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40-643: Epachthosaurus (meaning "heavy lizard") was a genus of titanosaurian sauropod dinosaur from the Late Cretaceous . It was a basal lithostrotian titanosaur. Its fossils have been found in Central and Northern Patagonia in South America . The type species , E. sciuttoi , was described by Powell in 1990. The bones assigned to it by Powell in 1990 were, originally, assigned to Antarctosaurus sp. , and then to Argyrosaurus superbus ? , before being named as

80-466: A basal titanosaur. Malawisaurus Paludititan Lohuecotitan Epachthosaurus Alamosaurus Opisthocoelicaudia Neuquensaurus Rocasaurus Saltasaurus Lirainosaurus Atsinganosaurus Ampelosaurus Bonatitan Rapetosaurus Nemegtosaurus Gondwanatitan Aeolosaurus Rinconsaurus Muyelensaurus Bonitasaura Mendozasaurus Futalognkosaurus Epachthosaurus

120-407: A genus is determined by taxonomists . The standards for genus classification are not strictly codified, so different authorities often produce different classifications for genera. There are some general practices used, however, including the idea that a newly defined genus should fulfill these three criteria to be descriptively useful: Moreover, genera should be composed of phylogenetic units of

160-643: A later homonym of a validly published name is a nomen illegitimum or nom. illeg. ; for a full list refer to the International Code of Nomenclature for algae, fungi, and plants and the work cited above by Hawksworth, 2010. In place of the "valid taxon" in zoology, the nearest equivalent in botany is " correct name " or "current name" which can, again, differ or change with alternative taxonomic treatments or new information that results in previously accepted genera being combined or split. Prokaryote and virus codes of nomenclature also exist which serve as

200-621: A long time and redescribed as new by a range of subsequent workers, or if a range of genera previously considered separate taxa have subsequently been consolidated into one. For example, the World Register of Marine Species presently lists 8 genus-level synonyms for the sperm whale genus Physeter Linnaeus, 1758, and 13 for the bivalve genus Pecten O.F. Müller, 1776. Within the same kingdom, one generic name can apply to one genus only. However, many names have been assigned (usually unintentionally) to two or more different genera. For example,

240-432: A new taxon. The holotype specimen is MACN -CH 1317, which consists of an incomplete posterior dorsal vertebra . Another specimen, the paratype MACN-CH 18689, consists of a cast of six articulated caudal vertebrae, the partial sacrum , and a fragmentary pubic peduncle from the right ilium . A nearly complete specimen referred to Epachthosaurus , UNPSJB-PV 920, was recovered during field research conducted as part of

280-409: A reference for designating currently accepted genus names as opposed to others which may be either reduced to synonymy, or, in the case of prokaryotes, relegated to a status of "names without standing in prokaryotic nomenclature". An available (zoological) or validly published (botanical) name that has been historically applied to a genus but is not regarded as the accepted (current/valid) name for

320-451: A right portion of the braincase (MMS/VBN.09.167) and a probable left pterygoid (MMS/VBN.09.158a), were described by Díez Díaz et al. (2018), and a few parts of the skull, several teeth, vertebrae from each section of the vertebral column, ribs, chevrons, shoulder and pelvic girdle elements, most of the bones of the arm, and most of the bones of the leg are also known from multiple individuals, including VBN.93.MHNA.99.21. The generic name

360-427: A taxon; however, the names published in suppressed works are made unavailable via the relevant Opinion dealing with the work in question. In botany, similar concepts exist but with different labels. The botanical equivalent of zoology's "available name" is a validly published name . An invalidly published name is a nomen invalidum or nom. inval. ; a rejected name is a nomen rejiciendum or nom. rej. ;

400-455: A total of c. 520,000 published names (including synonyms) as at end 2019, increasing at some 2,500 published generic names per year. "Official" registers of taxon names at all ranks, including genera, exist for a few groups only such as viruses and prokaryotes, while for others there are compendia with no "official" standing such as Index Fungorum for fungi, Index Nominum Algarum and AlgaeBase for algae, Index Nominum Genericorum and

