Misplaced Pages

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44-420: According to the most probable theory, the name has its origin in the word jasan , i.e. ' ash '. Jeseník (respectively Jesenný potok ) was first the name of a stream that flowed through an ash forest in a valley. The name was Germanized to Gesenke (i.e. 'slope') and used as a name of a small town that was founded in the valley (but later disappeared), and then it was transferred first to the valley, and then to

88-458: A biological control for the management of EAB populations in the United States. The public is being cautioned to avoid transporting unfinished wood products, such as firewood, to slow the spread of this insect pest. Damage occurs when emerald ash borer larvae feed on the inner bark, phloem , inside branches and tree trunks. Feeding on the phloem prevents nutrients and water transportation. If

132-763: A zygote . The definition avoids reference to male and female reproductive structures, which are rare in fungi. An individual of a dioecious fungal species not only requires a partner for mating, but performs only one of the roles in nuclear transfer, as either the donor or the recipient. A monoecious fungal species can perform both roles, but may not be self-compatible. Dioecy has the demographic disadvantage compared with hermaphroditism that only about half of reproductive adults are able to produce offspring. Dioecious species must therefore have fitness advantages to compensate for this cost through increased survival, growth, or reproduction. Dioecy excludes self-fertilization and promotes allogamy (outcrossing), and thus tends to reduce

176-671: A considerable litter problem with their seeds. Rowans or mountain ashes have leaves and buds superficially similar to those of true ashes, but belong to the unrelated genus Sorbus in the rose family . The tree's common English name, "ash", traces back to the Old English æsc , which relates to the Proto-Indo-European for the tree, while the generic name originated in Latin from a Proto-Indo-European word for birch. Both words are also used to mean " spear " in their respective languages, as

220-410: A female gamete then fuse to produce a new diploid sporophyte. In bryophytes ( mosses , liverworts and hornworts ), the gametophytes are fully independent plants. Seed plant gametophytes are dependent on the sporophyte and develop within the spores, a condition known as endospory . In flowering plants, the male gametophytes develop within pollen grains produced by the sporophyte's stamens , and

264-498: A few species. The seeds, popularly known as "keys" or "helicopter seeds", are a type of fruit known as a samara . Some Fraxinus species are dioecious , having male and female flowers on separate plants but sex in ash is expressed as a continuum between male and female individuals, dominated by unisexual trees. With age, ash may change their sexual function from predominantly male and hermaphrodite towards femaleness ; if grown as an ornamental and both sexes are present, ashes can cause

308-436: A frequency about half of that for the forward transition. In Silene , since there is no monoecy, it is suggested that dioecy evolved through gynodioecy. Very few dioecious fungi have been discovered. Monoecy and dioecy in fungi refer to the donor and recipient roles in mating, where a nucleus is transferred from one haploid hypha to another, and the two nuclei then present in the same cell merge by karyogamy to form

352-517: A good food source for the frogs, but also reduces its resistance to the ash borer . Species with higher leaf tannin levels (including maples and non-native ash species) are taking the place of native ash, thanks to their greater resistance to the ash borer. They produce much less suitable food for the tadpoles, resulting in poor survival rates and small frog sizes. Ash species native to North America also provide important habitat and food for various other creatures native to North America. This includes

396-477: A reduction in the supply of healthy trees, especially in Europe, is making ash an increasingly expensive option. Ash was commonly used for the structural members of the bodies of cars made by carriage builders. Early cars had frames which were intended to flex as part of the suspension system to simplify construction. The Morgan Motor Company of Great Britain still manufactures sports cars with frames made from ash. It

440-467: A species, such as the colonies of Siphonophorae (Portuguese man-of-war), which may be either dioecious or monoecious . Land plants ( embryophytes ) differ from animals in that their life cycle involves alternation of generations . In animals, typically an individual produces gametes of one kind, either sperm or egg cells . The gametes have half the number of chromosomes of the individual producing them, so are haploid . Without further dividing,

484-431: A sperm and an egg cell fuse to form a zygote that develops into a new individual. In land plants, by contrast, one generation – the sporophyte generation – consists of individuals that produce haploid spores rather than haploid gametes . Spores do not fuse, but germinate by dividing repeatedly by mitosis to give rise to haploid multicellular individuals, the gametophytes , which produce gametes. A male gamete and

