79-472: Eusauropoda (meaning "True Lizard Foot") is a derived clade of sauropod dinosaurs . Eusauropods represent the node-based group that includes all descendant sauropods starting with the basal eusauropods of Shunosaurus , and possibly Barapasaurus , and Amygdalodon , but excluding Vulcanodon and Rhoetosaurus . The Eusauropoda was coined in 1995 by Paul Upchurch to create a monophyletic new taxonomic group that would include all sauropods, except for
158-525: A clade (from Ancient Greek κλάδος (kládos) 'branch'), also known as a monophyletic group or natural group , is a grouping of organisms that are monophyletic – that is, composed of a common ancestor and all its lineal descendants – on a phylogenetic tree . In the taxonomical literature, sometimes the Latin form cladus (plural cladi ) is used rather than the English form. Clades are
237-479: A "ladder", with supposedly more "advanced" organisms at the top. Taxonomists have increasingly worked to make the taxonomic system reflect evolution. When it comes to naming , this principle is not always compatible with the traditional rank-based nomenclature (in which only taxa associated with a rank can be named) because not enough ranks exist to name a long series of nested clades. For these and other reasons, phylogenetic nomenclature has been developed; it
316-623: A clade can be described based on two different reference points, crown age and stem age. The crown age of a clade refers to the age of the most recent common ancestor of all of the species in the clade. The stem age of a clade refers to the time that the ancestral lineage of the clade diverged from its sister clade. A clade's stem age is either the same as or older than its crown age. Ages of clades cannot be directly observed. They are inferred, either from stratigraphy of fossils , or from molecular clock estimates. Viruses , and particularly RNA viruses form clades. These are useful in tracking
395-545: A distally elongated anterior ramus of the quadratojugal. Separating the anteroventral process of the nasal from the posterolateral process of the premaxilla, eusauropods also have a long maxilla that forms the posteroventral margin of the external naris. Eusauporoda are also hypothesized to have a semi-digitigrade foot posturem demonstrated by footprint evidence. Paleontologist Jeffrey Wilson explains that eusauropods differ from theropods and prosauropods that have digitigrade pes where their heel and metatarsals are lifted off
474-669: A fleet to reach Peru by sea, and sought the military support of various rebels from the Viceroyalty of Peru. In 1822 the two men met at the Guayaquil Conference in Ecuador, where they failed to agree on governance strategies for the liberated nations. Two years later Bolívar's forces beat the Spanish at the Battle of Ayacucho , securing the independence of Peru and the rest of South America. In
553-454: A long history of colonialism , the overwhelming majority of South Americans speak Spanish or Portuguese , and societies and states are rich in Western traditions . Relative to Africa, Asia, and Europe, post-1900 South America has been a peaceful continent with few wars, although high rates of violent crime remain a concern in some countries. South America occupies the southern portion of
632-653: A part of Oceania, also administered by Chile), the Galápagos Islands (administered by Ecuador , sometimes considered part of Oceania), and Tierra del Fuego (split between Argentina and Chile). In the Atlantic Ocean, Brazil administers Fernando de Noronha , Trindade and Martim Vaz , and the Saint Peter and Saint Paul Archipelago , while the Falkland Islands (Spanish: Islas Malvinas ) and South Georgia and
711-403: A retracted position of the external nares. Unlike prosauropods and theropods , which have a snout with smooth, unprotruding alveolar and subnarial regions, eusauropods have snouts with “stepped anterior margins”. Further distinguishing features of eusauropods include the absence of the contact between the squamosal and the quadratojugal, the absence of the anterior process of the prefrontal, and
790-422: A revised taxonomy based on a concept strongly resembling clades, although the term clade itself would not be coined until 1957 by his grandson, Julian Huxley . German biologist Emil Hans Willi Hennig (1913–1976) is considered to be the founder of cladistics . He proposed a classification system that represented repeated branchings of the family tree, as opposed to the previous systems, which put organisms on
