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Ecuadorian capuchin

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108-406: The Ecuadorian capuchin ( Cebus aequatorialis ), or Ecuadorian white-fronted capuchin is a species of gracile capuchin monkey of the family Cebidae . It was formerly classified as a subspecies of the white-fronted capuchin ( C. albifrons ).  Mittermeier and Rylands elevated it to a separate species in 2013. The primary physical distinction between C. albifrons and C. aequatorialis

216-473: A 2008 referendum , a new constitution implemented leftist reforms. In December 2008, Correa declared Ecuador's national debt illegitimate, based on the argument that it was odious debt contracted by prior corrupt and despotic regimes. He announced that the country would default on over $ 3 billion worth of bonds, and he succeeded in reducing the price of outstanding bonds by more than 60% by fighting creditors in international courts . He brought Ecuador into

324-598: A developing country whose economy is highly dependent on exports of commodities, primarily petroleum and agricultural products. The country is a founding member of the United Nations , Organization of American States , Mercosur , PROSUR , and the Non-Aligned Movement . According to the Center for Economic and Policy Research , between 2006 and 2016, poverty decreased from 36.7% to 22.5% and annual per capita GDP growth

432-501: A Plan to return to the constitutional system through universal elections. This plan enabled the new democratically elected president to assume the duties of the executive office. Elections were held on 29 April 1979, under a new constitution. Jaime Roldós Aguilera was elected president, garnering over one million votes, the most in Ecuadorian history. He took office on 10 August as the first constitutionally elected president, after nearly

540-652: A brief war erupted between Colombia and Peru, over Peru's claims to the Caquetá region, which ended with Peru reluctantly signing the Salomon-Lozano Treaty on 24 March 1922. Ecuador protested this secret treaty, since Colombia gave away Ecuadorian claimed land to Peru that Ecuador had given to Colombia in 1916. On 21 July 1924, the Ponce-Castro Oyanguren Protocol was signed between Ecuador and Peru where both agreed to hold direct negotiations and to resolve

648-408: A civil war. The army stationed north headed by Atahualpa marched south to Cuzco and massacred the royal family associated with his brother. In 1532, a small band of Spaniards headed by Francisco Pizarro reached Cajamarca and lured Atahualpa into a trap ( battle of Cajamarca ). Pizarro promised to release Atahualpa if he made good his promise of filling a room full of gold. But, after a mock trial,

756-524: A decade of civilian and military dictatorships. In 1980, he founded the Partido Pueblo, Cambio y Democracia (People, Change, and Democracy Party) after withdrawing from the Concentración de Fuerzas Populares (Popular Forces Concentration). He governed until 24 May 1981, when he died, along with his wife and the minister of defense Marco Subia Martinez , when his Air Force plane crashed in heavy rain near

864-499: A high priority for conservation. Through the combination of the IUCN’s map and the study of Campos and Jack, the Ecuadorian capuchin has lost over 90% of its habitat. The Ecuadorian Capuchin habitat faces various threats from habitat fragmentation , deforestation , and anthropogenic factors (farmers burning bushes, grazing , and illegal mining/ logging ). Habitat fragmentation is a severe issue for primates as it restricts movement, there

972-417: A probe. White-fronted capuchins have been observed using leaves as a cup to drink water. Intrasexual selection, or male-male competition, occurs when males invoke contests in order to gain the opportunity to reproduce with a female and maximize their reproductive success. Often males are adorned with weaponry, which can be used in order to increase their chances of winning contests for possible mates. In

1080-597: A range of 5-20. In an observational study, Campos and Jack found that female capuchins outnumbered the males and that there is a high immature to adult female proportion. New behaviour has been recorded for the Ecuadorian Capuchin, Guerrero-Casado et al. state that this species was found in a new location of Ecuador within low tree cover which is unusual behaviour, as this species prefers high tree cover. From this study, results showed that during fall/winter when trees lost their leaves and when fruits have fallen, it forces

1188-550: A range that is disjoint from the other gracile capuchins, living in northern Brazil within the states of Pará and Maranhão . The only species to inhabit the Caribbean islands is the Trinidad white-fronted capuchin. Some gracile capuchins are known to use tools . These include white-headed capuchins rubbing secretions from leaves over their bodies, using leaves as gloves when rubbing fruit or caterpillar secretions and using tools as

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1296-480: A rapid succession of rulers. The first president of Ecuador was the Venezuelan-born Juan José Flores , who was ultimately deposed. Leaders who followed him included Vicente Rocafuerte ; José Joaquín de Olmedo ; José María Urbina ; Diego Noboa ; Pedro José de Arteta ; Manuel de Ascásubi ; and Flores's own son, Antonio Flores Jijón , among others. The conservative Gabriel García Moreno unified

1404-430: A separate species. The gracile capuchins, like all capuchins, are members of the family Cebidae , which also includes the squirrel monkeys. The evolution of the squirrel monkeys and capuchin monkeys is believed to have diverged about 13 million years ago. According to genetic studies led by Lynch Alfaro in 2011, the gracile and robust capuchins diverged approximately 6.2 million years ago. Lynch Alfaro suspects that

