158-567: The common starling ( Sturnus vulgaris ), also known as the European starling in North America and simply as the starling in Great Britain and Ireland, is a medium-sized passerine bird in the starling family, Sturnidae . It is about 20 cm (8 in) long and has glossy black plumage with a metallic sheen, which is speckled with white at some times of the year. The legs are pink and
316-502: A tui -sized bird) and several bones of at least one species of saddleback -sized bird have recently been described. These date from the Early to Middle Miocene ( Awamoan to Lillburnian , 19–16 mya). In Europe, perching birds are not too uncommon in the fossil record from the Oligocene onward, belonging to several lineages: That suboscines expanded much beyond their region of origin
474-475: A Bantu language with many Arabic , Persian and other Middle Eastern and South Asian loan words , developed as a lingua franca for trade between the different peoples in southeastern Africa. In the Kalahari Desert of Southern Africa, the distinct people known as Bushmen (also "San", closely related to, but distinct from " Hottentots ") have long been present. The San evince unique physical traits, and are
632-410: A Business June 2008 through May 2009) , PFI (Press Freedom Index 2009) Sub-Saharan Africa contains over 1,500 languages. With the exception of the extinct Sumerian (a language isolate ) of Mesopotamia , Afroasiatic has the oldest documented history of any language family in the world. Egyptian was recorded as early as 3200 BCE. The Semitic branch was recorded as early as 2900 BCE in
790-461: A clearer picture of passerine origins and evolution that reconciles molecular affinities, the constraints of morphology, and the specifics of the fossil record. The first passerines are now thought to have evolved in the Southern Hemisphere in the late Paleocene or early Eocene , around 50 million years ago. The initial diversification of passerines coincides with the separation of
948-417: A colony, in which case several other nests may occupy the same or nearby trees. Males may mate with a second female while the first is still on the nest. The reproductive success of the bird is poorer in the second nest than it is in the primary nest and is better when the male remains monogamous. Breeding takes place during the spring and summer. Following copulation, the female lays eggs on a daily basis over
1106-434: A complex song is also useful in defending a territory and deterring less experienced males from encroaching. Along with having adaptions of the skull and muscles for singing, male starlings also have a much larger syrinx than females. This is due to increased muscle mass and enlarged elements of the syringeal skeleton. The male starling's syrinx is around 35% larger than its female counterpart. However, this sexual dimorphism
1264-406: A deformed bill was heavily infested with Mallophaga lice, presumably due to its inability to remove vermin. The hen flea ( Ceratophyllus gallinae ) is the most common flea in their nests. The small, pale house-sparrow flea C. fringillae , is also occasionally found there and probably arises from the habit of its main host of taking over the nests of other species. This flea does not occur in
1422-471: A distinct super-family Certhioidea . This list is in taxonomic order, placing related families next to one another. The families listed are those recognised by the International Ornithologists' Union (IOC). The order and the division into infraorders, parvorders, and superfamilies follows the phylogenetic analysis published by Carl Oliveros and colleagues in 2019. The relationships between
1580-469: A dozen and other species around five or six. The family Viduidae do not build their own nests, instead, they lay eggs in other birds' nests. The Passeriformes contain several groups of brood parasites such as the viduas , cuckoo-finches , and the cowbirds . The evolutionary history of the passerine families and the relationships among them remained rather mysterious until the late 20th century. In many cases, passerine families were grouped together on
1738-399: A dry winter season and a wet summer season. According to paleontology , early hominid skull anatomy was similar to that of their close cousins, the great African forest apes , gorilla and chimpanzee . However, they had adopted a bipedal locomotion and freed hands, giving them a crucial advantage enabling them to live in both forested areas and on the open savanna at a time when Africa
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#17330863374201896-493: A flock call, threat call, attack call, snarl call and copulation call. The alarm call is a harsh scream, and while foraging together common starlings squabble incessantly. They chatter while roosting and bathing, making a great deal of noise that can cause irritation to people living nearby. When a flock of common starlings is flying together, the synchronised movements of the birds' wings make a distinctive whooshing sound that can be heard hundreds of metres away. The common starling
2054-416: A given time of year. The throat feathers of males are long and loose and are used in display while those of females are smaller and more pointed. The legs are stout and pinkish- or greyish-red. The bill is narrow and conical with a sharp tip; in the winter it is brownish-black but in summer, females have lemon yellow beaks with pink bases while males have yellow bills with blue-grey bases. Moulting occurs once
2212-666: A grouping counterpart to North Africa , which is instead grouped with the definition of MENA (i.e. Middle East and North Africa) as it is part of the Arab world , and most North African states are likewise members of the Arab League . However, while they are also member states of the Arab League , the Comoros , Djibouti , Mauritania , and Somalia (and sometimes Sudan ) are all geographically considered to be part of sub-Saharan Africa. Overall,
2370-450: A habitat only rarely used by the nominate form. Nests are typically made out of straw, dry grass and twigs with an inner lining made up of feathers, wool and soft leaves. Construction usually takes four or five days and may continue through incubation. Common starlings are both monogamous and polygamous ; although broods are generally brought up by one male and one female, occasionally the pair may have an extra helper. Pairs may be part of
2528-508: A large variety of habitats has allowed them to disperse and establish themselves in diverse locations around the world resulting in a habitat range from coastal wetlands to alpine forests, from sea cliffs to mountain ranges 1,900 m (6,200 ft) above sea level. The common starling has been introduced to and has successfully established itself in New Zealand, Australia, South Africa, North America, Fiji and several Caribbean islands. As
2686-931: A marked increase in numbers throughout Europe from the 19th century to around the 1950s and 60s. In about 1830, S. v. vulgaris expanded its range in the British Isles, spreading into Ireland and areas of Scotland where it had formerly been absent, although S. v. zetlandicus was already present in Shetland and the Outer Hebrides. The common starling has bred in northern Sweden from 1850 and in Iceland from 1935. The breeding range spread through southern France to northeastern Spain, and there were other range expansions particularly in Italy, Austria and Finland. It started breeding in Iberia in 1960, while
2844-467: A mate. The scent of plants such as yarrow acts as an olfactory attractant to females. The males sing throughout much of the construction and even more so when a female approaches his nest. Following copulation , the male and female continue to build the nest. Nests may be in any type of hole, common locations include inside hollowed trees, buildings, tree stumps and man-made nest-boxes. S. v. zetlandicus typically breeds in crevices and holes in cliffs,
3002-899: A migration pattern, vacating much of Canada in winter. Birds in the east of the country move southwards, and those from farther west winter in the southwest of the US. Common starlings prefer urban or suburban areas where artificial structures and trees provide adequate nesting and roosting sites. Reedbeds are also favoured for roosting and the birds commonly feed in grassy areas such as farmland, grazing pastures, playing fields, golf courses and airfields where short grass makes foraging easy. They occasionally inhabit open forests and woodlands and are sometimes found in shrubby areas such as Australian heathland . Common starlings rarely inhabit dense, wet forests (i.e. rainforests or wet sclerophyll forests) but are found in coastal areas, where they nest and roost on cliffs and forage amongst seaweed . Their ability to adapt to
3160-481: A more uniform colour. At close range it can be seen that the latter has longer throat feathers, a fact particularly noticeable when it sings. The common starling is noisy, its song consisting of a wide variety of both melodic and mechanical-sounding noises as part of a ritual succession of sounds. The male is the main songster and engages in bouts of song lasting for a minute or more. Each of these typically includes four varieties of song type, which follow each other in