869-481: A site called Chavín de Huantar in modern Peru at an elevation of 3,177 meters (10,423 ft). Chavín civilization spanned 900 BC to 300 BC. In the central coast of Peru, around the beginning of the 1st millennium AD, Moche (100 BC – 700 AD, at the northern coast of Peru), Paracas and Nazca (400 BC – 800 AD, Peru) cultures flourished with centralized states with permanent militia improving agriculture through irrigation and new styles of ceramic art. At
SECTION 10
#1732881335502948-670: A subtropical climate of the dinaric type. At the highest points of the Andean region, climates are colder than the ones occurring at the highest point of the Norwegian fjords. In the Andean plateaus, the warm climate prevails, although it is tempered by the altitude, while in the coastal strip, there is an equatorial climate of the Guinean type. From this point until the north of the Chilean coast appear, successively, Mediterranean oceanic climate , temperate of
1027-429: A suffix added should be e.g. "dracohortian". A clade is by definition monophyletic , meaning that it contains one ancestor which can be an organism, a population, or a species and all its descendants. The ancestor can be known or unknown; any and all members of a clade can be extant or extinct. The science that tries to reconstruct phylogenetic trees and thus discover clades is called phylogenetics or cladistics ,
1106-431: A “bulk-browsing mode of feeding”. They describe the development of lateral plates on the alveolar margins of tooth-bearing bones. These plates can be used to strip foliage, the eusauropod's “U-shaped” jaws create a wide bite, and their loss of “fleshy cheeks” increased the gape. The crowns of eusauropod teeth also have “wrinkled enamel textures”, but it is unclear what this meant for their feeding habits. The skull length of
1185-499: Is also used with a similar meaning in other fields besides biology, such as historical linguistics ; see Cladistics § In disciplines other than biology . The term "clade" was coined in 1957 by the biologist Julian Huxley to refer to the result of cladogenesis , the evolutionary splitting of a parent species into two distinct species, a concept Huxley borrowed from Bernhard Rensch . Many commonly named groups – rodents and insects , for example – are clades because, in each case,
1264-515: Is because the most extensive part of the continent is in the equatorial zone (the region has more areas of equatorial plains than any other region), therefore giving the Southern Cone more oceanic influence, which moderates year round temperatures. The average annual temperatures in the Amazon basin oscillate around 27 °C (81 °F), with low thermal amplitudes and high rainfall indices. Between
1343-460: Is by far the most populous South American country, with almost half of the continent's population, followed by Colombia, Argentina, Venezuela, and Peru. In recent decades, Brazil has also generated half of the continent's GDP and has become the continent's first regional power. Most of the population lives near the continent's western or eastern coasts while the interior and the far south are sparsely populated. The geography of western South America
1422-597: Is dominated by the Andes mountains; in contrast, the eastern part contains both highland regions and vast lowlands where rivers such as the Amazon , Orinoco and Paraná flow. Most of the continent lies in the tropics, except for a large part of the Southern Cone located in the middle latitudes . The continent's cultural and ethnic outlook has its origin with the interaction of Indigenous peoples with European conquerors and immigrants and, more locally, with African slaves . Given
1501-471: Is in turn included in the mammal, vertebrate and animal clades. The idea of a clade did not exist in pre- Darwinian Linnaean taxonomy , which was based by necessity only on internal or external morphological similarities between organisms. Many of the better known animal groups in Linnaeus's original Systema Naturae (mostly vertebrate groups) do represent clades. The phenomenon of convergent evolution
1580-595: Is now Colombia. They established the Muisca Confederation of many clans, or cacicazgos , that had a free trade network among themselves. Many were goldsmiths and farmers. Other important Pre-Columbian cultures include: the Cañaris (in south central Ecuador), Chimú Empire (1300–1470, Peruvian northern coast), Chachapoyas , and the Aymaran kingdoms (1000–1450, Western Bolivia and southern Peru). Holding their capital at
1659-476: Is one of the most biodiverse continents on Earth. It is home to many unique species of animals including the llama , anaconda , piranha , jaguar , vicuña , and tapir , and to one of the largest known insects in the world, the Titan beetle . The Amazon rainforests possess high biodiversity , with Brazil estimated to contain 10% of Earth's species . 83% of South America's large mammals ( megafauna ) became extinct at