1512-535: A series of peaceful treaties, and Peru after a short war in which the Protocol of Rio de Janeiro was signed in 1942. During the struggle for independence , before Peru or Ecuador became independent, areas of the former Vice Royalty of New Granada declared themselves independent from Spain. A few months later, a part of the Peruvian liberation army of San Martín decided to occupy the independent cities of Tumbez and Jaén, with

1620-455: Is Guayaquil . The territories of modern-day Ecuador were once home to a variety of indigenous peoples that were gradually incorporated into the Inca Empire during the 15th century. The territory was colonized by Spanish Empire during the 16th century, achieving independence in 1820 as part of Gran Colombia , from which it emerged as a sovereign state in 1830. The legacy of both empires

1728-428: Is a higher energy expenditure for the primate to find food, and leads to inbreeding or a decrease in gene flow . Over 98% of the primary forest of western Ecuador is lost and the rest is broken into fragments that are at risk for further removal. In the case of the Ecuadorian capuchin, this species thrives off large tropical dry forests and because of fragmentation it becomes harder to forage for food, as well as there

1836-440: Is a robust capuchin and is now included in the genus Sapajus . The other three Cebus species included in that taxonomy were the gracile capuchin species Cebus albifrons , Cebus nigrivittatus and the type species Cebus capucinus . Cebus nigrivittatus was subsequently renamed Cebus olivaceus . Cebus kaapori had been considered a subspecies of C. olivaceus but Groves (2001 and 2005) and Silva (2001) regarded it as

1944-492: Is an arboreal primate but spends 70% of its time on the ground, making parasite transmission easier since the ground increases the possibility of parasite transmission when larvae come into contact with the skin. Other contributing factors to higher parasite transmission were associated with monkeys drinking from a creek that was contaminated with human feces which came from a nearby village. This study and other studies have demonstrated human interaction with this species increase

2052-626: Is combined with agriculture and arboriculture . Many civilizations arose in Ecuador, such as the Valdivia Culture and Machalilla Culture on the coast, the Quitus (near present-day Quito), and the Cañari (near present-day Cuenca ). In the highland Andes mountains, where life was more sedentary, groups of tribes cooperated and formed villages; thus the first nations based on agricultural resources and

2160-453: Is less room for energy expenditure. In response to habitat fragmentation, a corridor (Three Forest Conservation Corridor) was put in to mitigate the disconnection between patches. A study by Tleimat showed that the Ecuadorian capuchin is utilizing the corridor to connect it to other fragmented forests, so similar conservation techniques can help sustain them. The forests of Ecuador and Peru are already facing rapid deforestation with only 4% of

2268-423: Is likely it would possess more canine dimorphism, like the wedge-capped capuchin, because of its similar social structure with a monopolizing alpha male and peripheral subordinate males. If a female is presented an opportunity to copulate with a male she will evaluate both the costs and benefits of that male. Females can obtain direct benefits from males she mates with, where the female gains an instant benefit from

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2376-585: Is rare in the genus Cebus , only in the white-headed capuchin is there some interaction between males and offspring. In white-headed capuchins males will often investigate, or at least tolerate, their offspring. Alpha males are also more likely to interact with their offspring than subordinate males. All gracile capuchin species except the Kaapori capuchin are rated as least concern by the International Union for Conservation of Nature . The Kaapori capuchin

2484-643: Is rated as critically endangered . Ecuador Ecuador , officially the Republic of Ecuador , is a country in northwestern South America , bordered by Colombia on the north, Peru on the east and south, and the Pacific Ocean on the west. It also includes the Galápagos Islands in the Pacific, about 1,000 kilometers (621 mi) west of the mainland. The country's capital is Quito and its largest city

2592-477: Is referred to as allomaternal care . In the wedge-capped capuchin, the mother will provide the infant care for the first three months, however for the next three months the infant relies on the care of other females. In agreement with kin selection theory , kin of the mother are more likely to provide care to the infant compared to other females in the group; siblings were four times as likely to provide infant care compared to other group females. Male parental care

2700-438: Is reflected in Ecuador's ethnically diverse population, with most of its 17.8 million people being mestizos , followed by large minorities of Europeans , Native American , African , and Asian descendants. Spanish is the official language spoken by a majority of the population, although 13 native languages are also recognized, including Quechua and Shuar . Ecuador is a representative democratic presidential republic and

2808-665: Is the most northern species, occurring in Central America from Honduras to Panama . The Colombian white-headed capuchin also has a northern distribution in Colombia and Ecuador west of the Andes . The white-fronted capuchin is found over large portions of Colombia , Peru and western Brazil , as well as into southern Venezuela and northern Bolivia . The weeper capuchin is found over much of Venezuela and over The Guianas , as well as part of northern Brazil. The Kaapori capuchin has

2916-436: Is their coloration. Due to low density and distribution researchers have not been able to make a confident molecular genetic assessment of the C. aequatorialis population, but assign it species status based on geographical isolation, morphological characteristics, and the phylogenetic species concept . The location range of the Ecuadorian Capuchin is from Western lowland Ecuador to North West Peru . The conservation status of

3024-567: The 20 km walk . Ecuador adopted the United States dollar on 13 April 2000 as its national currency and on 11 September, the country eliminated the Ecuadorian sucre , in order to stabilize the country's economy . The US Dollar has been the only official currency of Ecuador since then. The emergence of the Amerindian population as an active constituency has added to the democratic volatility of

