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A feral (from Latin fera  'a wild beast') animal or plant is one that lives in the wild but is descended from domesticated individuals. As with an introduced species , the introduction of feral animals or plants to non-native regions may disrupt ecosystems and has, in some cases, contributed to extinction of indigenous species . The removal of feral species is a major focus of island restoration .

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121-437: A feral animal is one that has escaped from a domestic or captive status and is living more or less as a wild animal, or one that is descended from such animals. Other definitions include animals that have changed from being domesticated to being wild, natural, or untamed. Some common examples of animals with feral populations are horses , dogs , goats , cats , rabbits , camels , and pigs . Zoologists generally exclude from

242-571: A clade , meaning that they arose from a single common ancestor . Over 1.5 million living animal species have been described , of which around 1.05 million are insects , over 85,000 are molluscs , and around 65,000 are vertebrates . It has been estimated there are as many as 7.77 million animal species on Earth. Animal body lengths range from 8.5 μm (0.00033 in) to 33.6 m (110 ft). They have complex ecologies and interactions with each other and their environments, forming intricate food webs . The scientific study of animals

363-515: A dominance hierarchy . This is maintained in several ways. Cattle often engage in mock fights where they test each other's strength in a non-aggressive way. Licking is primarily performed by subordinates and received by dominant animals. Mounting is a playful behavior shown by calves of both sexes and by bulls and sometimes by cows in estrus, however, this is not a dominance related behavior as has been found in other species. Dominance-associated aggressiveness does not correlate with rank position, but

484-449: A feedlot for "45 days or more in a 12-month period". Historically, the cattle population of Britain rose from 9.8 million in 1878 to 11.7 million in 1908, but beef consumption rose much faster. Britain became the "stud farm of the world" exporting livestock to countries where there were no indigenous cattle. In 1929 80% of the meat trade of the world was products of what were originally English breeds. There were nearly 70 million cattle in

605-399: A gastrula with a digestive chamber and two separate germ layers , an external ectoderm and an internal endoderm . In most cases, a third germ layer, the mesoderm , also develops between them. These germ layers then differentiate to form tissues and organs. Repeated instances of mating with a close relative during sexual reproduction generally leads to inbreeding depression within

726-423: A noxious weed . The adaptive and ecological variables seen in plants that go wild closely resemble those of animals. Feral populations of crop plants, along with hybridization between crop plants and their wild relatives, brings a risk that genetically engineered characteristics such as pesticide resistance could be transferred to weed plants. The unintended presence of genetically modified crop plants or of

847-547: A clade Xenambulacraria for the Xenacoelamorpha + Ambulacraria; this is either within Deuterostomia, as sister to Chordata, or the Deuterostomia are recovered as paraphyletic, and Xenambulacraria is sister to the proposed clade Centroneuralia , consisting of Chordata + Protostomia. Eumetazoa , a clade which contains Ctenophora and ParaHoxozoa , has been proposed as a sister group to Porifera . A competing hypothesis

968-415: A cow is about nine months long. The ratio of male to female offspring at birth is approximately 52:48. A cow's udder has two pairs of mammary glands or teats. Farms often use artificial insemination , the artificial deposition of semen in the female's genital tract ; this allows farmers to choose from a wide range of bulls to breed their cattle. Estrus too may be artificially induced to facilitate

1089-414: A dairy, which may be onsite at the farm or the milk may be shipped to a dairy plant for eventual sale of a dairy product. Lactation is induced in heifers and spayed cows by a combination of physical and psychological stimulation, by drugs, or by a combination of those methods. For mother cows to continue producing milk, they give birth to one calf per year. If the calf is male, it is generally slaughtered at

1210-404: A diurnal pattern, with a stronger preference for clover in the morning, and the proportion of grass increasing towards the evening. When grazing, cattle vary several aspects of their bite, i.e. tongue and jaw movements, depending on characteristics of the plant they are eating. Bite area decreases with the density of the plants but increases with their height. Bite area is determined by the sweep of

1331-606: A domestic state. Such observations can provide useful information for the stock breeders or other owners of the domesticated conspecifics (i.e. animals of the same species). American mustangs have been protected since 1971 in part due to their romance and connection to the history of the American West . A similar situation is that of the Danube Delta horse from the Letea Forest in the Danube Delta . The Romanian government

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1452-520: A dominant cockerel, several hens, and subordinate cocks. Sometimes the dominant cockerel is designated by a fight between cocks. A feral population can have a significant impact on an ecosystem by predation on vulnerable plants or animals, or by competition with indigenous species. Feral plants and animals constitute a significant share of invasive species , and can be a threat to endangered species . However, they may also replace species lost from an ecosystem on initial human arrival to an area, or increase

1573-530: A feral state. However, in places where there are few predators, they may thrive, for example in the case of the Soay sheep . Both goats and sheep were sometimes intentionally released and allowed to go feral on island waypoints frequented by mariners, to serve as a ready food source. The dromedary camel, which has been domesticated for over 3,000 years, will also readily go feral. A substantial population of feral dromedaries, descended from pack animals that escaped in

1694-619: A learning experiment had higher heart rates, indicating an emotional reaction to their own learning. After separation from their mothers, Holstein calves react, indicating low mood. Similarly, after hot-iron dehorning , calves react to the post-operative pain. The position of the ears has been used as an indicator of emotional state. Cattle can tell when other cattle are stressed by the chemicals in their urine. Cattle are gregarious , and even short-term isolation causes psychological stress . When heifers are isolated, vocalizations, heart rate and plasma cortisol all increase. When visual contact

1815-504: A longer period of time than horses . Oxen are used worldwide, especially in developing countries . There are some 11 million draft oxen in sub-Saharan Africa, while in 1998 India had over 65 million oxen. At the start of the 21st century, about half the world's crop production depended on land preparation by draft animals. Cattle are not often kept solely for hides, and they are usually a by-product of beef production. Hides are used mainly for leather products such as shoes. In 2012, India

