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Cyphophthalmi

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133-573: Cyphophthalmi is a suborder of harvestmen , colloquially known as mite harvestmen . Cyphophthalmi comprises 36 genera, and more than two hundred described species. The six families are currently grouped into three infraorders: the Boreophthalmi, Scopulophthalmi, and Sternophthalmi. Cyphophthalmi are smaller than the more familiar "daddy long-legs" harvestmen, with adults ranging from 1 to 7 mm in length. Moreover, their legs are relatively short compared to most other harvestmen, typically shorter than

266-728: A general armistice ended the fighting. In the German Revolution (November 1918), Wilhelm II and the ruling princes abdicated their positions, and Germany was declared a federal republic . Germany's new leadership signed the Treaty of Versailles in 1919, accepting defeat by the Allies . Germans perceived the treaty as humiliating, which was seen by historians as influential in the rise of Adolf Hitler . Germany lost around 13% of its European territory and ceded all of its colonial possessions in Africa and

399-451: A pre-Linnaean work). In England, the Opiliones are called harvestmen, not because they appear at that season, but from a superstitious belief that if one is killed there will be a bad harvest that year. The interfamilial relationships within Opiliones are not yet fully resolved, although significant strides have been made in recent years to determine these relationships. The following list

532-505: A sustainable energy supply, the debt brake for balanced budgets, measures to increase the fertility rate ( pronatalism ), and high-tech strategies for the transition of the German economy, summarised as Industry 4.0 . During the 2015 European migrant crisis , the country took in over a million refugees and migrants. Germany is the seventh-largest country in Europe . It borders Denmark to

665-555: A temperate climate, ranging from oceanic in the north and west to continental in the east and southeast. Winters range from the cold in the Southern Alps to cool and are generally overcast with limited precipitation, while summers can vary from hot and dry to cool and rainy. The northern regions have prevailing westerly winds that bring in moist air from the North Sea, moderating the temperature and increasing precipitation. Conversely,

798-405: A 2012 phylogenetic analysis, a new Cyphophthalmi taxonomy was proposed. The Cyphophthalmi are currently represented by the following taxa, which represent approximately 200 species, although there are a significant number of undescribed species that have been collected, and the current number of species is believed to be a severe under-representation. The monophyly of most subgroups of Cyphophthalmi

931-455: A NATO effort to provide security in that country after the ousting of the Taliban . In the 2005 elections , Angela Merkel became the first female chancellor. In 2009, the German government approved a €50 billion stimulus plan. Among the major German political projects of the early 21st century are the advancement of European integration , the energy transition ( Energiewende ) for

1064-618: A broad range is unusual in arachnids, which are typically pure predators. Most hunting harvestmen ambush their prey, although active hunting is also found. Because their eyes cannot form images, they use their second pair of legs as antennae to explore their environment. Unlike most other arachnids, harvestmen do not have a sucking stomach or a filtering mechanism. Rather, they ingest small particles of their food, thus making them vulnerable to internal parasites such as gregarines . Although parthenogenetic species do occur, most harvestmen reproduce sexually . Except from small fossorial species in

1197-638: A centralised totalitarian state , withdrew from the League of Nations , and dramatically increased the country's rearmament . A government-sponsored programme for economic renewal focused on public works, the most famous of which was the Autobahn . In 1935, the regime withdrew from the Treaty of Versailles and introduced the Nuremberg Laws which targeted Jews and other minorities. Germany also reacquired control of

1330-407: A distance. The Opiliones are known for having exceptionally long legs relative to their body size; however, some species are short-legged. As in all Arachnida, the body in the Opiliones has two tagmata , the anterior cephalothorax or prosoma , and the posterior 10-segmented abdomen or opisthosoma . The most easily discernible difference between harvestmen and spiders is that in harvestmen,

1463-599: A framework laid out in the 1949 constitution known as the Grundgesetz (Basic Law). Amendments generally require a two-thirds majority of both the Bundestag and the Bundesrat ; the fundamental principles of the constitution, as expressed in the articles guaranteeing human dignity, the separation of powers, the federal structure, and the rule of law , are valid in perpetuity. The president , currently Frank-Walter Steinmeier ,

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1596-456: A further defence from predators because they can twitch for 60 seconds to an hour after detachment. This can also potentially serve as deflection from an attack and deceive a predator from attacking the animal. It has been shown to be successful against ants and spiders. The legs continue to twitch after they are detached because 'pacemakers' are located in the ends of the first long segment (femur) of their legs. These pacemakers send signals via

1729-529: A global power in industrial, scientific and technological sectors, it is both the world's third-largest exporter and importer . As a developed country , it offers social security , a universal health care system , and tuition-free university education . Germany is a member of the United Nations , Council of Europe , NATO and OECD , and a founding member of the European Union, G7 and G20 . It has

1862-658: A joint responsibility of the international community. It was the world's second-biggest aid donor in 2019 after the United States. Germany's military, the Bundeswehr (Federal Defence), is organised into the Heer (Army and special forces KSK ), Marine (Navy), Luftwaffe (Air Force) and Cyber- und Informationsraum (Cyber and Information Domain Service) branches. In absolute terms, German military spending in 2023

