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66-404: The park encompasses Cerro del Cimatario, a volcanic mountain which reaches 2400 meters elevation. The mountain offers panoramic views of Santiago de Querétaro and the surrounding countryside, and is home to several radio and television transmitting towers. Soils in the park are mostly volcanic in origin, formed during Pliocene eruptions. A red porous volcanic rock called tezontle is common, and

132-469: A monotreme , appeared. The predatory South American phorusrhacids were rare in this time; among the last was Titanis , a large phorusrhacid that migrated to North America and rivaled mammals as top predator. Other birds probably evolved at this time, some modern (such as the genera Cygnus , Bubo , Struthio and Corvus ), some now extinct. Alligators and crocodiles died out in Europe as

198-406: A large neocortex relative to body mass and stone tool manufacture. Improvements in dating methods and in the use of climate proxies have provided scientists with the means to test hypotheses of the evolution of human ancestors. Early hypotheses of the evolution of human traits emphasized the selective pressures produced by particular habitats. For example, many scientists have long favored

264-429: A particular habitat. The change to a cooler, drier, more seasonal climate had considerable impacts on Pliocene vegetation, reducing tropical species worldwide. Deciduous forests proliferated, coniferous forests and tundra covered much of the north, and grasslands spread on all continents (except Antarctica). Eastern Africa in particular saw a huge expansion of C 4 grasslands. Tropical forests were limited to

330-551: A series of continental glaciations in the Quaternary Period that followed. In ICS terminology, from upper (later, more recent) to lower (earlier): The Pliocene Epoch is subdivided into two ages: The Miocene Epoch is subdivided into six ages: In different geophysical regions of the world, other regional names are also used for the same or overlapping ages and other timeline subdivisions. The terms Neogene System (formal) and Upper Tertiary System (informal) describe

396-953: A tight band around the equator, and in addition to dry savannahs , deserts appeared in Asia and Africa. Both marine and continental faunas were essentially modern, although continental faunas were a bit more primitive than today. The land mass collisions meant great migration and mixing of previously isolated species, such as in the Great American Interchange . Herbivores got bigger, as did specialized predators. In North America, rodents , large mastodons and gomphotheres , and opossums continued successfully, while hoofed animals ( ungulates ) declined, with camel , deer , and horse all seeing populations recede. Three-toed horses ( Nannippus ), oreodonts , protoceratids , and chalicotheres became extinct. Borophagine dogs and Agriotherium became extinct, but other carnivores including

462-448: Is composed of trees from 4 to 12 meters high which typically lose their leaves during the dry season. Multi-trunked trees are common, and several species have bright-colored exfoliating bark. Typical dry forest tree species in the park include Bursera fagaroides , Bursera palmeri , Cedrela dugesii , Ipomoea murucoides, Erythrina coralloides , Lysiloma divaricatum , Senna polyantha , and Zanthoxylum fagara . The dry forest understory

528-475: Is due to the comparatively fine divisibility of time units as time approaches the present, and due to geological preservation that causes the youngest sedimentary geological record to be preserved over a much larger area and to reflect many more environments than the older geological record. By dividing the Cenozoic Era into three (arguably two) periods ( Paleogene , Neogene, Quaternary ) instead of seven epochs,

594-484: Is mostly herbs, including Rivina humilis . Climbing vines are common, and the epiphytes Tillandsia recurvata and Tillandsia calothyrsus grow in the tree canopy. Induced pasture grows in disturbed areas, including areas that were overgrazed by livestock and quarried for tezontle . Grasses up to 30 cm high are the predominant plants, with Melinis repens , Chloris gayana , Cynodon dactylon , Bromus carinatus , Setaria grisebachii , and Sporobolus atrovirens

660-675: Is often seen as an analogous climate to the projected climate of the near future as a result of anthropogenic global warming . Towards the end of the Pliocene, decreased heat transport towards the Antarctic resulting from a weakening of the Indonesian Throughflow (ITF) cooled the Earth, a process that exacerbated itself in a positive feedback as sea levels dropped and the ITF diminished and further limited

726-651: Is the most widespread plant community, found mostly in lower-elevation areas of the park. Shrubs up to four meters in height are the characteristic plants. Common cactus species include Myrtillocactus geometrizans , Opuntia leucotricha , Opuntia imbricata and, Nyctocereus serpentinus . Small trees, which typically have small leaves and spines, include Acacia schaffneri , Acacia farnesiana , Ipomoea mucuroides , Karwinskia humboldtiana , Anisacanthus quadrifidus , Anisacanthus pumilus , Calliandra eriophylla , Condalia velutina , Croton ciliatoglandulifer , and Zaluzania augusta var. rzedowskii . Tropical dry forest

