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61-532: Colugos ( / k ə ˈ l uː ɡ oʊ / ), flying lemurs , or cobegos ( / k ə ˈ b iː ɡ oʊ / ), are arboreal gliding euarchontogliran mammals that are native to Southeast Asia . Their closest evolutionary relatives are primates . There are just two living species of colugos: the Sunda flying lemur ( Galeopterus variegatus ) and the Philippine flying lemur ( Cynocephalus volans ). These two species make up

122-457: A anomodont synapsid from Russia dating to the Late Permian , about 260 million years ago, was also likely a specialised climber. IUCN The International Union for Conservation of Nature ( IUCN ) is an international organization working in the field of nature conservation and sustainable use of natural resources . Founded in 1948, IUCN has become the global authority on

183-555: A seawall ), to adapt to and mitigate climate change effects, while improving sustainable livelihoods and protecting natural ecosystems and biodiversity. At the IUCN World Conservation Congress 2016, IUCN Members agreed on a definition of nature-based solutions. Members also called for governments to include nature-based solutions in strategies to combat climate change . Some key dates in the growth and development of IUCN: According to its website, IUCN works on

244-444: A Conservation for Development Centre within its secretariat. The Centre undertook projects to ensure that nature conservation was integrated in development aid and in the economic policies of developing countries. Over the years, it supported the development of national conservation strategies in 30 countries. Several European countries began to channel considerable amounts of bilateral aid via IUCN's projects. Management of these projects

305-402: A Malayan colugo ( Galeopterus variegatus ) individual having been observed traveling about 150 m (490 ft) in one glide. Their ability to glide is possible because of a large membrane of skin that extends between their paired limbs. This gliding membrane, or patagium , runs from the shoulder blades to the fore paws, from the tip of the rear-most fingers to the tip of the toes, and from

366-545: A branch of a tree, can create special difficulties for animals who are not adapted to deal with balancing on small diameter substrates . During locomotion on the ground, the location of the center of mass may swing from side to side. But during arboreal locomotion, this would result in the center of mass moving beyond the edge of the branch, resulting in a tendency to topple over and fall. Not only do some arboreal animals have to be able to move on branches of varying diameter, but they also have to eat on these branches, resulting in

427-415: A diagonal sequence gait . Brachiation is a specialized form of arboreal locomotion, used by primates to move very rapidly while hanging beneath branches. Arguably the epitome of arboreal locomotion, it involves swinging with the arms from one handhold to another. Only a few species are brachiators , and all of these are primates; it is a major means of locomotion among spider monkeys and gibbons , and

488-568: A formal act constituting the International Union for the Protection of Nature (IUPN). The initiative to set up the new organisation came from UNESCO and especially from its first director general, the British biologist Julian Huxley . At the time of its founding IUCN was the only international organisation focusing on the entire spectrum of nature conservation (an international organisation for

549-519: A nocturnal species, they developed night vision . Colugos spend their days resting in tree holes and are active at night time; traveling around 1.7 km at night. Colugos may also be a territorial species. Although they are placental mammals, colugos raise their young in a manner similar to marsupials . Newborn colugos are underdeveloped and weigh only 35 g (1.2 oz). They spend the first six months of life clinging to their mother's belly. The mother colugo curls her tail and folds her patagium into

610-533: A partnership with the World Business Council for Sustainable Development . IUCN renewed a multi-year MOU ( Memorandum of understanding ) with WBCSD in December 2015. In 1996, after decades of seeking to address specific business issues, IUCN's Members asked for a comprehensive approach to engaging the business sector. Resolution 1.81 of the IUCN World Conservation Congress held that year "urged IUCN Members and

671-762: A stable financial basis for its work, IUCN participated in setting up the World Wildlife Fund (1961) (now the World Wide Fund for Nature WWF) to work on fundraising to cover part of the operational costs of IUCN. Also in 1961, the IUCN headquarters moved from Belgium to Morges in Switzerland. During the 1960s, IUCN lobbied the UN General Assembly to create a new status for NGOs . Resolution 1296, adopted in 1968, granted 'consultative' status to NGOs. IUCN itself