1995-732: A long period. Indeed, one species discovered in China, Mesobunus martensi , fossilized by fine-grained volcanic ash around 165 million years ago, is hardly discernible from modern-day harvestmen and has been placed in the extant family Sclerosomatidae . The Swedish naturalist and arachnologist Carl Jakob Sundevall (1801–1875) honored the naturalist Martin Lister (1638–1712) by adopting Lister's term Opiliones for this order, known in Lister's days as "harvest spiders" or "shepherd spiders", from Latin opilio , "shepherd"; Lister characterized three species from England (although not formally describing them, being

2128-534: A modified patch of cuticle near the adenostyle known as the Rambla's organ. The exact function of these glands and structures is currently unknown. Like most harvestmen, Cyphophthalmi rely heavily on chemical secretions for defense. This defensive character is particularly well developed in Cyphophthalmi, when compared to the more familiar Eupnoi and Dyspnoi. Cyphophthalmi possess more elaborate musculature associated with

2261-434: A number of key ways. Like all harvestmen, they have a pair of ozopores located on the sides of the prosoma. Unlike other harvestmen, however, the ozopores in Cyphophthalmi are located on elevated cone-shaped structures known as ozophores. Although members of most families are eyeless, most members of the families Pettalidae and Stylocellidae do have eyes located on or near the ozophores. These eyes, however, are not homologous to

2394-428: A pair median eyes also have two sets of vestigial eyes: one median pair (homologous to those of horseshoe crabs and sea spiders ), and one lateral pair (homologous to facetted eyes of horseshoe crabs and insects). This discovery suggests that the neuroanatomy of harvestmen is more primitive than derived arachnid groups, like spiders and scorpions. It also showed that the four-eyed fossil harvestman previously discovered

2527-418: A predator after direct or indirect contact, including thanatosis , freezing , bobbing, autotomy , fleeing, stridulation , retaliation and chemical secretions. Some animals respond to attacks by simulating an apparent death to avoid either detection or further attacks. Arachnids such as spiders practise this mechanism when threatened or even to avoid being eaten by female spiders after mating. Thanatosis

2660-611: A predator. Armored harvestmen in Laniatores can often use their modified morphology as weapons. Many have spines on their pedipalps, back legs, or bodies. By pinching with their chelicerae and pedipalps, they can cause harm to a potential predator. Also this has been proven to increase survival against recluse spiders by causing injury, allowing the harvestman to escape from predation. Harvestmen are well known for being chemically protected. They exude strongly odored secretions from their scent glands, called ozopores , that act as

2793-537: A provisional capital, to emphasise its stance that the two-state solution was temporary. West Germany was established as a federal parliamentary republic with a " social market economy ". Starting in 1948 West Germany became a major recipient of reconstruction aid under the American Marshall Plan . Konrad Adenauer was elected the first federal chancellor of Germany in 1949. The country enjoyed prolonged economic growth ( Wirtschaftswunder ) beginning in

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2926-510: A scientifically neglected group. Description of new taxa has always been dependent on the activity of a few dedicated taxonomists. Carl Friedrich Roewer described about a third (2,260) of today's known species from the 1910s to the 1950s, and published the landmark systematic work Die Weberknechte der Erde (Harvestmen of the World) in 1923, with descriptions of all species known to that time. Other important taxonomists in this field include: Since

3059-516: A shield against predators; this is the most effective defense they use which creates a strong and unpleasant taste. In Cyphophthalmi the scent glands release naphthoquinones, chloro-naphthoquinones and aliphatic methyl ketones, Insidiatores use nitrogen-containing substances, terpenes, aliphatic ketones, and phenolics, while Grassatores use alkylated phenolics and benzoquinones, and Palpatores use substances like naphthoquinones, methyl- and ethyl-ketones. These secretions have successfully protected

3192-615: A state, the East German authorities eased border restrictions, but this actually led to an acceleration of the Wende reform process culminating in the Two Plus Four Treaty under which Germany regained full sovereignty. This permitted German reunification on 3 October 1990, with the accession of the five re-established states of the former GDR. The fall of the Wall in 1989 became a symbol of

3325-692: A symbol of the Cold War . Tensions between East and West Germany were reduced in the late 1960s by Chancellor Willy Brandt 's Ostpolitik . In 1989, Hungary decided to dismantle the Iron Curtain and open its border with Austria , causing the emigration of thousands of East Germans to West Germany via Hungary and Austria. This had devastating effects on the GDR, where regular mass demonstrations received increasing support. In an effort to help retain East Germany as

3458-515: A total population of over 82 million in an area of 357,596 km (138,069 sq mi), making it the most populous member state of the European Union . It borders Denmark to the north, Poland and the Czech Republic to the east, Austria and Switzerland to the south, and France , Luxembourg , Belgium , and the Netherlands to the west. The nation's capital and most populous city

3591-492: A variety of primary and secondary defences against predation, ranging from morphological traits such as body armour to behavioral responses to chemical secretions. Some of these defences have been attributed and restricted to specific groups of harvestmen. Primary defences help the harvestmen avoid encountering a potential predator and include crypsis , aposematism , and mimicry . Particular patterns or colour markings on harvestmen's bodies can reduce detection by disrupting

3724-637: Is Berlin and its main financial centre is Frankfurt ; the largest urban area is the Ruhr . Settlement in the territory of modern Germany began in the Lower Paleolithic , with various tribes inhabiting it from the Neolithic onward, chiefly the Celts . Various Germanic tribes have inhabited the northern parts of modern Germany since classical antiquity . A region named Germania was documented before AD 100. In 962,