1936-633: A misfolded brain protein , in contaminated meat. Among the intestinal parasites of cattle are Paramphistomum flukes, affecting the rumen, and hookworms in the small intestine. Climate change is expected to exacerbate heat stress in cattle, and for longer periods. Heat-stressed cattle may experience accelerated breakdown of adipose tissue by the liver, causing lipidosis . Cattle eat less when heat stressed, resulting in ruminal acidosis , which can lead to laminitis . Cattle can attempt to deal with higher temperatures by panting more often ; this rapidly decreases carbon dioxide concentrations at

2057-668: A mix of taurine cattle, zebu, and yak. The aurochs originally ranged throughout Europe, North Africa, and much of Asia. In historical times, its range became restricted to Europe, and the last known individual died in Mazovia , Poland, around 1627. Breeders have attempted to recreate a similar appearance to the aurochs by crossing traditional types of domesticated cattle, producing the Heck breed . A group of taurine-type cattle exist in Africa; they either represent an independent domestication event or were

2178-1105: A population due to the increased prevalence of harmful recessive traits. Animals have evolved numerous mechanisms for avoiding close inbreeding . Some animals are capable of asexual reproduction , which often results in a genetic clone of the parent. This may take place through fragmentation ; budding , such as in Hydra and other cnidarians ; or parthenogenesis , where fertile eggs are produced without mating , such as in aphids . Animals are categorised into ecological groups depending on their trophic levels and how they consume organic material . Such groupings include carnivores (further divided into subcategories such as piscivores , insectivores , ovivores , etc.), herbivores (subcategorized into folivores , graminivores , frugivores , granivores , nectarivores , algivores , etc.), omnivores , fungivores , scavengers / detritivores , and parasites . Interactions between animals of each biome form complex food webs within that ecosystem . In carnivorous or omnivorous species, predation

2299-768: A powerful greenhouse gas, as a byproduct of enteric fermentation , with each cow belching out 100kg a year. Additional methane is produced by anaerobic fermentation of stored manure . The FAO estimates that in 2015 around 7% of global greenhouse gas emissions were due to cattle, but this is uncertain. Reducing methane emissions quickly helps limit climate change . Concentrated animal feeding operations in particular produce substantial amounts of wastewater and manure, which can cause environmental harms such as soil erosion, human and animal exposure to toxic chemicals, development of antibiotic resistant bacteria and an increase in E. coli contamination. In many world regions, overgrazing by cattle has reduced biodiversity of

2420-497: A quarter of the world's meat comes from cattle. World cattle meat production in 2021 was 72.3 million tons. Certain breeds of cattle, such as the Holstein-Friesian , are used to produce milk , much of which is processed into dairy products such as butter , cheese , and yogurt . Dairy cattle are usually kept on specialized dairy farms designed for milk production. Most cows are milked twice per day, with milk processed at

2541-428: A range of production and behavioral characteristics for both dairy and beef cattle. Cattle have played a key role in human history , having been domesticated since at least the early neolithic age. Archaeozoological and genetic data indicate that cattle were first domesticated from wild aurochs ( Bos primigenius ) approximately 10,500 years ago. There were two major areas of domestication: one in central Anatolia ,

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2662-400: A similar phenomenon of the genetic mixing of feral domestic cats and their wild counterparts. Feral animals compete with domestic livestock, and may degrade fences, water sources, and vegetation (by overgrazing or introducing seeds of invasive plants). Although hotly disputed, some cite as an example the competition between feral horses and cattle in the western United States. Another example

2783-399: A single opening, which serves as both mouth and anus. Animals in both phyla have distinct tissues, but these are not organised into discrete organs . They are diploblastic , having only two main germ layers, ectoderm and endoderm. Cattle Cattle ( Bos taurus ) are large, domesticated , bovid ungulates widely kept as livestock . They are prominent modern members of

2904-461: A third independent domestication from the North African aurochs. Whether there have been two or three domestications, European, African, and Asian cattle share much of their genomes both through their species ancestry and through repeated migrations of livestock and genetic material between species, as shown in the diagram. Cattle were originally identified as three separate species: Bos taurus ,

3025-456: A wider study mapped a further 4.2% of the cattle genome. Behavioral traits of cattle can be as heritable as some production traits, and often, the two can be related. The heritability of temperament (response to isolation during handling) has been calculated as 0.36 and 0.46 for habituation to handling. Rangeland assessments show that the heritability of aggressiveness in cattle is around 0.36. Quantitative trait loci have been found for

3146-506: A young age to produce veal . Cows produce milk until three weeks before birth. Over the last fifty years, dairy farming has become more intensive to increase the yield of milk produced by each cow. The Holstein-Friesian is the breed of dairy cow most common in the UK, Europe and the United States. It has been bred selectively to produce the highest yields of milk of any cow. The average in the UK

3267-464: Is a consumer–resource interaction where the predator feeds on another organism, its prey , who often evolves anti-predator adaptations to avoid being fed upon. Selective pressures imposed on one another lead to an evolutionary arms race between predator and prey, resulting in various antagonistic/ competitive coevolutions . Almost all multicellular predators are animals. Some consumers use multiple methods; for example, in parasitoid wasps ,

3388-725: Is a diurnal rhythm in suckling, peaking at roughly 6am, 11:30am, and 7pm. Under natural conditions, calves stay with their mother until weaning at 8 to 11 months. Heifer and bull calves are equally attached to their mothers in the first few months of life. Cattle have a variety of cognitive abilities. They can memorize the locations of multiple food sources, and can retain memories for at least 48 days. Young cattle learn more quickly than adults, and calves are capable of discrimination learning, distinguishing familiar and unfamiliar animals, and between humans, using faces and other cues. Calves prefer their own mother's vocalizations to those of an unfamiliar cow. Vocalizations provide information on

3509-433: Is a stage in embryonic development that is unique to animals, allowing cells to be differentiated into specialised tissues and organs. All animals are composed of cells, surrounded by a characteristic extracellular matrix composed of collagen and elastic glycoproteins . During development, the animal extracellular matrix forms a relatively flexible framework upon which cells can move about and be reorganised, making

3630-471: Is around 22 litres per day. Dairy is a large industry worldwide. In 2023, the 27 European Union countries produced 143 million tons of cow's milk; the United States 104.1 million tons; and India 99.5 million tons. India further produces 94.4 million tons of buffalo milk , making it (in 2023) the world's largest milk producer; its dairy industry employs some 80 million people. Oxen are cattle trained as draft animals . Oxen can pull heavier loads and for