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792-748: Is the second and most recent epoch of the Neogene Period in the Cenozoic Era . The Pliocene follows the Miocene Epoch and is followed by the Pleistocene Epoch. Prior to the 2009 revision of the geologic time scale, which placed the four most recent major glaciations entirely within the Pleistocene, the Pliocene also included the Gelasian Stage, which lasted from 2.59 to 1.81 Ma, and is now included in

858-567: The Hardangervidda plateau to 1200 m in the Early Pliocene. In Southern Sweden similar movements elevated the South Swedish highlands leading to a deflection of the ancient Eridanos river from its original path across south-central Sweden into a course south of Sweden. The Pliocene is bookended by two significant events in the evolution of human ancestors. The first is the appearance of

924-471: The International Commission on Stratigraphy (ICS) stages is not established. During the Pliocene epoch (5.3 to 2.6 million years ago (Ma)), the Earth's climate became cooler and drier, as well as more seasonal, marking a transition between the relatively warm Miocene to the cooler Pleistocene . However, the beginning of the Pliocene was marked by an increase in global temperatures relative to

990-676: The Isthmus of Panama during the Pliocene, making possible the Great American Interchange and bringing a nearly complete end to South America's distinctive native ungulate fauna, though other South American lineages like its predatory mammals were already extinct by this point and others like xenarthrans continued to do well afterwards. The formation of the Isthmus had major consequences on global temperatures, since warm equatorial ocean currents were cut off and an Atlantic cooling cycle began, with cold Arctic and Antarctic waters decreasing temperatures in

1056-745: The Paleogene . The Early Miocene was relatively cool; Early Miocene mid-latitude seawater and continental thermal gradients were already very similar to those of the present. During the Middle Miocene , Earth entered a warm phase known as the Middle Miocene Climatic Optimum (MMCO), which was driven by the emplacement of the Columbia River Basalt Group . Around 11 Ma, the Middle Miocene Warm Interval gave way to

1122-489: The hominin Australopithecus anamensis in the early Pliocene, around 4.2 million years ago. The second is the appearance of Homo , the genus that includes modern humans and their closest extinct relatives, near the end of the Pliocene at 2.6 million years ago. Key traits that evolved among hominins during the Pliocene include terrestrial bipedality and, by the end of the Pliocene, encephalized brains (brains with

1188-634: The savannah hypothesis . This proposes that the evolution of terrestrial bipedality and other traits was an adaptive response to Pliocene climate change that transformed forests into more open savannah . This was championed by Grafton Elliot Smith in his 1924 book, The Evolution of Man , as "the unknown world beyond the trees", and was further elaborated by Raymond Dart as the killer ape theory . Other scientists, such as Sherwood L. Washburn , emphasized an intrinsic model of hominin evolution. According to this model, early evolutionary developments triggered later developments. The model placed little emphasis on

1254-493: The weasel family diversified, and dogs and short-faced bears did well. Ground sloths , huge glyptodonts , and armadillos came north with the formation of the Isthmus of Panama. The latitudinal diversity gradient among terrestrial North American mammals became established during this epoch some time after 4 Ma. In Eurasia rodents did well, while primate distribution declined. Elephants , gomphotheres and stegodonts were successful in Asia (the largest land mammals of

1320-594: The Austrian palaeontologist Moritz Hörnes (1815–1868). The earlier term Tertiary Period was used to define the span of time now covered by Paleogene and Neogene and, despite no longer being recognized as a formal stratigraphic term , "Tertiary" still sometimes remains in informal use. During this period, mammals and birds continued to evolve into modern forms, while other groups of life remained relatively unchanged. The first humans ( Homo habilis ) appeared in Africa near

1386-545: The Cretaceous and the early Cenozoic . This may have contributed to further cooling of the oceans worldwide. The Pliocene seas were alive with sea cows , seals , sea lions , sharks and whales . Neogene The Neogene ( / ˈ n iː . ə dʒ iː n / NEE -ə-jeen , ) is a geologic period and system that spans 20.45 million years from the end of the Paleogene Period 23.03 million years ago ( Mya ) to