3857-558: Is a compilation of interfamilial relationships recovered from several recent phylogenetic studies, although the placement and even monophyly of several taxa are still in question. The family Stygophalangiidae (one species, Stygophalangium karamani ) from underground waters in North Macedonia is sometimes misplaced in the Phalangioidea. It is not a harvestman. Despite their long history, few harvestman fossils are known. This

3990-405: Is also known from the unrelated family Sandokanidae . Like most animals, Cyphophthalmi express morphological distinctions between male and female individuals. Male Cyphophthalmi possess a structure on their fourth pair of tarsi known as an adenostyle. The adenostyle usually appears as a small hornlike projection, but can take a variety of shapes, depending on species. The function of the adenostyle

4123-409: Is currently unknown, but is presumably associated with chemical glands. Aside from the adenostyle, males in many species possess glandular pores on either the underside of the opisthosoma or the anal region, although these glands are not present in all species. Males in several families also possess structural modifications of the anal plate and males of certain species in the family Stylocellidae possess

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4256-637: Is devoted to agriculture, while 30% is forested and 14% is covered by settlements or infrastructure. Plants and animals include those generally common to Central Europe. According to the National Forest Inventory, beeches , oaks , and other deciduous trees constitute just over 40% of the forests; roughly 60% are conifers , particularly spruce and pine . There are many species of ferns , flowers , fungi , and mosses . Wild animals include roe deer , wild boar , mouflon (a subspecies of wild sheep), fox , badger , hare , and small numbers of

4389-432: Is formed by extensions of the coxae of the pedipalps and the first pair of legs. Most Opiliones, except for Cyphophthalmi, have long been thought to have a single pair of camera-type eyes in the middle of the head, oriented sideways. Eyes in Cyphophthalmi, when present, are located laterally, near the ozopores. A 305-million-year-old fossilized harvestman with two pairs of eyes was reported in 2014. This find suggested that

4522-510: Is in rainforest patches in northwest Argentina which are in an area being dramatically destroyed by humans. The cave-living Picunchenops spelaeus is apparently endangered through human action. So far, no harvestman has been included in any kind of a Red List in Argentina, so they receive no protection. Maiorerus randoi has only been found in one cave in the Canary Islands . It is included in

4655-599: Is largely autonomous in regard to its internal organisation. As of 2017 , Germany is divided into 401 districts ( Kreise ) at a municipal level; these consist of 294 rural districts and 107 urban districts . Germany has a civil law system based on Roman law with some references to Germanic law . The Bundesverfassungsgericht (Federal Constitutional Court) is the German Supreme Court responsible for constitutional matters, with power of judicial review . Germany's specialized supreme court system includes

4788-508: Is mainly due to their delicate body structure and terrestrial habitat, making them unlikely to be found in sediments. As a consequence, most known fossils have been preserved within amber . The oldest known harvestman, from the 410-million-year-old Devonian Rhynie chert, displayed almost all the characteristics of modern species, placing the origin of harvestmen in the Silurian , or even earlier. A recent molecular study of Opiliones, however, dated

4921-404: Is most likely a member of the suborder Eupnoi (true daddy-longlegs). Harvestmen have a pair of prosomatic defensive scent glands ( ozopores ) that secrete a peculiar-smelling fluid when disturbed. In some species, the fluid contains noxious quinones . They do not have book lungs , and breathe through tracheae . A pair of spiracles is located between the base of the fourth pair of legs and

5054-490: Is seen with multiple long-legged species in the Leiobunum clade that either drop and run, or drop and remain motionless. This is also seen when disturbing an aggregation of multiple individuals, where they all scatter. Multiple species within the Laniatores and Dyspnoi possess stridulating organs, which are used as intraspecific communication and have also been shown to be used as a second line of defense when restrained by

5187-627: Is the head of state and invested primarily with representative responsibilities and powers. He is elected by the Bundesversammlung (federal convention), an institution consisting of the members of the Bundestag and an equal number of state delegates. The second-highest official in the German order of precedence is the Bundestagspräsident ( President of the Bundestag ), who is elected by

5320-452: Is the voluntary amputation of an appendage and is employed to escape when restrained by a predator. Eupnoi individuals, more specifically sclerosomatid harvestmen, commonly use this strategy in response to being captured. This strategy can be costly because harvestmen do not regenerate their legs, and leg loss reduces locomotion, speed, climbing ability, sensory perception, food detection, and territoriality. Autotomised legs provide

5453-617: Is thought to be commonly used during daylight, when they could be easily seen by any predators. Other harvestmen may exhibit mimicry to resemble other species' appearances. Some Gonyleptidae individuals that produce translucid secretions have orange markings on their carapaces . This may have an aposematic role by mimicking the colouration of glandular emissions of two other quinone-producing species. Mimicry ( Müllerian mimicry ) occurring between Brazilian harvestmen that resemble others could be explained by convergent evolution . Secondary defences allow for harvestmen to escape and survive from

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5586-594: Is used as a second line of defence when detected by a potential predator and is commonly observed within the Dyspnoi and Laniatores suborders, with individuals becoming rigid with legs either retracted or stretched. Freezing – or the complete halt of movement – has been documented in the family Sclerosomatidae. While this can mean an increased likelihood of immediate survival, it also leads to reduced food and water intake. To deflect attacks and enhance escape, long-legged species – commonly known as daddy long-legs – from

5719-483: Is vested in the parliament consisting of the Bundestag (Federal Diet) and Bundesrat (Federal Council), which together form the legislative body. The Bundestag is elected through direct elections using the mixed-member proportional representation system. The members of the Bundesrat represent and are appointed by the governments of the sixteen federated states. The German political system operates under