3751-474: Is better than that of horses, but worse at localising sounds than goats, and much worse than dogs or humans. They can distinguish between live and recorded human speech. Olfaction probably plays a large role in their social life, indicating social and reproductive status. Cattle can tell when other animals are stressed by smelling the alarm chemicals in their urine. Cattle can be trained to recognise conspecific individuals using olfaction only. Cattle live in

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3872-428: Is closely related to rank distance between individuals. The horns of cattle are honest signals used in mate selection. Horned cattle attempt to keep greater distances between themselves and have fewer physical interactions than hornless cattle, resulting in more stable social relationships. In calves, agonistic behavior becomes less frequent as space allowance increases, but not as group size changes, whereas in adults,

3993-536: Is common in India. Many feral animals can sometimes be captured at little cost and thus constitute a significant resource. Throughout most of Polynesia and Melanesia feral pigs constitute the primary sources of animal protein. Prior to the Wild and Free-Roaming Horses and Burros Act of 1971 , American mustangs were routinely captured and sold for horsemeat. In Australia, feral goats, pigs, horses, and dromedaries are harvested for

4114-461: Is considering the protection of the feral horses and transforming them into a tourist attraction, after it first approved the killing of the entire population. Due to the intervention of numerous organizations and widespread popular disapproval of the Romanians the horses have been saved, but still have an uncertain fate as their legal status is unclear and local people continue to claim the right to use

4235-496: Is contentious, and authorities such as the American Society of Mammalogists treat these taxa as separate species. Complicating the matter is the ability of cattle to interbreed with other closely related species. Hybrid individuals and even breeds exist, not only between taurine cattle and zebu (such as the sanga cattle ( Bos taurus africanus x Bos indicus ), but also between one or both of these and some other members of

4356-458: Is disputed, as they might be water-escape or other structures. Animals are monophyletic , meaning they are derived from a common ancestor. Animals are the sister group to the choanoflagellates , with which they form the Choanozoa . The dates on the phylogenetic tree indicate approximately how many millions of years ago ( mya ) the lineages split. Ros-Rocher and colleagues (2021) trace

4477-450: Is known as zoology , and the study of animal behaviour is known as ethology . The animal kingdom is divided into five infrakingdoms / superphyla , namely Porifera , Ctenophora , Placozoa , Cnidaria and Bilateria . Most living animal species belong to the infrakingdom Bilateria, a highly proliferative clade whose members have a bilaterally symmetric body plan , and the vast majority of bilaterians belong to two large superphyla :

4598-488: Is now northern Syria. Although European cattle are largely descended from the taurine lineage, gene flow from African cattle (partially of indicine origin) contributed substantial genomic components to both southern European cattle breeds and their New World descendants. A study on 134 breeds showed that modern taurine cattle originated from Africa, Asia, North and South America, Australia, and Europe. Some researchers have suggested that African taurine cattle are derived from

4719-496: Is of goats competing with cattle in Australia, or goats that degrade trees and vegetation in environmentally-stressed regions of Africa. Accidental crossbreeding by feral animals may result in harm to breeding programs of pedigreed animals; their presence may also excite domestic animals and push them to escape. Feral populations can also pass on transmissible infections to domestic herds. Loss to farmers by aggressive feral dog population

4840-808: Is often used to refer only to nonhuman animals. The term metazoa is derived from Ancient Greek μετα ( meta ) 'after' (in biology, the prefix meta- stands for 'later') and ζῷᾰ ( zōia ) 'animals', plural of ζῷον zōion 'animal'. Animals have several characteristics that set them apart from other living things. Animals are eukaryotic and multicellular . Unlike plants and algae , which produce their own nutrients , animals are heterotrophic , feeding on organic material and digesting it internally. With very few exceptions, animals respire aerobically . All animals are motile (able to spontaneously move their bodies) during at least part of their life cycle , but some animals, such as sponges , corals , mussels , and barnacles , later become sessile . The blastula

4961-422: Is only produced by sponges and pelagophyte algae. Its likely origin is from sponges based on molecular clock estimates for the origin of 24-ipc production in both groups. Analyses of pelagophyte algae consistently recover a Phanerozoic origin, while analyses of sponges recover a Neoproterozoic origin, consistent with the appearance of 24-ipc in the fossil record. The first body fossils of animals appear in

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5082-496: Is re-instated, vocalizations rapidly decline; heart rate decreases more rapidly if the returning cattle are familiar to the previously isolated individual. Mirrors have been used to reduce stress in isolated cattle. The average sleep time of a domestic cow is about 4 hours a day. Cattle do have a stay apparatus , but do not sleep standing up; they lie down to sleep deeply. In 2009, the National Institutes of Health and

5203-499: Is some 25–30 years. Beef cattle go to slaughter at around 18 months, and dairy cows at about five years. Cattle are ruminants , meaning their digestive system is highly specialized for processing plant material such as grass rich in cellulose , a tough carbohydrate polymer which many animals cannot digest. They do this in symbiosis with micro-organisms – bacteria , fungi , and protozoa – that possess cellulases , enzymes that split cellulose into its constituent sugars . Among

5324-409: Is spread by midges . Psoroptic mange is a disabling skin condition caused by mites . Bovine tuberculosis is caused by a bacterium; it causes disease in humans and in wild animals such as deer and badgers. Foot-and-mouth disease is caused by a virus, affects a range of hoofed livestock and is highly contagious. Bovine spongiform encephalopathy is a neurodegenerative disease spread by a prion ,

5445-537: Is the Benthozoa clade, which would consist of Porifera and ParaHoxozoa as a sister group of Ctenophora . Several animal phyla lack bilateral symmetry. These are the Porifera (sea sponges), Placozoa , Cnidaria (which includes jellyfish , sea anemones , and corals), and Ctenophora (comb jellies). Sponges are physically very distinct from other animals, and were long thought to have diverged first, representing