SECTION 20
#17328813355021738-515: Is responsible for many cases of misleading similarities in the morphology of groups that evolved from different lineages. With the increasing realization in the first half of the 19th century that species had changed and split through the ages, classification increasingly came to be seen as branches on the evolutionary tree of life . The publication of Darwin's theory of evolution in 1859 gave this view increasing weight. In 1876 Thomas Henry Huxley , an early advocate of evolutionary theory, proposed
1817-489: Is still controversial. As an example, see the full current classification of Anas platyrhynchos (the mallard duck) with 40 clades from Eukaryota down by following this Wikispecies link and clicking on "Expand". The name of a clade is conventionally a plural, where the singular refers to each member individually. A unique exception is the reptile clade Dracohors , which was made by haplology from Latin "draco" and "cohors", i.e. "the dragon cohort "; its form with
1896-571: Is the largest country in South America, covering a little less than half of the continent's land area and encompassing around half of the continent's population. The remaining countries and territories are divided among four subregions : the Andean states , Caribbean South America , The Guianas , and the Southern Cone . Physiographically, South America also includes some of the nearby islands. The Dutch ABC islands ( Aruba , Bonaire , and Curaçao ),
1975-510: The Altiplano , Tiahuanaco or Tiwanaku (100 BC – 1200 AD, Bolivia) managed a large commercial network based on religion. Around the 7th century, both Tiahuanaco and Wari or Huari Empire (600–1200, Central and northern Peru) expanded its influence to all the Andean region, imposing the Huari urbanism and Tiahuanaco religious iconography. The Muisca were the main indigenous civilization in what
2054-572: The Archipelago of San Andrés, Providencia and Santa Catalina ( San Andrés Island , Providencia Island , and Santa Catalina Island etc., which are administered by Colombia ) are politically parts of South American countries but physiographically parts of North America. Other islands often associated with geopolitical South America are the Chiloé Archipelago and Robinson Crusoe Island (both administered by Chile), Easter Island (culturally
2133-802: The Atacama Desert ; the wettest place on earth, López de Micay in Colombia; the largest rainforest, the Amazon rainforest ; the highest capital city, La Paz , Bolivia ; the highest commercially navigable lake in the world, Lake Titicaca ; and, excluding research stations in Antarctica , the world's southernmost permanently inhabited community, Puerto Toro , Chile. South America's major mineral resources are gold, silver, copper, iron ore, tin , and petroleum. These resources have brought high income to its countries, especially in times of war or of rapid economic growth by industrialized countries elsewhere. However,
2212-686: The Bering Land Bridge (now the Bering Strait ) at least 15,000 years ago from the territory that is present-day Russia. They migrated south through North America, and eventually reached South America through the Isthmus of Panama . Amongst the oldest evidence for human presence in South America is the Monte Verde II site in Chile, suggested to date to around 14,500 years ago. From around 13,000 years ago,
2291-619: The Caribbean colonies and Brazil, as European nations built up economically slave-dependent colonies in the New World . Nearly 40% of all African slaves trafficked to the Americas went to Brazil. An estimated 4.9 million slaves from Africa came to Brazil during the period from 1501 to 1866. In contrast to other European colonies in the Americas which mainly used the labor of African slaves, Spanish colonists mainly enslaved indigenous Americans. In 1750,
2370-1000: The Falkland Islands and South Georgia and the South Sandwich Islands ; and one internal territory : French Guiana . The ABC islands of the Dutch Caribbean (north of Venezuela in the Leeward Antilles ), Ascension Island (a part of Saint Helena, Ascension and Tristan da Cunha ), Bouvet Island (a dependency of Norway ), Panama , and the Caribbean island country of Trinidad and Tobago may also be considered parts of South America. South America has an area of 17,840,000 square kilometers (6,890,000 sq mi). Its population as of 2021 has been estimated at more than 434 million. South America ranks fourth in area (after Asia, Africa, and North America) and fifth in population (after Asia, Africa, Europe, and North America). Brazil