5852-400: Is well supported, although both Boreophthalmi and Sironidae have been recovered as paraphyletic in recent analyses. Harvestmen The Opiliones (formerly Phalangida) are an order of arachnids , colloquially known as harvestmen , harvesters , harvest spiders , or daddy longlegs . As of July 2024 , over 6,650 species of harvestmen have been discovered worldwide, although

5985-457: The Gründerzeit period following the unification of Germany, Bismarck's foreign policy as chancellor of Germany secured Germany's position as a great nation by forging alliances and avoiding war. However, under Wilhelm II , Germany took an imperialistic course, leading to friction with neighbouring countries. A dual alliance was created with the multinational realm of Austria-Hungary ;

6118-484: The Bundestag and responsible for overseeing the daily sessions of the body. The third-highest official and the head of government is the chancellor, who is appointed by the Bundespräsident after being elected by the party or coalition with the most seats in the Bundestag . The chancellor, currently Olaf Scholz , is the head of government and exercises executive power through his Cabinet . Since 1949,

6251-916: The Allies de jure abolished the German state and partitioned Berlin and Germany's remaining territory into four occupation zones. The western sectors, controlled by France, the United Kingdom , and the United States , were merged on 23 May 1949 to form the Federal Republic of Germany ( German : Bundesrepublik Deutschland ); on 7 October 1949, the Soviet Zone became the German Democratic Republic (GDR) ( German : Deutsche Demokratische Republik ; DDR). They were informally known as West Germany and East Germany. East Germany selected East Berlin as its capital, while West Germany chose Bonn as

6384-607: The Battle of Britain in the same year. In 1941, German troops invaded Yugoslavia , Greece and the Soviet Union . By 1942, Germany and its allies controlled most of continental Europe and North Africa , but following the Soviet victory at the Battle of Stalingrad , the Allied reconquest of North Africa and invasion of Italy in 1943, German forces suffered repeated military defeats. In 1944,

6517-765: The Battle of the Teutoburg Forest . The outcome of this battle dissuaded the Romans from their ambition of conquering Germania and is thus considered one of the most important events in European history . By 100 AD, when Tacitus wrote Germania , Germanic tribes had settled along the Rhine and the Danube (the Limes Germanicus ), occupying most of modern Germany. However, Baden-Württemberg , southern Bavaria , southern Hesse and

6650-578: The Berchtesgaden National Park . In addition, there are 17 Biosphere Reserves , and 105 nature parks . More than 400 zoos and animal parks operate in Germany. The Berlin Zoo , which opened in 1844, is the oldest in Germany, and claims the most comprehensive collection of species in the world. Germany is a federal , parliamentary , representative democratic republic. Federal legislative power

6783-678: The Black Death of 1348–1350. The Golden Bull issued in 1356 provided the constitutional structure of the Empire and codified the election of the emperor by seven prince-electors . Johannes Gutenberg introduced moveable-type printing to Europe, laying the basis for the democratization of knowledge . In 1517, Martin Luther incited the Protestant Reformation and his translation of the Bible began

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6916-802: The Carolingian Empire in 800; it was divided in 843 . The eastern successor kingdom of East Francia stretched from the Rhine in the west to the Elbe river in the east and from the North Sea to the Alps. Subsequently, the Holy Roman Empire emerged from it. The Ottonian rulers (919–1024) consolidated several major duchies . In 996, Gregory V became the first German Pope, appointed by his cousin Otto III , whom he shortly after crowned Holy Roman Emperor. The Holy Roman Empire absorbed northern Italy and Burgundy under

7049-624: The Celtic , Iranian , Baltic , and Slavic tribes. Southern Germany was inhabited by Celtic-speaking peoples, who belonged to the wider La Tène culture . They were later assimilated by the Germanic conquerors. Under Augustus , the Roman Empire began to invade lands inhabited by the Germanic tribes, creating a short-lived Roman province of Germania between the Rhine and Elbe rivers. In 9 AD, three Roman legions were defeated by Arminius in

7182-840: The Cenozoic , three from the Mesozoic , and at least seven from the Paleozoic . Germany – in Europe  (light green & dark grey) – in the European Union  (light green) Germany , officially the Federal Republic of Germany , is a country in Central Europe . It lies between the Baltic and North Sea to the north and the Alps to the south. Its sixteen constituent states have

7315-452: The Devonian Rhynie chert , 410 million years ago, already show characteristics like tracheae and sexual organs, indicating that the group has lived on land since that time. Despite being similar in appearance to, and often confused with, spiders, they are probably closely related to the scorpions , pseudoscorpions , and solifuges ; these four orders form the clade Dromopoda . The Opiliones have remained almost unchanged morphologically over

7448-558: The Eupnoi suborder, use two mechanisms. One is bobbing, for which these particular individuals bounce their bodies. It potentially serves to confuse and deflect any identification of the exact location of their bodies. This can be a deceiving mechanism to avoid predation when they are in a large aggregation of individuals, which are all trembling at the same time. Cellar spiders ( Pholcidae ) that are commonly mistaken for daddy long-legs (Opiliones) also exhibit this behavior when their webs are disturbed or even during courtship. Autotomy

7581-430: The Eurasian beaver . The blue cornflower was once a German national symbol . The 16 national parks in Germany include the Jasmund National Park , the Vorpommern Lagoon Area National Park , the Müritz National Park , the Wadden Sea National Parks , the Harz National Park , the Hainich National Park , the Black Forest National Park , the Saxon Switzerland National Park , the Bavarian Forest National Park and