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528-469: Is a tonewood commonly used in the manufacture of electric guitars . It exhibits a pronounced bright tone with a scooped midrange. It is lightweight, easy to work and sand, accepts glue, stain , paint and finish very well and is inexpensive. All this has made it a favourite of large factories mass-producing instruments. The Fender musical instrument company has been continuously and uninterruptedly using Ash to make electric guitars since 1956. Swamp ash

572-457: Is a characteristic of certain species that have distinct unisexual individuals, each producing either male or female gametes , either directly (in animals ) or indirectly (in seed plants ). Dioecious reproduction is biparental reproduction. Dioecy has costs, since only the female part of the population directly produces offspring. It is one method for excluding self-fertilization and promoting allogamy (outcrossing), and thus tends to reduce

616-562: Is a wood-boring beetle accidentally introduced to North America from eastern Asia via solid wood packing material in the late 1980s to early 1990s. It has killed tens of millions of trees in 22 states in the United States and adjacent Ontario and Quebec in Canada. It threatens some seven billion ash trees in North America. Research is being conducted to determine whether three native Asian wasps that are natural predators of EAB could be used as

660-408: Is also used for making drum shells . Woodworkers generally consider ash a "poor cousin" to the other major open pore wood, oak, but it is useful in any furniture application. Ash veneers are extensively used in office furniture. Ash is not used much outdoors due to the heartwood having a low durability to ground contact, meaning it will typically perish within five years. The F. japonica species

704-788: Is called " dioecious " when each sporophyte plant has only one kind of spore-producing organ, all of whose spores give rise either to male gametophytes, which produce only male gametes (sperm), or to female gametophytes, which produce only female gametes (egg cells). For example, a single flowering plant sporophyte of a fully dioecious species like holly has either flowers with functional stamens producing pollen containing male gametes (staminate or 'male' flowers), or flowers with functional carpels producing female gametes (carpellate or 'female' flowers), but not both. (See Plant reproductive morphology for further details, including more complex cases, such as gynodioecy and androdioecy .) Slightly different terms, dioicous and monoicous , may be used for

748-444: Is favored as a material for making baseball bats by Japanese sporting-goods manufacturers. Its robust structure, good looks, and flexibility combine to make ash ideal for staircases. Ash stairs are extremely hard-wearing, which is particularly important for treads. Due to its elasticity, ash can also be steamed and bent to produce curved stair parts such as volutes (curled sections of handrail) and intricately shaped balusters. However,

792-454: Is more common in woody plants , and heterotrophic species. In most dioecious plants, whether male or female gametophytes are produced is determined genetically, but in some cases it can be determined by the environment, as in Arisaema species. Certain algae , such as some species of Polysiphonia , are dioecious. Dioecy is prevalent in the brown algae ( Phaeophyceae ) and may have been

836-406: Is unlikely that mutations for male and female sterility occurred at the same time. In angiosperms unisexual flowers evolve from bisexual ones. Dioecy occurs in almost half of plant families, but only in a minority of genera, suggesting recent evolution. For 160 families that have dioecious species, dioecy is thought to have evolved more than 100 times. In the family Caricaceae , dioecy is likely

880-444: Is used a lot in guitar building because of its figure. It is a choice of material for electric guitar bodies and, less commonly, for acoustic guitar bodies, known for its bright, cutting edge and sustaining quality. Some Fender Stratocasters and Telecasters are made of ash, (such as Bruce Springsteen 's Telecaster on the Born to Run album cover), as an alternative to alder. Ash

924-638: Is widely planted as a street tree in the United States. The inner bark of the blue ash ( F. quadrangulata ) has been used as a source for blue dye . In Sicily, Italy, sugars are obtained by evaporating the sap of the manna ash , extracted by making small cuts in the bark. The manna ash, native to southern Europe and southwest Asia, produces a blue-green sap, which has medicinal value as a mild laxative, demulcent, and weak expectorant. The young seedpods of Ash trees, also known as "keys", are edible for human consumption. In Britain, they are traditionally pickled with vinegar, sugar and spices. In Greek mythology ,

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968-807: The Czech Republic . There are 56 peaks with an altitude of at least 1,000 m, which are spread evenly throughout the territory. It is bordered with the Nízký Jeseník in the southeast, with the Zlatohorská Highlands in the northeast, with the Golden Mountains in the northwest, and with the Hanušovice Highlands in the southwest. Hrubý Jeseník is further subdivided into the microregions of Keprník Mountains, Medvědí Mountains and Praděd Mountains. The highest peaks are: A distinctive feature of