3132-559: The Axis Powers in World War II favoring Peru with the territory they occupied at the time the war came to an end. The 1944 Glorious May Revolution followed a military-civilian rebellion and a subsequent civic strike which successfully removed Carlos Arroyo del Río as a dictator from Ecuador's government. However, a post-Second World War recession and popular unrest led to a return to populist politics and domestic military interventions in

3240-641: The Bolivarian Alliance for the Americas in June 2009. Correa's administration reduced the high levels of poverty and unemployment in Ecuador. Correa's three consecutive terms (from 2007 to 2017) were followed by his former Vice President Lenín Moreno 's four years as president (2017–21). After being elected in 2017, President Moreno's government adopted economically liberal policies, such as reduction of public spending , trade liberalization , and flexibility of

3348-658: The Cayapas of Coastal Ecuador were the only groups to resist both Inca and Spanish domination, maintaining their languages and cultures well into the 21st century. Before the arrival of the Spaniards, the Inca Empire was involved in a civil war . The untimely death of both the heir Ninan Cuyochi and the Emperor Huayna Capac, from a European disease that spread into Ecuador, created a power vacuum between two factions and led to

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3456-508: The Guayas and Manabí provinces of Ecuador, this area is chosen due to having a greater extent of continuous and/or undisturbed forest, and the success rate of conservation is favourable as it already has some kind of protection already. There has been a long history of capuchin and human relationships as humans kept them in captivity , like zoos , because of their highly amicable social behaviours and manipulative skills. In addition to captivity,

3564-724: The Spanish Empire and the Portuguese Empire . Moreover, to add legitimacy to his claims, on 16 February 1840, Flores signed a treaty with Spain, whereby Flores convinced Spain to officially recognize Ecuadorian independence and its sole rights to colonial titles over Spain's former colonial territory known anciently to Spain as the Kingdom and Presidency of Quito. Ecuador during its long and turbulent history has lost most of its contested territories to each of its more powerful neighbors, such as Colombia in 1832 and 1916, Brazil in 1904 through

3672-742: The Treaty of Guayaquil (1829) which Peru reluctantly signed, after the overwhelmingly outnumbered Gran Colombian force led by Antonio José de Sucre defeated President and General La Mar's Peruvian invasion force in the Battle of Tarqui . In addition, Ecuador's eastern border with the Portuguese colony of Brazil in the Amazon Basin was modified before the Wars of Independence by the First Treaty of San Ildefonso (1777) between

3780-556: The Tumbes and Piura regions of northern Peru. Campos and Jack’s study revealed that there is 5208km2 of suitable habitat that should be conserved for the wellbeing of the Ecuadorian capuchin and other species, further conservation should focus on the Ecuadorian and Tumbes-Piura tropical dry forests. The area deemed the most important to conserve for the Ecuadorian capuchin is located along the Chongón-Colonche range of coastal mountains in

3888-471: The Viceroyalty of Peru and later the Viceroyalty of New Granada . The 1797 Riobamba earthquake , which caused up to 40,000 casualties, was studied by Alexander von Humboldt , when he visited the area in 1801–1802. After nearly 300 years of Spanish rule, Quito still remained small with a population of 10,000 people. On 10 August 1809, the city's criollos called for independence from Spain (first among

3996-544: The "base of epiphytic bromeliad leaves because they are tender" (p. 181). However the most important foods to the Ecuadorian capuchin diet is mainly composed of fruits and invertebrates . Suitable habitat for the Ecuadorian Capuchin is determined to be a high percentage of tree cover (most important factor and maximized when over 25%), land cover (as deciduous , evergreen broad leaved, or forested cropland mosaic), mild temperature seasonality, low annual precipitation , and low human population density. According to Tleimat,

4104-575: The 1960s, while foreign companies developed oil resources in the Ecuadorian Amazon. In 1972, construction of the Andean pipeline was completed. The pipeline brought oil from the east side of the Andes to the coast, making Ecuador South America's second largest oil exporter. In 1978, the city of Quito and the Galápagos Islands were inscribed as UNESCO World Heritage Sites , making the first two properties in

4212-665: The Caqueta river and the Marañon-Amazon river. Peru ignored these protests and created the Department of Loreto in 1853 with its capital in Iquitos. Peru briefly occupied Guayaquil again in 1860, since Peru thought that Ecuador was selling some of the disputed land for development to British bond holders, but returned Guayaquil after a few months. The border dispute was then submitted to Spain for arbitration from 1880 to 1910, but to no avail. In

4320-614: The Caquetá river. Later, Ecuador contended that the Republic of Colombia, while reorganizing its government, unlawfully made its eastern border provisional and that Colombia extended its claims south to the Napo River because it said that the Government of Popayán extended its control all the way to the Napo River. When Ecuador seceded from the Gran Colombia, Peru contested Ecuador's claims with

4428-537: The Cedula of 1802 an ecclesiastical instrument, which had nothing to do with political borders. Peru began its de facto occupation of disputed Amazonian territories, after it signed a secret 1851 peace treaty in favor of Brazil. This treaty disregarded Spanish rights that were confirmed during colonial times by a Spanish-Portuguese treaty over the Amazon regarding territories held by illegal Portuguese settlers. Peru began occupying