3318-671: A natural or artificial cavity in which four or five glossy, pale blue eggs are laid. These take two weeks to hatch and the young remain in the nest for another three weeks. There are normally one or two breeding attempts each year. This species is omnivorous, taking a wide range of invertebrates , as well as seeds and fruit. It is hunted by various mammals and birds of prey , and is host to a range of external and internal parasites. Large flocks typical of this species can be beneficial to agriculture by controlling invertebrate pests ; however, starlings can also be pests themselves when they feed on fruit and sprouting crops. Common starlings may also be
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#17330863374203476-487: A nuisance through the noise and mess caused by their large urban roosts. Introduced populations in particular have been subjected to a range of controls, including culling , but these have had limited success, except in preventing the colonisation of Western Australia. The species has declined in numbers in parts of northern and western Europe since the 1980s due to fewer grassland invertebrates being available as food for growing chicks. Despite this, its huge global population
3634-526: A period of several days. If an egg is lost during this time, she will lay another to replace it. There are normally four or five eggs that are ovoid in shape and pale blue or occasionally white, and they commonly have a glossy appearance. The colour of the eggs seems to have evolved through the relatively good visibility of blue at low light levels. The egg size is 26.5–34.5 mm (1.04–1.36 in) in length and 20.0–22.5 mm (0.79–0.89 in) in maximum diameter. Incubation lasts thirteen days, although
3792-402: A population of ancestral S. vulgaris that survived in an Iberian refugium during an Ice Age retreat, and mitochondrial gene studies suggest that it could be considered a subspecies of the common starling. There is more genetic variation between common starling populations than between the nominate common starling and the spotless starling. Although common starling remains are known from
3950-432: A regular order without pause. The bout starts with a series of pure-tone whistles and these are followed by the main part of the song, a number of variable sequences that often incorporate snatches of song mimicked from other species of bird and various naturally occurring or man-made noises. The structure and simplicity of the sound mimicked is of greater importance than the frequency with which it occurs. In some instances,
4108-424: A result of the movement of its closest neighbours. Very large roosts, up to 1.5 million birds, form in city centres, woodlands and reedbeds, causing problems with their droppings. These may accumulate up to 30 cm (12 in) deep, killing trees by their concentration of chemicals. In smaller amounts, the droppings act as a fertiliser , and therefore woodland managers may try to move roosts from one area of
4266-505: A result, it has also been able to migrate to Thailand , Southeast Asia and New Guinea . Five individuals conveyed on a ship from England alighted near Lago de Maracaibo in Venezuela in November 1949, but subsequently vanished. In 1987, a small population of common starlings was observed nesting in gardens in the city of Buenos Aires . Since then, despite some initial attempts at eradication,
4424-650: A spiritual religious guardian. This imperial cult or system of divine kings was spread to most of central Africa by rivals in kingship migrating and forming new states. Many new states received legitimacy by claiming descent from the Lunda dynasties. The Kingdom of Kongo existed from the Atlantic west to the Kwango river to the east. During the 15th century, the Bakongo farming community was united with its capital at M'banza-Kongo , under
4582-477: A wild starling has been observed to mimic a sound it has heard only once. Each sound clip is repeated several times before the bird moves on to the next. After this variable section comes a number of types of repeated clicks followed by a final burst of high-frequency song, again formed of several types. Each bird has its own repertoire with more proficient birds having a range of up to 35 variable song types and as many as 14 types of clicks. Males sing constantly as
4740-561: A wingspan of 31–44 cm (12–17 in) and a weight of 58–101 g (2.0–3.6 oz). Among standard measurements, the wing chord is 11.8 to 13.8 cm (4.6 to 5.4 in), the tail is 5.8 to 6.8 cm (2.3 to 2.7 in), the culmen is 2.5 to 3.2 cm (0.98 to 1.26 in) and the tarsus is 2.7 to 3.2 cm (1.1 to 1.3 in). The plumage is iridescent black, glossed purple or green, and spangled with white, especially in winter. The underparts of adult male common starlings are less spotted than those of adult females at
4898-530: A wood to another to benefit from the soil enhancement and avoid large toxic deposits. Flocks of more than a million common starlings may be observed just before sunset in spring in southwestern Jutland , Denmark, over the seaward marshlands of Tønder and Esbjerg municipalities between Tønder and Ribe . They gather in March until northern Scandinavian birds leave for their breeding ranges by mid-April. Their swarm behaviour creates complex shapes silhouetted against
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5056-459: A year- in late summer after the breeding season has finished; the fresh feathers are prominently tipped white (breast feathers) or buff (wing and back feathers), which gives the bird a speckled appearance. The reduction in the spotting in the breeding season is achieved through the white feather tips largely wearing off. Juveniles are grey-brown and by their first winter resemble adults though often retaining some brown juvenile feathering, especially on
5214-520: Is a highly gregarious species, especially in autumn and winter. Although flock size is highly variable, huge, noisy flocks (murmurations) may form near roosts. These dense concentrations of birds are thought to be a defence against attacks by birds of prey such as peregrine falcons or Eurasian sparrowhawks . Flocks form a tight sphere -like formation in flight, frequently expanding and contracting and changing shape, seemingly without any sort of leader. Each common starling changes its course and speed as
5372-427: Is absent from regions where the ground is baked so dry that it cannot probe for insects. It may compete with native birds for crevice nesting sites, but the indigenous species are probably more disadvantaged by destruction of their natural habitat than they are by inter-specific competition. It breeds from September to December and outside the breeding season may congregate in large flocks, often roosting in reedbeds . It
5530-460: Is any bird of the order Passeriformes ( / ˈ p æ s ə r ɪ f ɔːr m iː z / ; from Latin passer 'sparrow' and formis '-shaped') which includes more than half of all bird species. Sometimes known as perching birds , passerines generally have an anisodactyl arrangement of their toes (three pointing forward and one back), which facilitates perching. With more than 140 families and some 6,500 identified species, Passeriformes
5688-493: Is common starling. The starling family, Sturnidae, is an entirely Old World group apart from introductions elsewhere, with the greatest numbers of species in Southeast Asia and sub-Saharan Africa . The genus Sturnus is polyphyletic and relationships between its members are not fully resolved. The closest relation of the common starling is the spotless starling . The non-migratory spotless starling may be descended from
5846-482: Is currently divided into three suborders: Acanthisitti (New Zealand wrens), Tyranni , (suboscines) and Passeri (oscines or songbirds). The Passeri is now subdivided into two major groups recognized now as Corvides and Passerida respectively containing the large superfamilies Corvoidea and Meliphagoidea , as well as minor lineages, and the superfamilies Sylvioidea , Muscicapoidea , and Passeroidea but this arrangement has been found to be oversimplified. Since
6004-577: Is fairly common but local in Jamaica, Grand Bahama and Bimini , and is rare in the rest of the Bahamas and eastern Cuba. The global population of the common starling is estimated to be more than 310 million individuals and its numbers are not thought to be declining significantly, so the bird is classified by the International Union for Conservation of Nature as being of least concern . It had shown
6162-405: Is itself preyed upon by the predatory mite Androlaelaps casalis . The presence of this control on numbers of the parasitic species may explain why birds are prepared to reuse old nests. Flying insects that parasitise common starlings include the louse-fly Omithomya nigricornis and the saprophagous fly Carnus hemapterus . The latter species breaks off the feathers of its host and lives on
6320-448: Is less pronounced than it is in songbird species like the zebra finch, where the male's syrinx is 100% larger than the female's syrinx. Singing also occurs outside the breeding season, taking place throughout the year apart from the moulting period. The songsters are more commonly male although females also sing on occasion. The function of such out-of-season song is poorly understood. Eleven other types of call have been described including