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732-616: A variety of anatomical, behavioral and ecological consequences as well as variations throughout different species. Furthermore, many of these same principles may be applied to climbing without trees, such as on rock piles or mountains. Some animals are exclusively arboreal in habitat, such as tree snails . Arboreal habitats pose numerous mechanical challenges to animals moving in them, which have been solved in diverse ways. These challenges include moving on narrow branches, moving up and down inclines, balancing, crossing gaps, and dealing with obstructions. Moving along narrow surfaces, such as

793-509: A warm, secure, quasipouch to protect and transport her young. The young do not reach maturity until they are two to three years old. In captivity, they live up to 15 years, but their lifespan in the wild is unknown. Both species are threatened by habitat destruction , and the Philippine flying lemur was once classified by the IUCN as vulnerable. In 1996, the IUCN declared the species vulnerable owing to destruction of lowland forests and hunting. It

854-458: Is estimated the ancestors of the colugos split from other mammals about 80 million years ago, leading to the present day forms that consist of 7 to 14 extant species. The Mixodectidae and Plagiomenidae appear to be fossil Dermoptera. Although other Paleogene mammals have been interpreted as related to dermopterans, the evidence for this association is uncertain and many of the fossils are no longer interpreted as being gliding mammals. At present,

915-425: Is linked to sustainable development and poverty reduction. IUCN states that it aims to have a solid factual base for its work and takes into account the knowledge held by indigenous groups and other traditional users of natural resources. The IUCN Programme 2017–2020 identifies three priority areas: IUCN does not itself aim to directly mobilize the general public. Education has been part of IUCN's work program since

976-415: Is occasionally used by female orangutans . Gibbons are the experts of this mode of locomotion, swinging from branch to branch distances of up to 15 m (50 ft), and traveling at speeds of as much as 56 km/h (35 mph). To bridge gaps between trees, many animals such as the flying squirrel have adapted membranes, such as patagia for gliding flight . Some animals can slow their descent in

1037-543: Is to expand the global network of national parks and other protected areas and promote good management of such areas. In particular, it focuses on greater protection of the oceans and marine habitats . IUCN has a growing program of partnerships with the corporate sector on a regional, national and international level to promote sustainable use of natural resources. On the national level, IUCN helps governments prepare national biodiversity policies. Internationally, IUCN provides advice to environmental conventions such as

1098-410: Is to increase the amount of contact their limbs are making with the substrate to increase friction and braking power. Due to the height of many branches and the potentially disastrous consequences of a fall, balance is of primary importance to arboreal animals. On horizontal and gently sloped branches, the primary problem is tipping to the side due to the narrow base of support. The narrower the branch,

1159-590: The Convention on Biological Diversity , CITES , and the Framework Convention on Climate Change . It advises UNESCO on natural world heritage . It has a formally accredited permanent observer mission to the United Nations . IUCN has official relations with the multiple other international bodies. As an organization, IUCN has three components: the member organizations, the six scientific commissions and

1220-482: The United Nations , and plays a role in the implementation of several international conventions on nature conservation and biodiversity . It was involved in establishing the World Wide Fund for Nature and the World Conservation Monitoring Centre . In the past, IUCN has been criticized for placing the interests of nature over those of indigenous peoples. In recent years, its closer relations with

1281-520: The spider monkey and crested gecko , the tip of the tail has either a bare patch or adhesive pad, which provides increased friction. Claws can be used to interact with rough substrates and re-orient the direction of the force the animal applies. This is what allows squirrels to climb tree trunks that are so large as to be essentially flat, from the perspective of such a small animal. However, claws can interfere with an animal's ability to grasp very small branches, as they may wrap too far around and prick