5566-624: Is the number of animals within a specified area. High stocking density can affect cattle health, welfare, productivity, and feeding behaviour. Densely-stocked cattle feed more rapidly and lie down sooner, increasing the risk of teat infection, mastitis, and embryo loss . The stress and negative health impacts induced by high stocking density such as in concentrated animal feeding operations or feedlots , auctions, and transport may be detrimental to cattle welfare. To produce milk from dairy cattle , most calves are separated from their mothers soon after birth and fed milk replacement in order to retain

5687-564: The American west, they are called mustangs . Other isolated feral populations exist, including the Chincoteague Pony and the Banker horse . They are often referred to as " wild horses ", but this is a misnomer. There are truly "wild" horses that have never been domesticated, most notably Przewalski's horse . While the horse was originally indigenous to North America, the wild ancestor died out at

5808-605: The Cambrian explosion , which began around 539  million years ago (Mya), and most classes during the Ordovician radiation 485.4 Mya. 6,331 groups of genes common to all living animals have been identified; these may have arisen from a single common ancestor that lived about 650 Mya during the Cryogenian period. Historically, Aristotle divided animals into those with blood and those without . Carl Linnaeus created

5929-614: The Ediacaran , represented by forms such as Charnia and Spriggina . It had long been doubted whether these fossils truly represented animals, but the discovery of the animal lipid cholesterol in fossils of Dickinsonia establishes their nature. Animals are thought to have originated under low-oxygen conditions, suggesting that they were capable of living entirely by anaerobic respiration , but as they became specialized for aerobic metabolism they became fully dependent on oxygen in their environments. Many animal phyla first appear in

6050-448: The Levant and Western Iran , giving rise to the taurine line, and a second in the area that is now Pakistan, resulting in the indicine line. Modern mitochondrial DNA variation indicates the taurine line may have arisen from as few as 80 aurochs tamed in the upper reaches of Mesopotamia near the villages of Çayönü Tepesi in what is now southeastern Turkey, and Dja'de el-Mughara in what

6171-914: The Levant and Western Iran . A separate domestication event occurred in the Indian subcontinent , which gave rise to zebu. There were over 940 million cattle in the world by 2022. Cattle are responsible for around 7% of global greenhouse gas emissions . They were one of the first domesticated animals to have a fully-mapped genome . The term cattle was borrowed from Anglo-Norman catel (replacing native Old English terms like kine , now considered archaic, poetic, or dialectal), itself from Medieval Latin capitale 'principal sum of money, capital', itself derived in turn from Latin caput 'head'. Cattle originally meant movable personal property , especially livestock of any kind, as opposed to real property (the land, which also included wild or small free-roaming animals such as chickens—they were sold as part of

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6292-459: The US Department of Agriculture reported having mapped the bovine genome . Cattle have some 22,000 genes, of which 80% are shared with humans; they have about 1000 genes that they share with dogs and rodents, but not with humans. Using this bovine "HapMap", researchers can track the differences between breeds that affect meat and milk yields. Early research focused on Hereford genetic sequences;

6413-482: The Wnt and TGF-beta signalling pathways which may have enabled animals to become multicellular by providing a pattern for the body's system of axes (in three dimensions), and another 7 are for transcription factors including homeodomain proteins involved in the control of development . Giribet and Edgecombe (2020) provide what they consider to be a consensus internal phylogeny of the animals, embodying uncertainty about

6534-491: The aurochs , were aggressive, similar to the modern Cape buffalo . Modern cattle, especially those raised on open range, are generally more docile, but when threatened can display aggression. Cattle, particularly those raised for beef , are often allowed to roam quite freely and have established long term independence in Australia , New Zealand and several Pacific Islands along with small populations of semi-feral animals roaming

6655-414: The bird , reptile , and mammal populations. A local population of feral cats living in an urban area and using a common food source is sometimes called a feral cat colony . As feral cats multiply quickly, it is difficult to control their populations. Animal shelters attempt to adopt out feral cats, especially kittens, but often are overwhelmed with sheer numbers and euthanasia is used. In rural areas of

6776-665: The evolutionary relationships between taxa . Humans make use of many other animal species for food (including meat , eggs , and dairy products ), for materials (such as leather , fur , and wool ), as pets and as working animals for transportation , and services . Dogs , the first domesticated animal, have been used in hunting , in security and in warfare , as have horses , pigeons and birds of prey ; while other terrestrial and aquatic animals are hunted for sports, trophies or profits. Non-human animals are also an important cultural element of human evolution , having appeared in cave arts and totems since

6897-468: The fossil record during the Cambrian explosion , starting about 539 million years ago, in beds such as the Burgess shale . Extant phyla in these rocks include molluscs , brachiopods , onychophorans , tardigrades , arthropods , echinoderms and hemichordates , along with numerous now-extinct forms such as the predatory Anomalocaris . The apparent suddenness of the event may however be an artifact of

7018-520: The genus Bos – yaks (the dzo or yattle ), banteng , and gaur . Hybrids such as the beefalo breed can even occur between taurine cattle and either species of bison , leading some authors to consider them part of the genus Bos , as well. The hybrid origin of some types may not be obvious – for example, genetic testing of the Dwarf Lulu breed, the only taurine-type cattle in Nepal, found them to be

7139-497: The lobe-finned fish Tiktaalik started to move on to land in the late Devonian , about 375 million years ago. Animals occupy virtually all of earth's habitats and microhabitats, with faunas adapted to salt water, hydrothermal vents, fresh water, hot springs, swamps, forests, pastures, deserts, air, and the interiors of other organisms. Animals are however not particularly heat tolerant ; very few of them can survive at constant temperatures above 50 °C (122 °F) or in

7260-525: The mallard duck , wild boar , the rock dove or pigeon, the red junglefowl ( Gallus gallus ) (ancestor of all chickens ), carp , and more recently salmon . Other examples of genetic swamping lie in the breeding history of dingoes . Dingoes are wild true dogs that will interbreed with dogs of other origins, thus leading to the proliferation of dingo hybrids and the possibility of the extinction of pure wild dingoes. Researches in Scotland have remarked on

