98-404: The Adephaga (from Greek ἀδηφάγος, adephagos , "gluttonous") are a suborder of beetles , and with more than 40,000 recorded species in 10 families, the second-largest of the four beetle suborders. Members of this suborder are collectively known as adephagans . The largest family is Carabidae (ground beetles) which comprises most of the suborder with over 40,000 species. Adephaga also includes
196-570: A basis for coinages: anthropology , photography , telephony , isomer , biomechanics , cinematography , etc. Together with Latin words , they form the foundation of international scientific and technical vocabulary ; for example, all words ending in -logy ('discourse'). There are many English words of Greek origin . Greek is an independent branch of the Indo-European language family. The ancient language most closely related to it may be ancient Macedonian , which, by most accounts,
294-647: A conduit for the dispersal of pollen grains, spores and seeds , the atmosphere can be considered to be a habitat-type in its own right. There are metabolically active microbes present that actively reproduce and spend their whole existence airborne, with hundreds of thousands of individual organisms estimated to be present in a cubic meter of air. The airborne microbial community may be as diverse as that found in soil or other terrestrial environments, however, these organisms are not evenly distributed, their densities varying spatially with altitude and environmental conditions. Aerobiology has not been studied much, but there
392-404: A direct result of human activities, such as deforestation , the plowing of ancient grasslands, the diversion and damming of rivers, the draining of marshland and the dredging of the seabed. The introduction of alien species can have a devastating effect on native wildlife – through increased predation , through competition for resources or through the introduction of pests and diseases to which
490-489: A downpour occurs and lays its eggs in the transient pools that form; the tadpoles develop with great rapidity, sometimes in as little as nine days, undergo metamorphosis , and feed voraciously before digging a burrow of their own. Other organisms cope with the drying up of their aqueous habitat in other ways. Vernal pools are ephemeral ponds that form in the rainy season and dry up afterwards. They have their specially-adapted characteristic flora, mainly consisting of annuals,
588-552: A fairly stable set of consonantal contrasts . The main phonological changes occurred during the Hellenistic and Roman period (see Koine Greek phonology for details): In all its stages, the morphology of Greek shows an extensive set of productive derivational affixes , a limited but productive system of compounding and a rich inflectional system. Although its morphological categories have been fairly stable over time, morphological changes are present throughout, particularly in
686-556: A faster, more convenient cursive writing style with the use of ink and quill . The Greek alphabet consists of 24 letters, each with an uppercase ( majuscule ) and lowercase ( minuscule ) form. The letter sigma has an additional lowercase form (ς) used in the final position of a word: In addition to the letters, the Greek alphabet features a number of diacritical signs : three different accent marks ( acute , grave , and circumflex ), originally denoting different shapes of pitch accent on
784-531: A foreign language. It is also often stated that the historical changes have been relatively slight compared with some other languages. According to one estimation, " Homeric Greek is probably closer to Demotic than 12-century Middle English is to modern spoken English ". Greek is spoken today by at least 13 million people, principally in Greece and Cyprus along with a sizable Greek-speaking minority in Albania near
882-413: A forest is divided into parts by logging, with strips of cleared land separating woodland blocks, and the distances between the remaining fragments exceeds the distance an individual animal is able to travel, that species becomes especially vulnerable. Small populations generally lack genetic diversity and may be threatened by increased predation, increased competition, disease and unexpected catastrophe. At
980-526: A home for both static organisms, anchored to the substrate , and for a large range of organisms crawling on or burrowing into the surface. Some creatures float among the waves on the surface of the water, or raft on floating debris, others swim at a range of depths, including organisms in the demersal zone close to the seabed, and myriads of organisms drift with the currents and form the plankton . Many animals and plants have taken up residence in urban environments. They tend to be adaptable generalists and use
1078-709: A mile below the ice of Antarctica; in the absence of sunlight, they must rely on organic material from elsewhere, perhaps decaying matter from glacier melt water or minerals from the underlying rock. Other bacteria can be found in abundance in the Mariana Trench , the deepest place in the ocean and on Earth; marine snow drifts down from the surface layers of the sea and accumulates in this undersea valley, providing nourishment for an extensive community of bacteria. Other microbes live in environments lacking in oxygen, and are dependent on chemical reactions other than photosynthesis . Boreholes drilled 300 m (1,000 ft) into
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#17330851169961176-444: A natural habitat is no longer able to support its native species. The organisms once living there have either moved to elsewhere or are dead, leading to a decrease in biodiversity and species numbers . Habitat destruction is in fact the leading cause of biodiversity loss and species extinction worldwide. The protection of habitat types is a necessary step in the maintenance of biodiversity because if habitat destruction occurs,