6478-669: Is located on the Swahili coast . Johannesburg is the largest city in South Africa . It is the provincial capital and largest city in Gauteng , which is the wealthiest province in South Africa. While Johannesburg is not one of South Africa's three capital cities , it is the seat of the Constitutional Court . The city is located in the mineral-rich Witwatersrand range of hills, and is
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6636-887: Is more scant before the Pleistocene, from which several still-existing families are documented. Apart from the indeterminable MACN -SC-1411 (Pinturas Early/Middle Miocene of Santa Cruz Province, Argentina), an extinct lineage of perching birds has been described from the Late Miocene of California, United States: the Palaeoscinidae with the single genus Palaeoscinis . "Palaeostruthus" eurius (Pliocene of Florida) probably belongs to an extant family, most likely passeroidean . Acanthisitti – New Zealand wrens (1 family containing 7 species, only 2 extant) Tyranni – suboscines (16 families containing 1,356 species) Passeri – oscines (125 families containing 5,158 species) The Passeriformes
6794-491: Is not thought to be declining significantly, so the common starling is classified as being of least concern by the International Union for Conservation of Nature . The common starling was first described by Carl Linnaeus in his Systema Naturae in 1758 under its current binomial name. Sturnus and vulgaris are derived from the Latin for "starling" and "common" respectively. The Old English staer , later stare , and
6952-801: Is now common in the southern Cape region, thinning out northwards to the Johannesburg area. It is present in the Western Cape , the Eastern Cape and the Free State provinces of South Africa and lowland Lesotho , with occasional sightings in KwaZulu-Natal , Gauteng and around the town of Oranjemund in Namibia . In Southern Africa populations appear to be resident and the bird is strongly associated with man and anthropogenic habitats . It favours irrigated land and
7110-478: Is one of the fastest-growing cities in the world, and also one of the most populous urban agglomerations . Lagos is a major financial centre in Africa; this megacity has the highest GDP, and also houses Apapa , one of the largest and busiest ports on the continent. Dar es Salaam is the former capital of, as well as the most populous city in, Tanzania ; it is a regionally important economic centre. It
7268-523: Is proven by several fossils from Germany such as a presumed broadbill ( Eurylaimidae ) humerus fragment from the Early Miocene (roughly 20 mya) of Wintershof , Germany, the Late Oligocene carpometacarpus from France listed above, and Wieslochia , among others. Extant Passeri super-families were quite distinct by that time and are known since about 12–13 mya when modern genera were present in
7426-546: Is quite strong and direct; their triangular wings beat very rapidly, and periodically the birds glide for a short way without losing much height before resuming powered flight. When in a flock, the birds take off almost simultaneously, wheel and turn in unison, form a compact mass or trail off into a wispy stream, bunch up again and land in a coordinated fashion. Common starling on migration can fly at 60–80 km/h (37–50 mph) and cover up to 1,000–1,500 km (620–930 mi). Several terrestrial starlings, including those in
7584-442: Is the long-tailed widowbird . The chicks of passerines are altricial : blind, featherless, and helpless when hatched from their eggs. Hence, the chicks require extensive parental care. Most passerines lay colored eggs, in contrast with nonpasserines, most of whose eggs are white except in some ground-nesting groups such as Charadriiformes and nightjars , where camouflage is necessary, and in some parasitic cuckoos , which match
7742-459: Is the short-tailed pygmy tyrant , at 6.5 cm (2.6 in) and 4.2 g (0.15 oz). The foot of a passerine has three toes directed forward and one toe directed backward, called anisodactyl arrangement. The hind toe ( hallux ) is long and joins the leg at approximately the same level as the front toes. This arrangement enables passerine birds to easily perch upright on branches. The toes have no webbing or joining, but in some cotingas ,
7900-508: Is the area and regions of the continent of Africa that lie south of the Sahara . These include Central Africa , East Africa , Southern Africa , and West Africa . Geopolitically, in addition to the African countries and territories that are situated fully in that specified region, the term may also include polities that only have part of their territory located in that region, per the definition of
8058-545: Is the largest order of birds and among the most diverse clades of terrestrial vertebrates , representing 60% of birds. Passerines are divided into three suborders : Acanthisitti (New Zealand wrens), Tyranni (composed mostly of South American suboscines), and Passeri (oscines or songbirds). Passerines originated in the Southern Hemisphere around 60 million years ago. Most passerines are insectivorous or omnivorous , and eat both insects and fruit or seeds. The terms "passerine" and "Passeriformes" are derived from
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#17330863374208216-462: Is the most common bird species in urban and agricultural areas. In 1901, the inhabitants of Saint Kitts petitioned the Colonial Secretary for a ″government grant of starlings to exterminate″ an outbreak of grasshoppers which was causing enormous damage to their crops. The common starling was introduced to Jamaica in 1903, and the Bahamas and Cuba were colonised naturally from the US. This bird
8374-641: Is thought that the camel was first brought to Egypt after the Persian Empire conquered Egypt in 525 BCE, although large herds did not become common enough in North Africa for camels to be the pack animal of choice for the trans-saharan trade. The Bantu expansion is a major migration movement that originated in West Central Africa (possibly around Cameroon) around 2500 BCE, reaching East and Central Africa by 1000 BCE and Southern Africa by
8532-414: Is typical, or just one north of 48°N. The young are born blind and naked. They develop light fluffy down within seven days of hatching and can see within nine days. As with other passerines, the nest is kept clean and the chicks' faecal sacs are removed by the adults. Once the chicks are able to regulate their body temperature, about six days after hatching, the adults largely cease removing droppings from
8690-655: The Baltic States , and smaller declines in much of the rest of northern and central Europe. The bird has been adversely affected in these areas by intensive agriculture, and in several countries it has been red-listed due to population declines of more than 50%. Numbers dwindled in the United Kingdom by more than 80% between 1966 and 2004; although populations in some areas such as Northern Ireland were stable or even increased, those in other areas, mainly England, declined even more sharply. The overall decline seems to be due to
8848-637: The Bantu expansion . In Southern Africa, their speakers are the Khoikhoi and San (Bushmen), in Southeast Africa, the Sandawe and Hadza . The Niger–Congo family is the largest in the world in terms of the number of languages (1,436) it contains. The vast majority of languages of this family are tonal , such as Yoruba and Igbo . However, others such as Fulani , Wolof and Kiswahili are not. A major branch of
9006-460: The Bosporus ; and S. v. heinrichi Stresemann , 1928, an intergrade between caucasicus and nobilior in northern Iran. S. v. persepolis Ticehurst , 1928 from southern Iran's ( Fars Province ) is very similar to vulgaris , and it is not clear whether it is a distinct resident population or simply migrants from southeastern Europe. The common starling is 19–23 cm (7.5–9.1 in) long, with
9164-652: The Corvida and numerous minor lineages make up songbird diversity today. Extensive biogeographical mixing happens, with northern forms returning to the south, southern forms moving north, and so on. Perching bird osteology , especially of the limb bones, is rather diagnostic. However, the early fossil record is poor because passerines are relatively small, and their delicate bones do not preserve well. Queensland Museum specimens F20688 ( carpometacarpus ) and F24685 ( tibiotarsus ) from Murgon, Queensland , are fossil bone fragments initially assigned to Passeriformes . However,
9322-737: The Dutch East India Company . For most of the 17th and 18th centuries, the slowly expanding settlement was a Dutch possession. In 1795, the Dutch colony was captured by the British during the French Revolutionary Wars . The British intended to use Cape Town as a major port on the route to Australia and India . It was later returned to the Dutch in 1803, but soon afterward the Dutch East India Company declared bankruptcy, and
