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40-539: The Douglas fir ( Pseudotsuga menziesii ) is an evergreen conifer species in the pine family, Pinaceae . It is native to western North America and is also known as Douglas-fir , Douglas spruce , Oregon pine , and Columbian pine . There are three varieties: coast Douglas-fir ( P. menziesii var. menziesii ), Rocky Mountain Douglas-fir ( P. menziesii var. glauca ) and Mexican Douglas-fir ( P. menziesii var. lindleyana ). Despite its common names , it

80-552: A cold season or dry season, evergreen plants are usually an adaptation of low nutrient levels. Additionally, they usually have hard leaves and have an excellent water economy due to scarce resources in the area in which they reside. The excellent water economy within the evergreen species is due to high abundance when compared to deciduous species. Whereas deciduous trees lose nutrients whenever they lose their leaves. In warmer areas, species such as some pines and cypresses grow on poor soils and disturbed ground. In Rhododendron ,

120-556: A few months to several decades (over 30 years in the Great Basin bristlecone pine ). Japanese umbrella pine is unique in that it has its own family of which it is the only species. Evergreen and deciduous species vary in a range of morphological and physiological characters. Generally, broad-leaved evergreen species have thicker leaves than deciduous species, with a larger volume of parenchyma and air spaces per unit leaf area. They have larger leaf biomass per unit leaf area, and hence

160-475: A genus with many broadleaf evergreens, several species grow in mature forests but are usually found on highly acidic soil where the nutrients are less available to plants. In taiga or boreal forests , it is too cold for the organic matter in the soil to decay rapidly, so the nutrients in the soil are less easily available to plants, thus favoring evergreens. In temperate climates, evergreens can reinforce their own survival; evergreen leaf and needle litter has

200-469: A higher carbon-nitrogen ratio than deciduous leaf litter , contributing to a higher soil acidity and lower soil nitrogen content. This is the case with Mediterranean evergreen seedlings, which have unique C and N storages that allow stored resources to determine fast growth within the species, limiting competition and bolstering survival. These conditions favor the growth of more evergreens and make it more difficult for deciduous plants to persist. In addition,

240-525: A lower specific leaf area . Construction costs do not differ between the groups. Evergreens have generally a larger fraction of total plant biomass present as leaves (LMF), but they often have a lower rate of photosynthesis. Deciduous trees shed their leaves usually as an adaptation to a cold or dry/wet season. Evergreen trees also lose leaves, but each tree loses its leaves gradually and not all at once. Most tropical rainforest plants are considered to be evergreens, replacing their leaves gradually throughout

280-549: A number of specifications including kiln dried and grade stamped, and can be supplied in very long lengths to 60 feet. West coast mills are sophisticated in their processing of timbers, making lead times predictable and availability reliable. Paints adhere well to Douglas fir. Stains perform well on Douglas fir timbers with the mild caution that the natural color of this species varies and care must be taken to ensure uniformity of color. Pitch pockets that may ooze resin can be present in timbers that have not been kiln dried. Because of

320-457: A single strait trunk up to 100 cm in diameter at the base. In mature trees the trunk is often clear of branches for 2/3 of the tree's height. The crown is pyramidal in young trees, becoming rounded in older trees, and varying from dense to open. In young trees and new branches the bark is smooth and red-brown. As trees mature the bark becomes rough and scaly, and divided into large irregular plates by deep fissures. Pinus douglasiana inhabits

360-421: A tangy citrus flavor and may serve in some recipes as a wild substitute for rosemary . Evergreen There are many different kinds of evergreen plants, including trees , shrubs , and vines. Evergreens include: The Latin binomial term sempervirens , meaning "always green", refers to the evergreen nature of the plant, for instance: The longevity of individual leaves in evergreen plants varies from

400-454: Is a significant component. The red vole nests almost exclusively in the foliage of the trees, typically 2–50 metres (5–165 ft) above the ground, and its diet consists chiefly of Douglas-fir needles. Douglas-fir needles are generally poor browse for ungulates , although in the winter when other food sources are lacking it can become important, and black-tailed deer browse new seedlings and saplings in spring and summer. The spring diet of