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1342-544: The Director General, based on the need to influence private sector policies in support of the Mission of IUCN, to expand dialogue and productive relationships with the private sector and find new ways to interact with members of the business community". The IUCN Global Business and Biodiversity Program (BBP) was established in 2003 to influence and support private partners in addressing environmental and social issues. In 2004,

1403-629: The IUCN expanded its relations with UN-agencies and established links with the Council of Europe . IUCN's best known publication, the Red Data Book on the conservation status of species, was first published in 1964. IUCN began to play a part in the development of international treaties and conventions, starting with the African Convention on the Conservation of Nature and Natural Resources . Africa

1464-604: The Primatomorpha. Scandentia (treeshrews) Lagomorpha (rabbits, hares, pikas) Rodentia (rodents) Dermoptera (colugos) † Plesiadapiformes Primates [REDACTED] Arboreal Arboreal locomotion is the locomotion of animals in trees . In habitats in which trees are present, animals have evolved to move in them. Some animals may scale trees only occasionally (scansorial), but others are exclusively arboreal. The habitats pose numerous mechanical challenges to animals moving through them and lead to

1525-465: The air using a method known as parachuting, such as Rhacophorus (a " flying frog " species) that has adapted toe membranes allowing it to fall more slowly after leaping from trees. Many species of snake are highly arboreal, and some have evolved specialized musculature for this habitat. While moving in arboreal habitats, snakes move slowly along bare branches using a specialized form of concertina locomotion , but when secondary branches emerge from

1586-451: The animal's own paw. Adhesion is an alternative to claws, which works best on smooth surfaces. Wet adhesion is common in tree frogs and arboreal salamanders , and functions either by suction or by capillary adhesion. Dry adhesion is best typified by the specialized toes of geckos , which use van der Waals forces to adhere to many substrates, even glass. Frictional gripping is used by primates, relying upon hairless fingertips. Squeezing

1647-551: The branch being moved on, snakes use lateral undulation , a much faster mode. As a result, snakes perform best on small perches in cluttered environments, while limbed organisms seem to do best on large perches in uncluttered environments. The earliest known climbing tetrapod is the varanopid amniote Eoscansor from the Late Carboniferous ( Pennsylvanian ) of North America which is clearly specialised with adaptations for grasping, likely onto tree trunks. Suminia ,

1708-626: The branch between the fingertips generates a frictional force that holds the animal's hand to the branch. However, this type of grip depends upon the angle of the frictional force; thus upon the diameter of the branch, with larger branches resulting in reduced gripping ability. Animals other than primates that use gripping in climbing include the chameleon, which has mitten-like grasping feet, and many birds that grip branches in perching or moving about. To control descent, especially down large diameter branches, some arboreal animals such as squirrels have evolved highly mobile ankle joints that permit rotating

1769-425: The branch. Both pitching and tipping become irrelevant, as the only method of failure would be losing their grip. Arboreal species have behaviors specialized for moving in their habitats, most prominently in terms of posture and gait. Specifically, arboreal mammals take longer steps, extend their limbs further forwards and backwards during a step, adopt a more 'crouched' posture to lower their center of mass, and use

1830-602: The business sector have caused controversy. IUCN was established in 1948. It was initially called the International Union for the Protection of Nature (1948–1956) and has also been formerly known as the World Conservation Union (1990–2008). IUCN was established on 5 October 1948, in Fontainebleau , France, when representatives of governments and conservation organizations spurred by UNESCO signed

1891-591: The day resting. At night, colugos spend most of their time up in the trees foraging, with gliding being used to either find another foraging tree or to find possible mates and protect territory. Colugos are shy, nocturnal , solitary animals found in the tropical forests of Southeast Asia. Consequently, very little is known about their behavior. They are herbivorous and eat leaves, shoots, flowers, sap, and fruit. They have well-developed stomachs and long intestines capable of extracting nutrients from leaves and other fibrous material. As part of Colugos' evolution into