440-741: Is considered to be the most basal titanosaurian known with procoelous caudal vertebrae. Below is a phylogenetical cladogram showing the position of Epachthosaurus within Titanosauria : Phuwiangosaurus Andesaurus Argyrosaurus Paralititan Janenschia Aeolosaurus Santa Rosa indet. Isisaurus Alamosaurus Opisthocoelicaudia Antarctosaurus Argentinosaurus Aegyptosaurus Epachthosaurus Nemegtosauridae Lirainosaurus Saltasauridae The cladogram below follows Mocho et al. (2019), placing Epachthosaurus within Lithostrotia, instead of

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480-583: Is derived from the Greek word "τσιγγάνος" or "αθίγγανος", both meaning "gypsy", which refers to the possible migration from east to west of the species. The specific name is named after its finding place, Velaux - La Bastide Neuve . A 2018 cladistic analysis of Titanosauria places Ampelosaurus , Atsinganosaurus , and Lirainosaurus in the new lithostrotian clade Lirainosaurinae . Díez Díaz et al. (2018) suggested that adults reached 8–12 metres (26–39 ft) long, possibly up to 14 metres (46 ft) long for

520-596: Is discouraged by both the International Code of Zoological Nomenclature and the International Code of Nomenclature for algae, fungi, and plants , there are some five thousand such names in use in more than one kingdom. For instance, A list of generic homonyms (with their authorities), including both available (validly published) and selected unavailable names, has been compiled by the Interim Register of Marine and Nonmarine Genera (IRMNG). The type genus forms

560-739: Is known from the early Late Cretaceous of the Bajo Barreal Formation . Other fauna from the formation include the basal chelid turtles Bonapartemys and Prochelidella , the abelisaurid Xenotarsosaurus , and an unidentified carnotaurine abelisaurid . The Bajo Barreal Formation dates back to the late Cenomanian and early Turonian of the Cretaceous . Other genera that lived alongside Epachthosaurus are Secernosaurus , Notohypsilophodon , Drusilasaura , Campylodoniscus , Aniksosaurus , and Sarmientosaurus . [REDACTED] [REDACTED] Genus The composition of

600-460: Is somewhat arbitrary. Although all species within a genus are supposed to be "similar", there are no objective criteria for grouping species into genera. There is much debate among zoologists about whether enormous, species-rich genera should be maintained, as it is extremely difficult to come up with identification keys or even character sets that distinguish all species. Hence, many taxonomists argue in favor of breaking down large genera. For instance,

640-474: Is the type species , and the generic name is permanently associated with the type specimen of its type species. Should the specimen turn out to be assignable to another genus, the generic name linked to it becomes a junior synonym and the remaining taxa in the former genus need to be reassessed. In zoological usage, taxonomic names, including those of genera, are classified as "available" or "unavailable". Available names are those published in accordance with

680-635: The Grès à Reptiles Formation of the Aix-en-Provence Basin. The type and only species is A. velauciensis . Between 2009 and 2010, excavations were carried out at the Grès à Reptiles Formation at Velaux-La Bastide Neuve by the Palaios Association and the University of Poitiers . During the excavations, the holotype of Atsinganosaurus , VBN.93.01.a-d, a set of four posterior dorsal vertebrae,

720-621: The International Code of Zoological Nomenclature ; the earliest such name for any taxon (for example, a genus) should then be selected as the " valid " (i.e., current or accepted) name for the taxon in question. Consequently, there will be more available names than valid names at any point in time; which names are currently in use depending on the judgement of taxonomists in either combining taxa described under multiple names, or splitting taxa which may bring available names previously treated as synonyms back into use. "Unavailable" names in zoology comprise names that either were not published according to

760-799: The International Plant Names Index for plants in general, and ferns through angiosperms, respectively, and Nomenclator Zoologicus and the Index to Organism Names for zoological names. Totals for both "all names" and estimates for "accepted names" as held in the Interim Register of Marine and Nonmarine Genera (IRMNG) are broken down further in the publication by Rees et al., 2020 cited above. The accepted names estimates are as follows, broken down by kingdom: The cited ranges of uncertainty arise because IRMNG lists "uncertain" names (not researched therein) in addition to known "accepted" names;