1274-410: A parasitic organism, its habitat is the particular part of the outside or inside of its host on or in which it is adapted to live. The life cycle of some parasites involves several different host species, as well as free-living life stages, sometimes within vastly different microhabitat types. One such organism is the trematode (flatworm) Microphallus turgidus , present in brackish water marshes in
1372-443: A particular community of plants and animals. The chief environmental factors affecting the distribution of living organisms are temperature, humidity, climate, soil and light intensity , and the presence or absence of all the requirements that the organism needs to sustain it. Generally speaking, animal communities are reliant on specific types of plant communities. Some plants and animals have habitat requirements which are met in
1470-557: A particular species well, but its presence or absence at any particular location depends to some extent on chance, on its dispersal abilities and its efficiency as a colonizer. Arid habitats are those where there is little available water. The most extreme arid habitats are deserts . Desert animals have a variety of adaptations to survive the dry conditions. Some frogs live in deserts, creating moist habitat types underground and hibernating while conditions are adverse. Couch's spadefoot toad ( Scaphiopus couchii ) emerges from its burrow when
1568-487: A similar way; their eggs hatch and the juvenile fish grow with great rapidity when the conditions are right, but the whole population of fish may end up as eggs in diapause in the dried up mud that was once a pond. Freshwater habitat types include rivers, streams, lakes, ponds, marshes and bogs. They can be divided into running waters (rivers, streams) and standing waters (lakes, ponds, marshes, bogs). Although some organisms are found across most of these habitat types,
1666-509: A similarly rich fauna of invertebrates as a more varied habitat. The monotypic habitat occurs in both botanical and zoological contexts. Some invasive species may create monocultural stands that prevent other species from growing there. A dominant colonization can occur from retardant chemicals exuded, nutrient monopolization, or from lack of natural controls, such as herbivores or climate, that keep them in balance with their native habitat types. The yellow starthistle, Centaurea solstitialis
1764-777: A tube leading to an opening valve. The secretory lobes differ structurally from one taxon to another; it may be elongated or oval, branched basally or apically , or unbranched. Secretion can occur in multiple manners: The secretions differ in the chemical constituents, according to the taxa. Gyrinids , for instance, secrete norsesquiterpenes such as gyrinidal, gyrinidione, or gyrinidone. Dytiscids discharge aromatic aldehydes , esters , and acids , especially benzoic acid . Carabids typically produce carboxylic acids , particularly formic acid , methacrylic acid , and tiglic acid , but also aliphatic ketones , saturated esters , phenols , aromatic aldehydes, and quinones . Accessory glands or modified structures are present in some taxa:
1862-488: A variety of aquatic beetles, such as predaceous diving beetles and whirligig beetles . Adephagans have simple antennae with no pectination or clubs. The galeae of the maxillae usually consist of two segments. Adult adephagans have visible notopleural sutures . The first visible abdominal sternum is completely separated by the hind coxae , which is one of the most easily recognizable traits of adephagans. Five segments are on each foot. The transverse fold of
1960-498: A variety of bacteria and fungi; and snowfields on which algae grow. Whether from natural processes or the activities of man, landscapes and their associated habitat types change over time. There are the slow geomorphological changes associated with the geologic processes that cause tectonic uplift and subsidence , and the more rapid changes associated with earthquakes, landslides, storms, flooding, wildfires, coastal erosion , deforestation and changes in land use. Then there are
2058-442: A very limited set of factors to survive. The habitat of a species is not necessarily found in a geographical area, it can be the interior of a stem, a rotten log, a rock or a clump of moss ; a parasitic organism has as its habitat the body of its host , part of the host's body (such as the digestive tract), or a single cell within the host's body. Habitat types are environmental categorizations of different environments based on
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#17330851169962156-595: A wide range of locations. The small white butterfly Pieris rapae for example is found on all the continents of the world apart from Antarctica . Its larvae feed on a wide range of Brassicas and various other plant species, and it thrives in any open location with diverse plant associations. The large blue butterfly Phengaris arion is much more specific in its requirements; it is found only in chalk grassland areas, its larvae feed on Thymus species, and because of complex life cycle requirements it inhabits only areas in which Myrmica ants live. Disturbance
2254-562: Is chemosynthesis , a process by which microbes convert such substances as hydrogen sulfide or ammonia into organic molecules. These bacteria and Archaea are the primary producers in these ecosystems and support a diverse array of life. About 350 species of organism, dominated by molluscs , polychaete worms and crustaceans , had been discovered around hydrothermal vents by the end of the twentieth century, most of them being new to science and endemic to these habitat types. Besides providing locomotion opportunities for winged animals and