7714-485: The Fall of Communism , the dissolution of the Soviet Union , German reunification and Die Wende ("the turning point"). United Germany was considered the enlarged continuation of West Germany so it retained its memberships in international organisations. Based on the Berlin/Bonn Act (1994), Berlin again became the capital of Germany, while Bonn obtained the unique status of a Bundesstadt (federal city) retaining some federal ministries. The relocation of

7847-513: The Imperial Estates . The legal system initiated by a series of Imperial Reforms (approximately 1495–1555) provided for considerable local autonomy and a stronger Imperial Diet . The House of Habsburg held the imperial crown from 1438 until the death of Charles VI in 1740. Following the War of the Austrian Succession and the Treaty of Aix-la-Chapelle , Charles VI's daughter Maria Theresa ruled as empress consort when her husband, Francis I , became emperor. From 1740, dualism between

7980-401: The Kingdom of Germany formed the bulk of the Holy Roman Empire . During the 16th century, northern German regions became the centre of the Protestant Reformation . Following the Napoleonic Wars and the dissolution of the Holy Roman Empire in 1806, the German Confederation was formed in 1815. Formal unification of Germany into the modern nation-state commenced on 18 August 1866 with

8113-521: The Laniatores , and more than 70,000 in certain Eupnoi . Gregarious behavior is likely a strategy against climatic odds, but also against predators, combining the effect of scent secretions, and reducing the probability of any particular individual being eaten. Harvestmen clean their legs after eating by drawing each leg in turn through their jaws. Predators of harvestmen include a variety of animals, including some mammals, amphibians, and other arachnids like spiders and scorpions. Opiliones display

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8246-467: The Molotov–Ribbentrop Pact that divided Eastern Europe into German and Soviet spheres of influence. On 1 September 1939, Germany invaded Poland , beginning World War II in Europe ; Britain and France declared war on Germany on 3 September. In the spring of 1940, Germany conquered Denmark and Norway , the Netherlands , Belgium , Luxembourg , and France , forcing the French government to sign an armistice . The British repelled German air attacks in

8379-399: The North German Confederation Treaty establishing the Prussia -led North German Confederation , which became the German Empire in 1871. After World War I and the German Revolution of 1918–1919 , the Empire was replaced by the Weimar Republic . The Nazi rise to power in 1933 led to the establishment of a totalitarian dictatorship , World War II , and the Holocaust . In 1949, after

8512-430: The Saarland in 1935, remilitarised the Rhineland in 1936, annexed Austria in 1938, annexed the Sudetenland in 1938 with the Munich Agreement , and in violation of the agreement occupied Czechoslovakia in March 1939. Kristallnacht (Night of Broken Glass) saw the burning of synagogues, the destruction of Jewish businesses, and mass arrests of Jewish people. In August 1939, Hitler's government negotiated

8645-423: The Salian emperors (1024–1125), although the emperors lost power through the Investiture controversy . Under the Hohenstaufen emperors (1138–1254), German princes encouraged German settlement to the south and east ( Ostsiedlung ). Members of the Hanseatic League , mostly north German towns, prospered in the expansion of trade. The population declined starting with the Great Famine in 1315, followed by

8778-543: The Swabian Jura , including 42,000-year-old flutes which are the oldest musical instruments ever found, the 40,000-year-old Lion Man , and the 41,000-year-old Venus of Hohle Fels . The Nebra sky disk , created during the European Bronze Age , has been attributed to a German site. The Germanic peoples are thought to have emerged from the Jastorf culture during the Nordic Bronze Age or early Iron Age . From southern Scandinavia and northern Germany , they expanded south, east, and west, coming into contact with

8911-450: The Triple Alliance of 1882 included Italy. Britain, France and Russia also concluded alliances to protect against Habsburg interference with Russian interests in the Balkans or German interference against France. At the Berlin Conference in 1884, Germany claimed several colonies including German East Africa , German South West Africa , Togoland , and Kamerun . Later, Germany further expanded its colonial empire to include holdings in

9044-417: The inquisitorial Federal Court of Justice for civil and criminal cases, along with the Federal Labour Court , Federal Social Court , Federal Fiscal Court , and Federal Administrative Court for other matters. Criminal and private laws are codified on the national level in the Strafgesetzbuch and the Bürgerliches Gesetzbuch respectively. The German penal system seeks the rehabilitation of

9177-473: The leaf litter found in their environments. Some hard-bodied harvestmen have epizoic cyanobacteria and liverworts growing on their bodies that suggest potential benefits for camouflage against large backgrounds to avoid detection by diurnal predators. Some harvestmen have elaborate and brightly coloured patterns or appendages which contrast with the body colouration, potentially serving as an aposematic warning to potential predators. This mechanism

9310-428: The revolutions of 1848 in the German states , raising the German question. King Frederick William IV of Prussia was offered the title of emperor, but with a loss of power; he rejected the crown and the proposed constitution, a temporary setback for the movement. King William I appointed Otto von Bismarck as the Minister President of Prussia in 1862. Bismarck successfully concluded the war with Denmark in 1864 ;

9443-573: The scutum , which in most such species is fused with the carapace . Some such Opiliones only have this shield in the males. In some species, the two posterior abdominal segments are reduced. Some of them are divided medially on the surface to form two plates beside each other. The second pair of legs is longer than the others and function as antennae or feelers. In short-legged species, this may not be obvious. The feeding apparatus ( stomotheca ) differs from most arachnids in that Opiliones can swallow chunks of solid food, not only liquids. The stomotheca