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4536-814: The Colombian government's representatives an agreement was reached and the Muñoz Vernaza-Suarez Treaty was signed 15 July 1916, in which Colombian rights to the Putumayo river were recognized as well as Ecuador's rights to the Napo river and the new border was a line that ran midpoint between those two rivers. In this way, Ecuador gave up the claims it had to the Amazonian territories between the Caquetá River and Napo River to Colombia, thus cutting itself off from Brazil. Later,

4644-629: The District of the South as far back as April 1830. Moreover, the Cauca region, throughout its long history, had very strong economic and cultural ties with the people of Ecuador. Also, the Cauca region, which included such cities as Pasto , Popayán , and Buenaventura , had always been dependent on the Presidencia or Audiencia of Quito. Fruitless negotiations continued between the governments of Bogotá and Quito, where

4752-450: The Ecuadorian Capuchin requires about 500ha to use their energy expenditure. The dispersal of the Ecuadorian capuchin has been restricted due to deforestation and habitat fragmentation, as a result the distribution range used to be from western Ecuador to northwestern Peru . As a result of Hurtado et al.'s study they state the Ecuadorian capuchin has a low encounter rate with low density through their distribution, and this should make them

4860-483: The Ecuadorian Capuchin to come down to forage on the ground. Another reason for this new behaviour can originate from a decrease in food sources from up in trees or due to habitat fragmentation. Habitat fragmentation and environmental factors are forcing the Ecuadorian capuchin to change its behaviours as a coping mechanism to survive. The Ecuadorian Capuchin has a varied diet, it includes insects, fruits, and other invertebrate species. In addition, they are known to eat

4968-542: The Ecuadorian Capuchin was originally near threatened but was revised in 2008 by the IUCN to critically endangered due to the population's rapid decline. Anthropogenic factors such as habitat fragmentation from rapid deforestation , creation of agricultural lands, and persecution from farmers are to blame for the species' critically endangered status. Ecuadorian Capuchins mainly live in multi-male, multi-female social groups. Social groups tend to hold around 13 individuals, with

5076-568: The Ecuadorian border troops. Finally, on 23 July 1941, the Peruvians launched a major invasion, crossing the Zarumilla river in force and advancing into the Ecuadorian province of El Oro . During the course of the Ecuadorian–Peruvian War , Peru gained control over part of the disputed territory and some parts of the province of El Oro, and some parts of the province of Loja , demanding that

5184-404: The Ecuadorian capuchin is highly targeted for animal trafficking which contributes to an immediate stop to its gene flow . A negative relationship between humans and the Ecuadorian capuchin is the threat from farmers as the capuchin forages their crops, the common crops that they forage are corn, bananas, plantain and cacao. Martin et al. studied captive and free-ranging groups of capuchins in

5292-587: The Ecuadorian government give up its territorial claims. The Peruvian Navy blocked the port of Guayaquil , almost cutting all supplies to the Ecuadorian troops. After a few weeks of war and under pressure by the United States and several Latin American nations, all fighting came to a stop. Ecuador and Peru came to an accord formalized in the Rio Protocol , signed on 29 January 1942, in favor of hemispheric unity against

5400-484: The Ecuadorian military shot down Peruvian aircraft and helicopters and Peruvian infantry marched into southern Ecuador. Each country blamed the other for the onset of hostilities, known as the Cenepa War . Sixto Durán Ballén , the Ecuadorian president, famously declared that he would not give up a single centimeter of Ecuador. Popular sentiment in Ecuador became strongly nationalistic against Peru: graffiti could be seen on

5508-541: The Gran Colombian federation on 13 May 1830. After Ecuador's separation, the Department of Cauca voluntarily decided to unite itself with Ecuador due to instability in the central government of Bogota. The Venezuelan born President of Ecuador, the general Juan José Flores , with the approval of the Ecuadorian congress annexed the Department of Cauca on 20 December 1830, since the government of Cauca had called for union with

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5616-468: The Pacific Ocean coastline. Even though their languages were unrelated, these groups developed similar groups of cultures, each based in different environments. The people of the coast combined agriculture with fishing, hunting, and gathering; the people of the highland Andes developed a sedentary agricultural way of life; and peoples of the Amazon basin relied on hunting and gathering; in some cases, this

5724-672: The Pacific Ocean known as Tumbes . In Ecuador's southern Andes Mountain region where the Marañon cuts across, Ecuador had claims to an area it called Jaén de Bracamoros . These areas were included as part of the territory of Gran Colombia by Bolivar on 17 December 1819, during the Congress of Angostura when the Republic of Gran Colombia was created. Tumbes declared itself independent from Spain on 17 January 1821, and Jaén de Bracamoros on 17 June 1821, without any outside help from revolutionary armies. However, that same year, Peruvian forces participating in

5832-473: The Peruvian border. Many people believe that he was assassinated by the CIA, given the multiple death threats against him because of his reformist agenda, the deaths in automobile crashes of two key witnesses before they could testify during the investigation, and the sometimes contradictory accounts of the incident. Roldos was immediately succeeded by Vice President Osvaldo Hurtado. In 1984, León Febres Cordero from