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1452-566: The Late Miocene and Pliocene, indicating increased aridification. Further aridification of Central Asia was caused by the development of Northern Hemisphere glaciation during the Late Pliocene. A sediment core from the northern South China Sea shows an increase in dust storm activity during the middle Pliocene. The South Asian Summer Monsoon (SASM) increased in intensity after 2.95 Ma, likely because of enhanced cross-equatorial pressure caused by

1518-497: The Miocene-Pliocene boundary. The first hominins , the ancestors of humans, may have appeared in southern Europe and migrated into Africa. The first humans (belonging to the species Homo habilis ) appeared in Africa near the end of the period. About 20 million years ago gymnosperms in the form of some conifer and cycad groups started to diversify and produce more species due to the changing conditions. In response to

1584-689: The Neogene and the Pliocene end at 2.58 Ma, that the Gelasian be transferred to the Pleistocene, and the Quaternary be recognized as the third period in the Cenozoic, citing key changes in Earth's climate, oceans, and biota that occurred 2.58 Ma and its correspondence to the Gauss-Matuyama magnetostratigraphic boundary . In 2006 ICS and INQUA reached a compromise that made Quaternary a sub-era, subdividing Cenozoic into

1650-761: The Piacenzian and Gelasian together) stages. As usual in stratigraphy, there are many other regional and local subdivisions in use. In Britain , the Pliocene is divided into the following stages (old to young): Gedgravian, Waltonian , Pre-Ludhamian, Ludhamian, Thurnian, Bramertonian or Antian, Pre-Pastonian or Baventian, Pastonian and Beestonian . In the Netherlands the Pliocene is divided into these stages (old to young): Brunssumian C, Reuverian A, Reuverian B, Reuverian C, Praetiglian , Tiglian A, Tiglian B, Tiglian C1-4b, Tiglian C4c, Tiglian C5, Tiglian C6 and Eburonian . The exact correlations between these local stages and

1716-570: The Pleistocene glacial-interglacial cycles. During the late Pliocene and early Pleistocene, 3.6 to 2.6 Ma, the Arctic was much warmer than it is at the present day (with summer temperatures some 8 °C warmer than today). That is a key finding of research into a lake-sediment core obtained in Eastern Siberia, which is of exceptional importance because it has provided the longest continuous late Cenozoic land-based sedimentary record thus far. During

1782-417: The Pleistocene. As with other older geologic periods, the geological strata that define the start and end are well-identified but the exact dates of the start and end of the epoch are slightly uncertain. The boundaries defining the Pliocene are not set at an easily identified worldwide event but rather at regional boundaries between the warmer Miocene and the relatively cooler Pleistocene. The upper boundary

1848-546: The Pliocene Warm Interval (PWI), interrupting the longer-term cooling trend. The Pliocene Thermal Maximum (PTM) occurred between 3.3 and 3.0 Ma. During the Pliocene, Green Sahara phases of wet conditions in North Africa were frequent and occurred about every 21 kyr, being especially intense when Earth's orbit's eccentricity was high. The PWI had similar levels of atmospheric carbon dioxide to contemporary times and

1914-429: The Pliocene may have accelerated on the disappearance of forests and the spread of grasslands and savannas. During the Pliocene the earth climate system response shifted from a period of high frequency-low amplitude oscillation dominated by the 41,000-year period of Earth's obliquity to one of low-frequency, high-amplitude oscillation dominated by the 100,000-year period of the orbital eccentricity characteristic of

1980-503: The Pliocene were such proboscideans as Deinotherium , Anancus , and Mammut borsoni , ) though proboscidean diversity declined significantly during the Late Pliocene. Hyraxes migrated north from Africa. Horse diversity declined, while tapirs and rhinos did fairly well. Bovines and antelopes were successful; some camel species crossed into Asia from North America. Hyenas and early saber-toothed cats appeared, joining other predators including dogs, bears, and weasels. Africa

2046-533: The Western Atlantic, assemblages of bivalves exhibited remarkable stasis with regards to their basal metabolic rates throughout the various climatic changes of the Pliocene. The Pliocene was a high water mark for species diversity among Caribbean corals. From 5 to 2 Ma, coral species origination rates were relatively high in the Caribbean, although a noticeable extinction event and drop in diversity occurred at

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2112-633: The Zanclean until around 3.5 Ma. During the Piacenzian, from about 3.5 to 2.6 Ma, the region was forested at irregular intervals and contained a significant Saharan palaeoriver until 3.35 Ma, when trade winds began to dominate over fluvial transport of pollen. Around 3.26 Ma, a strong aridification event that was followed by a return to more humid conditions, which was itself followed by another aridification around 2.7 Ma. From 2.6 to 2.4 Ma, vegetation zones began repeatedly shifting latitudinally in response to glacial-interglacial cycles. The climate of eastern Africa