440-592: Is also one of the most common lumber trees used in forestry alongside Radiata pine with large plantations throughout the country. The species was introduced in the 1900s for its wood. Douglas-fir prefers acidic or neutral soils. However, it exhibits considerable morphological plasticity, and on drier sites P. menziesii var. menziesii will generate deeper taproots. Pseudotsuga menziesii var. glauca exhibits even greater plasticity, occurring in stands of interior temperate rainforest in British Columbia, as well as at

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480-507: Is extensively used in forestry management as a plantation tree for softwood timber . Douglas-fir is one of the world's best timber-producing species and yields more timber than any other species in North America, making the forestlands of western Oregon, Washington, and British Columbia the most productive on the continent. In 2011, Douglas-fir represented 34.2% of US lumber exports, to a total of 1.053 billion board-feet. Douglas fir timber

520-998: Is found in the Klamath and California Coast Ranges as far south as the Santa Lucia Range , with a small stand as far south as the Purisima Hills in Santa Barbara County. One of the last remaining old growth stands of conifers is in the Mattole Watershed, and is under threat of logging. In the Sierra Nevada , it ranges as far south as the Yosemite region. It occurs from sea level along the coast to elevations of 1,500 m (4,900 ft) or higher, and inland in some cases up to 2,100 m (6,900 ft). Another variety exists further inland, Pseudotsuga menziesii var. glauca ,

560-411: Is misleading since it is not a true fir, i.e., not a member of the genus Abies . For this reason, the name is often written as Douglas-fir (a name also used for the genus Pseudotsuga as a whole). The specific epithet menziesii is after Archibald Menzies , a Scottish physician and rival naturalist to David Douglas. Menzies first documented the tree on Vancouver Island in 1791. Colloquially,

600-552: Is more shade tolerant than some associated fire-dependent species, such as western larch and ponderosa pine, and often replaces these species further inland. Fungal diseases such as laminated root rot and shoestring root-rot can cause significant damage, and in plantation settings dominated by Douglas-fir monocultures may cause extreme damage to vast swathes of trees. Interplanting with resistant or nonhost species such as western redcedar and beaked hazelnut can reduce this risk. Other threats to Douglas-fir include red ring rot and

640-530: Is not a true fir ( genus Abies ), spruce (genus Picea ), or pine (genus Pinus ). It is also not a hemlock ; the genus name Pseudotsuga means "false hemlock". Douglas-firs are medium-size to extremely large evergreen trees, 20–100 metres (70–330 feet) tall (although only coast Douglas-firs , reach heights near 100 m) and commonly reach 2.4 m (8 ft) in diameter, although trees with diameters of almost 5 metres (16 feet) exist. The largest coast Douglas-firs regularly live over 500 years, with

680-453: Is often considered a variety of P. menziesii . Fossils (wood, pollen) of Pseudotsuga are recorded from the Miocene and Pliocene of Europe ( Siebengebirge , Gleiwitz , Austria). It is also naturalised throughout Europe, Argentina and Chile (called Pino Oregón ). In New Zealand it is considered to be an invasive species, called a wilding conifer , and is subject to control measures. But

720-457: Is often found with other pines, including Pinus pseudostrobus , Pinus herrerae , Pinus leiophylla , Pinus lawsonii , and Pinus ayacahuite in the southern part of its range. At lower elevations it is sometimes found with Pinus oocarpa , and with Pinus devoniana on drier sites. In some areas oaks ( Quercus spp.) are co-dominant canopy trees, especially in areas which have been logged of pines. In high-elevation, high-rainfall sites in

760-477: Is the Douglas-fir dwarf mistletoe ( Arceuthobium douglasii ). Epiphytes such as crustose lichens and mosses are common sights on Douglas-firs. As it is only moderately shade tolerant , undisturbed Douglas-fir stands in humid areas will eventually give way to later successional, more shade-tolerant associates such as the western redcedar and western hemlock—though this process may take a thousand years or more. It

800-414: Is used for timber frame construction and timber trusses using traditional joinery, veneer , and flooring due to its strength, hardness and durability. As of 2024, the only wooden ships still currently in use by the U.S. Navy in conventional naval operations are Avenger-class minesweepers , made of Douglas-fir. Douglas fir sees wide use in heavy timber structures, as its wood is strong, available in