5940-454: The Putomayo and Marañon Rivers. In July 1941, troops were mobilized in both countries. Peru had an army of 11,681 troops who faced a poorly supplied and inadequately armed Ecuadorian force of 2,300, of which only 1,300 were deployed in the southern provinces. Hostilities erupted on 5 July 1941, when Peruvian forces crossed the Zarumilla river at several locations, testing the strength and resolve of

6048-665: The Social Christian Party was elected president. Rodrigo Borja Cevallos of the Democratic Left (Izquierda Democrática, or ID) party won the presidency in 1988, winning the runoff election against Abdalá Bucaram (brother in law of Jaime Roldos and founder of the Ecuadorian Roldosist Party). His government was committed to improving human rights protection and carried out some reforms, notably an opening of Ecuador to foreign trade. The Borja government negotiated

6156-547: The Spaniards executed Atahualpa by strangulation. New infectious diseases such as smallpox , endemic to the Europeans, caused high fatalities among the Amerindian population during the first decades of Spanish rule, as they had no immunity . At the same time, the natives were forced into the encomienda labor system for the Spanish. In 1563, Quito became the seat of a real audiencia (administrative district) of Spain and part of

6264-536: The Spanish Royalist forces at the Battle of Pichincha , near Quito . Following the battle, Ecuador joined Simón Bolívar 's Republic of Gran Colombia , also including modern-day Colombia , Venezuela , and Panama . In 1830, Ecuador separated from Gran Colombia and became an independent republic. Two years later, it annexed the Galapagos Islands . The 19th century was marked by instability for Ecuador with

6372-450: The Trujillo revolution occupied both Jaén and Tumbes. Peruvian generals, without any legal titles backing them up and with Ecuador still federated with the Gran Colombia, had the desire to annex Ecuador to the Republic of Peru at the expense of the Gran Colombia, feeling that Ecuador was once part of the Inca Empire. On 28 July 1821, Peruvian independence was proclaimed in Lima by San Martín, and Tumbes and Jaén, which were included as part of

6480-437: The United States of America) ruled that the border of the undelineated zone was to be set at the line of the Cordillera del Cóndor . While Ecuador had to give up its decades-old territorial claims to the eastern slopes of the Cordillera, as well as to the entire western area of Cenepa headwaters, Peru was compelled to give to Ecuador, in perpetual lease but without sovereignty, 1 km (0.39 sq mi) of its territory, in

6588-399: The Western Amazon, Ecuador to determine how they are affected by gastrointestinal parasites . The results indicated that 6 parasites were detected across 26 animals where Strongyloides were the most prevalent parasite. According to their study, both groups were in contact with humans who provided food to the primates which forced the primates to spend more time on the ground. This species

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6696-399: The Zarumilla River, war broke out with Peru. Peru claimed that Ecuador's military presence in Peruvian-claimed territory was an invasion; Ecuador, for its part, claimed that Peru had recently invaded Ecuador around the Zarumilla River and that Peru since Ecuador's independence from Spain has systematically occupied Tumbez, Jaén, and most of the disputed territories in the Amazonian Basin between

6804-406: The aforementioned study, and the ITIS classifies it as a subspecies of C. albifrons , also due to debate over the aforementioned study. Philip Hershkovitz and William Charles Osman Hill published taxonomies of the capuchin monkeys in 1949 and 1960, respectively. These taxonomies established four species of capuchin monkey in the genus Cebus . One of those species, Cebus apella ,

6912-420: The amount of resource competition and increase the amount of male protection for her offspring. Capuchin infants are born in an altricial state, which means they need a lot of parental care in order to survive. The majority of parental care in the genus Cebus is provided by the mother, but in the case of the wedge-capped capuchin, parental care is also provided by other conspecific females; this type of care

7020-422: The area where the Ecuadorian base of Tiwinza – focal point of the war – had been located within Peruvian soil and which the Ecuadorian Army held during the conflict. The final border demarcation came into effect on 13 May 1999, and the multi-national MOMEP (Military Observer Mission for Ecuador and Peru) troop deployment withdrew on 17 June 1999. In 1972, a "revolutionary and nationalist" military junta overthrew

7128-410: The country in recent years. The population has been motivated by government failures to deliver on promises of land reform, lower unemployment and provision of social services, and the historical exploitation by the land-holding elite. Their movement, along with the continuing destabilizing efforts by both the elite and leftist movements, has led to a deterioration of the executive office. The populace and

7236-435: The country in the 1860s with the support of the Roman Catholic Church. In the late 19th century, world demand for cocoa tied the economy to commodity exports and led to migrations from the highlands to the agricultural frontier on the coast. Ecuador abolished slavery in 1851. The descendants of enslaved Ecuadorians are among today's Afro-Ecuadorian population. The Liberal Revolution of 1895 under Eloy Alfaro reduced

7344-521: The department and republic, is located only about 40 kilometers (25 mi), 1 ⁄ 4 of a degree , south of the equator. Various peoples had settled in the area of future Ecuador before the arrival of the Incas . The archeological evidence suggests that the Paleo-Indians ' first dispersal into the Americas occurred near the end of the last glacial period , around 16,500–13,000 years ago. The first people who reached Ecuador may have journeyed by land from North and Central America or by boat down