2352-404: Is a concept sometimes used in conservation biology , in which a single species of animal or plant is the only species of its type to be found in a specific habitat and forms a monoculture . Even though it might seem such a habitat type is impoverished in biodiversity as compared with polytypic habitat types, this is not necessarily the case. Monocultures of the exotic plant Hydrilla support
2450-509: Is a vigorous grass from Europe which has been introduced to the United States where it has become invasive. It is highly adapted to fire, producing large amounts of flammable detritus and increasing the frequency and intensity of wildfires. In areas where it has become established, it has altered the local fire regimen to such an extant that native plants cannot survive the frequent fires, allowing it to become even more dominant. A marine example
2548-766: Is also found in Bulgaria near the Greek-Bulgarian border. Greek is also spoken worldwide by the sizable Greek diaspora which has notable communities in the United States , Australia , Canada , South Africa , Chile , Brazil , Argentina , Russia , Ukraine , the United Kingdom , and throughout the European Union , especially in Germany . Historically, significant Greek-speaking communities and regions were found throughout
2646-742: Is an Indo-European language, constituting an independent Hellenic branch within the Indo-European language family. It is native to Greece , Cyprus , Italy (in Calabria and Salento ), southern Albania , and other regions of the Balkans , Caucasus , the Black Sea coast, Asia Minor , and the Eastern Mediterranean . It has the longest documented history of any Indo-European language, spanning at least 3,400 years of written records. Its writing system
2744-474: Is an Indo-European language, but also includes a number of borrowings from the languages of the populations that inhabited Greece before the arrival of Proto-Greeks, some documented in Mycenaean texts ; they include a large number of Greek toponyms . The form and meaning of many words have changed. Loanwords (words of foreign origin) have entered the language, mainly from Latin, Venetian , and Turkish . During
2842-494: Is evidence of nitrogen fixation in clouds , and less clear evidence of carbon cycling, both facilitated by microbial activity. There are other examples of extreme habitat types where specially adapted lifeforms exist; tar pits teeming with microbial life; naturally occurring crude oil pools inhabited by the larvae of the petroleum fly ; hot springs where the temperature may be as high as 71 °C (160 °F) and cyanobacteria create microbial mats ; cold seeps where
2940-559: Is important in the creation of biodiverse habitat types. In the absence of disturbance, a climax vegetation cover develops that prevents the establishment of other species. Wildflower meadows are sometimes created by conservationists but most of the flowering plants used are either annuals or biennials and disappear after a few years in the absence of patches of bare ground on which their seedlings can grow. Lightning strikes and toppled trees in tropical forests allow species richness to be maintained as pioneering species move in to fill
3038-414: Is more appropriate. The physical factors may include (for example): soil , moisture , range of temperature , and light intensity . Biotic factors include the availability of food and the presence or absence of predators . Every species has particular habitat requirements, habitat generalist species are able to thrive in a wide array of environmental conditions while habitat specialist species require
Adephaga - Misplaced Pages Continue
3136-510: Is protected and promoted officially as a regional and minority language in Armenia, Hungary , Romania, and Ukraine. It is recognized as a minority language and protected in Turkey by the 1923 Treaty of Lausanne . The phonology , morphology , syntax , and vocabulary of the language show both conservative and innovative tendencies across the entire attestation of the language from the ancient to
3234-787: Is sometimes called aljamiado , as when Romance languages are written in the Arabic alphabet. Article 1 of the Universal Declaration of Human Rights in Greek: Transcription of the example text into Latin alphabet : Article 1 of the Universal Declaration of Human Rights in English: Proto-Greek Mycenaean Ancient Koine Medieval Modern Habitat (ecology) In ecology , habitat refers to
3332-518: Is spoken by at least 13.5 million people today in Greece, Cyprus, Italy, Albania, Turkey , and the many other countries of the Greek diaspora . Greek roots have been widely used for centuries and continue to be widely used to coin new words in other languages; Greek and Latin are the predominant sources of international scientific vocabulary . Greek has been spoken in the Balkan peninsula since around
3430-758: Is still used internationally for the writing of Ancient Greek . In Greek, the question mark is written as the English semicolon, while the functions of the colon and semicolon are performed by a raised point (•), known as the ano teleia ( άνω τελεία ). In Greek the comma also functions as a silent letter in a handful of Greek words, principally distinguishing ό,τι ( ó,ti , 'whatever') from ότι ( óti , 'that'). Ancient Greek texts often used scriptio continua ('continuous writing'), which means that ancient authors and scribes would write word after word with no spaces or punctuation between words to differentiate or mark boundaries. Boustrophedon , or bi-directional text,
3528-624: Is suggested to represent a subgroup of Carabidae rather than a distinct family, with Cicindelidae often being treated as a distinct family from Carabidae. Cladogram of the relationships of living adephagan families after Vasilikopoulos et al. 2021 and Baca et al. 2021: Gyrinidae Trachypachidae Cicindelidae Carabidae Haliplidae Noteridae Hygrobiidae Aspidytidae Amphizoidae Dytiscidae Greek language Greek ( Modern Greek : Ελληνικά , romanized : Elliniká , [eliniˈka] ; Ancient Greek : Ἑλληνική , romanized : Hellēnikḗ )