9480-691: The Ethiopian Empire , and the Geledi Sultanate , whose military dominance forced governors of the Omani empire north of the city of Lamu to pay tribute to the Somali Sultan Ahmed Yusuf . According to the theory of recent African origin of modern humans , the mainstream position held within the scientific community, all humans originate from either Southeast Africa or the Horn of Africa. During
9638-459: The H. erectus species. The fossil and genetic evidence shows Homo sapiens developed in Southern and East Africa by around 350,000 to 260,000 years ago and gradually migrated across the continent in waves. Between 50,000 and 60,000 years ago, their expansion out of Africa launched the colonisation of the planet by modern humans. By 10,000 BCE, Homo sapiens had spread to all corners of
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#17330863374209796-586: The Kermadec Islands where it is abundant, that group being roughly equidistant between New Zealand and Fiji. Its spread in Fiji has been limited, and there are doubts about the population's viability. Tonga was colonised at about the same date and the birds there have been slowly spreading north through the group. In South Africa, the common starling was introduced in 1897 by Cecil Rhodes . It spread slowly, and by 1954, had reached Clanwilliam and Port Elizabeth . It
9954-578: The Mali Empire , Songhai Empire , the Kanem Empire and the subsequent Bornu Empire . They built stone structures like in Tichit , but mainly constructed in adobe . The Great Mosque of Djenne is most reflective of Sahelian architecture and is the largest adobe building in the world. In the forest zone, several states and empires such as Bono State , Akwamu and others emerged. The Ashanti Empire arose in
10112-586: The Middle Pleistocene , part of the problem in resolving relationships in the Sturnidae is the paucity of the fossil record for the family as a whole. There are several subspecies of the common starling, which vary clinally in size and the colour tone of the adult plumage. The gradual variation over geographic range and extensive intergradation means that acceptance of the various subspecies varies between authorities. Birds from Fair Isle , St. Kilda and
10270-621: The Old World warblers and Old World babblers have turned out to be paraphyletic and are being rearranged. Several taxa turned out to represent highly distinct lineages, so new families had to be established, some of theirs – like the stitchbird of New Zealand and the Eurasian bearded reedling – monotypic with only one living species. In the Passeri alone, a number of minor lineages will eventually be recognized as distinct superfamilies. For example,
10428-564: The Outer Hebrides are intermediate in size between S. v. zetlandicus and the nominate form, and their subspecies placement varies according to the authority. The dark juveniles typical of these island forms are occasionally found in mainland Scotland and elsewhere, indicating some gene flow from faroensis or zetlandicus , subspecies formerly considered to be isolated. Several other subspecies have been named, but are generally no longer considered valid. Most are intergrades that occur where
10586-578: The Palearctic to western Mongolia, and it has been introduced as an invasive species to Australia , New Zealand , Canada , the United States , Mexico , Argentina , South Africa and Fiji . This bird is resident in western and southern Europe and southwestern Asia, while northeastern populations migrate south and west in the winter within the breeding range and also further south to Iberia and North Africa. The common starling builds an untidy nest in
10744-650: The Sabaeans , Parthians and Axumites for the wealthy Indo – Greco – Roman trade. In the Middle Ages several powerful Somali empires dominated the region's trade, including the Ajuran Sultanate , which excelled in hydraulic engineering and fortress building, the Sultanate of Adal , whose General Ahmed Gurey was the first African commander in history to use cannon warfare on the continent during Adal's conquest of
10902-643: The Songhai , Zarma and Kanuri . The Old Nubian language is also a member of this family. Major languages of Africa by region, family and number of primary language speakers in millions: Sub-Saharan Africa has several large cities. Lagos is a city in the Nigerian state of Lagos . The city, with its adjoining conurbation , is the most populous in Nigeria , and the second-most populous in Africa after Cairo , Egypt . It
11060-621: The UN Development Programme applies the "sub-Saharan" classification to 46 of Africa's 55 countries, excluding Djibouti, SADR , Somalia, and Sudan. Since around 3900 BCE , the Saharan and sub-Saharan regions of Africa have been separated by the extremely harsh climate of the sparsely populated Sahara, forming an effective barrier that is interrupted only by the Nile in Sudan, though navigation on
11218-481: The United Nations (UN). This is considered a non-standardised geographical region with the number of countries included varying from 46 to 48 depending on the organisation describing the region (e.g. UN , WHO , World Bank , etc.). The African Union (AU) uses a different regional breakdown, recognising all 55 member states on the continent—grouping them into five distinct and standard regions. The term serves as
11376-748: The Walls of Benin , which is the largest man-made structure in the world. In the 18th century, the Oyo and the Aro confederacy were responsible for most of the slaves exported from modern-day Nigeria, selling them to European slave traders . Following the Napoleonic Wars , the British expanded their influence into the Nigerian interior. In 1885, British claims to a West African sphere of influence received international recognition, and in
11534-631: The crows , do not sound musical to human beings. Some, such as the lyrebird , are accomplished mimics. The New Zealand wrens are tiny birds restricted to New Zealand , at least in modern times; they were long placed in Passeri. Most passerines are smaller than typical members of other avian orders. The heaviest and altogether largest passerines are the thick-billed raven and the larger races of common raven , each exceeding 1.5 kg (3.3 lb) and 70 cm (28 in). The superb lyrebird and some birds-of-paradise , due to very long tails or tail coverts, are longer overall. The smallest passerine
11692-435: The genus Sturnus , have adaptations of the skull and muscles that help with feeding by probing. This adaptation is most strongly developed in the common starling (along with the spotless and white-cheeked starlings ), where the protractor muscles responsible for opening the jaw are enlarged and the skull is narrow, allowing the eye to be moved forward to peer down the length of the bill. This technique involves inserting
11850-445: The kinglets constitute a single genus with less than 10 species today but seem to have been among the first perching bird lineages to diverge as the group spread across Eurasia. No particularly close relatives of theirs have been found among comprehensive studies of the living Passeri, though they might be fairly close to some little-studied tropical Asian groups. Nuthatches , wrens , and their closest relatives are currently grouped in
12008-611: The medieval period used the general term bilâd as-sûdân ("Land of the Blacks") for the vast Sudan region (an expression denoting Central and West Africa ), or sometimes extending from the coast of West Africa to Western Sudan . Its equivalent in Southeast Africa was Zanj ("Country of the Blacks"), which was situated in the vicinity of the Great Lakes region. The geographers drew an explicit ethnographic distinction between
12166-460: The northern goshawk ( Accipiter gentilis ) and Eurasian sparrowhawk ( Accipiter nisus ), and falcons including the peregrine falcon ( Falco peregrinus ), Eurasian hobby ( Falco subbuteo ) and common kestrel ( Falco tinnunculus ). Slower raptors like black and red kites ( Milvus migrans & milvus ), eastern imperial eagle ( Aquila heliaca ), common buzzard ( Buteo buteo ) and Australasian harrier ( Circus approximans ) tend to take
12324-450: The rook and the common starling are the most infested wild birds. Other recorded internal parasites include the spiny-headed worm Prosthorhynchus transverses . Common starlings may contract avian tuberculosis , avian malaria and retrovirus -induced lymphomas . Captive starlings often accumulate excess iron in the liver, a condition that can be prevented by adding black tea-leaves to the food. The global population of common starlings
12482-456: The scientific name of the house sparrow , Passer domesticus , and ultimately from the Latin term passer , which refers to sparrows and similar small birds. The order is divided into three suborders, Tyranni (suboscines), Passeri (oscines or songbirds), and the basal Acanthisitti . Oscines have the best control of their syrinx muscles among birds, producing a wide range of songs and other vocalizations, though some of them, such as
12640-402: The superb lyrebird has 16, and several spinetails in the family Furnariidae have 10, 8, or even 6, as is the case of Des Murs's wiretail . Species adapted to tree trunk climbing such as treecreepers and woodcreeper have stiff tail feathers that are used as props during climbing. Extremely long tails used as sexual ornaments are shown by species in different families. A well-known example