7452-431: The differences in social structure of these three groups. The alpha male of the wedge-capped capuchin tends to monopolize mating, therefore engaging in more male-male competition, while in the white-faced capuchin and in the white-fronted capuchin the alpha male does not monopolize mating and allows subordinate males to mate with females. While not much is known about the Kaapori capuchin , due to its low population size, it

7560-459: The disbanding of the small terrorist group, " ¡Alfaro Vive, Carajo! " ("Alfaro Lives, Dammit!"), named after Eloy Alfaro . However, continuing economic problems undermined the popularity of the ID party, and opposition parties gained control of Congress in 1999. A notable event was the Cenepa War fought between Ecuador and Peru in 1995. Ecuador won its first Olympic medal in the 1996 Summer Olympics in Atlanta when Jefferson Pérez won gold in

7668-469: The dispute in an equitable manner and to submit the differing points of the dispute to the United States for arbitration. Negotiations between the Ecuadorian and Peruvian representatives began in Washington on 30 September 1935. The negotiations turned into arguments during the next 7 months and finally on 29 September 1937, the Peruvian representatives decided to break off the negotiations. In 1941, amid fast-growing tensions within disputed territories around

7776-806: The divergence was triggered by the creation of the Amazon River , which separated the monkeys in the Amazon north of the Amazon River, which evolved into the gracile capuchins, from those in the Atlantic Forest south of the river, which evolved into the robust capuchins. Gracile capuchins have longer limbs relative to their body size compared with robust capuchins. Gracile capuchins also have rounder skulls and other differences in skull morphology. Gracile capuchins lack certain adaptations for opening hard nuts which robust capuchins have. These include differences in

7884-505: The domestication of animals formed. Eventually, through wars and marriage alliances of their leaders , groups of nations formed confederations. When the Incas arrived, they found that these confederations were so developed that it took the Incas two generations of rulers— Topa Inca Yupanqui and Huayna Capac —to absorb them into the Inca Empire . People belonging to the confederations that gave them

7992-478: The early part of the 20th century, Ecuador made an effort to peacefully define its eastern Amazonian borders with its neighbours through negotiation. On 6 May 1904, Ecuador signed the Tobar-Rio Branco Treaty recognizing Brazil's claims to the Amazon in recognition of Ecuador's claim to be an Amazonian country to counter Peru's earlier Treaty with Brazil back on 23 October 1851. Then after a few meetings with

8100-462: The environment and indigenous people more hostile. Moreover, when the Incas tried to subdue them, these indigenous people withdrew to the interior and resorted to guerrilla tactics. As a result, Inca expansion into the Amazon Basin and the Pacific coast of Ecuador was hampered. The indigenous people of the Amazon jungle and coastal Ecuador remained relatively autonomous until the Spanish soldiers and missionaries arrived in force. The Amazonian people and

8208-468: The female compared to other subordinate males. In the white-faced capuchin the alpha male fathers 70-90% of the offspring produced by females in his group. It is hypothesized that females are mating with alpha males while they are ovulating and then mating with subordinate males after they are no longer conceptive. Some female primates, like in the white-fronted capuchin, will mate will subordinate males while they are no longer conceptive in order to decrease

8316-592: The forest left. The study conducted by Campos and Jack revealed four important areas that should highly be conserved for the survival of the Ecuadorian capuchin. The Areas identified as the most suitable habitat are the Chongon-Colonche hills near the coast of south-central Ecuador and west of Guayaquil , the northern coast of the Manabí province in Ecuador, the foothills of the Andes Mountains in southern Ecuador, and

8424-419: The genus Cebus . At one time all capuchin monkeys were included within the genus Cebus . In 2011, Jessica Lynch Alfaro et al. proposed splitting the genus between the robust capuchin monkeys , such as the tufted capuchin , and the gracile capuchins. The gracile capuchins retain the genus name Cebus , while the robust species have been transferred to Sapajus . Following Groves (2005), taxa within

8532-579: The genus Cebus include: Subsequent revisions have split some of these into additional species: The placement of the Trinidad white-fronted capuchin is controversial; the American Society of Mammalogists classifies it as conspecific with C. brunneus based on a 2012 study later found to be flawed, while the IUCN Red List classifies it as a distinct species ( Cebus triniatis ) due to debate over

8640-451: The genus Cebus, there is a large amount of dimorphism in canine size between males and females. Canines are hypothesized to be larger in males because canine dimorphism is generally correlated to male-male competition. In the wedge-capped capuchin there is a larger amount of canine dimorphism compared to the white-faced capuchin and the white-fronted capuchin . The difference in canine dimorphism between these species can be correlated to

8748-680: The government of Bogotá did not recognize the separation of Ecuador or that of Cauca from the Gran Colombia until war broke out in May 1832. In five months, New Granada defeated Ecuador due to the fact that the majority of the Ecuadorian Armed Forces were composed of rebellious angry unpaid veterans from Venezuela and Colombia that did not want to fight against their fellow countrymen. Seeing that his officers were rebelling, mutinying, and changing sides, President Flores had no option but to reluctantly make peace with New Granada. The Treaty of Pasto of 1832