7381-590: The miniature Zebu are kept as pets . Taurine cattle are widely distributed across Europe and temperate areas of Asia, the Americas, and Australia. Zebus are found mainly in India and tropical areas of Asia, America, and Australia. Sanga cattle are found primarily in sub-Saharan Africa . These types, sometimes classified as separate species or subspecies, are further divided into over 1,000 recognized breeds . Around 10,500 years ago, taurine cattle were domesticated from wild aurochs progenitors in central Anatolia ,

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7502-495: The protostomes , which includes organisms such as arthropods , molluscs , flatworms , annelids and nematodes ; and the deuterostomes , which include echinoderms , hemichordates and chordates , the latter of which contains the vertebrates . The much smaller basal phylum Xenacoelomorpha have an uncertain position within Bilateria. Animals first appear in the fossil record in the late Cryogenian period and diversified in

7623-413: The rumen , reticulum , omasum , and abomasum . The rumen is the largest compartment and it harbours the most important parts of the microbiome. The reticulum, the smallest compartment, is known as the "honeycomb". The omasum's main function is to absorb water and nutrients from the digestible feed. The abomasum has a similar function to the human stomach. Cattle regurgitate and re-chew their food in

7744-510: The 19th and early 20th centuries, thrives in the Australian interior today. Water buffalo run rampant in Western and Northern Australia. The Australian government encourages the hunting of feral water buffalo because of their large numbers. Cattle have been domesticated since the neolithic era, but can survive on open range for months or years with little or no supervision. Their ancestors,

7865-698: The Cambrian explosion, possibly as early as 1 billion years ago. Early fossils that might represent animals appear for example in the 665-million-year-old rocks of the Trezona Formation of South Australia . These fossils are interpreted as most probably being early sponges . Trace fossils such as tracks and burrows found in the Tonian period (from 1 gya) may indicate the presence of triploblastic worm-like animals, roughly as large (about 5 mm wide) and complex as earthworms. However, similar tracks are produced by

7986-712: The Cnidaria) never grow larger than 20 μm , and one of the smallest species ( Myxobolus shekel ) is no more than 8.5 μm when fully grown. The following table lists estimated numbers of described extant species for the major animal phyla, along with their principal habitats (terrestrial, fresh water, and marine), and free-living or parasitic ways of life. Species estimates shown here are based on numbers described scientifically; much larger estimates have been calculated based on various means of prediction, and these can vary wildly. For instance, around 25,000–27,000 species of nematodes have been described, while published estimates of

8107-529: The European or "taurine" cattle (including similar types from Africa and Asia); Bos indicus , the Indicine or "zebu" ; and the extinct Bos primigenius , the aurochs . The aurochs is ancestral to both zebu and taurine cattle. They were later reclassified as one species, Bos taurus , with the aurochs ( B. t. primigenius ), zebu ( B. t. indicus ), and taurine ( B. t. taurus ) cattle as subspecies. However, this taxonomy

8228-567: The Pacific Islands. Pigs were introduced to the Melanesian and Polynesian regions by humans from several thousand to 500 years ago, and to the Americas within the past 500 years. In Australia, domesticated pigs escaped in the 18th century, and now cover 40 percent of Australia, with a population estimated at 30 million. While pigs are thought to have been brought to New Zealand by the original Polynesian settlers, this population had become extinct by

8349-452: The US by the early 1930s. Cattle have the largest biomass of any animal species on Earth, at roughly 400 million tonnes, followed closely by Antarctic krill at 379 million tonnes and humans at 373 million tonnes. In 2023, the countries with the most cattle were India with 307.5 million (32.6% of the total), Brazil with 194.4 million, and China with 101.5 million, out of a total of 942.6 million in

8470-455: The US, excessive numbers of feral cats are often shot. The " trap-neuter-return " method has been used in many locations as an alternative means of managing the feral cat population. The goat is one of the oldest domesticated creatures, yet readily returns to a feral state. Sheep are close contemporaries and cohorts of goats in the history of domestication, but the domestic sheep is vulnerable to predation and injury, and thus rarely seen in

8591-457: The age, sex, dominance status and reproductive status of the caller, and may indicate estrus in cows and competitive display in bulls. Cows can categorize images as familiar and unfamiliar individuals. Cloned calves from the same donor form subgroups, suggesting that kin discrimination may be a basis of grouping behaviour. Cattle use visual/brain lateralisation when scanning novel and familiar stimuli. They prefer to view novel stimuli with

8712-676: The available food; foraging velocity decreases and intake rate increases in areas of abundant palatable forage. Cattle avoid grazing areas contaminated by the faeces of other cattle more strongly than they avoid areas contaminated by sheep, but they do not avoid pasture contaminated by rabbits. In cattle, temperament or behavioral disposition can affect productivity, overall health, and reproduction. Five underlying categories of temperament traits have been proposed: shyness–boldness, exploration–avoidance, activity, aggressiveness , and sociability. There are many indicators of emotion in cattle. Holstein–Friesian heifers that had made clear improvements in

8833-547: The biodiversity of a human-altered area by being able to survive in it in ways local species cannot. Feral zebu have been reintroduced in Kuno Wildlife Sanctuary in order to replace their ancestor, the aurochs . Animals of domestic origin sometimes can produce fertile hybrids with native, wild animals which leads to genetic pollution (not a clear term itself) in the naturally evolved wild gene pools, many times threatening rare species with extinction . Cases include

8954-474: The bulls weigh 1,000–1,200 kg (2,200–2,600 lb). Before 1790, beef cattle averaged only 160 kg (350 lb) net. Thereafter, weights climbed steadily. Cattle breeds vary widely in size; the tallest and heaviest is the Chianina , where a mature bull may be up to 1.8 m (5 ft 11 in) at the shoulder, and may reach 1,280 kg (2,820 lb) in weight. The natural life of domestic cattle

9075-481: The climate warms, increasing the risk of tick-borne diseases. Both beef and milk production are likely to experience declines due to climate change. Cattle health is at once a veterinary issue (for animal welfare and productivity), a public health issue (to limit the spread of disease), and a food safety issue (to ensure meat and dairy products are safe to eat). These concerns are reflected in farming regulations. These rules can become political matters, as when it