9576-542: The third-highest number of UNESCO World Heritage Sites , 54 , of which 51 are cultural. The English word Germany derives from the Latin Germania , which came into use after Julius Caesar adopted it for the peoples east of the Rhine . The German term Deutschland , originally diutisciu land ('the German lands'), is derived from deutsch ( cf. Dutch ), descended from Old High German diutisc 'of

9709-479: The 1990s, study of the biology and ecology of harvestmen has intensified, especially in South America . Early work on the developmental biology of Opiliones from the mid-20th century was resurrected by Prashant P. Sharma , who established Phalangium opilio as a model system for the study of arachnid comparative genomics and evolutionary-developmental biology . Harvestmen are ancient arachnids . Fossils from

9842-543: The Arachnida is disputed; their closest relatives may be camel spiders (Solifugae) or a larger clade comprising horseshoe crabs, Ricinulei, and Arachnopulmonata (scorpions, pseudoscorpions, and Tetrapulmonata). Although superficially similar to and often misidentified as spiders (order Araneae ), the Opiliones are a distinct order that is not closely related to spiders. They can be easily distinguished from long-legged spiders by their fused body regions and single pair of eyes in

9975-865: The Austrian Habsburg monarchy and the Kingdom of Prussia dominated German history. In 1772, 1793, and 1795, Prussia and Austria, along with the Russian Empire , agreed to the Partitions of Poland . During the period of the French Revolutionary Wars , the Napoleonic era and the subsequent final meeting of the Imperial Diet , most of the Free Imperial Cities were annexed by dynastic territories;

10108-551: The Brazilian national list of endangered species, all of them cave-dwelling: Giupponia chagasi , Iandumoema uai , Pachylospeleus strinatii and Spaeleoleptes spaeleus . Several Opiliones in Argentina appear to be vulnerable, if not endangered. These include Pachyloidellus fulvigranulatus , which is found only on top of Cerro Uritorco , the highest peak in the Sierras Chicas chain (provincia de Cordoba) and Pachyloides borellii

10241-764: The Catálogo Nacional de especies amenazadas (National catalog of threatened species) from the Spanish government. Texella reddelli and Texella reyesi are listed as endangered species in the United States. Both are from caves in central Texas . Texella cokendolpheri from a cave in central Texas and Calicina minor , Microcina edgewoodensis , Microcina homi , Microcina jungi , Microcina leei , Microcina lumi , and Microcina tiburona from around springs and other restricted habitats of central California are being considered for listing as endangered species, but as yet receive no protection. An urban legend claims that

10374-496: The German Bundestag , though they have never been part of the federal government. In the 2017 German federal election , the right-wing populist Alternative for Germany gained enough votes to attain representation in the parliament for the first time. Germany is a federation and comprises sixteen constituent states which are collectively referred to as Länder . Each state ( Land ) has its own constitution, and

10507-597: The German Democratic Republic was a communist Eastern Bloc state and member of the Warsaw Pact . After the fall of the communist led-government in East Germany, German reunification saw the former East German states join the Federal Republic of Germany on 3 October 1990 . Germany has been described as a great power with a strong economy ; it has the largest economy in Europe by nominal GDP . As

10640-538: The Holocaust , the German government persecuted minorities , including interning them in concentration and death camps across Europe. The regime systematically murdered 6 million Jews, at least 130,000 Romani , 275,000 disabled , thousands of Jehovah's Witnesses , thousands of homosexuals , and hundreds of thousands of political and religious opponents . Nazi policies in German-occupied countries resulted in

10773-617: The Pacific and China. The colonial government in South West Africa (present-day Namibia ), from 1904 to 1907, carried out the annihilation of the local Herero and Namaqua peoples as punishment for an uprising; this was the 20th century's first genocide . The assassination of Austria's crown prince on 28 June 1914 provided the pretext for Austria-Hungary to attack Serbia and trigger World War I . After four years of warfare, in which approximately two million German soldiers were killed,

10906-664: The Pacific. On 11 August 1919, President Friedrich Ebert signed the democratic Weimar Constitution . In the subsequent struggle for power, communists seized power in Bavaria , but conservative elements elsewhere attempted to overthrow the Republic in the Kapp Putsch . Street fighting in the major industrial centres, the occupation of the Ruhr by Belgian and French troops, and a period of hyperinflation followed. A debt restructuring plan and

11039-583: The Reichstag after a special election in 1932 and Hindenburg appointed Hitler as chancellor of Germany on 30 January 1933. After the Reichstag fire , a decree abrogated basic civil rights and the first Nazi concentration camp opened. On 23 March 1933, the Enabling Act gave Hitler unrestricted legislative power, overriding the constitution, and marked the beginning of Nazi Germany. His government established

11172-653: The Soviets pushed into Eastern Europe ; the Western allies landed in France and entered Germany despite a final German counteroffensive . Following Hitler's suicide during the Battle of Berlin , Germany signed the surrender document on 8 May 1945, ending World War II in Europe and Nazi Germany. Following the end of the war, surviving Nazi officials were tried for war crimes at the Nuremberg trials . In what later became known as