3626-630: Is the Greek alphabet , which has been used for approximately 2,800 years; previously, Greek was recorded in writing systems such as Linear B and the Cypriot syllabary . The alphabet arose from the Phoenician script and was in turn the basis of the Latin , Cyrillic , Coptic , Gothic , and many other writing systems. The Greek language holds a very important place in the history of the Western world. Beginning with
3724-416: Is the small-scale physical requirements of a particular organism or population. Every habitat includes large numbers of microhabitat types with subtly different exposure to light, humidity, temperature, air movement, and other factors. The lichens that grow on the north face of a boulder are different from those that grow on the south face, from those on the level top, and those that grow on the ground nearby;
3822-503: Is very challenging to research, and as well as being little-studied, it is vast, with 79% of the Earth's biosphere being at depths greater than 1,000 m (3,300 ft). With no plant life, the animals in this zone are either detritivores , reliant on food drifting down from surface layers, or they are predators, feeding on each other. Some organisms are pelagic , swimming or drifting in mid-ocean, while others are benthic, living on or near
3920-468: Is when sea urchin populations " explode " in coastal waters and destroy all the macroalgae present. What was previously a kelp forest becomes an urchin barren that may last for years and this can have a profound effect on the food chain . Removal of the sea urchins, by disease for example, can result in the seaweed returning, with an over-abundance of fast-growing kelp. Habitat destruction (also termed habitat loss and habitat reduction) occurs when
4018-559: The Dytiscidae and Hygrobiidae also possess paired prothoracic glands secreting steroids ; and the Gyrinidae are unique in the extended shape of the external opening of the pygidial gland. The function of many compounds remain unknown, yet several hypotheses have been advanced: Habitats range from caves to rainforest canopy and alpine habitats. The body forms of some are structurally modified for adaptation to habitats: members of
Adephaga - Misplaced Pages Continue
4116-486: The Eastern Mediterranean , in what are today Southern Italy , Turkey , Cyprus , Syria , Lebanon , Israel , Palestine , Egypt , and Libya ; in the area of the Black Sea , in what are today Turkey, Bulgaria , Romania , Ukraine , Russia , Georgia , Armenia , and Azerbaijan ; and, to a lesser extent, in the Western Mediterranean in and around colonies such as Massalia , Monoikos , and Mainake . It
4214-510: The Indo-Iranian languages (see Graeco-Aryan ), but little definitive evidence has been found. In addition, Albanian has also been considered somewhat related to Greek and Armenian, and it has been proposed that they all form a higher-order subgroup along with other extinct languages of the ancient Balkans; this higher-order subgroup is usually termed Palaeo-Balkan , and Greek has a central position in it. Linear B , attested as early as
4312-441: The abdomen , which are used for secreting chemicals. The glands consist of complex invaginations of the cuticle lined with epidermal cells contiguous with the integument . The glands have no connection with the rectum and open on the eighth abdominal tergum . Secretions pass from the secretory lobes , which are aggregations of secretory cells, through a tube to a reservoir lined with muscles. This reservoir then narrows to
4410-476: The hind wing is near the wing tip. The median nervure ends at this fold, where it is joined by a cross nervure . Adephagans have four Malpighian tubules . Unlike the genetical structures of other beetles, yolk chambers alternate with egg chambers in the ovarian tubes of adephagans. The coiled, tubular testes consist of a single follicle, and the ovaries are polytrophic . All families of adephagan have paired pygidial glands located posterodorsally in
4508-502: The methane and hydrogen sulfide issue from the ocean floor and support microbes and higher animals such as mussels which form symbiotic associations with these anaerobic organisms ; salt pans that harbour salt-tolerant bacteria , archaea and also fungi such as the black yeast Hortaea werneckii and basidiomycete Wallemia ichthyophaga ; ice sheets in Antarctica which support fungi Thelebolus spp., glacial ice with
4606-492: The nominal and verbal systems. The major change in the nominal morphology since the classical stage was the disuse of the dative case (its functions being largely taken over by the genitive ). The verbal system has lost the infinitive , the synthetically -formed future, and perfect tenses and the optative mood . Many have been replaced by periphrastic ( analytical ) forms. Pronouns show distinctions in person (1st, 2nd, and 3rd), number (singular, dual , and plural in
4704-467: The photic zone in the oceans are relatively familiar habitat types. However the vast bulk of the ocean is inhospitable to air-breathing humans, with scuba divers limited to the upper 50 m (160 ft) or so. The lower limit for photosynthesis is 100 to 200 m (330 to 660 ft) and below that depth the prevailing conditions include total darkness, high pressure, little oxygen (in some places), scarce food resources and extreme cold. This habitat
4802-465: The 3rd millennium BC, or possibly earlier. The earliest written evidence is a Linear B clay tablet found in Messenia that dates to between 1450 and 1350 BC, making Greek the world's oldest recorded living language . Among the Indo-European languages, its date of earliest written attestation is matched only by the now-extinct Anatolian languages . The Greek language is conventionally divided into