9196-500: The cows' milk for human consumption. Animal welfare advocates are critical of this practice , stating that this breaks the natural bond between the mother and her calf. The welfare of veal calves is also a concern. Two sports involving cattle are thought to be cruel by animal welfare groups: rodeos and bullfighting . Such groups oppose rodeo activities including bull riding , calf roping and steer roping , stating that rodeos are unnecessary and cause stress, injury, and death to

9317-456: The dark sea floor consume organic matter produced through chemosynthesis (via oxidizing inorganic compounds such as hydrogen sulfide ) by archaea and bacteria . Animals evolved in the sea. Lineages of arthropods colonised land around the same time as land plants , probably between 510 and 471 million years ago during the Late Cambrian or Early Ordovician . Vertebrates such as

9438-450: The earliest times, and are frequently featured in mythology , religion , arts , literature , heraldry , politics , and sports . The word animal comes from the Latin noun animal of the same meaning, which is itself derived from Latin animalis 'having breath or soul'. The biological definition includes all members of the kingdom Animalia. In colloquial usage, the term animal

9559-538: The end of the last ice age . In both Australia and the Americas, modern "wild" horses descended from domesticated horses brought by European explorers and settlers that escaped, spread, and thrived. Australia hosts a feral donkey population , as do the Virgin Islands and the American southwest. The pig has established feral populations worldwide, including in Australia, New Zealand, the United States, New Guinea and

9680-426: The export for their meat trade. At certain times, animals were sometimes deliberately left to go feral, typically on islands, in order to be later recovered for profit or food use for travellers (particularly sailors) at the end of a few years. Populations of feral animals present good sources for studies of population dynamics, and especially of ecology and behavior (ethology) in a wide state of species known mainly in

9801-413: The feral category animals that were genuinely wild before they escaped from captivity: neither lions escaped from a zoo nor the white-tailed eagles re-introduced to the UK are regarded as feral. Domesticated plants that revert to wild are referred to as escaped, introduced, naturalized , or sometimes as feral crops. Individual plants are known as volunteers. Large numbers of escaped plants may become

9922-565: The first hierarchical biological classification for animals in 1758 with his Systema Naturae , which Jean-Baptiste Lamarck expanded into 14 phyla by 1809. In 1874, Ernst Haeckel divided the animal kingdom into the multicellular Metazoa (now synonymous with Animalia) and the Protozoa , single-celled organisms no longer considered animals. In modern times, the biological classification of animals relies on advanced techniques, such as molecular phylogenetics , which are effective at demonstrating

10043-448: The formation of complex structures possible. This may be calcified, forming structures such as shells , bones , and spicules . In contrast, the cells of other multicellular organisms (primarily algae, plants, and fungi ) are held in place by cell walls, and so develop by progressive growth. Animal cells uniquely possess the cell junctions called tight junctions , gap junctions , and desmosomes . With few exceptions—in particular,

10164-520: The fossil record, rather than showing that all these animals appeared simultaneously. That view is supported by the discovery of Auroralumina attenboroughii , the earliest known Ediacaran crown-group cnidarian (557–562 mya, some 20 million years before the Cambrian explosion) from Charnwood Forest , England. It is thought to be one of the earliest predators , catching small prey with its nematocysts as modern cnidarians do. Some palaeontologists have suggested that animals appeared much earlier than

10285-669: The giant single-celled protist Gromia sphaerica , so the Tonian trace fossils may not indicate early animal evolution. Around the same time, the layered mats of microorganisms called stromatolites decreased in diversity, perhaps due to grazing by newly evolved animals. Objects such as sediment-filled tubes that resemble trace fossils of the burrows of wormlike animals have been found in 1.2 gya rocks in North America, in 1.5 gya rocks in Australia and North America, and in 1.7 gya rocks in Australia. Their interpretation as having an animal origin

10406-489: The grazed plants and of animals at different trophic levels in the ecosystem . A well documented consequence of overgrazing is woody plant encroachment in rangelands, which significantly reduces the carrying capacity of the land over time. Cattle husbandry practices including branding , castration , dehorning , ear tagging , nose ringing , restraint, tail docking , the use of veal crates, and cattle prods have raised welfare concerns. Stocking density

10527-436: The horses in their own interest. Animal Animals are multicellular , eukaryotic organisms in the biological kingdom Animalia ( / ˌ æ n ɪ ˈ m eɪ l i ə / ). With few exceptions, animals consume organic material , breathe oxygen , have myocytes and are able to move , can reproduce sexually , and grow from a hollow sphere of cells, the blastula , during embryonic development . Animals form

10648-685: The land). The word is a variant of chattel (a unit of personal property) and closely related to capital in the economic sense. The word cow came via Anglo-Saxon cū (plural cȳ ), from Common Indo-European gʷōus ( genitive gʷowés ) 'a bovine animal', cf. Persian : gâv , Sanskrit : go- . In older English sources such as the King James Version of the Bible, cattle often means livestock, as opposed to deer , which are wild. Cattle are large artiodactyls , mammals with cloven hooves , meaning that they walk on two toes,

10769-441: The larvae feed on the hosts' living tissues, killing them in the process, but the adults primarily consume nectar from flowers. Other animals may have very specific feeding behaviours , such as hawksbill sea turtles which mainly eat sponges . Most animals rely on biomass and bioenergy produced by plants and phytoplanktons (collectively called producers ) through photosynthesis . Herbivores, as primary consumers , eat

10890-402: The left eye (using the right brain hemisphere), but the right eye for familiar stimuli. Individual cattle have also been observed to display different personality traits, such as fearfulness and sociability. Vision is the dominant sense; cattle obtain almost half of their information visually. Being prey animals, cattle evolved to look out for predators almost all around, with eyes that are on

11011-440: The many bacteria that contribute are Fibrobacter succinogenes , Ruminococcus flavefaciens , and Ruminococcus albus . Cellulolytic fungi include several species of Neocallimastix , while the protozoa include the ciliates Eudiplodinium maggie and Ostracodinium album . If the animal's feed changes over time, the composition of this microbiome changes in response. Cattle have one large stomach with four compartments;