2178-532: The atmosphere, C 4 plants expanded and reached ecological dominance in grasslands during the last 10 million years. Also Asteraceae (daisies) went through a significant adaptive radiation . Eucalyptus fossil leaves occur in the Miocene of New Zealand, where the genus is not native today, but have been introduced from Australia. The Neogene traditionally ended at the end of the Pliocene Epoch, just before

2244-573: The beginning of the present Quaternary Period 2.58 million years ago. It is the second period of the Cenozoic and the eleventh period of the Phanerozoic . The Neogene is sub-divided into two epochs , the earlier Miocene and the later Pliocene . Some geologists assert that the Neogene cannot be clearly delineated from the modern geological period, the Quaternary . The term "Neogene" was coined in 1853 by

2310-718: The climate cooled. Venomous snake genera continued to increase as more rodents and birds evolved. Rattlesnakes first appeared in the Pliocene. The modern species Alligator mississippiensis , having evolved in the Miocene, continued into the Pliocene, except with a more northern range; specimens have been found in very late Miocene deposits of Tennessee . Giant tortoises still thrived in North America, with genera like Hesperotestudo . Madtsoid snakes were still present in Australia. The amphibian order Allocaudata became extinct. In

2376-490: The cooler Messinian . This increase was related to the 1.2 million year obliquity amplitude modulation cycle . By 3.3-3.0 Ma, during the Mid-Piacenzian Warm Period (mPWP), global average temperature was 2–3 °C higher than today, while carbon dioxide levels were the same as today (400 ppm). Global sea level was about 25 m higher, though its exact value is uncertain. The northern hemisphere ice sheet

2442-459: The cooler, seasonal climate, tropical plant species gave way to deciduous ones and grasslands replaced many forests. Grasses therefore greatly diversified, and herbivorous mammals evolved alongside it, creating the many grazing animals of today such as horses , antelope , and bison . Ice age mammals like the mammoths and woolly rhinoceros were common in Pliocene . With lower levels of CO 2 in

2508-475: The early-mid Pliocene (5 Ma – 3 Ma), after three-million-year intervals with modern or glacial ice volume became longer and collapse occurs only at times when warmer global temperature coincide with strong austral summer insolation anomalies. Continents continued to drift , moving from positions possibly as far as 250 km from their present locations to positions only 70 km from their current locations. South America became linked to North America through

2574-631: The end of the period. Some continental movements took place, the most significant event being the connection of North and South America at the Isthmus of Panama , late in the Pliocene. This cut off the warm ocean currents from the Pacific to the Atlantic Ocean, leaving only the Gulf Stream to transfer heat to the Arctic Ocean . The global climate cooled considerably throughout the Neogene, culminating in

2640-506: The end of this interval. Oceans continued to be relatively warm during the Pliocene, though they continued cooling. The Arctic ice cap formed , drying the climate and increasing cool shallow currents in the North Atlantic. Deep cold currents flowed from the Antarctic. The formation of the Isthmus of Panama about 3.5 million years ago cut off the final remnant of what was once essentially a circum-equatorial current that had existed since

2706-543: The first time since the Cretaceous , with North American rodents and primates mixing with southern forms. Litopterns and the notoungulates , South American natives, were mostly wiped out, except for the macrauchenids and toxodonts , which managed to survive. Small weasel-like carnivorous mustelids , coatis and short-faced bears migrated from the north. Grazing glyptodonts , browsing giant ground sloths and smaller caviomorph rodents , pampatheres , and armadillos did

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2772-460: The heat transported southward by the Leeuwin Current . By the end of the period the first of a series of glaciations of the current Ice Age began. Marine and continental flora and fauna have a modern appearance. The reptile group Choristodera went extinct in the early part of the period, while the amphibians known as Allocaudata disappeared at the end of it. Neogene also marked the end of

2838-413: The influence of seasonal variation. More recent research has emphasized the variability selection hypothesis, which proposes that variability in climate fostered development of hominin traits. Improved climate proxies show that the Pliocene climate of east Africa was highly variable, suggesting that adaptability to varying conditions was more important in driving hominin evolution than the steady pressure of

2904-460: The late Zanclean, Italy remained relatively warm and humid. Central Asia became more seasonal during the Pliocene, with colder, drier winters and wetter summers, which contributed to an increase in the abundance of C 4 plants across the region. In the Loess Plateau , δ13C values of occluded organic matter increased by 2.5% while those of pedogenic carbonate increased by 5% over the course of