4900-486: The Greek alphabet since approximately the 9th century BC. It was created by modifying the Phoenician alphabet , with the innovation of adopting certain letters to represent the vowels. The variant of the alphabet in use today is essentially the late Ionic variant, introduced for writing classical Attic in 403 BC. In classical Greek, as in classical Latin, only upper-case letters existed. The lower-case Greek letters were developed much later by medieval scribes to permit
4998-425: The Greek language are often emphasized. Although Greek has undergone morphological and phonological changes comparable to those seen in other languages, never since classical antiquity has its cultural, literary, and orthographic tradition been interrupted to the extent that one can speak of a new language emerging. Greek speakers today still tend to regard literary works of ancient Greek as part of their own rather than
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#17330851169965096-491: The Greek language was the Cypriot syllabary (also a descendant of Linear A via the intermediate Cypro-Minoan syllabary ), which is closely related to Linear B but uses somewhat different syllabic conventions to represent phoneme sequences. The Cypriot syllabary is attested in Cyprus from the 11th century BC until its gradual abandonment in the late Classical period, in favor of the standard Greek alphabet. Greek has been written in
5194-629: The Greek verb have likewise remained largely the same over the course of the language's history but with significant changes in the number of distinctions within each category and their morphological expression. Greek verbs have synthetic inflectional forms for: Many aspects of the syntax of Greek have remained constant: verbs agree with their subject only, the use of the surviving cases is largely intact (nominative for subjects and predicates, accusative for objects of most verbs and many prepositions, genitive for possessors), articles precede nouns, adpositions are largely prepositional, relative clauses follow
5292-661: The Greek-Albanian border. A significant percentage of Albania's population has knowledge of the Greek language due in part to the Albanian wave of immigration to Greece in the 1980s and '90s and the Greek community in the country. Prior to the Greco-Turkish War and the resulting population exchange in 1923 a very large population of Greek-speakers also existed in Turkey , though very few remain today. A small Greek-speaking community
5390-404: The acute during the late 20th century, and it has only been retained in typography . After the writing reform of 1982, most diacritics are no longer used. Since then, Greek has been written mostly in the simplified monotonic orthography (or monotonic system), which employs only the acute accent and the diaeresis. The traditional system, now called the polytonic orthography (or polytonic system),
5488-402: The ancient language; singular and plural alone in later stages), and gender (masculine, feminine, and neuter), and decline for case (from six cases in the earliest forms attested to four in the modern language). Nouns, articles, and adjectives show all the distinctions except for a person. Both attributive and predicative adjectives agree with the noun. The inflectional categories of
5586-436: The animals and plants reliant on that habitat suffer. Many countries have enacted legislation to protect their wildlife. This may take the form of the setting up of national parks, forest reserves and wildlife reserves, or it may restrict the activities of humans with the objective of benefiting wildlife. The laws may be designed to protect a particular species or group of species, or the legislation may prohibit such activities as
5684-409: The array of resources, physical and biotic factors that are present in an area, such as to support the survival and reproduction of a particular species . A species habitat can be seen as the physical manifestation of its ecological niche . Thus "habitat" is a species-specific term, fundamentally different from concepts such as environment or vegetation assemblages, for which the term "habitat-type"
5782-416: The changes in habitat types brought on by alterations in farming practices, tourism, pollution, fragmentation and climate change. Loss of habitat is the single greatest threat to any species. If an island on which an endemic organism lives becomes uninhabitable for some reason, the species will become extinct . Any type of habitat surrounded by a different habitat is in a similar situation to an island. If
5880-424: The characteristics of a given geographical area, particularly vegetation and climate. Thus habitat types do not refer to a single species but to multiple species living in the same area. For example, terrestrial habitat types include forest , steppe , grassland , semi-arid or desert . Fresh-water habitat types include marshes , streams , rivers , lakes , and ponds ; marine habitat types include salt marshes,
5978-570: The coast, the intertidal zone , estuaries , reefs , bays, the open sea, the sea bed, deep water and submarine vents . Habitat types may change over time. Causes of change may include a violent event (such as the eruption of a volcano , an earthquake , a tsunami , a wildfire or a change in oceanic currents); or change may occur more gradually over millennia with alterations in the climate , as ice sheets and glaciers advance and retreat, and as different weather patterns bring changes of precipitation and solar radiation . Other changes come as
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#17330851169966076-455: The collecting of bird eggs, the hunting of animals or the removal of plants. A general law on the protection of habitat types may be more difficult to implement than a site specific requirement. A concept introduced in the United States in 1973 involves protecting the critical habitat of endangered species , and a similar concept has been incorporated into some Australian legislation. International treaties may be necessary for such objectives as