1012-553: The Meliae are nymphs associated with the ash, perhaps specifically of the manna ash ( Fraxinus ornus ), as dryads were nymphs associated with the oak . They appear in Hesiod 's Theogony, which states that they were born when drops of Ouranos's blood fell on the earth (Gaia). In Norse mythology , a vast, evergreen ash tree Yggdrasil ("the steed ( gallows ) of Odin "), watered by three magical springs, serves as axis mundi , sustaining

1056-625: The fungus Hymenoscyphus fraxineus , causing ash dieback in a large number of trees since the mid-1990s, particularly in eastern and northern Europe. The disease has infected about 90% of Denmark's ash trees. At the end of October 2012 in the UK, the Department for Environment, Food and Rural Affairs (Defra) reported that ash dieback had been discovered in mature woodland in Suffolk ; previous occurrences had been on young trees imported from Europe. In 2016,

1100-435: The gametophyte generation of non-vascular plants, although dioecious and monoecious are also used. A dioicous gametophyte either produces only male gametes (sperm) or produces only female gametes (egg cells). About 60% of liverworts are dioicous. Dioecy occurs in a wide variety of plant groups. Examples of dioecious plant species include ginkgos , willows , cannabis and African teak . As its specific name implies,

1144-612: The Javorový vrch National Nature Monument. Fraxinus Fraxinus ( / ˈ f r æ k s ɪ n ə s / ), commonly called ash , is a genus of plants in the olive and lilac family, Oleaceae , and comprises 45–65 species of usually medium-to-large trees , most of which are deciduous trees, although some subtropical species are evergreen trees. The genus is widespread throughout much of Europe, Asia, and North America. The leaves are opposite (rarely in whorls of three), and mostly pinnately compound , though simple in

1188-458: The ancestral sexual system. Dioecious flowering plants can evolve from monoecious ancestors that have flowers containing both functional stamens and functional carpels. Some authors argue monoecy and dioecy are related. In the genus Sagittaria , since there is a distribution of sexual systems, it has been postulated that dioecy evolved from monoecy through gynodioecy mainly from mutations that resulted in male sterility. However, since

1232-568: The ancestral state in that group. In plants, dioecy has evolved independently multiple times either from hermaphroditic species or from monoecious species. A previously untested hypothesis is that this reduces inbreeding; dioecy has been shown to be associated with increased genetic diversity and greater protection against deleterious mutations. Regardless of the evolutionary pathway the intermediate states need to have fitness advantages compared to cosexual flowers in order to survive. Dioecy evolves due to male or female sterility, although it

1276-597: The ancestral state is unclear, more work is needed to clarify the evolution of dioecy via monoecy. Dioecy usually evolves from hermaphroditism through gynodioecy but may also evolve through androdioecy , through distyly or through heterostyly . In the Asteraceae , dioecy may have evolved independently from hermaphroditism at least 5 or 9 times. The reverse transition, from dioecy back to hermaphroditism has also been observed, both in Asteraceae and in bryophytes, with

1320-487: The ash is attacked, the branches can die and eventually the whole tree can as well. Ways to detect emerald ash borer infestation include seeing bark peeling off, vertical cracks in the bark, seeing galleries within the tree that contain powdery substance, and D-shaped exit holes on the branches or trunk. Not all of these may be present, but any of these warning signs could be an indication of possible infestation. The European ash, Fraxinus excelsior , has been affected by

1364-401: The ash tree was reported as in danger of extinction in Europe. Ash is a hardwood and is dense (within 20% of 670 kg/m for Fraxinus americana , and higher at 710 kg/m for Fraxinus excelsior ), tough and very strong but elastic , extensively used for making bows , tool handles, baseball bats , hurleys , and other uses demanding high strength and resilience . Ash

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1408-650: The entire territory of Hrubý Jeseník is protected within the Jeseníky Protected Landscape Area . The Jeseníky PLA then extends further to the north into the Zlatohorská Highlands and to the south into the Hanušovice Highlands . Furthermore, small-scaled protected areas are defined. There are 32 small-scaled protected areas in the Jeseníky PLA. The most important are the national nature reserves Praděd , Šerák-Keprník , Rejvíz and Skřítek (Skřítek and Rejvíz extend only marginally into Hrubý Jeseník), and

1452-585: The expression of recessive deleterious mutations present in a population. In trees, compensation is realized mainly through increased seed production by females. This in turn is facilitated by a lower contribution of reproduction to population growth, which results in no demonstrable net costs of having males in the population compared to being hermaphroditic. Dioecy may also accelerate or retard lineage diversification in angiosperms . Dioecious lineages are more diversified in certain genera, but less in others. An analysis suggested that dioecy neither consistently places