12798-581: The 14th and 15th centuries, large medieval Southeast African kingdoms and states emerged, such as the Buganda , Bunyoro and Karagwe kingdoms of Uganda and Tanzania. During the early 1960s, the Southeast African nations achieved independence from colonial rule. Settlements of Bantu -speaking peoples, who were iron -using agriculturists and herdsmen, were already present south of the Limpopo River by
12956-594: The 1870s. As part of a nationwide effort, about 60 common starlings were released in 1890 into New York's Central Park by Eugene Schieffelin , president of the American Acclimatization Society . It has been widely reported that he had tried to introduce every bird species mentioned in the works of William Shakespeare into North America, but this claim has been traced to an essay in 1948 by naturalist Edwin Way Teale , whose notes appear to indicate that it
13114-585: The 18th century in modern-day Ghana . The Kingdom of Nri , was established by the Igbo in the 11th century. Nri was famous for having a priest-king who wielded no military power. Nri was a rare African state which was a haven for freed slaves and outcasts who sought refuge in their territory. Other major states included the kingdoms of Ifẹ and Oyo in the western block of Nigeria which became prominent about 700–900 and 1400 respectively, and center of Yoruba culture. The Yoruba built massive mud walls around their cities,
13272-520: The 1950s and early 1960s, most sub-Saharan African nations achieved independence from colonial rule. According to the 2022 revision of the World Population Prospects , the population of sub-Saharan Africa was 1.1 billion in 2019. The current growth rate is 2.3%. The UN predicts for the region a population between 2 and 2.5 billion by 2050 with a population density of 80 per km compared to 170 for Western Europe, 140 for Asia and 30 for
13430-526: The 4th or 5th century displacing and absorbing the original Khoisan speakers. They slowly moved south, and the earliest ironworks in modern-day KwaZulu-Natal Province are believed to date from around 1050. The southernmost group was the Xhosa people , whose language incorporates certain linguistic traits from the earlier Khoisan inhabitants. They reached the Fish River in today's Eastern Cape Province . Monomotapa
13588-767: The Americas. Sub-Saharan African countries top the list of countries and territories by fertility rate with 40 of the highest 50, all with TFR greater than 4 in 2008. All are above the world average except South Africa and Seychelles . More than 40% of the population in sub-Saharan countries is younger than 15 years old, as well as in Sudan , with the exception of South Africa. GDP per Capita (PPP) (2016, 2017 (PPP, US$ )) , Life (Exp.) (Life Expectancy 2006) , Literacy (Male/Female 2006) , Trans (Transparency 2009) , HDI (Human Development Index) , EODBR (Ease of Doing Business Rank June 2008 through May 2009) , SAB ( Starting
13746-516: The Dutch (now under French control) and the British found themselves at war again. The British captured the Dutch possession yet again at the Battle of Blaauwberg , commanded by Sir David Blair . The Zulu Kingdom was a Southern African tribal state in what is now KwaZulu-Natal in southeastern South Africa. The small kingdom gained world fame during and after their defeat in the Anglo-Zulu War . During
13904-652: The Eastern Baribah or Barbaroi , as the ancestors of the Somalis were referred to by medieval Arab and ancient Greek geographers, respectively. In the 19th and 20th centuries, the populations south of the Sahara were divided into three broad ancestral groups: Hamites and Semites in the Horn of Africa and Sahel related to those in North Africa, who spoke languages belonging to the Afroasiatic family; Negroes in most of
14062-617: The Horn of Africa (Ethiopia, Eritrea). The Chadic branch is distributed in Central and West Africa. Hausa , its most widely spoken language, serves as a lingua franca in West Africa (Niger, Ghana, Togo, Benin, Cameroon, and Chad). The several families lumped under the term Khoi-San include languages indigenous to Southern Africa and Tanzania , though some, such as the Khoi languages , appear to have moved to their current locations not long before
14220-598: The Latin sturnus are both derived from an unknown Indo-European root dating back to the second millennium BC. "Starling" was first recorded in the 11th century, when it referred to the juvenile of the species, but by the 16th century it had already largely supplanted "stare" to refer to birds of all ages. The older name is referenced in William Butler Yeats' poem "The Stare's Nest by My Window". The International Ornithological Congress 's preferred English vernacular name
14378-640: The Mediterranean. Growing from the proto-Aksumite Iron Age period ( c. 4th century BCE), it rose to prominence by the 1st century CE. The Aksumites constructed monolithic stelae to cover the graves of their kings, such as King Ezana's Stele . The later Zagwe dynasty , established in the 12th century, built churches out of solid rock. These rock-hewn structures include the Church of St. George at Lalibela . In ancient Somalia , city-states flourished such as Opone , Mosyllon and Malao that competed with
14536-539: The Niger–Congo languages is Bantu , which covers a greater geographic area than the rest of the family. Bantu speakers represent the majority of inhabitants in southern, central and southeastern Africa, though San , Pygmy , and Nilotic groups, respectively, can also be found in those regions. Bantu-speakers can also be found in parts of Central Africa such as the Gabon , Equatorial Guinea and southern Cameroon . Swahili ,
14694-620: The Nile was blocked by the Sudd and the river's cataracts . There is also an evident genetic divide between North Africa and sub-Saharan Africa that dates back to the Neolithic . The Sahara pump theory explains how flora and fauna (including Homo sapiens ) left Africa to penetrate Eurasia and beyond. African pluvial periods are associated with a " Wet Sahara " phase, during which larger lakes and more rivers existed. Geographers historically divided
14852-670: The Sahel (northern Mauritania, northern Mali, northern Niger). The Cushitic branch of Afroasiatic is centered in the Horn, and is also spoken in the Nile Valley and parts of the African Great Lakes region. Additionally, the Semitic branch of the family, in the form of Arabic , is widely spoken in the parts of Africa that are within the Arab world. South Semitic languages are also spoken in parts of
15010-465: The Southeast coast of Kenya and Tanzania encountered by these later Arab and Persian settlers have been variously identified with the trading settlements of Rhapta , Azania and Menouthias referenced in early Greek and Chinese writings from 50 CE to 500 CE. These early writings perhaps document the first wave of Bantu settlers to reach Southeast Africa during their migration. Between
15168-687: The Sudan region and its analogue Zanj, from the area to their extreme east on the Red Sea coast in the Horn of Africa . In modern-day Ethiopia and Eritrea was Al-Habash or Abyssinia, which was inhabited by the Habash or Abyssinians, who were the forebears of the Habesha . In northern Somalia was Barbara or the Bilad al-Barbar ("Land of the Berbers "), which was inhabited by
15326-412: The US, even on house sparrows . Lice include Menacanthus eurystemus , Brueelia nebulosa and Stumidoecus sturni . Other arthropod parasites include Ixodes ticks and mites such as Analgopsis passerinus , Boydaia stumi , Dermanyssus gallinae , Ornithonyssus bursa , O. sylviarum , Proctophyllodes species, Pteronyssoides truncatus and Trouessartia rosteri . The hen mite D. gallinae
15484-448: The adult survival rate is closer to 60%. The average life span is about 2–3 years, with a longevity record of 22 years 11 months. A majority of starling predators are avian. The typical response of starling groups is to take flight, with a common sight being undulating flocks of starling flying high in quick and agile patterns. Their abilities in flight are seldom matched by birds of prey. Adult common starlings are hunted by hawks such as
15642-421: The application of animal husbandry towards carrying water, food, and supplies across the desert. Prior to the introduction of the camel , the use of oxen, mule, and horses for desert crossing was common, and trade routes followed chains of oases that were strung across the desert. The trans-saharan trade was in full motion by 500 BCE with Carthage being a major economic force for its establishment. It
15800-411: The art of making fire. They were the first hominids to leave Africa, going on to colonise the entire Old World , and perhaps later on giving rise to Homo floresiensis . Although some recent writers suggest that H. georgicus , a H. habilis descendant , was the first and most primitive hominid to ever live outside Africa, many scientists consider H. georgicus to be an early and primitive member of