2970-507: The most common species, along with scattered shrubs of Eysenhardtia polystachya . There are also reforested areas, which include many introduced species from elsewhere in Mexico and around the world. Pliocene The Pliocene ( / ˈ p l aɪ . ə s iː n , ˈ p l aɪ . oʊ -/ PLY -ə-seen, PLY -oh- ; also Pleiocene ) is the epoch in the geologic time scale that extends from 5.33 to 2.58 million years ago (Ma). It

3036-477: The much cooler Late Miocene. The ice caps on both poles began to grow and thicken, a process enhanced by positive feedbacks from increased formation of sea ice. Between 7 and 5.3 Ma, a decrease in global temperatures termed the Late Miocene Cooling (LMC) ensued, driven by decreases in carbon dioxide concentrations. During the Pliocene, from about 5.3 to 2.7 Ma, another warm interval occurred, being known as

3102-724: The now-separated Atlantic Ocean. Africa 's collision with Europe formed the Mediterranean Sea , cutting off the remnants of the Tethys Ocean . The border between the Miocene and the Pliocene is also the time of the Messinian salinity crisis . During the Late Pliocene, the Himalayas became less active in their uplift, as evidenced by sedimentation changes in the Bengal Fan . The land bridge between Alaska and Siberia ( Beringia )

3168-623: The official timescale of the ICS , the Pliocene is subdivided into two stages . From youngest to oldest they are: The Piacenzian is sometimes referred to as the Late Pliocene, whereas the Zanclean is referred to as the Early Pliocene. In the system of In the Paratethys area (central Europe and parts of western Asia) the Pliocene contains the Dacian (roughly equal to the Zanclean) and Romanian (roughly equal to

3234-595: The old classical Tertiary and Quaternary, a compromise that was rejected by International Union of Geological Sciences because it split both Neogene and Pliocene in two. Following formal discussions at the 2008 International Geological Congress in Oslo, Norway, the ICS decided in May 2009 to make the Quaternary the youngest period of the Cenozoic Era with its base at 2.58 Mya and including

3300-503: The older definition of the beginning of the Quaternary Period; many time scales show this division. However, there was a movement amongst geologists (particularly marine geologists ) to also include ongoing geological time (Quaternary) in the Neogene, while others (particularly terrestrial geologists) insist the Quaternary to be a separate period of distinctly different record. The somewhat confusing terminology and disagreement amongst geologists on where to draw what hierarchical boundaries

3366-504: The onset of continental glaciation in the Arctic, suggesting that vegetation changes in Australia during the Pliocene behaved similarly to during the Late Pleistocene and were likely characterised by comparable cycles of aridity and humidity. The equatorial Pacific Ocean sea surface temperature gradient was considerably lower than it is today. Mean sea surface temperatures in the east were substantially warmer than today but similar in

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3432-400: The opposite, migrating to the north and thriving there. The marsupials remained the dominant Australian mammals, with herbivore forms including wombats and kangaroos , and the huge Diprotodon . Carnivorous marsupials continued hunting in the Pliocene, including dasyurids , the dog-like thylacine and cat-like Thylacoleo . The first rodents arrived in Australia. The modern platypus ,

3498-616: The periods are more closely comparable to the duration of periods in the Mesozoic and Paleozoic Eras. The International Commission on Stratigraphy (ICS) once proposed that the Quaternary be considered a sub-era (sub-erathem) of the Neogene, with a beginning date of 2.58 Ma, namely the start of the Gelasian Stage . In the 2004 proposal of the ICS, the Neogene would have consisted of the Miocene and Pliocene Epochs. The International Union for Quaternary Research (INQUA) counterproposed that

3564-776: The reorganisation of the Indonesian Throughflow. In the south-central Andes , an arid period occurred from 6.1 to 5.2 Ma, with another occurring from 3.6 to 3.3 Ma. These arid periods are coincident with global cold periods, during which the position of the Southern Hemisphere westerlies shifted northward and disrupted the South American Low Level Jet, which brings moisture to southeastern South America. From around 3.8 Ma to about 3.3 Ma, North Africa experienced an extended humid period. In northwestern Africa, tropical forests extended up to Cape Blanc during