1496-536: The expression of recessive deleterious mutations present in a population. Plants have several other methods of preventing self-fertilization including, for example, dichogamy , herkogamy , and self-incompatibility . In zoology, dioecy means that an animal is either male or female, in which case the synonym gonochory is more often used. For example, most animal species are gonochoric, almost all vertebrate species are gonochoric, and all bird and mammal species are gonochoric. Dioecy may also describe colonies within

1540-428: The female gametophytes develop within ovules produced by the sporophyte's carpels . The sporophyte generation of a seed plant is called " monoecious " when each sporophyte plant has both kinds of spore-producing organ but in separate flowers or cones. For example, a single flowering plant of a monoecious species has both functional stamens and carpels, in separate flowers. The sporophyte generation of seed plants

1584-417: The larvae of multiple long-horn beetles , as well as other insects including those in the genus Tropidosteptes , lace bugs , aphids , larvae of gall flies , and caterpillars. Birds are also interested in black, green, and white ash trees. The black ash alone supports wood ducks , wild turkey , cardinals , pine grosbeaks , cedar waxwings , and yellow-bellied sapsuckers , with habitat and food (such as

1628-548: The mountain range. The most important are the Desná and the sources of the Opava River. The only notable body of water is the Dlouhé stráně Reservoir . Due to the mountainous character of the landscape, there are no larger settlements here. The largest settlement in the territory is Vrbno pod Pradědem . The town of Jeseník is located just beyond the borders of the mountain range. Almost

1672-487: The nine worlds of the cosmos in its roots and branches. Askr , the first man in Norse myth, literally means 'ash'. In Italian folklore , an ash stake could be used to kill a vampire . Dioecy Dioecy ( / d aɪ ˈ iː s i / dy- EE -see ; from Ancient Greek διοικία dioikía  'two households'; adj. dioecious , / d aɪ ˈ iː ʃ ( i ) ə s / dy- EE -sh(ee-)əs )

1716-657: The perennial stinging nettle Urtica dioica is dioecious, while the annual nettle Urtica urens is monoecious. Dioecious flora are predominant in tropical environments. About 65% of gymnosperm species are dioecious, but almost all conifers are monoecious. In gymnosperms, the sexual systems dioecy and monoecy are strongly correlated with the mode of pollen dispersal, monoecious species are predominantly wind dispersed ( anemophily ) and dioecious species animal-dispersed ( zoophily ). About 6 percent of flowering plant species are entirely dioecious and about 7% of angiosperm genera contain some dioecious species. Dioecy

1760-483: The relief is also Mt. Medvědí vrch, which at 1,216 m (3,990 ft) is the highest mountain of the Medvědí Mountains microregion and is among the most prominent mountains of Hrubý Jeseník. The territory has an approximately circular shape. Hrubý Jeseník has an area of 530 square kilometres (200 sq mi) and an average elevation of 887.6 metres (2,912 ft). Several rivers and streams originate in

1804-401: The sap being of interest to the sapsucker) among others. Many mammalian species from meadow voles eating the seeds to white-tailed deer eating the foliage to silver-haired bats nesting will also make use of ash trees. Ash is used as a food plant by the larvae of some Lepidoptera species ( butterflies and moths ). The emerald ash borer ( Agrilus planipennis ), also called EAB,

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1848-592: The whole mountain range. Later the name was changed back to Czech Jeseník . Jeseníky (plural form of Jeseník) is a collective term for an area that includes the mountain ranges of Hrubý Jeseník (literally 'rough Jeseník') and Nízký Jeseník (i.e. 'low Jeseník'). Hrubý Jeseník is a mesoregion of the Eastern Sudetes , which is part of the Sudetes within the Bohemian Massif . It is the second-highest mountain range in

1892-460: The wood is good for shafts. Species are arranged into sections supported by phylogenetic analysis: North American native ash tree species are a critical food source for North American frogs, as their fallen leaves are particularly suitable for tadpoles to feed upon in ponds (both temporary and permanent), large puddles, and other water bodies. Lack of tannins in the American ash makes their leaves

1936-402: Was also widely used by early aviation pioneers for aircraft construction. It lights and burns easily, so is used for starting fires and barbecues , and is usable for maintaining a fire, though it produces only a moderate heat. The two most economically important species for wood production are white ash, in eastern North America, and European ash in Europe. The green ash ( F. pennsylvanica )

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