2449-465: The Fishtail projectile point style became widespread across South America, with its disappearance around 11,000 years ago coincident with the disappearance of South America's megafauna. Maize was present in northern South America by around 6,000 years ago. By 2000 BC, many agrarian communities had been settled throughout the Andes and the surrounding regions. Fishing became a widespread practice along
Eusauropoda - Misplaced Pages Continue
2528-467: The Fortaleza area, contemporaneous with the pyramids of Ancient Egypt , one of the oldest-known civilizations in the Americas and one of the six sites where civilization separately originated in the ancient world. Caral–Supe governing class established a trade network and developed agriculture then followed by Chavín by 900 BC, according to some estimates and archaeological finds. Artifacts were found at
2607-688: The Maracaibo Lake and the mouth of the Orinoco, predominates an equatorial climate of the type Congolese, that also includes parts of the Brazilian territory. The east-central Brazilian plateau has a humid and warm tropical climate. The northern and eastern parts of the Argentine pampas have a humid subtropical climate with dry winters and humid summers of the Chinese type, while the western and eastern ranges have
2686-579: The Northern Hemisphere , and titanosaurs being found in Southern Hemisphere . However, basal eusauropods that do not fall into either group are fairly well represented. Early eusauropods such as Volkeimeria and Amygdalodon , and more derived eusauropods such as Patagosaurus have been found in South America . Volkeimeria is classified as a basal eusauropod, though in 2004 Paul Upchurch
2765-707: The Northern Hemisphere . It can also be described as the southern subregion of the Americas . South America is bordered on the west by the Pacific Ocean and on the north and east by the Atlantic Ocean ; North America and the Caribbean Sea lie to the northwest. The continent includes twelve sovereign states : Argentina , Bolivia , Brazil , Chile , Colombia , Ecuador , Guyana , Paraguay , Peru , Suriname , Uruguay , and Venezuela ; two dependent territories :
2844-585: The Orinoco Llanos and in the Guianas Plateau , the precipitation levels go from moderate to high. The Pacific coast of Colombia and northern Ecuador are rainy regions, with Chocó in Colombia being the rainiest place in the world along with the northern slopes of Indian Himalayas. The Atacama Desert, along this stretch of coast, is one of the driest regions in the world. The central and southern parts of Chile are subject to extratropical cyclones , and most of
2923-539: The Portuguese Crown abolished the enslavement of indigenous peoples in colonial Brazil , under the belief that they were unfit for labor and less effective than enslaved Africans. Enslaved Africans were brought to the Americas on slave ships , under inhuman conditions and ill-treatment, and those who survived were sold in slave markets . After independence, all South American countries maintained slavery for some time. The first South American country to abolish slavery
3002-705: The Portuguese Kingdom of Brazil , Dom Pedro I (also Pedro IV of Portugal), son of the Portuguese King Dom João VI , proclaimed the independent Kingdom of Brazil in 1822, which later became the Empire of Brazil . Despite the Portuguese loyalties of garrisons in Bahia , Cisplatina and Pará , independence was diplomatically accepted by Portugal in 1825, on condition of a high compensation paid by Brazil mediated by
3081-641: The South American Plate . South America is home to several superlatives, including the world's highest uninterrupted waterfall , Angel Falls in Venezuela; the highest single-drop waterfall Kaieteur Falls in Guyana; the largest river by volume, the Amazon River ; the longest mountain range, the Andes (whose highest mountain is Aconcagua at 6,962 m or 22,841 ft); the driest non-polar place on earth,
3160-588: The Spanish American wars of independence between the patriots, who promoted such autonomy, and the royalists , who supported Spanish authority over the Americas. The Juntas, in both Spain and the Americas, promoted the ideas of the Enlightenment . Five years after the beginning of the war, Ferdinand VII returned to the throne and began the Absolutist Restoration , as the royalists got the upper hand in