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1952-463: The early days but the focus is on stakeholder involvement and strategic communication rather than mass-campaigns. IUCN runs field projects for habitat and species conservation around the world. It produces the IUCN Red List of Threatened Species and the IUCN Red List of Ecosystems . The IUCN Red List of Ecosystems is applicable at local, national, regional, and global levels. IUCN's stated goal

2013-679: The entire family Cynocephalidae ( / ˌ s aɪ n oʊ ˌ s ɛ f ə ˈ l aɪ d i , - ˌ k ɛ -/ ) and order Dermoptera . Colugos are nocturnal, tree-dwelling mammals. They reach lengths of 35 to 40 cm (14 to 16 in) and weigh 1 to 2 kg (2.2 to 4.4 lb). They have long, slender front and rear limbs, a medium-length tail, and a relatively light build. The head is small, with large, front-focused eyes for excellent binocular vision, and small rounded ears. The incisor teeth of colugos are highly distinctive; they are comb-like in shape with up to 20 tines on each tooth. The incisors are analogous in appearance and function to

2074-793: The fact that conservation of nature by banning human presence no longer worked. The Strategy was followed in 1982 by the World Charter for Nature , which was adopted by the United Nations General Assembly , after preparation by IUCN. In 1980, IUCN and WWF moved into shared new offices in Gland, Switzerland . This marked a phase of closer cooperation with WWF, but the close ties between IUCN and WWF were severed in 1985 when WWF decided to take control of its own field projects, which so far had been run by IUCN. Sustainable development and regionalisation: 1985 to present day In 1982, IUCN set up

2135-483: The first IUCN Private Sector Engagement Strategy was developed (in response to Council Decision C/58/41). Most prominent in the Business and Biodiversity Program is the five-year collaboration IUCN started with the energy company Shell International in 2007. IUCN has been involved in minimum energy consumption and zero-carbon construction since 2005 by integrating energy-saving materials, developed by Jean-Luc Sandoz in

2196-681: The focus of power was still with the Headquarters in Gland but the regional offices and regional members' groups gradually got a bigger say in operations. In 1991, IUCN (together with UNEP and WWF) published Caring for the Earth , a successor to the World Conservation Strategy. Social aspects of conservation were now integrated in IUCN's work; at the General Assembly in 1994 the IUCN mission

2257-472: The following themes: business, climate change , economics, ecosystems , environmental law , forest conservation , gender , global policy, marine and polar , protected areas , science and knowledge, social policy , species , water, and world heritage . IUCN works on the basis of four-year programs, determined by the membership. In the IUCN Programme for 2017–2020 conserving nature and biodiversity

2318-408: The foot into a 'reversed' posture. This allows the claws to hook into the rough surface of the bark, opposing the force of gravity. Many arboreal species lower their center of mass to reduce pitching and toppling movement when climbing. This may be accomplished by postural changes, altered body proportions, or smaller size. Small size provides many advantages to arboreal species: such as increasing

2379-547: The footsteps of Julius Natterer . Today, the Business and Biodiversity Programme continues to set the strategic direction, coordinate IUCN's overall approach and provide institutional quality assurance in all business engagements. The Programme ensures that the Business Engagement Strategy is implemented through IUCN's global thematic and regional programmes as well as helps guide the work of IUCN's six Commissions. Nature-based solutions (NbS) use ecosystems and

2440-522: The force of gravity to raise its body, making the movement more difficult. To get past this difficulty, many animals have to grasp the substrate with all four limbs and increase the frequency of their gait sequence. Conversely, as the animal descends, it must also fight gravity to control its descent and prevent falling. Descent can be particularly problematic for many animals, and highly arboreal species often have specialized methods for controlling their descent. One way animals prevent falling while descending

2501-405: The form of branches emerging from the one being moved on and other branches impinging on the space the animal needs to move through. These obstructions may impede locomotion, or may be used as additional contact points to enhance it. While obstructions tend to impede limbed animals, they benefit snakes by providing anchor points. Arboreal organisms display many specializations for dealing with