6174-404: The early 19th century that was used for literary and official purposes in the newly formed Greek state. In 1976, Dimotiki was declared the official language of Greece, after having incorporated features of Katharevousa and thus giving birth to Standard Modern Greek , used today for all official purposes and in education . The historical unity and continuing identity between the various stages of
6272-420: The edge of each forest fragment, increased light encourages secondary growth of fast-growing species and old growth trees are more vulnerable to logging as access is improved. The birds that nest in their crevices, the epiphytes that hang from their branches and the invertebrates in the leaf litter are all adversely affected and biodiversity is reduced. Habitat fragmentation can be ameliorated to some extent by
6370-602: The epics of Homer , ancient Greek literature includes many works of lasting importance in the European canon . Greek is also the language in which many of the foundational texts in science and philosophy were originally composed. The New Testament of the Christian Bible was also originally written in Greek. Together with the Latin texts and traditions of the Roman world , the Greek texts and Greek societies of antiquity constitute
6468-695: The family Gyrinidae live at the air-water interface , Rhysodinae live in heartwood , and Paussinae carabids inhabit ant nests . Most species are predators . Other less-typical forms of feeding include: eating algae (family Haliplidae ), seed-feeding ( harpaline carabids), fungus-feeding (rhysodine carabids), and snail-feeding ( licinine and cychrine carabids). Some species are ectoparasitoids of insects ( brachinine and lebiine carabids) or of millipedes ( peleciine carabids). Some species are ovoviviparous , such as pseudomorphine carabids. The larvae are active, with well- chitinized cuticle, often with elongated cerci and five-segmented legs,
6566-452: The following periods: In the modern era, the Greek language entered a state of diglossia : the coexistence of vernacular and archaizing written forms of the language. What came to be known as the Greek language question was a polarization between two competing varieties of Modern Greek: Dimotiki , the vernacular form of Modern Greek proper, and Katharevousa , meaning 'purified', a compromise between Dimotiki and Ancient Greek developed in
6664-589: The foot-segment carrying two claws. Larvae have a fused labrum and no mandibular molae . Adephagans diverged from their sister group in the Late Permian , the most recent common ancestor of living adephagans probably existing in the early Triassic , around 240 million years ago. Both aquatic and terrestrial representatives of the suborder appear in fossil records of the late Triassic. The Jurassic fauna consisted of trachypachids , carabids, gyrinids, and haliplid -like forms. The familial and tribal diversification of
6762-745: The gaps created. Similarly, coastal habitat types can become dominated by kelp until the seabed is disturbed by a storm and the algae swept away, or shifting sediment exposes new areas for colonisation. Another cause of disturbance is when an area may be overwhelmed by an invasive introduced species which is not kept under control by natural enemies in its new habitat. Terrestrial habitat types include forests, grasslands, wetlands and deserts. Within these broad biomes are more specific habitat types with varying climate types, temperature regimes, soils, altitudes and vegetation. Many of these habitat types grade into each other and each one has its own typical communities of plants and animals. A habitat-type may suit
6860-696: The group spans the Mesozoic , with a few tribes radiating explosively during the Tertiary . The adephagans were formerly grouped into the Geadephaga with the two terrestrial families Carabidae and Trachypachidae and the Hydradephaga, for the aquatic families. However this is no longer used as the Hydradephaga are not a monophyletic group. Modern analysis has supported the clade Dytiscoidea instead, which includes many aquatic adephagans, notably excluding Gyrinidae. Rhysodidae
6958-419: The indigenous species have no immunity. The word "habitat" has been in use since about 1755 and derives from the Latin habitāre , to inhabit, from habēre , to have or to hold. Habitat can be defined as the natural environment of an organism , the type of place in which it is natural for it to live and grow. It is similar in meaning to a biotope ; an area of uniform environmental conditions associated with
7056-439: The infinitive entirely (employing a raft of new periphrastic constructions instead) and uses participles more restrictively. The loss of the dative led to a rise of prepositional indirect objects (and the use of the genitive to directly mark these as well). Ancient Greek tended to be verb-final, but neutral word order in the modern language is VSO or SVO. Modern Greek inherits most of its vocabulary from Ancient Greek, which in turn
7154-463: The late 15th century BC, was the first script used to write Greek. It is basically a syllabary , which was finally deciphered by Michael Ventris and John Chadwick in the 1950s (its precursor, Linear A , has not been deciphered and most likely encodes a non-Greek language). The language of the Linear B texts, Mycenaean Greek , is the earliest known form of Greek. Another similar system used to write