3239-712: The native populations had no immune resistance – caused large-scale depopulation of the native population under Spanish control. Systems of forced labor, such as the haciendas and mining industry's mit'a also contributed to the depopulation. After this, enslaved Africans , who had developed immunities to these diseases, were quickly brought in to replace them. The Spaniards were committed to converting their native subjects to Christianity and were quick to purge any native cultural practices that hindered this end; however, many initial attempts at this were only partially successful, as native groups simply blended Catholicism with their established beliefs and practices. Furthermore,
Eusauropoda - Misplaced Pages Continue
3318-732: The Americas. The continent is generally delimited on the northwest by the Darién watershed along the Colombia–Panama border , although some may consider the border instead to be the Panama Canal . Geopolitically and geographically, all of Panama – including the segment east of the Panama Canal in the isthmus – is typically included in North America alone and among the countries of Central America . Almost all of mainland South America sits on
3397-666: The Argentine Patagonia is desert . In the Pampas of Argentina, Uruguay and South of Brazil the rainfall is moderate, with rains well distributed during the year. The moderately dry conditions of the Chaco oppose the intense rainfall of the eastern region of Paraguay. In the semiarid coast of the Brazilian Northeast the rains are linked to a monsoon regime. Important factors in the determination of climates are sea currents, such as
3476-726: The Breton type and, already in Tierra del Fuego , cold climate of the Siberian type. The distribution of rainfall is related to the regime of winds and air masses. In most of the tropical region east of the Andes, winds blowing from the northeast, east and southeast carry moisture from the Atlantic, causing abundant rainfall. However, due to a consistently strong wind shear and a weak Intertropical Convergence Zone , South Atlantic tropical cyclones are rare. In
3555-1171: The Canadon Asfalto and Canadon Calcereo formations reveal a more diverse and widespread peleobiology of eusauropods in the Late Jurassic period. The following cladogram demonstrates hypothesized relationships within the Eusauropoda. The basal eusauropods include the Turiasauria ( Turiasaurus , Zby and others). Shunosaurus Barapasaurus Patagosaurus Omeisaurus Mamenchisaurus Cetiosaurus Jobaria Haplocanthosaurus Limaysaurus Nigersaurus Amargasaurus Dicraeosaurus Apatosaurus Brontosaurus Barosaurus Diplodocus Camarasaurus Brachiosaurus Phuwiangosaurus Malawisaurus Rapetosaurus Isisaurus Opisthocoelicaudia Saltasaurus [REDACTED] [REDACTED] [REDACTED] [REDACTED] [REDACTED] [REDACTED] [REDACTED] Clade In biological phylogenetics ,
3634-495: The South Sandwich Islands ( biogeographically and hydrologically associated with Antarctica) have been administered as two British Overseas Territories under the Crown , whose sovereignty over the islands is disputed by Argentina . An isolated volcanic island on the South American Plate , Ascension Island is geologically a part of South America. Administered as a dependency of Saint Helena, Ascension and Tristan da Cunha ,
3713-995: The Spaniards brought their language to the degree they did with their religion, although the Roman Catholic Church 's evangelization in Quechua , Aymara , and Guaraní actually contributed to the continuous use of these native languages albeit only in the oral form. Eventually, the natives and the Spaniards interbred, forming a mestizo class. At the beginning, many mestizos of the Andean region were offspring of Amerindian mothers and Spanish fathers. After independence, most mestizos had native fathers and European or mestizo mothers. Many native artworks were considered pagan idols and destroyed by Spanish explorers; this included many gold and silver sculptures and other artifacts found in South America, which were melted down before their transport to Spain or Portugal. Spaniards and Portuguese brought
3792-530: The Spanish and Portuguese colonies. First, Napoleon invaded Portugal, but the House of Braganza avoided capture by escaping to Brazil . Napoleon captured King Ferdinand VII of Spain, and appointed his own brother instead. This appointment provoked popular resistance, which created Juntas to rule in the name of the captured king. Many cities in the Spanish colonies, however, considered themselves equally authorized to appoint local Juntas like those of Spain. This began