8856-710: The government of Velasco Ibarra. The coup d'état was led by General Guillermo Rodríguez and executed by navy commander Jorge Queirolo G. The new president exiled José María Velasco to Argentina. He remained in power until 1976, when he was removed by another military government. That military junta was led by Admiral Alfredo Poveda , who was declared chairman of the Supreme Council. The Supreme Council included two other members: General Guillermo Durán Arcentales and General Luis Pintado. The civil society more and more insistently called for democratic elections. Colonel Richelieu Levoyer , Government Minister, proposed and implemented

8964-425: The government restored fuel subsidies and withdrew an austerity package, which ended nearly two weeks of protests. In the 11 April 2021 election , conservative former banker Guillermo Lasso took 52.4% of the vote, compared to 47.6% for left-wing economist Andrés Aráuz , who was supported by exiled former president Correa. Lasso had finished second in the 2013 and 2017 presidential elections. On 24 May 2021, Lasso

9072-564: The government waived the right to tax increases in raw material prices and foreign exchange repatriations. In October 2018, Moreno cut diplomatic relations with the Maduro administration of Venezuela, a close ally of Correa. The relations with the United States improved significantly under Moreno. In June 2019, Ecuador agreed to allow US military planes to operate from an airport on the Galapagos Islands. In February 2020, his visit to Washington

9180-501: The intention of using them as springboards to occupy the independent city of Guayaquil and then liberate the rest of the Audiencia de Quito (Ecuador). It was common knowledge among officers of the liberation army from the south that their leader San Martín wished to liberate present-day Ecuador and add it to the future republic of Peru, since it had been part of the Inca Empire before the Spaniards conquered it. However, Bolívar 's intention

9288-480: The labour code. Ecuador also left the left-wing Bolivarian Alliance for the Americas (Alba) in August 2018. The Productive Development Act introduced an austerity policy, and reduced the previous development and redistribution policies. Regarding taxes, the authorities aimed to "encourage the return of investors" by granting amnesty to fraudsters and proposing measures to reduce tax rates for large companies . In addition,

9396-454: The male to herself. Direct benefits that would apply to females of the genus Cebus would include; vigilance from males, protection from predators and conspecifics, and increased resources. Females can also benefit indirectly from males, in the form of phenotypic and genotypic benefits to her offspring as well as male protection of those offspring. Alpha males are more fit, and therefore more likely, to provide direct and indirect benefits to

9504-572: The missionary villages in the Mainas or Maynas region, which it began calling Loreto, with its capital in Iquitos . During its negotiations with Brazil, Peru claimed Amazonian Basin territories up to Caqueta River in the north and toward the Andes Mountain range. Colombia protested stating that its claims extended south toward the Napo and Amazon Rivers. Ecuador protested that it claimed the Amazon Basin between

9612-430: The most problems were deported to distant areas of Peru, Bolivia, and north Argentina. Similarly, a number of loyal Inca subjects from Peru and Bolivia were brought to Ecuador to prevent rebellion. Thus, the region of highland Ecuador became part of the Inca Empire in 1463 sharing the same language. In contrast, when the Incas made incursions into coastal Ecuador and the eastern Amazon jungles of Ecuador, they found both

9720-531: The newly discovered Real Cedula of 1802, by which Peru claims the King of Spain had transferred these lands from the Viceroyalty of New Granada to the Viceroyalty of Peru. During colonial times this was to halt the ever-expanding Portuguese settlements into Spanish domains, which were left vacant and in disorder after the expulsion of Jesuit missionaries from their bases along the Amazon Basin. Ecuador countered by labeling

9828-549: The other branches of government give the president very little political capital, as illustrated by the most recent removal of President Lucio Gutiérrez from office by Congress in April 2005. Vice President Alfredo Palacio took his place In the election of 2006 , Rafael Correa gained the presidency. In January 2007, several left-wing political leaders of Latin America, his future allies, attended his swearing-in ceremony. Endorsed in

9936-453: The peoples of Latin America). They were led by Juan Pío Montúfar, Quiroga, Salinas, and Bishop Cuero y Caicedo. Quito's nickname, " Luz de América " ("Light of America"), is based on its leading role in trying to secure an independent, local government. Although the new government lasted no more than two months, it had important repercussions and was an inspiration for the independence movement of

10044-483: The possibility of parasite transmission. Gracile capuchin monkey Cebus aequatorialis Cebus albifrons Cebus brunneus Cebus capucinus Cebus castaneus Cebus cesarae Cebus cuscinus Cebus imitator Cebus kaapori Cebus leucocephalus Cebus malitiosus Cebus olivaceus Cebus trinitatis Cebus unicolor Cebus versicolor Cebus yuracus Gracile capuchin monkeys are capuchin monkeys in

10152-433: The power of the clergy and the conservative land owners. This liberal wing retained power until the military "Julian Revolution" of 1925. The 1930s and 1940s were marked by instability and emergence of populist politicians, such as five-time President José María Velasco Ibarra . Brasilia Presidential Act After Ecuador's separation from Colombia on 13 May 1830, its first President, General Juan José Flores , laid claim to

10260-505: The rest of Spanish America. Today, 10 August is celebrated as Independence Day, a national holiday . On 9 October 1820, the Department of Guayaquil became the first territory in Ecuador to gain its independence from Spain, and it spawned most of the Ecuadorian coastal provinces, establishing itself as an independent state. Its inhabitants celebrated what is now Ecuador's official Independence Day on 24 May 1822. The rest of Ecuador gained its independence after Antonio José de Sucre defeated