7252-720: The lichens growing in the grooves and on the raised surfaces are different from those growing on the veins of quartz. Lurking among these miniature "forests" are the microfauna , species of invertebrate , each with its own specific habitat requirements. There are numerous different microhabitat types in a wood; coniferous forest, broad-leafed forest, open woodland, scattered trees, woodland verges, clearings, and glades; tree trunk, branch, twig, bud, leaf, flower, and fruit; rough bark, smooth bark, damaged bark, rotten wood, hollow, groove, and hole; canopy, shrub layer, plant layer, leaf litter , and soil; buttress root, stump, fallen log, stem base, grass tussock, fungus, fern, and moss. The greater
7350-531: The majority have more specific requirements. The water velocity, its temperature and oxygen saturation are important factors, but in river systems, there are fast and slow sections, pools, bayous and backwaters which provide a range of habitat types. Similarly, aquatic plants can be floating, semi-submerged, submerged or grow in permanently or temporarily saturated soils besides bodies of water. Marginal plants provide important habitat for both invertebrates and vertebrates, and submerged plants provide oxygenation of
7448-621: The membership of Greece and Cyprus in the European Union, Greek is one of the organization's 24 official languages . Greek is recognized as a minority language in Albania, and used co-officially in some of its municipalities, in the districts of Gjirokastër and Sarandë . It is also an official minority language in the regions of Apulia and Calabria in Italy. In the framework of the European Charter for Regional or Minority Languages , Greek
7546-441: The modern period. The division into conventional periods is, as with all such periodizations, relatively arbitrary, especially because, in all periods, Ancient Greek has enjoyed high prestige, and the literate borrowed heavily from it. Across its history, the syllabic structure of Greek has varied little: Greek shows a mixed syllable structure, permitting complex syllabic onsets but very restricted codas. It has only oral vowels and
7644-515: The needs of the local residents for food, fuel and other resources. Faced with hunger and destitution, a farmer is likely to plough up a level patch of ground despite it being the last suitable habitat for an endangered species such as the San Quintin kangaroo rat , and even kill the animal as a pest. In the interests of ecotourism it is desirable that local communities are educated on the uniqueness of their flora and fauna. A monotypic habitat type
7742-399: The noun they modify and relative pronouns are clause-initial. However, the morphological changes also have their counterparts in the syntax, and there are also significant differences between the syntax of the ancient and that of the modern form of the language . Ancient Greek made great use of participial constructions and of constructions involving the infinitive, and the modern variety lacks
7840-521: The objects of study of the discipline of Classics . During antiquity , Greek was by far the most widely spoken lingua franca in the Mediterranean world . It eventually became the official language of the Byzantine Empire and developed into Medieval Greek . In its modern form , Greek is the official language of Greece and Cyprus and one of the 24 official languages of the European Union . It
7938-706: The older periods of Greek, loanwords into Greek acquired Greek inflections, thus leaving only a foreign root word. Modern borrowings (from the 20th century on), especially from French and English, are typically not inflected; other modern borrowings are derived from Albanian , South Slavic ( Macedonian / Bulgarian ) and Eastern Romance languages ( Aromanian and Megleno-Romanian ). Greek words have been widely borrowed into other languages, including English. Example words include: mathematics , physics , astronomy , democracy , philosophy , athletics , theatre, rhetoric , baptism , evangelist , etc. Moreover, Greek words and word elements continue to be productive as
8036-545: The proteins the flexibility they need. There are also unsaturated fats in their membranes which prevent them from solidifying at low temperatures. Hydrothermal vents were first discovered in the ocean depths in 1977. They result from seawater becoming heated after seeping through cracks to places where hot magma is close to the seabed. The under-water hot springs may gush forth at temperatures of over 340 °C (640 °F) and support unique communities of organisms in their immediate vicinity. The basis for this teeming life
8134-428: The provision of wildlife corridors connecting the fragments. These can be a river, ditch, strip of trees, hedgerow or even an underpass to a highway. Without the corridors, seeds cannot disperse and animals, especially small ones, cannot travel through the hostile territory, putting populations at greater risk of local extinction . Habitat disturbance can have long-lasting effects on the environment. Bromus tectorum
8232-573: The rocky seabed have found microbial communities apparently based on the products of reactions between water and the constituents of rocks. These communities have not been studied much, but may be an important part of the global carbon cycle . Rock in mines two miles deep also harbour microbes; these live on minute traces of hydrogen produced in slow oxidizing reactions inside the rock. These metabolic reactions allow life to exist in places with no oxygen or light, an environment that had previously been thought to be devoid of life. The intertidal zone and
8330-565: The seabed. Their growth rates and metabolisms tend to be slow, their eyes may be very large to detect what little illumination there is, or they may be blind and rely on other sensory inputs. A number of deep sea creatures are bioluminescent ; this serves a variety of functions including predation, protection and social recognition. In general, the bodies of animals living at great depths are adapted to high pressure environments by having pressure-resistant biomolecules and small organic molecules present in their cells known as piezolytes , which give