11305-406: The abdomen, with one opening on each side. In more active species, spiracles are also found upon the tibia of the legs. They have a gonopore on the ventral cephalothorax, and the copulation is direct as male Opiliones have a penis , unlike other arachnids. All species lay eggs . Typical body length does not exceed 7 mm (0.28 in), and some species are smaller than 1 mm, although

11438-500: The animals' outlines or providing camouflage. Markings on legs can cause an interruption of the leg outline and loss of leg proportion recognition. Darker colourations and patterns function as camouflage when they remain motionless. Within the genus Leiobunum are multiple species with cryptic colouration that changes over ontogeny to match the microhabitat used at each life stage. Many species have also been able to camouflage their bodies by covering with secretions and debris from

11571-583: The body. Some superficially resemble mites , which is where they get their common name. Their coloration is almost always some shade of brown, with a heavily sclerotized body, and they are quite inconspicuous, residing in leaf litter or in caves . Many Cyphophthalmi are eyeless, and presumably rely on olfactory cues to find food and mates. Very little is known about their behavior, though they likely subsist mostly by scavenging and preying on minute arthropods. They have low dispersal rates and consequently high endemism . Cyphophthalmi differ from other harvestmen in

11704-522: The communist-controlled Socialist Unity Party of Germany , supported by the Stasi , an immense secret service. While East German propaganda was based on the benefits of the GDR's social programmes and the alleged threat of a West German invasion, many of its citizens looked to the West for freedom and prosperity. The Berlin Wall , built in 1961, prevented East German citizens from escaping to West Germany, becoming

11837-421: The connection between the cephalothorax and abdomen is broad, so that the body appears to be a single oval structure. Other differences include the fact that Opiliones have no venom glands in their chelicerae and thus pose no danger to humans. They also have no silk glands and therefore do not build webs. In some highly derived species, the first five abdominal segments are fused into a dorsal shield called

11970-416: The crane flies of the superfamily Tipuloidea , and the cellar spiders of the family Pholcidae, (commonly referred to as "daddy long-leg spiders") most likely because of their similar appearance. Harvestmen are also referred to as "shepherd spiders" in reference to how their unusually long legs reminded observers of the ways that some European shepherds used stilts to better observe their wandering flocks from

12103-462: The creation of a new currency in 1924 ushered in the Golden Twenties , an era of artistic innovation and liberal cultural life. The worldwide Great Depression hit Germany in 1929. Chancellor Heinrich Brüning 's government pursued a policy of fiscal austerity and deflation which caused unemployment of nearly 30% by 1932. The Nazi Party led by Adolf Hitler became the largest party in

12236-466: The creation of a more unified European political, economic and security apparatus. The governments of Germany and the United States are close political allies . Cultural ties and economic interests have crafted a bond between the two countries resulting in Atlanticism . After 1990, Germany and Russia worked together to establish a "strategic partnership" in which energy development became one of

12369-536: The criminal and the protection of the public. With the exceptions of petty crimes, tried by a single professional judge, and of serious political crimes, all charges are adjudicated by mixed tribunals where lay judges ( Schöffen ) and professional judges preside together. As of 2016, Germany's murder rate stood at a low of 1.18 murders per 100,000. In 2018, the overall crime rate fell to its lowest since 1992. Same-sex marriage has been legal in Germany since 2017, and LGBT rights are generally protected in

12502-461: The deaths of an estimated 2.7 million Poles , 1.3 million Ukrainians , 1 million Belarusians and 3.5 million Soviet prisoners of war . German military casualties have been estimated at 5.3 million, and around 900,000 German civilians died. Around 12 million ethnic Germans were expelled from across Eastern Europe, and Germany lost roughly one-quarter of its pre-war territory. After Nazi Germany surrendered,

12635-544: The defensive glands than other harvestmen, and the secretions, which primarily consist of a variety of naphthoquinones and ketones, are thought to be more potent than in Eupnoi and Dyspnoi. Cyphophthalmids are found on all continents, with the exception of Antarctica , where they probably also once lived. As they did not disperse onto any oceanic islands, and it is believed that they did not travel between separate landmasses, they make an interesting subject for biogeography . Each of

12768-600: The diversification of the modern Cyphophthalmi lineages to have occurred more recently, during the Jurassic, with the emergence of the suborder as a whole at less than 300 MYA. Cyphophthalmi is one of the two major lineages of harvestmen; the other, containing the Laniatores , Dyspnoi and Eupnoi , is known as Phalangida . The extinct suborder, Tetrophthalmi , shares several features in common with Cyphophthalmi, and these two suborders may represent sister taxa. They are grouped into

12901-566: The early 1950s. West Germany joined NATO in 1955 and was a founding member of the European Economic Community . On 1 January 1957, the Saarland joined West Germany. East Germany was an Eastern Bloc state under political and military control by the Soviet Union via occupation forces and the Warsaw Pact . Although East Germany claimed to be a democracy, political power was exercised solely by leading members ( Politbüro ) of

13034-681: The ecclesiastical territories were secularised and annexed. In 1806 the Imperium was dissolved; France, Russia, Prussia, and the Habsburgs (Austria) competed for hegemony in the German states during the Napoleonic Wars . Following the fall of Napoleon , the Congress of Vienna founded the German Confederation, a loose league of 39 sovereign states . The appointment of the emperor of Austria as

13167-594: The eyes in Cyphophthalmi are not homologous to the eyes of other harvestmen. Many cave-adapted species are eyeless, such as the Brazilian Caecobunus termitarum ( Grassatores ) from termite nests, Giupponia chagasi ( Gonyleptidae ) from caves, most species of Cyphophthalmi, and all species of the Guasiniidae . However, recent work studying the embryonic development of the species Phalangium opilio and some Laniatores revealed that harvestman in addition to