10368-417: The revolution of Trujillo by the Peruvian occupying force, had the whole region swear allegiance to the new Peruvian flag and incorporated itself into Peru. Gran Colombia had always protested Peru for the return of Jaén and Tumbes for almost a decade, then finally Bolivar after long and futile discussion over the return of Jaén, Tumbes, and part of Mainas, declared war. President and General José de La Mar , who

10476-703: The signing of the Treaty of Guayaquil in September 1829, whereby Peru and its Congress recognized Gran Colombian rights over Tumbes, Jaén, and Maynas. Through meetings between Peru and Gran Colombia, the border was set as Tumbes river in the west, and in the east, the Maranon and Amazon rivers were to be followed toward Brazil as the most natural borders between them. According to the peace negotiations Peru agreed to return Guayaquil, Tumbez, and Jaén; despite this, Peru returned Guayaquil, but failed to return Tumbes and Jaén, alleging that it

10584-485: The teeth and jaws, and the lack of a sagittal crest . Exterior differences include the fact that, although some females have tufts on their head ( Humboldt's white-fronted capuchin and Guianan weeper capuchin ), no male gracile capuchin has tufts, while all robust capuchins have tufts. Also, no gracile capuchins have beards . Gracile capuchin monkeys have a wide range over Central America and north and north-west South America . The Panamanian white-headed capuchin

10692-557: The territory that had belonged to the Real Audiencia of Quito , also referred to as the Presidencia of Quito. He supported his claims with Spanish Royal decrees, or real cedulas , that delineated the borders of Spain's former overseas colonies. In the case of Ecuador, Flores based Ecuador's de jure claims on the Real Cedulas of 1563, 1739, and 1740; with modifications in the Amazon Basin and Andes Mountains that were introduced through

10800-609: The walls of Quito referring to Peru as the " Cain de Latinoamérica ", a reference to the murder of Abel by his brother Cain in the Book of Genesis . Ecuador and Peru signed the Brasilia Presidential Act peace agreement on 26 October 1998, which ended hostilities, and effectively put an end to the Western Hemisphere's longest running territorial dispute. The Guarantors of the Rio Protocol (Argentina, Brazil, Chile, and

10908-610: The world to become listed sites. The Rio Protocol failed to precisely resolve the border along a little river in the remote Cordillera del Cóndor region in southern Ecuador. This caused a long-simmering dispute between Ecuador and Peru, which ultimately led to fighting between the two countries; first a border skirmish in January–February 1981 known as the Paquisha Incident , and ultimately full-scale warfare in January 1995 where

11016-517: The world to recognize legally enforceable rights of nature . The country's name means " Equator " in Spanish, truncated from the Spanish official name, República del Ecuador ( lit. "Republic of the Equator"), derived from the former Ecuador Department of Gran Colombia established in 1824 as a division of the former territory of the Royal Audience of Quito . Quito, which remained the capital of

11124-407: Was 1.5 percent (as compared to 0.6 percent over the prior two decades). At the same time, the country's Gini index of economic inequality improved from 0.55 to 0.47. One of 17 megadiverse countries in the world, Ecuador hosts many endemic plants and animals, such as those of the Galápagos Islands . In recognition of its unique ecological heritage, the new constitution of 2008 is the first in

11232-508: Was born in Ecuador, believing his opportunity had come to annex the District of Ecuador to Peru, personally, with a Peruvian force, invaded and occupied Guayaquil and a few cities in the Loja region of southern Ecuador on 28 November 1828. The war ended when an outnumbered southern Gran Colombian army at Battle of Tarqui on 27 February 1829, led by Antonio José de Sucre , defeated the Peruvian invasion force led by President La Mar. This defeat led to

11340-568: Was not obligated to follow the agreements, since the Gran Colombia ceased to exist when it divided itself into three different nations – Ecuador, Colombia, and Venezuela. The Central District of the Gran Colombia, known as Cundinamarca or New Granada (modern Colombia) with its capital in Bogota, did not recognize the separation of the Southern District of the Gran Colombia, with its capital in Quito, from

11448-580: Was signed by which the Department of Cauca was turned over to New Granada (modern Colombia), the government of Bogotá recognized Ecuador as an independent country and the border was to follow the Ley de División Territorial de la República de Colombia (Law of the Division of Territory of the Gran Colombia) passed on 25 June 1824. This law set the border at the river Carchi and the eastern border that stretched to Brazil at

11556-408: Was the first meeting between an Ecuadorian and U.S. president in 17 years. A series of protests began on 3 October 2019 against the end of fuel subsidies and austerity measures adopted by Moreno. On 10 October, protesters overran Quito, the capital, causing the Government of Ecuador to relocate temporarily to Guayaquil . The government eventually returned to Quito in 2019. On 14 October 2019,

11664-482: Was to form a new republic known as the Gran Colombia , out of the liberated Spanish territory of New Granada which consisted of Colombia, Venezuela, and Ecuador. San Martín's plans were thwarted when Bolívar, descended from the Andes mountains and occupied Guayaquil; they also annexed the newly liberated Audiencia de Quito to the Republic of Gran Colombia. In the south, Ecuador had claims to a small piece of land beside

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