15958-545: The basis of morphological similarities that, it is now believed, are the result of convergent evolution , not a close genetic relationship. For example, the wrens of the Americas and Eurasia , those of Australia , and those of New Zealand look superficially similar and behave in similar ways, yet belong to three far-flung branches of the passerine family tree; they are as unrelated as it is possible to be while remaining Passeriformes. Advances in molecular biology and improved paleobiogeographical data gradually are revealing
16116-424: The bill into the ground and opening it as a way of searching for hidden food items. Common starlings have the physical traits that enable them to use this feeding technique, which has undoubtedly helped the species spread far and wide. In Iberia, the western Mediterranean and northwest Africa, the common starling may be confused with the closely related spotless starling, the plumage of which, as its name implies, has
16274-594: The bill is black in winter and yellow in summer; young birds have browner plumage than the adults. It is a noisy bird, especially in communal roosts and other gregarious situations, with an unmusical but varied song. Its gift for mimicry has been noted in literature including the Mabinogion and the works of Pliny the Elder and William Shakespeare . The common starling has about 12 subspecies breeding in open habitats across its native range in temperate Europe and across
16432-478: The bird has been expanding its breeding range at an average rate of 7.5 km (4.7 mi) per year, keeping within 30 km (19 mi) of the Atlantic coast. In Argentina, the species makes use of a variety of natural and man-made nesting sites, particularly woodpecker holes. The common starling was introduced to Australia to consume insect pests of farm crops. Early settlers looked forward to their arrival, believing that common starlings were also important for
16590-565: The breeding period approaches and perform less often once pairs have bonded. In the presence of a female, a male sometimes flies to his nest and sings from the entrance, apparently attempting to entice the female in. Older birds tend to have a wider repertoire than younger ones. Those males that engage in longer bouts of singing and that have wider repertoires attract mates earlier and have greater reproductive success than others. Females appear to prefer mates with more complex songs, perhaps because this indicates greater experience or longevity. Having
16748-476: The carcasses of animals killed by other predators, both for their meat and their marrow. In hunting, H. habilis was most likely not capable of competing with large predators and was more prey than hunter, although H. habilis likely did steal eggs from nests and may have been able to catch small game and weakened larger prey such as cubs and older animals. The tools were classed as Oldowan . Roughly 1.8 million years ago, Homo ergaster first appeared in
16906-547: The common starling eats the eggs of the endangered roseate tern . Measures are being introduced to reduce common starling populations by culling before the terns return to their breeding colonies in spring. There are several methods by which common starlings obtain their food, but, for the most part, they forage close to the ground, taking insects from the surface or just underneath. Generally, common starlings prefer foraging amongst short-cropped grasses and eat with grazing animals or perch on their backs, where they will also feed on
17064-653: The common starling was introduced from Europe along with the house sparrow to control the pests. It was first brought over in 1862 by the Nelson Acclimatisation Society and other introductions followed. The birds soon became established and are now found all over the country including the subtropical Kermadec Islands to the north and the equally distant Macquarie Island far to the south. Various acclimatisation society records mention instances of starlings being introduced in Cincinnati, Quebec and New York in
17222-530: The common starling. "Probing" involves the bird plunging its beak into the ground randomly and repetitively until an insect has been found, and is often accompanied by bill gaping where the bird opens its beak in the soil to enlarge a hole. This behaviour, first described by Konrad Lorenz and given the German term zirkeln , is also used to create and widen holes in plastic garbage bags. It takes time for young common starlings to perfect this technique, and because of this
17380-427: The consumption of invertebrates is necessary for successful breeding, common starlings are omnivorous and can also eat grains , seeds , fruits , nectar and food waste if the opportunity arises. The Sturnidae differ from most birds in that they cannot easily metabolise foods containing high levels of sucrose , although they can cope with other fruits such as grapes and cherries. The isolated Azores subspecies of
17538-502: The corvoidean and basal songbirds. The modern diversity of Passerida genera is known mostly from the Late Miocene onward and into the Pliocene (about 10–2 mya). Pleistocene and early Holocene lagerstätten (<1.8 mya) yield numerous extant species, and many yield almost nothing but extant species or their chronospecies and paleosubspecies. In the Americas , the fossil record
17696-644: The country and the pathways of these different streams of bird may cross. Of the 15,000 birds ringed as nestlings in Merseyside , England, individuals have been recovered at various times of year as far afield as Norway, Sweden, Finland, Russia, Ukraine, Poland, Germany and the Low Countries . Small numbers of common starlings have sporadically been observed in Japan and Hong Kong but it is unclear whence these birds originated. In North America, northern populations have developed
17854-442: The deposition of fat. Unpaired males find a suitable cavity and begin to build nests in order to attract single females, often decorating the nest with ornaments such as flowers and fresh green material, which the female later disassembles upon accepting him as a mate. The amount of green material is not important, as long as some is present, but the presence of herbs in the decorative material appears to be significant in attracting
18012-448: The diet of young birds will often contain fewer insects. " Hawking " is the capture of flying insects directly from the air, and "lunging" is the less common technique of striking forward to catch a moving invertebrate on the ground. Earthworms are caught by pulling from soil. Common starlings that have periods without access to food, or have a reduction in the hours of light available for feeding, compensate by increasing their body mass by
18170-464: The early centuries CE. The Djenné-Djenno city-state flourished from 250 BCE to 900 CE and was influential to the development of the Ghana Empire . The Nok culture of Nigeria (lasting from 1,500 BCE to 200 CE) is known from a type of terracotta figure. There were a number of medieval empires of the southern Sahara and the Sahel, based on trans-Saharan trade , including the Ghana Empire and
18328-404: The families in the suborder Tyranni (suboscines) were all well determined but some of the nodes in Passeri (oscines or songbirds) were unclear owing to the rapid splitting of the lineages. Infraorder Eurylaimides : Old World suboscines Infraorder Tyrannides : New World suboscines Parvorder Furnariida Parvorder Tyrannida Relationships between living Passeriformes families based on
18486-451: The fats produced by growing plumage. Larvae of the moth Hofmannophila pseudospretella are nest scavengers, which feed on animal material such as faeces or dead nestlings. Protozoan blood parasites of the genus Haemoproteus have been found in common starlings, but a better known pest is the brilliant scarlet nematode Syngamus trachea . This worm moves from the lungs to the trachea and may cause its host to suffocate. In Britain,
18644-618: The first millennium BCE. Trade and improved agricultural techniques supported more sophisticated societies, leading to the early civilisations of Sao , Kanem , Bornu , Shilluk , Baguirmi , and Wadai . Following the Bantu Migration into Central Africa, during the 14th century, the Luba Kingdom in southeast Congo came about under a king whose political authority derived from religious, spiritual legitimacy. The kingdom controlled agriculture and regional trade of salt and iron from
18802-534: The first millennium CE, Nilotic and Bantu -speaking peoples moved into the region , and the latter now account for three-quarters of Kenya's population. On the coastal section of Southeast Africa, a mixed Bantu community developed through contact with Muslim Arab and Persian traders, leading to the development of the mixed Arab, Persian and African Swahili City States . The Swahili culture that emerged from these exchanges evinces many Arab and Islamic influences not seen in traditional Bantu culture, as do
18960-558: The following year the Royal Niger Company was chartered under the leadership of Sir George Goldie . In 1900, the company's territory came under the control of the British government, which moved to consolidate its hold over the area of modern Nigeria. On 1 January 1901, Nigeria became a British protectorate as part of the British Empire , the foremost world power at the time. Nigeria was granted its independence in 1960 during