8428-400: The seeds of which survive the drought, but also some uniquely adapted perennials. Animals adapted to these extreme habitat types also exist; fairy shrimps can lay "winter eggs" which are resistant to desiccation , sometimes being blown about with the dust, ending up in new depressions in the ground. These can survive in a dormant state for as long as fifteen years. Some killifish behave in
8526-519: The setting up of marine reserves. Another international agreement, the Convention on the Conservation of Migratory Species of Wild Animals , protects animals that migrate across the globe and need protection in more than one country. Even where legislation protects the environment, a lack of enforcement often prevents effective protection. However, the protection of habitat types needs to take into account
8624-528: The southeastern United States. Its first intermediate host is a snail and the second, a glass shrimp . The final host is the waterfowl or mammal that consumes the shrimp. Although the vast majority of life on Earth lives in mesophyllic (moderate) environments, a few organisms, most of them microbes , have managed to colonise extreme environments that are unsuitable for more complex life forms. There are bacteria , for example, living in Lake Whillans , half
8722-437: The stressed vowel; the so-called breathing marks ( rough and smooth breathing ), originally used to signal presence or absence of word-initial /h/; and the diaeresis , used to mark the full syllabic value of a vowel that would otherwise be read as part of a diphthong. These marks were introduced during the course of the Hellenistic period. Actual usage of the grave in handwriting saw a rapid decline in favor of uniform usage of
8820-554: The structural diversity in the wood, the greater the number of microhabitat types that will be present. A range of tree species with individual specimens of varying sizes and ages, and a range of features such as streams, level areas, slopes, tracks, clearings, and felled areas will provide suitable conditions for an enormous number of biodiverse plants and animals. For example, in Britain it has been estimated that various types of rotting wood are home to over 1700 species of invertebrate. For
8918-455: The town's features to make their homes. Rats and mice have followed man around the globe, pigeons , peregrines , sparrows , swallows and house martins use the buildings for nesting, bats use roof space for roosting, foxes visit the garbage bins and squirrels , coyotes , raccoons and skunks roam the streets. About 2,000 coyotes are thought to live in and around Chicago . A survey of dwelling houses in northern European cities in
9016-418: The twentieth century found about 175 species of invertebrate inside them, including 53 species of beetle, 21 flies, 13 butterflies and moths, 13 mites, 9 lice, 7 bees, 5 wasps, 5 cockroaches, 5 spiders, 4 ants and a number of other groups. In warmer climates, termites are serious pests in the urban habitat; 183 species are known to affect buildings and 83 species cause serious structural damage. A microhabitat
9114-460: The water, absorb nutrients and play a part in the reduction of pollution. Rivers Ponds Lakes Marine habitats include brackish water, estuaries, bays, the open sea, the intertidal zone, the sea bed, reefs and deep / shallow water zones. Further variations include rock pools , sand banks , mudflats , brackish lagoons, sandy and pebbly beaches, and seagrass beds, all supporting their own flora and fauna. The benthic zone or seabed provides
9212-568: Was a distinct dialect of Greek itself. Aside from the Macedonian question, current consensus regards Phrygian as the closest relative of Greek, since they share a number of phonological, morphological and lexical isoglosses , with some being exclusive between them. Scholars have proposed a Graeco-Phrygian subgroup out of which Greek and Phrygian originated. Among living languages, some Indo-Europeanists suggest that Greek may be most closely related to Armenian (see Graeco-Armenian ) or
9310-549: Was also used as the official language of government and religion in the Christian Nubian kingdoms , for most of their history. Greek, in its modern form, is the official language of Greece, where it is spoken by almost the entire population. It is also the official language of Cyprus (nominally alongside Turkish ) and the British Overseas Territory of Akrotiri and Dhekelia (alongside English ). Because of
9408-546: Was also used in Ancient Greek. Greek has occasionally been written in the Latin script , especially in areas under Venetian rule or by Greek Catholics . The term Frankolevantinika / Φραγκολεβαντίνικα applies when the Latin script is used to write Greek in the cultural ambit of Catholicism (because Frankos / Φράγκος is an older Greek term for West-European dating to when most of (Roman Catholic Christian) West Europe
9506-566: Was under the control of the Frankish Empire ). Frankochiotika / Φραγκοχιώτικα (meaning 'Catholic Chiot') alludes to the significant presence of Catholic missionaries based on the island of Chios . Additionally, the term Greeklish is often used when the Greek language is written in a Latin script in online communications. The Latin script is nowadays used by the Greek-speaking communities of Southern Italy . The Yevanic dialect
9604-653: Was written by Romaniote and Constantinopolitan Karaite Jews using the Hebrew Alphabet . In a tradition, that in modern time, has come to be known as Greek Aljamiado , some Greek Muslims from Crete wrote their Cretan Greek in the Arabic alphabet . The same happened among Epirote Muslims in Ioannina . This also happened among Arabic-speaking Byzantine rite Christians in the Levant ( Lebanon , Palestine, and Syria ). This usage
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