3871-588: The United Kingdom. The newly independent nations began a process of fragmentation, with several civil and international wars. However, it was not as strong as in Central America. Some countries created from provinces of larger countries stayed as such up to modern times (such as Paraguay or Uruguay), while others were reconquered and reincorporated into their former countries (such as the Republic of Entre Ríos and
3950-430: The basal eusauropod, Patagosaurus is about 60 centimetres (24 in). One of the most basal eusauropods, Shunosaurus , has two characteristic features of the eusauropod elongated neck: the incorporation of the equivalent of the first dorsal vertebra into the cervical region of the spine, and the addition of two cervical vertebra in the middle of the cervical vertebrae. Other synapomorphies of Eusauropoda includes
4029-437: The coast, helping establish fish as a primary source of food. Irrigation systems were also developed at this time, which aided in the rise of an agrarian society . South American cultures began domesticating llamas , vicuñas , guanacos , and alpacas in the highlands of the Andes circa 3500 BC. Besides their use as sources of meat and wool, these animals were used for transportation of goods. The rise of plant growing and
SECTION 50
#17328813355024108-461: The concentration in producing one, or few, major export commodities has often hindered the development and diversification of its economies. The fluctuation in the price of commodities in international markets has led historically to major highs and lows, booms and busts, in the economies of South American states, often causing political instability. This has led for calls to diversify production and increase trade within South America itself. Brazil
4187-424: The conflict. The independence of South America was secured by Simón Bolívar (Venezuela) and José de San Martín (Argentina), the two most important Libertadores . Bolívar led a great uprising in the north, then led his army south towards Lima , the capital of the Viceroyalty of Peru . Meanwhile, San Martín led an army across the Andes Mountains, along with Chilean expatriates, and liberated Chile. He organized
4266-405: The current Humboldt and Falklands . The equatorial current of the South Atlantic strikes the coast of the Northeast and there is divided into two others: the current of Brazil and a coastal current that flows to the northwest towards the Antilles , where there it moves towards northeast course thus forming the most Important and famous ocean current in the world, the Gulf Stream . South America
4345-507: The distribution of some of these shared derived traits that distinguish Eusauropoda is still completely clear. However the discovery of the vulcanodont, Tazoudasaurus , which does preserve cranial material did help refine some of the characteristics. Eusauropods are long-necked, strictly herbivorous , obligate quadrupedals . They have a highly specialized set of skeletal adaptions due to their large size, and are graviportal . Yates and Upchurch described eusauropod evolution as moving towards
4424-427: The end of the Pleistocene around 11,000 years ago as part of the Quaternary extinction event , among the highest of any continent, with the casualties including saber-toothed cats , ground sloths , glyptodonts , gomphotheres , the equines Hippidion and Equus neogeus , and all remaining South American native ungulates . South America is thought to have been first inhabited by humans when people were crossing
4503-451: The fundamental unit of cladistics , a modern approach to taxonomy adopted by most biological fields. The common ancestor may be an individual, a population , or a species ( extinct or extant ). Clades are nested, one in another, as each branch in turn splits into smaller branches. These splits reflect evolutionary history as populations diverged and evolved independently. Clades are termed monophyletic (Greek: "one clan") groups. Over
4582-400: The genus Shunosaurus is likely one of the most basal eusauropods. The genus Barapasaurus has been found in India, and may represent a cetiosaurid, a basal eusauropod, or a genus outside Eusauropoda. The data around sauropods evolution, as Novas points out, is largely based on a few formations mostly in the Northern Hemisphere. However, other beds in places such as Tanzania, specifically
4661-502: The great city of Cusco , the Inca civilization dominated the Andes region from 1438 to 1533. Known as Tawantin suyu , and "the land of the four regions", in Quechua , the Inca Empire was highly distinct and developed. Inca rule extended to nearly a hundred linguistic or ethnic communities, some nine to fourteen million people connected by a 25,000 kilometer road system . Cities were built with precise, unmatched stonework, constructed over many levels of mountain terrain. Terrace farming