11132-532: The modified traits in other plants as a result of cross-breeding is known as "adventitious presence (AP)". Certain familiar animals go feral easily and successfully, while others are much less inclined to wander and usually fail promptly outside domestication. Some species will detach readily from humans and pursue their own devices, but do not stray far or spread readily. Others depart and are gone, seeking out new territory or range to exploit and displaying active invasiveness. Whether they leave readily and venture far,

11253-748: The most extreme cold deserts of continental Antarctica . The blue whale ( Balaenoptera musculus ) is the largest animal that has ever lived, weighing up to 190 tonnes and measuring up to 33.6 metres (110 ft) long. The largest extant terrestrial animal is the African bush elephant ( Loxodonta africana ), weighing up to 12.25 tonnes and measuring up to 10.67 metres (35.0 ft) long. The largest terrestrial animals that ever lived were titanosaur sauropod dinosaurs such as Argentinosaurus , which may have weighed as much as 73 tonnes, and Supersaurus which may have reached 39 meters. Several animals are microscopic; some Myxozoa ( obligate parasites within

11374-647: The number of agonistic encounters increases with group size. Dominance relationships in semi-wild highland cattle are very firm, with few overt aggressive conflicts: most disputes are settled by agonistic (non-aggressive, competitive) behaviors with no physical contact between opponents, reducing the risk of injury. Dominance status depends on age and sex, with older animals usually dominant to young ones and males dominant to females. Young bulls gain superior dominance status over adult cows when they reach about 2 years of age. Cattle eat mixed diets, but prefer to eat approximately 70% clover and 30% grass. This preference has

11495-620: The oldest animal phylum and forming a sister clade to all other animals. Despite their morphological dissimilarity with all other animals, genetic evidence suggests sponges may be more closely related to other animals than the comb jellies are. Sponges lack the complex organization found in most other animal phyla; their cells are differentiated, but in most cases not organised into distinct tissues, unlike all other animals. They typically feed by drawing in water through pores, filtering out small particles of food. The comb jellies and Cnidaria are radially symmetric and have digestive chambers with

11616-525: The origins of animals to unicellular ancestors, providing the external phylogeny shown in the cladogram. Uncertainty of relationships is indicated with dashed lines. Holomycota (inc. fungi) [REDACTED] Ichthyosporea [REDACTED] Pluriformea [REDACTED] Filasterea [REDACTED] [REDACTED] [REDACTED] The most basal animals, the Porifera , Ctenophora , Cnidaria , and Placozoa , have body plans that lack bilateral symmetry . Their relationships are still disputed;

11737-519: The plant material directly to digest and absorb the nutrients, while carnivores and other animals on higher trophic levels indirectly acquire the nutrients by eating the herbivores or other animals that have eaten the herbivores. Animals oxidize carbohydrates , lipids , proteins and other biomolecules, which allows the animal to grow and to sustain basal metabolism and fuel other biological processes such as locomotion . Some benthic animals living close to hydrothermal vents and cold seeps on

11858-438: The price of increasing pH , respiratory alkalosis . To deal with this, cattle are forced to shed bicarbonate through urination , at the expense of rumen buffering. These two pathologies can both cause lameness . Another specific risk is mastitis . This worsens as Calliphora blowflies increase in number with continued warming, spreading mastitis-causing bacteria. Ticks too are likely to increase in temperate zones as

11979-437: The process of chewing the cud , like most ruminants. While feeding, cows swallow their food without chewing; it goes into the rumen for storage. Later, the food is regurgitated to the mouth, a mouthful at a time, where the cud is chewed by the molars, grinding down the coarse vegetation to small particles. The cud is then swallowed again and further digested by the micro-organisms in the cow's stomach. The gestation period for

12100-483: The process. Copulation lasts several seconds and consists of a single pelvic thrust . Cows seek secluded areas for calving. Semi-wild Highland cattle heifers first give birth at 2 or 3 years of age, and the timing of birth is synchronized with increases in natural food quality. Average calving interval is 391 days, and calving mortality within the first year of life is 5%. Beef calves suckle an average of 5 times per day, spending some 46 minutes suckling. There

12221-472: The result of crossing taurines domesticated elsewhere with local aurochs, but they are genetically distinct; some authors name them as a separate subspecies, Bos taurus africanus . The only pure African taurine breeds remaining are the N'Dama , Kuri and some varieties of the West African Shorthorn. Feral cattle are those that have been allowed to go wild. Populations exist in many parts of

12342-495: The sides of their head rather than the front. This gives them a field of view of 330°, but limits binocular vision (and therefore stereopsis ) to some 30° to 50°, compared to 140° in humans. They are dichromatic , like most mammals. Cattle avoid bitter-tasting foods, selecting sweet foods for energy. Their sensitivity to sour-tasting foods helps them to maintain optimal ruminal pH . They seek out salty foods by taste and smell to maintain their electrolyte balance. Their hearing

12463-731: The sister group to all other animals could be the Porifera or the Ctenophora, both of which lack hox genes , which are important for body plan development . Hox genes are found in the Placozoa, Cnidaria, and Bilateria. 6,331 groups of genes common to all living animals have been identified; these may have arisen from a single common ancestor that lived 650 million years ago in the Precambrian . 25 of these are novel core gene groups, found only in animals; of those, 8 are for essential components of

12584-407: The smaller, motile gametes are spermatozoa and the larger, non-motile gametes are ova . These fuse to form zygotes , which develop via mitosis into a hollow sphere, called a blastula. In sponges, blastula larvae swim to a new location, attach to the seabed, and develop into a new sponge. In most other groups, the blastula undergoes more complicated rearrangement. It first invaginates to form

12705-414: The southwestern United States and northern Mexico . Such cattle are variously called mavericks , scrubbers or cleanskins. Most free roaming cattle, however untamed, are generally too valuable not to be eventually rounded up and recovered in closely settled regions. Horses and donkeys , domesticated about 5000 BCE, are feral in open grasslands worldwide. In Australia , they are known as Brumbies ; in

12826-474: The sponges and placozoans —animal bodies are differentiated into tissues . These include muscles , which enable locomotion, and nerve tissues , which transmit signals and coordinate the body. Typically, there is also an internal digestive chamber with either one opening (in Ctenophora, Cnidaria, and flatworms) or two openings (in most bilaterians). Nearly all animals make use of some form of sexual reproduction. They produce haploid gametes by meiosis ;