19118-440: The following year. Intraspecific brood parasites are common in common starling nests. Female "floaters" (unpaired females during the breeding season) present in colonies often lay eggs in another pair's nest. Fledglings have also been reported to invade their own or neighbouring nests and evict a new brood. Common starling nests have a 48% to 79% rate of successful fledging, although only 20% of nestlings survive to breeding age;
19276-517: The form of the Akkadian language of Mesopotamia ( Assyria and Babylonia ) and circa 2500 BCE in the form of the Eblaite language of northeastern Syria . The distribution of the Afroasiatic languages within Africa is principally concentrated in North Africa and the Horn of Africa. Languages belonging to the family's Berber branch are mainly spoken in the north, with its speech area extending into
19434-502: The fossil record in Africa. From Homo ergaster , Homo erectus (upright man) evolved 1.5 million years ago. Some of the earlier representatives of this species were small-brained and used primitive stone tools , much like H. habilis . The brain later grew in size, and H. erectus eventually developed a more complex stone tool technology called the Acheulean . Potentially the first hominid to engage in hunting, H. erectus mastered
19592-403: The ground its strange, somewhat waddling gait is also characteristic. The colouring and build usually distinguish this bird from other starlings, although the closely related spotless starling may be physically distinguished by the lack of iridescent spots in adult breeding plumage. Like most terrestrial starlings the common starling moves by walking or running, rather than hopping. Their flight
19750-707: The head. They can usually be sexed by the colour of the irises, rich brown in males, mouse-brown or grey in females. Estimating the contrast between an iris and the central always-dark pupil is 97% accurate in determining sex, rising to 98% if the length of the throat feathers is also considered. The common starling is mid-sized by both starling standards and passerine standards. It is readily distinguished from other mid-sized passerines, such as thrushes , icterids or small corvids , by its relatively short tail, sharp, blade-like bill, round-bellied shape and strong, sizeable (and rufous-coloured) legs. In flight, its strongly pointed wings and dark colouration are distinctive, while on
19908-558: The indigenous people of southern Africa. Pygmies are the pre-Bantu indigenous peoples of Central Africa. The Nilo-Saharan languages are concentrated in the upper parts of the Chari and Nile rivers of Central Africa and Southeast Africa. They are principally spoken by Nilotic peoples and are also spoken in Sudan among the Fur , Masalit , Nubian and Zaghawa peoples and in West and Central Africa among
20066-666: The king title, Manikongo . Other significant states and peoples included the Kuba Kingdom , producers of the famous raffia cloth, the Eastern Lunda , Bemba , Burundi , Rwanda , and the Kingdom of Ndongo . Nubia , covered by present-day northern Sudan and southern Egypt , was referred to as " Aethiopia " ("land of the burnt face") by the Greeks . Nubia in her greatest phase is considered sub-Saharan Africa's oldest urban civilisation. Nubia
20224-399: The last egg laid may take 24 hours longer than the first to hatch. Both parents share the responsibility of brooding the eggs, but the female spends more time incubating them than does the male, and is the only parent to do so at night when the male returns to the communal roost. A pair can raise up to three broods per year, frequently reusing and relining the same nest, although two broods
20382-402: The low survival rate of young birds, which may be caused by changes in agricultural practices. The intensive farming methods used in northern Europe mean there is less pasture and meadow habitat available, and the supply of grassland invertebrates needed for the nestlings to thrive is correspondingly reduced. Passerine and see text A passerine ( / ˈ p æ s ə r aɪ n / )
20540-445: The mammal's external parasites. Large flocks may engage in a practice known as "roller-feeding", where the birds at the back of the flock continually fly to the front where the feeding opportunities are best. The larger the flock, the nearer individuals are to one another while foraging. Flocks often feed in one place for some time, and return to previous successfully foraged sites. There are three types of foraging behaviours observed in
20698-511: The many Afro-Arab members of the Bantu Swahili people . With its original speech community centered on the coastal parts of Tanzania (particularly Zanzibar ) and Kenya – a seaboard referred to as the Swahili Coast – the Bantu Swahili language contains many Arabic loan-words as a consequence of these interactions. The earliest Bantu inhabitants of
20856-649: The material is too fragmentary and their affinities have been questioned. Several more recent fossils from the Oligocene of Europe, such as Wieslochia , Jamna , Resoviaornis , and Crosnoornis , are more complete and definitely represent early passeriforms, and have been found to belong to a variety of modern and extinct lineages. From the Bathans Formation at the Manuherikia River in Otago , New Zealand, MNZ S42815 (a distal right tarsometatarsus of
21014-524: The mid-2000s, studies have investigated the phylogeny of the Passeriformes and found that many families from Australasia traditionally included in the Corvoidea actually represent more basal lineages within oscines. Likewise, the traditional three-superfamily arrangement within the Passeri has turned out to be far more complex and will require changes in classification. Major " wastebin " families such as
21172-448: The more easily caught fledglings or juveniles. While perched in groups by night, they can be vulnerable to owls, including the little owl ( Athene noctua ), long-eared owl ( Asio otus ), short-eared owl ( Asio flammeus ), barn owl ( Tyto alba ), tawny owl ( Strix aluco ) and Eurasian eagle-owl ( Bubo bubo ). More than twenty species of hawk , owl and falcon are known to occasionally predate starlings in North America, though
21330-506: The most famous being Sungbo's Eredo . Another prominent kingdom in southwestern Nigeria was the Kingdom of Benin , whose power lasted between the 15th and 19th century. Their dominance reached as far as the well-known city of Eko which was named Lagos by the Portuguese traders and other early European settlers. The Edo -speaking people of Benin are known for their famous bronze casting and rich coral, wealth, ancient science and technology and
21488-671: The most regular predators of adults are likely to be urban-living peregrine falcons or merlins ( Falco columbarius ). Common mynas ( Acridotheres tristis ) sometimes evict eggs, nestlings and adult common starlings from their nests, and the lesser honeyguide ( Indicator minor ), a brood parasite , uses the common starling as a host. Starlings are more commonly the culprits rather than victims of nest eviction however, especially towards other starlings and woodpeckers . Nests can be raided by mammals capable of climbing to them, such as small mustelids ( Mustela spp.), raccoons ( Procyon lotor ) and squirrels ( Sciurus spp.), and cats may catch
21646-496: The nest. Prior to that, the fouling would wet both the chicks' plumage and the nest material, thereby reducing their effectiveness as insulation and increasing the risk of chilling the hatchlings. Nestlings remain in the nest for three weeks, where they are fed continuously by both parents. Fledglings continue to be fed by their parents for another one or two weeks. Within two months, most juveniles will have moulted and gained their first basic plumage. They acquire their adult plumage
21804-691: The north and copper from the Zambian/Congo copper belt. Rival kingship factions which split from the Luba Kingdom later moved among the Lunda people, marrying into its elite and laying the foundation of the Lunda Empire in the 16th century. The ruling dynasty centralised authority among the Lunda under the Mwata Yamyo or Mwaant Yaav. The Mwata Yamyo's legitimacy, like that of the Luba king, came from being viewed as
21962-509: The passerine host's egg. The vinous-throated parrotbill has two egg colors, white and blue, to deter the brood parasitic common cuckoo . Clutches vary considerably in size: some larger passerines of Australia such as lyrebirds and scrub-robins lay only a single egg, most smaller passerines in warmer climates lay between two and five, while in the higher latitudes of the Northern Hemisphere, hole-nesting species like tits can lay up to
22120-678: The period of decoloniszation . Archeological finds in Central Africa provide evidence of human settlement that may date back over 10,000 years. According to Zangato and Holl, there is evidence of iron-smelting in the Central African Republic and Cameroon that may date back to 3,000 to 2,500 BCE. Extensive walled sites and settlements have recently been found in Zilum , Chad. The area is located approximately 60 km (37 mi) southwest of Lake Chad, and has been radiocarbon dated to
