall without being self-supporting like trees by climbing on other plants in
2688-489: The summer. Members of Ixora prefer acidic soil, and are suitable choices for bonsai . It is also a popular choice for hedges in parts of South East Asia. In tropical climates, they flower year round and are commonly used in Hindu worship, as well as in ayurveda and Indian folk medicine. In Brazil, fungal species Pseudocercospora ixoricola was found to be causing leaf spots on Ixora coccinea . Then in 2018, in Taiwan, during
2744-628: The voyage to the rest of the Pacific. It reached Austroasiatic and Kra-Dai -speakers in Mainland Southeast Asia and southern China by 5000 years ago. Rice spread around the rest of the world through cultivation, migration and trade, eventually to the Americas as part of the Columbian exchange after 1492. The now less common Oryza glaberrima (African rice) was independently domesticated in Africa around 3,000 years ago, and introduced to
2800-785: The wet season, while thrips outbreaks are associated with drought. Rice blast , caused by the fungus Magnaporthe grisea , is the most serious disease of growing rice. It and bacterial leaf streak (caused by Xanthomonas oryzae pv. oryzae ) are perennially the two worst rice diseases worldwide; they are both among the ten most important diseases of all crop plants. Other major rice diseases include sheath blight (caused by Rhizoctonia solani ), false smut ( Ustilaginoidea virens ), and bacterial panicle blight ( Burkholderia glumae ). Viral diseases include rice bunchy stunt, rice dwarf, rice tungro , and rice yellow mottle. Crop protection scientists are developing sustainable techniques for managing rice pests. Sustainable pest management
2856-505: The world's staple calorie intake, and all three plants are cereals from the Poaceae family (colloquially known as grasses). Other families provide important industrial plant products such as wood , paper and cotton , and supply numerous ingredients for beverages , sugar production , traditional medicine and modern pharmaceuticals . Flowering plants are also commonly grown for decorative purposes , with certain flowers playing significant cultural roles in many societies. Out of
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#17328915133532912-539: The year provided that sufficient water is made available. It is normally an annual, but in the tropics it can survive as a perennial , producing a ratoon crop. Like all crops, rice depends for its growth on both biotic and abiotic environmental factors. The principal biotic factors are crop variety, pests , and plant diseases . Abiotic factors include the soil type, whether lowland or upland, amount of rain or irrigation water, temperature, day length , and intensity of sunlight. Rice grains can be planted directly into
2968-483: Was coined in the form "Angiospermae" by Paul Hermann in 1690, including only flowering plants whose seeds were enclosed in capsules. The term angiosperm fundamentally changed in meaning in 1827 with Robert Brown , when angiosperm came to mean a seed plant with enclosed ovules. In 1851, with Wilhelm Hofmeister 's work on embryo-sacs, Angiosperm came to have its modern meaning of all the flowering plants including Dicotyledons and Monocotyledons. The APG system treats
3024-465: Was first domesticated in China 9,000 years ago, by people of Neolithic cultures in the Upper and Lower Yangtze , associated with Hmong-Mien -speakers and pre-Austronesians , respectively. The functional allele for nonshattering , the critical indicator of domestication in grains, as well as five other single-nucleotide polymorphisms , is identical in both indica and japonica . This implies
3080-559: Was introduced early into Sino-Tibetan cultures in northern China by around 6000 to 5600 years ago, and to the Korean peninsula and Japan by around 5500 to 3200 years ago. It was also carried into Taiwan by the Dapenkeng culture by 5500 to 4000 years ago, before spreading southwards via the Austronesian migrations to Island Southeast Asia , Madagascar , and Guam , but did not survive
3136-439: Was stored by rural households. The dry grain is milled to remove the outer layers, namely the husk and bran . These can be removed in a single step, in two steps, or as in commercial milling in a multi-step process of cleaning, dehusking, separation, polishing, grading, and weighing. Brown rice only has the inedible husk removed. Further milling removes bran and the germ to create successively whiter products. Parboiled rice
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