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2562-629: The fossil record of definitive dermopterans is limited to two species of the Eocene and Oligocene cynocephalid genus Dermotherium . Molecular phylogenetic studies have demonstrated that colugos emerged as a basal Primatomorpha clade – which, in turn, is a basal Euarchontoglires clade. Scandentia are widely considered to be the closest relatives of Primatomorpha, within Euarchonta . Some studies, however, place Scandentia as sister of Glires ( lagomorphs and rodents ), in an unnamed sister clade of

2623-818: The greater the difficulty in balancing a given animal faces. On steep and vertical branches, tipping becomes less of an issue, and pitching backwards or slipping downwards becomes the most likely failure. In this case, large-diameter branches pose a greater challenge since the animal cannot place its forelimbs closer to the center of the branch than its hindlimbs. Some arboreal animals need to be able to move from tree to tree in order to find food and shelter. To be able to get from tree to tree, animals have evolved various adaptations. In some areas trees are close together and can be crossed by simple brachiation . In other areas, trees are not close together and animals need to have specific adaptations to jump far distances or glide. Arboreal habitats often contain many obstructions, both in

2684-454: The hind legs to the tip of the tail. The spaces between the colugo's fingers and toes are webbed. As a result, colugos were once considered to be close relatives of bats . Today, on account of genetic data, they are considered to be more closely related to primates . Colugos are unskilled climbers; they lack opposable thumbs. They progress up trees in a series of slow hops, gripping onto the bark with their small, sharp claws. They spend most of

2745-417: The incisor suite in strepsirrhines , which is used for grooming. The second upper incisors have two roots, another unique feature among mammals. The dental formula of colugos is: 2.1.2.3 3.1.2.3 Colugos are proficient gliders, and thought better adapted for flight than any other gliding mammal. They can travel as far as 70 m (230 ft) from one tree to another without losing much altitude, with

2806-579: The mechanical challenges of moving through their habitats. Arboreal animals frequently have elongated limbs that help them cross gaps, reach fruit or other resources, test the firmness of support ahead, and in some cases, to brachiate . However, some species of lizard have reduced limb size that helps them avoid limb movement being obstructed by impinging branches. Many arboreal species, such as howler monkeys , green tree pythons , emerald tree boas , chameleons , silky anteaters , spider monkeys , and possums , use prehensile tails to grasp branches. In

2867-407: The need for the ability to balance while using their hands to feed themselves. This resulted in various types of grasping such as pedal grasping in order to clamp themselves onto small branches for better balance. Branches are frequently oriented at an angle to gravity in arboreal habitats, including being vertical, which poses special problems. As an animal moves up an inclined branch, it must fight

2928-576: The organisation on relatively sound financial footing for the first time since 1948. This period saw the beginning of a gradual change in IUCN's approach to conservation in which it tried to become more appealing to the developing world. In 1975 IUCN started work on the World Conservation Strategy (1980). The drafting process, and the discussions with the UN agencies involved, led to an evolution in thinking within IUCN and growing acceptance of

2989-508: The protection of birds, now BirdLife International , had been established in 1922). IUCN (International Union for conservation of Nature) started out with 65 members in Brussels and was closely associated to UNESCO. They jointly organized the 1949 Conference on Protection of Nature Lake Success , US and drafted the first list of gravely endangered species. In the early years of its existence IUCN depended almost entirely on UNESCO funding and

3050-544: The public in support of nature conservation. It tries to influence the actions of governments, business and other stakeholders by providing information and advice and through building partnerships. The organization is best known to the wider public for compiling and publishing the IUCN Red List of Threatened Species , which assesses the conservation status of species worldwide. IUCN has a membership of over 1,400 governmental and non-governmental organizations from over 170 countries. Some 16,000 scientists and experts participate in

3111-424: The relative size of branches to the animal, lower center of mass, increased stability, lower mass (allowing movement on smaller branches), and the ability to move through more cluttered habitat. Size relating to weight affects gliding animals such as the reduced weight per snout-vent length for 'flying' frogs . Some species of primate , bat , and all species of sloth achieve passive stability by hanging beneath