22278-919: The phylogenetic analysis of Oliveros et al (2019). Some terminals have been renamed to reflect families recognised by the IOC but not in that study. The IOC families Alcippeidae and Teretistridae were not sampled in this study. Acanthisittidae (New Zealand wrens) Eurylaimidae (eurylaimid broadbills) Philepittidae (asites) Calyptomenidae (African and green broadbills) Pittidae (pittas) Sapayoidae (sapayoa) Melanopareiidae (crescent chests) Conopophagidae (gnateaters) Thamnophilidae (antbirds) Grallariidae (antpittas) Rhinocryptidae (tapaculos) Formicariidae (antthrushes) Scleruridae (leaftossers) Dendrocolaptidae (woodcreepers) Furnariidae (ovenbirds) Pipridae (manakins) Cotingidae (cotingas) Tityridae (tityras, becards) Sub-Saharan Africa Sub-Saharan Africa or Subsahara
22436-688: The pollination of flax , a major agricultural product. Nest-boxes for the newly released birds were placed on farms and near crops. The common starling was introduced to Melbourne in 1857 and Sydney two decades later. By the 1880s, established populations were present in the southeast of the country thanks to the work of acclimatisation committees. By the 1920s, common starlings were widespread throughout Victoria , Queensland and New South Wales , but by then they were considered to be pests. Although common starlings were first sighted in Albany, Western Australia in 1917, they have been largely prevented from spreading to
22594-415: The ranges of various subspecies meet. These include: S. v. ruthenus Menzbier , 1891 and S. v. jitkowi Buturlin, 1904, which are intergrades between vulgaris and poltaratskyi from western Russia; S. v. graecus Tschusi , 1905 and S. v. balcanicus Buturlin and Harms, 1909, which are intergrades between vulgaris and tauricus from the southern Balkans to central Ukraine and throughout Greece to
22752-557: The region into several distinct ethnographic sections based on each area's respective inhabitants. The concept has been criticised as a racist construction intended to separate North Africa or "European/White Africa" and "Black Africa" or "Africa noire". Critics from various countries have provided arguments supporting the interconnectedness of continental Africa, pointing to historical and cultural connections, as well as trade between North, West, and East Africa . Commentators in Arabic in
22910-732: The rest of the subcontinent (hence, the toponym Black Africa for Africa south of the Sahara ), who spoke languages belonging to the Niger-Congo and Nilo-Saharan families; and Khoisan in Southern Africa , who spoke languages belonging to the Khoisan family. Sub-Saharan Africa has a wide variety of climate zones or biomes . South Africa and the Democratic Republic of the Congo in particular are considered megadiverse countries . It has
23068-509: The second and third toes are united at their basal third. The leg of passerine birds contains an additional special adaptation for perching. A tendon in the rear of the leg running from the underside of the toes to the muscle behind the tibiotarsus will automatically be pulled and tighten when the leg bends, causing the foot to curl and become stiff when the bird lands on a branch. This enables passerines to sleep while perching without falling off. Most passerine birds have 12 tail feathers but
23226-722: The sky, a phenomenon known locally as sort sol ("black sun"). Flocks of anything from five to fifty thousand common starlings form in areas of the UK just before sundown during mid-winter. These flocks are commonly called murmurations. The common starling is largely insectivorous and feeds on both pest and other arthropods . The food range includes spiders , crane flies , moths , mayflies , dragonflies , damsel flies , grasshoppers , earwigs , lacewings , caddisflies , flies , beetles , sawflies , bees , wasps and ants . Prey are consumed in both adult and larvae stages of development, and common starlings will also feed on earthworms , snails , small amphibians and lizards . While
23384-487: The south and west of Europe and south of latitude 40°N are mainly resident , although other populations migrate from regions where the winter is harsh, the ground frozen and food scarce. Large numbers of birds from northern Europe, Russia and Ukraine migrate south westwards or south eastwards. In the autumn, when immigrants are arriving from eastern Europe, many of Britain's common starlings are setting off for Iberia and North Africa. Other groups of birds are in passage across
23542-603: The southern continents in the early Eocene . The New Zealand wrens are the first to become isolated in Zealandia , and the second split involved the origin of the Tyranni in South America and the Passeri in the Australian continent . The Passeri experienced a great radiation of forms in Australia. A major branch of the Passeri, the parvorder Passerida , dispersed into Eurasia and Africa about 40 million years ago, where they experienced further radiation of new lineages. This eventually led to three major Passerida lineages comprising about 4,000 species, which in addition to
23700-499: The spotless starling's range had been expanding northward since the 1950s. The low rate of advance, about 4.7 km (2.9 mi) per year for both species, is due to the suboptimal mountain and woodland terrain. Expansion has since slowed even further due to direct competition between the two similar species where they overlap in southwestern France and northwestern Spain. Major declines in populations have been observed from 1980 onward in Sweden, Finland, northern Russia ( Karelia ) and
23858-532: The state. The wide and arid Nullarbor Plain provides a natural barrier and control measures have been adopted that have killed 55,000 birds over three decades. The common starling has also colonised Kangaroo Island , Lord Howe Island , Norfolk Island and Tasmania . The early settlers in New Zealand cleared the bush and found their newly planted crops were invaded by hordes of caterpillars and other insects deprived of their previous food sources. Native birds were not habituated to living in close proximity to man so
24016-404: The unwary. Common starlings are hosts to a wide range of parasites. A survey of three hundred common starlings from six US states found that all had at least one type of parasite; 99% had external fleas, mites or ticks, and 95% carried internal parasites, mostly various types of worm. Blood-sucking species leave their host when it dies, but other external parasites stay on the corpse. A bird with
24174-462: The world. This dispersal of the human species is suggested by linguistic, cultural and genetic evidence. During the 11th millennium BP , pottery was independently invented in West Africa, with the earliest pottery there dating to about 9,400 BC from central Mali. It spread throughout the Sahel and southern Sahara . After the Sahara became a desert, it did not present a totally impenetrable barrier for travelers between north and south because of
24332-423: Was a major source of gold for the ancient world. Nubians built famous structures and numerous pyramids. Sudan, the site of ancient Nubia, has more pyramids than anywhere else in the world. The Axumite Empire spanned the southern Sahara, south Arabia and the Sahel along the western shore of the Red Sea . Located in northern Ethiopia and Eritrea , Aksum was deeply involved in the trade network between India and
24490-432: Was a medieval kingdom (c. 1250–1629), which existed between the Zambezi and Limpopo rivers of Southern Africa in the territory of modern-day Zimbabwe and Mozambique . Its old capital was located at Great Zimbabwe . In 1487, Bartolomeu Dias became the first European to reach the southernmost tip of Africa. In 1652, a victualling station was established at the Cape of Good Hope by Jan van Riebeeck on behalf of
24648-441: Was drying up, with savanna encroaching on forested areas. This occurred 10 million to 5 million years ago. By 3 million years ago several australopithecine hominid species had developed throughout Southern , East , and Central Africa . They were tool users rather than tool manufacturers. The next major evolutionary step occurred around 2.3 million BCE, when primitive stone tools were used to scavenge
24806-504: Was estimated to be 310 million individuals in 2004, occupying a total area of 8,870,000 km (3,420,000 sq mi). Widespread throughout the Northern Hemisphere, the bird is native to Eurasia and is found throughout Europe, northern Africa (from Morocco to Egypt ), India (mainly in the north but regularly extending farther south and extending into the Maldives) Nepal , the Middle East including Israel , Syria , Iran , and Iraq , and northwestern China. Common starlings in
24964-587: Was speculation. About the same date, the Portland Song Bird Club released 35 pairs of common starlings in Portland, Oregon. Earlier introductions are recorded to have died out within a few years, with the 1890 New York and Portland introductions reported as being the most successful. Population of the birds is estimated to have grown to 150 million, occupying an area extending from southern Canada and Alaska to Central America. The common starling appears to have arrived in Fiji in 1925 on Ono-i-lau and Vatoa islands. It may have colonised from New Zealand via Raoul in
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