800-419: The nomenclature codes , which allow each species a single unique name that, for animals (including protists ), plants (also including algae and fungi ) and prokaryotes ( bacteria and archaea ), is Latin and binomial in form; this contrasts with common or vernacular names , which are non-standardized, can be non-unique, and typically also vary by country and language of usage. Except for viruses ,

840-404: The platypus belongs to the genus Ornithorhynchus although George Shaw named it Platypus in 1799 (these two names are thus synonyms ) . However, the name Platypus had already been given to a group of ambrosia beetles by Johann Friedrich Wilhelm Herbst in 1793. A name that means two different things is a homonym . Since beetles and platypuses are both members of the kingdom Animalia,

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880-442: The base for higher taxonomic ranks, such as the family name Canidae ("Canids") based on Canis . However, this does not typically ascend more than one or two levels: the order to which dogs and wolves belong is Carnivora ("Carnivores"). The numbers of either accepted, or all published genus names is not known precisely; Rees et al., 2020 estimate that approximately 310,000 accepted names (valid taxa) may exist, out of

920-446: The form "author, year" in zoology, and "standard abbreviated author name" in botany. Thus in the examples above, the genus Canis would be cited in full as " Canis Linnaeus, 1758" (zoological usage), while Hibiscus , also first established by Linnaeus but in 1753, is simply " Hibiscus L." (botanical usage). Each genus should have a designated type , although in practice there is a backlog of older names without one. In zoology, this

960-727: The generic name (or its abbreviated form) still forms the leading portion of the scientific name, for example, Canis lupus lupus for the Eurasian wolf subspecies, or as a botanical example, Hibiscus arnottianus ssp. immaculatus . Also, as visible in the above examples, the Latinised portions of the scientific names of genera and their included species (and infraspecies, where applicable) are, by convention, written in italics . The scientific names of virus species are descriptive, not binomial in form, and may or may not incorporate an indication of their containing genus; for example,

1000-465: The hyposphene; a strongly developed intraprezygapophyseal lamina, and processes projecting laterally from the dorsal portion of the spinodiapophyseal lamina; hyposphene-hypantrum articulations in caudals 1–14; and a pedal phalangeal formula of 2-2-3-2-0. The genus shares the following apomorphies with various titanosaurians: caudal vertebrae with ventrally expanded posterior centrodiapophyseal laminae; six sacral vertebrae; an ossified ligament or tendon above

1040-628: The largest component, with 23,236 ± 5,379 accepted genus names, of which 20,845 ± 4,494 are angiosperms (superclass Angiospermae). By comparison, the 2018 annual edition of the Catalogue of Life (estimated >90% complete, for extant species in the main) contains currently 175,363 "accepted" genus names for 1,744,204 living and 59,284 extinct species, also including genus names only (no species) for some groups. The number of species in genera varies considerably among taxonomic groups. For instance, among (non-avian) reptiles , which have about 1180 genera,

1080-481: The lizard genus Anolis has been suggested to be broken down into 8 or so different genera which would bring its ~400 species to smaller, more manageable subsets. Atsinganosaurus Atsinganosaurus is a genus of titanosaurian sauropod dinosaur which existed in what is now France during the Late Cretaceous . Well-preserved remains (and the only known) of Atsinganosaurus were collected from

1120-403: The most (>300) have only 1 species, ~360 have between 2 and 4 species, 260 have 5–10 species, ~200 have 11–50 species, and only 27 genera have more than 50 species. However, some insect genera such as the bee genera Lasioglossum and Andrena have over 1000 species each. The largest flowering plant genus, Astragalus , contains over 3,000 species. Which species are assigned to a genus

1160-428: The name could not be used for both. Johann Friedrich Blumenbach published the replacement name Ornithorhynchus in 1800. However, a genus in one kingdom is allowed to bear a scientific name that is in use as a generic name (or the name of a taxon in another rank) in a kingdom that is governed by a different nomenclature code. Names with the same form but applying to different taxa are called "homonyms". Although this