4740-626: The ground. Eusauropods show asymmetry in their metatarsal shaft diameters where metatarsal I is broader than the others, suggesting that their weight was mostly assumed by their inner feet. According to Steven Salisbury and Jay Nair, basal eusauropods retain four pedal unguals but reduce their phalangeal number in their fourth digit to three units. The metatarsus in eusauropods is less than a quarter of their tibial length, unlike sauropod outgroups that have long hindlimbs and metatarsus that are almost half of their tibial length. Eusauropods are found on all major continents, with diplocodoids being widespread in
4819-546: The group consists of a common ancestor with all its descendant branches. Rodents, for example, are a branch of mammals that split off after the end of the period when the clade Dinosauria stopped being the dominant terrestrial vertebrates 66 million years ago. The original population and all its descendants are a clade. The rodent clade corresponds to the order Rodentia, and insects to the class Insecta. These clades include smaller clades, such as chipmunk or ant , each of which consists of even smaller clades. The clade "rodent"
SECTION 60
#17328813355024898-417: The island is geopolitically a part of Africa. All of the world's major climate zones are present in South America. The distribution of the average temperatures in the region presents a constant regularity from the 30° of latitude south , when the isotherms tend, more and more, to be confused with the degrees of latitude. In temperate latitudes, winters and summers are milder than in North America. This
4977-487: The islands of Trinidad and Tobago ( Trinidad Island and Tobago Island etc.), the State of Nueva Esparta , and the Federal Dependencies of Venezuela sit on the northern portion of the South American continental shelf and are sometimes considered parts of the continent. Geopolitically, all the island countries and territories in the Caribbean have generally been grouped as a subregion of North America instead. By contrast, Aves Island (administered by Venezuela ) and
5056-466: The land outside Europe should be an exclusive duopoly between the two countries. The treaty established an imaginary line along a north–south meridian 370 leagues west of the Cape Verde Islands , roughly 46° 37' W. In terms of the treaty, all land to the west of the line (known to comprise most of the South American soil) would belong to Spain, and all land to the east, to Portugal. As accurate measurements of longitude were impossible at that time,
5135-590: The last few decades, the cladistic approach has revolutionized biological classification and revealed surprising evolutionary relationships among organisms. Increasingly, taxonomists try to avoid naming taxa that are not clades; that is, taxa that are not monophyletic . Some of the relationships between organisms that the molecular biology arm of cladistics has revealed include that fungi are closer relatives to animals than they are to plants, archaea are now considered different from bacteria , and multicellular organisms may have evolved from archaea. The term "clade"
5214-518: The latter term coined by Ernst Mayr (1965), derived from "clade". The results of phylogenetic/cladistic analyses are tree-shaped diagrams called cladograms ; they, and all their branches, are phylogenetic hypotheses. Three methods of defining clades are featured in phylogenetic nomenclature : node-, stem-, and apomorphy-based (see Phylogenetic nomenclature§Phylogenetic definitions of clade names for detailed definitions). The relationship between clades can be described in several ways: The age of
5293-476: The line was not strictly enforced, resulting in a Portuguese expansion of Brazil across the meridian. Beginning in the 1530s, the people and natural resources of South America were repeatedly exploited by foreign conquistadors , first from Spain and later from Portugal. These competing colonial nations claimed the land and resources as their own and divided it into colonies. European infectious diseases ( smallpox , influenza , measles , and typhus ) – to which
5372-402: The mid-17th century. It became a Dutch colony in 1667. The indigenous peoples of the Americas in various European colonies were forced to work in European plantations and mines; along with enslaved Africans who were also introduced in the proceeding centuries via the slave trade . European colonists were heavily dependent on indigenous labor during the initial phases of settlement to maintain