840-501: The Douglas-fir beetle . Many different Native American groups used the bark, resin, and needles to make herbal treatments for various diseases. Native Hawaiians built waʻa kaulua ( double-hulled canoes ) from coast Douglas-fir logs that had drifted ashore. The wood has historically been favored as firewood , especially from the coastal variety. In addition early settlers used Douglas fir for all forms of building construction, including floors, beams, and fine carving. The species

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880-533: The Lynn Valley Tree and the Nooksack Giant . The leaves are flat, soft, linear needles 2–4 centimetres ( 3 ⁄ 4 – 1 + 1 ⁄ 2  in) long, generally resembling those of the firs, occurring singly rather than in fascicles ; they completely encircle the branches, which can be useful in recognizing the species. As the trees grow taller in denser forest, they lose their lower branches, such that

920-539: The Umpqua River in Oregon. It is most dominant in areas with a more frequent fire regime that suppresses less fire-resistant conifers. Douglas-fir seeds are an extremely important food source for small mammals such as moles , shrews , and chipmunks , which consume an estimated 65% of each annual seed crop. The Douglas squirrel harvests and hoards great quantities of Douglas-fir cones, and also consumes mature pollen cones,

960-467: The blue grouse features Douglas-fir needles prominently. The leaves are also used by the woolly conifer aphid Adelges cooleyi ; this 0.5 mm-long sap -sucking insect is conspicuous on the undersides of the leaves by the small white "fluff spots" of protective wax that it produces. It is often present in large numbers, and can cause the foliage to turn yellowish from the damage it causes. Exceptionally, trees may be partially defoliated by it, but

1000-614: The Rocky Mountain Douglas-fir or interior Douglas-fir. Interior Douglas-fir intergrades with coast Douglas-fir in the Cascades of northern Washington and southern British Columbia, and from there ranges northward to central British Columbia and southeastward to the Mexican border, becoming increasingly disjunct as latitude decreases and altitude increases. Mexican Douglas-fir ( P. lindleyana ), which ranges as far south as Oaxaca ,

1040-409: The back half of a mouse, with two feet and a tail. The massive mega- genome of Douglas fir was sequenced in 2017 by the large PineRefSeq consortium, revealing a specialized photosynthetic apparatus in the light-harvesting complex of genes. The common name honors David Douglas , a Scottish botanist and collector who first reported the extraordinary nature and potential of the species. The common name

1080-593: The damage is rarely this severe. Among Lepidoptera , apart from some that feed on Pseudotsuga in general, the gelechiid moths Chionodes abella and C. periculella as well as the cone scale-eating tortrix moth Cydia illutana have been recorded specifically on P. menziesii . The inner bark is the primary winter food for the North American porcupine . Poriol is a flavanone, a type of flavonoid, produced by P. menziesii in reaction to infection by Poria weirii . A parasitic plant which uses P. menziesii

1120-884: The edge of semi-arid sagebrush steppe throughout much of its range, where it generates even deeper taproots still. The coast Douglas-fir variety is the dominant tree west of the Cascade Mountains in the Pacific Northwest . It occurs in nearly all forest types and competes well on most parent materials, aspects, and slopes. Adapted to a more moist, mild climate than the interior subspecies, it grows larger and faster than Rocky Mountain Douglas-fir. Associated trees include western hemlock , Sitka spruce , sugar pine , western white pine , ponderosa pine , grand fir , coast redwood , western redcedar , California incense-cedar , Lawson's cypress , tanoak , bigleaf maple and several others. Pure stands are also common, particularly north of

1160-421: The foliage may start as high as 34 m (110 ft) off the ground. Douglas-firs in environments with more light may have branches much closer to the ground. The bark on young trees is thin, smooth, grey, and contains numerous resin blisters. On mature trees, usually exceeding 80 years, it is very thick and corky, growing up to 36 cm (14 in) thick with distinctive, deep vertical fissures caused by