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3172-447: The secretariat. IUCN Members are States, government agencies, international nongovernmental organizations, national nongovernmental organizations and indigenous peoples' organisations. In 2017, IUCN had 1400 members. The members can organize themselves in national or regional committees to promote cooperation. In 2016, there were 62 national committees and 7 regional committees. The seven IUCN Commissions involve volunteer experts from

3233-498: The services they provide to address societal challenges such as climate change, food security or natural disasters. The emergence of the NbS concept in environmental sciences and nature conservation contexts came as international organisations, such as IUCN and the World Bank, searched for solutions to work with ecosystems rather than relying on conventional engineering interventions (such as

3294-569: The status of the natural world and the measures needed to safeguard it. It is involved in data gathering and analysis , research, field projects, advocacy, and education. IUCN's mission is to "influence, encourage and assist societies throughout the world to conserve nature and to ensure that any use of natural resources is equitable and ecologically sustainable". Over the past decades, IUCN has widened its focus beyond conservation ecology and now incorporates issues related to sustainable development in its projects. IUCN does not itself aim to mobilize

3355-447: The work of IUCN commissions on a voluntary basis. It employs over 900 full-time staff in more than 50 countries. Its headquarters are in Gland , Switzerland. Every four years, IUCN convenes for the IUCN World Conservation Congress where IUCN Members set the global conservation agenda by voting on recommendations and guide the secretariat's work by passing resolutions and the IUCN Programme. IUCN has observer and consultative status at

3416-556: Was downlisted to least-concern status in 2008 but still faces the same threats. In addition to the ongoing clearing of its rainforest habitat, it is hunted for its meat and fur. It is also a favorite prey item for the critically endangered Philippine eagle ; some studies suggest colugos account for 90% of the eagle's diet. Their family name Cynocephalidae comes from the Greek words κύων kyōn "dog" and κεφαλή kephalē "head" because their heads are broad with short snouts like dogs. It

3477-776: Was eventually accredited with six UN organizations. IUCN was one of the few environmental organisations formally involved in the preparations of the United Nations Conference on the Human Environment (Stockholm, 1972). The Stockholm Conference eventually led to three new international conventions, with IUCN involved in their drafting and implementation: IUCN entered into an agreement with the United Nations Environment Programme UNEP to provide regular reviews of world conservation. The income this generated, combined with growing revenue via WWF, put

3538-548: Was forced to temporarily scale down activities when this ended unexpectedly in 1954. IUCN was successful in engaging prominent scientists and identifying important issues such as the harmful effects of pesticides on wildlife but not many of the ideas it developed were turned into action. This was caused by unwillingness to act on the part of governments, uncertainty about the IUCN mandate and lack of resources. In 1956, IUCN changed its name to International Union for Conservation of Nature and Natural Resources. During this period,

3599-410: Was primarily done by IUCN staff, often working from the new regional and country offices IUCN set up around the world. This marked a shift within the organisation. Previously, the volunteer Commissions had been very influential, now the secretariat and its staff began to play a more dominant role. In 1989, IUCN moved into a separate building in Gland, close to the offices it had shared with WWF. Initially,

3660-471: Was redrafted to its current wording to include the equitable and ecologically use of natural resources. Closer to business: 2000 to present day Since the creation of IUCN in 1948, IUCN Members have passed more than 300 resolutions that include or focus on business related activities. The increased attention on sustainable development as a means to protect nature brought IUCN closer to the corporate sector. The members decided against this, but IUCN did forge

3721-431: Was the focus of many of the early IUCN conservation field projects. IUCN supported the 'Yellowstone model' of protected area management, which severely restricted human presence and activity in order to protect nature. The IUCN also suffered from restricted financing in its early years. For this reason, Tracy Philipps , secretary-general from 1955 to 1958, did not draw a salary during his period in office. To establish

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