1200-748: The project Los vertebrados de la Formación Bajo Barreal, Provincia de Chubut, Patagonia, Argentina by researchers from the Laboratorio de Paleontologia de Vertebrados of the Universidad Nacional de la Patagonia. The specimen, which is one of the most complete titanosaurian skeletons known, consists of a complete, well-preserved, and articulated skeleton only missing the skull, neck, four or five cranial dorsal vertebrae, and several distal caudals. The autapomorphies that distinguish Epachthosaurus from other genera are: middle and caudal dorsal vertebrae with unique articular processes extending ventrolaterally from

1240-526: The provisions of the ICZN Code, e.g., incorrect original or subsequent spellings, names published only in a thesis, and generic names published after 1930 with no type species indicated. According to "Glossary" section of the zoological Code, suppressed names (per published "Opinions" of the International Commission of Zoological Nomenclature) remain available but cannot be used as the valid name for

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1280-474: The sacral neural spines; procoelous proximal, middle, and distal caudal centra with well-developed distal articular condyles; semilunar sternal plates with cranioventral ridges; humeri with squared proximolateral margins and proximolateral processes; unossified carpals; greatly reduced manual phalanges; nearly horizontal, craniolaterally expanded iliac preacetabular processes; pubes proximodistally longer than ischia; and transversely expanded ischia. Epachthosaurus

1320-466: The same kind as other (analogous) genera. The term "genus" comes from Latin genus , a noun form cognate with gignere ('to bear; to give birth to'). The Swedish taxonomist Carl Linnaeus popularized its use in his 1753 Species Plantarum , but the French botanist Joseph Pitton de Tournefort (1656–1708) is considered "the founder of the modern concept of genera". The scientific name (or

1360-408: The scientific epithet) of a genus is also called the generic name ; in modern style guides and science, it is always capitalised. It plays a fundamental role in binomial nomenclature , the system of naming organisms , where it is combined with the scientific name of a species : see Botanical name and Specific name (zoology) . The rules for the scientific names of organisms are laid down in

1400-497: The specific name particular to the wolf. A botanical example would be Hibiscus arnottianus , a particular species of the genus Hibiscus native to Hawaii. The specific name is written in lower-case and may be followed by subspecies names in zoology or a variety of infraspecific names in botany . When the generic name is already known from context, it may be shortened to its initial letter, for example, C. lupus in place of Canis lupus . Where species are further subdivided,

1440-412: The standard format for a species name comprises the generic name, indicating the genus to which the species belongs, followed by the specific epithet, which (within that genus) is unique to the species. For example, the gray wolf 's scientific name is Canis lupus , with Canis ( Latin for 'dog') being the generic name shared by the wolf's close relatives and lupus (Latin for 'wolf') being

1480-403: The taxon is termed a synonym ; some authors also include unavailable names in lists of synonyms as well as available names, such as misspellings, names previously published without fulfilling all of the requirements of the relevant nomenclatural code, and rejected or suppressed names. A particular genus name may have zero to many synonyms, the latter case generally if the genus has been known for

1520-566: The values quoted are the mean of "accepted" names alone (all "uncertain" names treated as unaccepted) and "accepted + uncertain" names (all "uncertain" names treated as accepted), with the associated range of uncertainty indicating these two extremes. Within Animalia, the largest phylum is Arthropoda , with 151,697 ± 33,160 accepted genus names, of which 114,387 ± 27,654 are insects (class Insecta). Within Plantae, Tracheophyta (vascular plants) make up

1560-429: The virus species " Salmonid herpesvirus 1 ", " Salmonid herpesvirus 2 " and " Salmonid herpesvirus 3 " are all within the genus Salmonivirus ; however, the genus to which the species with the formal names " Everglades virus " and " Ross River virus " are assigned is Alphavirus . As with scientific names at other ranks, in all groups other than viruses, names of genera may be cited with their authorities, typically in

1600-410: Was discovered along with the remains of Garrigatitan , another titanosaurian. Atsinganosaurus was first described by Géraldine Garcia, Sauveur Amico, Francois Fournier, Eudes Thouand and Xavier Valentin in 2010 , and the type species is A. velauciensis . The holotype now resides in the collection of the University of Poitiers . More specimens, including an occipital condyle (MMS/VBN.09.41),

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