12947-411: The structure at the base of the tree (dashed lines). Porifera [REDACTED] Ctenophora [REDACTED] Placozoa [REDACTED] Cnidaria [REDACTED] Xenacoelomorpha [REDACTED] Ambulacraria [REDACTED] Chordata [REDACTED] Ecdysozoa [REDACTED] Spiralia [REDACTED] An alternative phylogeny, from Kapli and colleagues (2021), proposes

13068-583: The subfamily Bovinae and the most widespread species of the genus Bos . Mature female cattle are called cows and mature male cattle are bulls . Young female cattle are called heifers , young male cattle are oxen or bullocks , and castrated male cattle are known as steers . Cattle are commonly raised for meat , for dairy products , and for leather . As draft animals , they pull carts and farm implements . In India , cattle are sacred animals within Hinduism, and may not be killed. Small breeds such as

13189-525: The subject of the documentary film, The Wild Parrots of Telegraph Hill ) being particularly successful outside of their native habitats and adapting well to suburban environments. Wild cocks are derived from domestic chickens ( Gallus gallus domesticus ) who have returned to the wild. Like the red junglefowl (the closest wild relative of domestic chickens), wild cocks will take flight and roost in tall trees and bushes in order to avoid predators at night. Wild cocks typically form social groups composed of,

13310-511: The subsequent Ediacaran period in what is known as the Avalon explosion . Earlier evidence of animals is still controversial; the sponge -like organism Otavia has been dated back to the Tonian period at the start of the Neoproterozoic , but its identity as an animal is heavily contested. Nearly all modern animal phyla became clearly established in the fossil record as marine species during

13431-416: The third and fourth digits. Like all bovid species, they can have horns, which are unbranched and are not shed annually. Coloration varies with breed; common colors are black, white, and red/brown, and some breeds are spotted or have mixed colors. Bulls are larger than cows of the same breed by up to a few hundred kilograms. British Hereford cows, for example, weigh 600–800 kg (1,300–1,800 lb), while

13552-525: The time of European colonization, and all feral pigs in New Zealand today are descendants of European stock. Many European wild boar populations are also partially descended from escaped domestic pigs and are thus feral animals within the native range of the ancestral species. Rock doves were formerly kept for their meat or more commonly as racing animals and have established feral populations in cities worldwide. Colonies of honey bees often escape into

13673-419: The tongue; in one study observing 750-kilogram (1,650 lb) steers, bite area reached a maximum of approximately 170 cm (30 sq in). Bite depth increases with the height of the plants. By adjusting their behavior, cattle obtain heavier bites in swards that are tall and sparse compared with short, dense swards of equal mass/area. Cattle adjust other aspects of their grazing behavior in relation to

13794-467: The total number of nematode species include 10,000–20,000; 500,000; 10 million; and 100 million. Using patterns within the taxonomic hierarchy, the total number of animal species—including those not yet described—was calculated to be about 7.77 million in 2011. 3,000–6,500 4,000–25,000 Evidence of animals is found as long ago as the Cryogenian period. 24-Isopropylcholestane (24-ipc) has been found in rocks from roughly 650 million years ago; it

13915-538: The ultimate criterion for success is longevity . Persistence depends on their ability to establish themselves and reproduce reliably in the new environment . Neither the duration nor the intensity with which a species has been domesticated offers a useful correlation with its feral potential. The cat returns readily to a feral state if it has not been socialized when young. Feral cats, especially if left to proliferate, are frequently considered to be pests in both rural and urban areas, and may be blamed for devastating

14036-470: The wild from managed apiaries when they swarm ; their behavior, however, is no different from their behavior in captivity, unless they breed with other feral honey bees of a different genetic stock, which may lead them to become more docile or more aggressive (see Africanized bees ). Large colonies of feral parrots are present in various parts of the world, with rose-ringed parakeets , monk parakeets and red-masked parakeets (the latter of which became

14157-721: The world, sometimes on small islands. Some, such as Amsterdam Island cattle , Chillingham cattle , and Aleutian wild cattle have become sufficiently distinct to be described as breeds. Cattle are often raised by allowing herds to graze on the grasses of large tracts of rangeland . Raising cattle extensively in this manner allows the use of land that might be unsuitable for growing crops. The most common interactions with cattle involve daily feeding , cleaning and milking . Many routine husbandry practices involve ear tagging , dehorning , loading, medical operations , artificial insemination , vaccinations and hoof care, as well as training for agricultural shows and preparations. Around

14278-565: The world, Fulani husbandry rests on behavioural techniques, whereas in Europe, cattle are controlled primarily by physical means, such as fences . Breeders use cattle husbandry to reduce tuberculosis susceptibility by selective breeding and maintaining herd health to avoid concurrent disease. In the United States, many cattle are raised intensively, kept in concentrated animal feeding operations , meaning there are at least 700 mature dairy cows or at least 1000 other cattle stabled or confined in

14399-406: The world. Cattle are kept on farms to produce meat, milk, and leather, and sometimes to pull carts or farm implements. The meat of adult cattle is known as beef , and that of calves as veal . Other body parts are used as food products, including blood, liver , kidney , heart and oxtail . Approximately 300 million cattle, including dairy animals, are slaughtered each year for food. About

14520-500: Was proposed in the UK in 2011 that milk from tuberculosis -infected cattle should be allowed to enter the food chain. Cattle disease attracted attention in the 1980s and 1990s when bovine spongiform encephalopathy (mad cow disease) broke out in the United Kingdom . BSE can cross into humans as the deadly variant Creutzfeldt–Jakob disease ; 178 people in the UK had died from it by 2010. The gut flora of cattle produce methane ,

14641-415: Was the world's largest producer of cattle hides. Cattle hides account for around 65% of the world's leather production. Cattle are subject to pests including arthropod parasites such as ticks (which can in turn transmit diseases caused by bacteria and protozoa), and diseases caused by pathogens including bacteria and viruses . Some viral diseases are spread by insects - i.e. bluetongue disease

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