3630-501: The reptilian genera Langstonia and Barinasuchus , terrestrial predators that were the last surviving members of Sebecosuchia , a group related to crocodiles. The oceans were dominated by large carnivores like megalodons and livyatans , and 19 million years ago about 70% of all pelagic shark species disappeared. Mammals and birds continued to be the dominant terrestrial vertebrates, and took many forms as they adapted to various habitats. An explosive radiation of ursids took place at

3696-610: The rocks deposited during the Neogene Period . The continents in the Neogene were very close to their current positions. The Isthmus of Panama formed, connecting North and South America . The Indian subcontinent continued to collide with Asia , forming the Himalayas . Sea levels fell, creating land bridges between Africa and Eurasia and between Eurasia and North America. The global climate became more seasonal and continued an overall drying and cooling trend which began during

3762-469: The scene. Bears, dogs and weasels (originally from North America) joined cats, hyenas and civets as the African predators, forcing hyenas to adapt as specialized scavengers. Most mustelids in Africa declined as a result of increased competition from the new predators, although Enhydriodon omoensis remained an unusually successful terrestrial predator. South America was invaded by North American species for

3828-400: The surrounding environment. Anthropologists tended to focus on intrinsic models while geologists and vertebrate paleontologists tended to put greater emphasis on habitats. Alternatives to the savanna hypothesis include the woodland/forest hypothesis, which emphasizes the evolution of hominins in closed habitats, or hypotheses emphasizing the influence of colder habitats at higher latitudes or

3894-495: The west. This condition has been described as a permanent El Niño state, or “El Padre.” Several mechanisms have been proposed for this pattern, including increased tropical cyclone activity . The extent of the West Antarctic Ice Sheet oscillated at the 40 kyr period of Earth's obliquity. Ice sheet collapse occurred when the global average temperature was 3 °C warmer than today and carbon dioxide concentration

3960-506: Was at 400 ppmv. This resulted in open waters in the Ross Sea . Global sea-level fluctuation associated with ice-sheet collapse was probably up to 7 meters for the west Antarctic and 3 meters for the east Antarctic. Model simulations are consistent with reconstructed ice-sheet oscillations and suggest a progression from a smaller to a larger West Antarctic ice sheet in the last 5 million years. Intervals of ice sheet collapse were much more common in

4026-480: Was dominated by hoofed animals, and primates continued their evolution, with australopithecines (some of the first hominins ) and baboon-like monkeys such as the Dinopithecus appearing in the late Pliocene. Rodents were successful, and elephant populations increased. Cows and antelopes continued diversification and overtook pigs in numbers of species. Early giraffes appeared. Horses and modern rhinos came onto

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4092-477: Was ephemeral before the onset of extensive glaciation over Greenland that occurred in the late Pliocene around 3 Ma. The formation of an Arctic ice cap is signaled by an abrupt shift in oxygen isotope ratios and ice-rafted cobbles in the North Atlantic and North Pacific Ocean beds. Mid-latitude glaciation was probably underway before the end of the epoch. The global cooling that occurred during

4158-452: Was first flooded near the start of the Pliocene, allowing marine organisms to spread between the Arctic and Pacific Oceans. The bridge would continue to be periodically flooded and restored thereafter. Pliocene marine formations are exposed in northeast Spain , southern California , New Zealand , and Italy . During the Pliocene parts of southern Norway and southern Sweden that had been near sea level rose. In Norway this rise elevated

4224-467: Was quarried to make bricks and construct baths and bread ovens. The climate is semi-arid and subtropical to temperate. Rainfall averages 549.3 mm and is seasonal, with a rainy season from June through September. Average annual temperature 18 to 19º C, with a maximum of 22º C in May. El Cimatario National Park includes portions of the Bajío dry forests and Central Mexican matorral ecoregions . Cactus scrub

4290-505: Was set at the start of the Pleistocene glaciations. Charles Lyell (later Sir Charles) gave the Pliocene its name in Principles of Geology (volume 3, 1833). The word pliocene comes from the Greek words πλεῖον ( pleion , "more") and καινός ( kainos , "new" or "recent") and means roughly "continuation of the recent", referring to the essentially modern marine mollusc fauna. In

4356-541: Was very similar to what it is today. Unexpectedly, the expansion of grasslands in eastern Africa during this epoch appears to have been decoupled from aridification and not caused by it, as evidenced by their asynchrony. Southwestern Australia hosted heathlands , shrublands , and woodlands with a greater species diversity compared to today during the Middle and Late Pliocene. Three different aridification events occurred around 2.90, 2.59, and 2.56 Ma, and may have been linked to

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