1200-559: The gradual expansion of the growing tree. Some of the mature bark is brown, while other parts are lighter colored with a cork-like texture; these develop in multiple layers. This thick bark makes the Douglas-fir one of the most fire-resistant tree native to the Pacific Northwest. The female cones are pendulous, with persistent scales, unlike those of true firs. They have distinctive long, trifid (three-pointed) bracts which protrude prominently above each scale and are said to resemble

1240-417: The inner bark, terminal shoots, and developing young needles. Mature or "old-growth" Douglas-fir forest is the primary habitat of the red tree vole ( Arborimus longicaudus ) and the spotted owl ( Strix occidentalis ). Home range requirements for breeding pairs of spotted owls are at least 400 hectares (4.0 km; 990 acres) of old growth. Red tree voles may also be found in immature forests if Douglas-fir

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1280-428: The most vulnerable aspect of many plants. Even though roots are insulated by soil, which tends to be warmer than average air temperatures, soil temperatures that drop too low can kill the plant. The exact temperature which evergreen roots can handle depends on the species, for example, Picea Glauca (White Spruce) roots are killed at −10 °F (−23 °C). In areas where there is a reason for being deciduous, e.g.

1320-483: The mountains of western and central Mexico, where it grows between 1100 and 2700 meters elevation. It is found in the southern Sierra Madre Occidental up to the crest of the mountains along the border of Sinaloa and Durango , in the western Trans-Mexican Volcanic Belt in the states of Nayarit, Jalisco , Michoacán , México and Morelos , and locally in the Sierra Madre del Sur of Guerrero and Oaxaca . It

1360-470: The northwest side of a structure. Pinus douglasiana Pinus douglasiana is a species of evergreen conifer in the family Pinaceae . It is found only in Mexico . A common name is Douglas pine , but that name is often applied to the more widespread species Pseudotsuga menziesii which is also known as Douglas fir. Pinus douglasiana is typically a tall tree, up to 45 meters in height, with

1400-457: The oldest specimens living for over 1,300 years. Rocky Mountain Douglas-firs , found further to the east, are less long-lived, usually not exceeding 400 years in age. There are records of former coast Douglas-firs exceeding 120 metres (390 feet) in height, which if alive today would make it the tallest tree species on Earth. Particular historical specimens with heights exceeding 400 feet include

1440-472: The shelter provided by existing evergreen plants can make it easier for younger evergreen plants to survive cold and/or drought. Evergreen plants can have decorative as well as functional uses. In months where most other plants are dormant, evergreens with their sturdy structure, and vibrant foliage are popular choices to beautify a landscape. Additionally, evergreens can serve as a windbreak , stopping heat loss from buildings during cold months when placed on

1480-461: The species is also known simply as Doug fir or Douglas pine (although the latter common name may also refer to Pinus douglasiana ). Other names for this tree have included Oregon pine , British Columbian pine , Puget Sound pine , Douglas spruce , false hemlock , red fir , or red pine (although again red pine may refer to a different tree species, Pinus resinosa , and red fir may refer to Abies magnifica ). One Coast Salish name for

1520-488: The timber sizes available, stamped timber grading, and relatively short lead times, Douglas fir sees wide use in both public and residential projects. The species has ornamental value in large parks and gardens. It has been commonly used as a Christmas tree since the 1920s, and the trees are typically grown on plantations. The buds have been used to flavor eau de vie , a clear, colorless fruit brandy. Douglas-fir pine leaves can be used to make pine needle tea . They possess

1560-677: The tree, used in the Halkomelem language, is lá:yelhp . In the Lushootseed language, the tree is called čəbidac . Pseudotsuga menziesii var. menziesii , the coast Douglas-fir, grows in the coastal regions from west-central British Columbia southward to central California . In Oregon and Washington , its range is continuous from the eastern edge of the Cascades west to the Pacific Coast Ranges and Pacific Ocean. In California, it

1600-518: The year as the leaves age and fall, whereas species growing in seasonally arid climates may be either evergreen or deciduous. Most warm temperate climate plants are also evergreen. In cool temperate climates, fewer plants are evergreen. In such climates, there is a predominance of conifers because few evergreen broadleaf plants can tolerate severe cold below about −26 °C (−15 °F). In addition, evergreen foliage experiences significant leaf damage in these cold, dry climates. Root systems are

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