5451-475: The possibly eusauropod genera Cardiodon and Oplosaurus , known only from teeth. The family Mamenchisauridae is found widespread throughout Asia. A majority of the genera are found in China, although a possible specimen of Mamenchisaurus has been found in Thailand . Also in China, the basal eusauropod Nebulasaurus taito was found to be a sister taxon to Spinophorosaurus , and more derived than Mamenchisauridae, but less derived than Patagosaurus , and
5530-430: The spread of viral infections . HIV , for example, has clades called subtypes, which vary in geographical prevalence. HIV subtype (clade) B, for example is predominant in Europe, the Americas and Japan, whereas subtype A is more common in east Africa. South America South America is a continent entirely in the Western Hemisphere and mostly in the Southern Hemisphere , with a considerably smaller portion in
5609-421: The subsequent appearance of permanent human settlements allowed for the multiple and overlapping beginnings of civilizations in South America. One of the earliest known South American civilizations was Caral–Supe , on the central Peruvian coast. Though a pre-ceramic culture, the monumental architecture of Caral–Supe created one of the first cities of the world, generally dated to around 3500 BC, at Huaricanga in
5688-411: The subsistence economy, and natives were often captured by expeditions. The importation of African slaves began midway through the 16th century, but the enslavement of indigenous peoples continued well into the 17th and 18th centuries. The Atlantic slave trade brought enslaved Africans primarily to South American colonies, beginning with the Portuguese since 1502. The main destinations of this phase were
5767-517: The vulcanodontids. Eusauropoda are herbivorous, quadrupedal, and have long necks. They have been found in South America, Europe, North America, Asia, Australia, and Africa. The temporal range of Eusauropoda ranges from the early Jurassic to the Latest Cretaceous periods. The most basal forms of eusauropods are not well known and because the cranial material for the Vulcanodon is not available, and
5846-568: The western European architectural style to the continent, and helped to improve infrastructures like bridges, roads, and the sewer system of the cities they discovered or conquered. They also significantly increased economic and trade relations, not just between the old and new world but between the different South American regions and peoples. Finally, with the expansion of the Portuguese and Spanish languages, many cultures that were previously separated became united through that of Latin American. Guyana
5925-465: Was Chile in 1823, Uruguay in 1830, Bolivia in 1831, Guyana in 1833, Colombia and Ecuador in 1851, Argentina in 1853, Peru and Venezuela in 1854, Suriname in 1863, Paraguay in 1869, and in 1888 Brazil was the last South American nation and the last country in western world to abolish slavery. The European Peninsular War (1807–14), a theater of the Napoleonic Wars , changed the political situation of
6004-665: Was a useful form of agriculture. The Mapuche in Central and Southern Chile resisted the European and Chilean settlers, waging the Arauco War for more than 300 years. In 1494, Portugal and Spain, the two great maritime European powers of that time, on the expectation of new lands being discovered in the west, signed the Treaty of Tordesillas , by which they agreed, with the support of the Pope, that all
6083-705: Was found in Morocco , and Jobaria was found in Niger. However, both genera have been found as possible Macronarians, but Atlasaurus was found to be a turiasaurian, and Jobaria a eusauropod, by a phylogenetic analysis of Xing in 2012. In Europe, the clade Turiasauria has been found in France, Spain, and possibly England, with multiple genera from the same locality in Spain. Cetiosaurus skeletons have also been found in England, along with
6162-499: Was initially colonized by the Dutch before coming under British control , though there was a brief period during the Napoleonic Wars when it was occupied by the French . The region was initially partitioned between the Dutch, French and British before fully coming under the control of Britain. Suriname was first explored by the Spanish in the 16th century and then settled by the English in
6241-400: Was suspicious of its placement, because of its “opisthoceolous cervical centra, the absence of a femoral anterior trochanter, and laterally projecting cnemial crest of the tibia”, and instead thought it may be a generic sauropod. African eusauropods may include Spinophorosaurus , from Niger, although that taxon may instead be closer to Vulcanodon and outside Eusauropoda. Also, Atlasaurus
#501498