13300-401: The eyes of other harvestmen, and are instead derived from ancestral lateral eyes, instead of ancestral median eyes, as is the case for other harvestmen. The male genitalia of Cyphophthalmi is unique among harvestmen, as males lack a true penis, and instead have a short spermatopositor , a structure which is not inserted inside the female, but used to deposit a spermatophore. In addition, they are

13433-519: The following infraorders: Boreophthalmi, Scopulophthalmi, and Sternophthalmi. The Boreophthalmi and Sternophthalmi together form a monophyletic clade, and comprise 5 of the 6 families of Cyphphthalmi. The Scopulophthalmi, with only 1 family, the Pettalidae, forms the sister group to all the other families. Within the Sternophthalmi, the families Ogoveidae and Neogoveidae are most closely related, and form

13566-509: The fossil record for Cyphophthalmi is so sparse, recent studies have attempted to predict the diversification time of the suborder by using molecular clocks and comparing gene sequences among living taxa. One 2012 study estimated the earliest diversification of the suborder at approximately 332 MYA, in the Carboniferous. Another study, in 2014, recovered a diversification age of approximately 340 MYA. A more recent study in 2017, however, recovered

13699-597: The government was completed in 1999, and modernisation of the East German economy was scheduled to last until 2019. Since reunification, Germany has taken a more active role in the European Union , signing the Maastricht Treaty in 1992 and the Lisbon Treaty in 2007, and co-founding the eurozone . Germany sent a peacekeeping force to secure stability in the Balkans and sent German troops to Afghanistan as part of

13832-490: The harvestman is the most venomous animal in the world but possesses fangs too short or a mouth too round and small to bite a human, rendering it harmless (the same myth applies to Pholcus phalangioides and the crane fly , which are both also called a "daddy longlegs"). None of the known species of harvestmen have venom glands; their chelicerae are not hollowed fangs but grasping claws that are typically very small and not strong enough to break human skin. Harvestmen are

13965-565: The harvestmen against wandering spiders ( Ctenidae ), wolf spiders (Lycosidae) and Formica exsectoides ants. However, these chemical irritants are not able to prevent four species of harvestmen being preyed upon by the black scorpion Bothriurus bonariensis ( Bothriuridae ). These secretions contain multiple volatile compounds that vary among individuals and clades. All troglobitic species (of all animal taxa) are considered to be at least threatened in Brazil . Four species of Opiliones are on

14098-568: The largest known species, Trogulus torosus ( Trogulidae ), grows as long as 22 mm (0.87 in). The leg span of many species is much greater than the body length and sometimes exceeds 160 mm (6.3 in) and to 340 mm (13 in) in Southeast Asia. Most species live for a year. Many species are omnivorous , eating primarily small insects and all kinds of plant material and fungi . Some are scavengers , feeding upon dead organisms, bird dung, and other fecal material. Such

14231-557: The male is solely responsible for guarding the eggs resulting from multiple partners, often against egg-eating females , and cleaning the eggs regularly. Paternal care has evolved at least three times independently: once in the clade Progonyleptoidellinae + Caelopyginae, once in the Gonyleptinae, and once in the Heteropachylinae. Maternal care in opiliones probably evolved due to natural selection, while paternal care appears to be

14364-435: The males defend territories. In some species, males also exhibit post-copulatory behavior in which the male specifically seeks out and shakes the female's sensory leg. This is believed to entice the female into mating a second time. The female lays her eggs shortly after mating to several months later. Some species build nests for this purpose. A unique feature of harvestmen is that some species practice parental care, in which

14497-413: The middle of the cephalothorax . Spiders have a distinct abdomen that is separated from the cephalothorax by a constriction, and they have three to four pairs of eyes, usually around the margins of the cephalothorax. English speakers may colloquially refer to species of Opiliones as "daddy longlegs" or "granddaddy longlegs", but this name is also used for two other distantly related groups of arthropods ,

14630-469: The most important factors. As a result of the cooperation, Germany imported most of its natural gas and crude oil from Russia. Germany's development policy functions as a distinct sector within its foreign policy framework. It is formulated by the Federal Ministry for Economic Cooperation and Development and carried out by the implementing organisations. The German government sees development policy as

14763-641: The nation. Germany has a network of 227 diplomatic missions abroad and maintains relations with more than 190 countries. Germany is a member of NATO , the OECD , the G7 , the G20 , the World Bank and the IMF . It has played an influential role in the European Union since its inception and has maintained a strong alliance with France and all neighbouring countries since 1990. Germany promotes

14896-424: The nerves to the muscles to extend the leg and then the leg relaxes between signals. While some harvestman's legs twitch for a minute, others have been recorded to twitch up to an hour. The twitching has been hypothesised to function as an evolutionary advantage by keeping the attention of a predator while the harvestman escapes. Individuals that are able to detect potential threats can flee rapidly from attack. This

15029-539: The north, Poland and Czechia to the east, Austria and Switzerland to the south, and France , Luxembourg , Belgium , and the Netherlands to the west. Germany is also bordered by the North Sea and, at the north-northeast, by the Baltic Sea. German territory covers 357,596 km (138,069 sq mi). Elevation ranges from the mountains of the Alps (highest point: the Zugspitze at 2,963 metres or 9,721 feet) in

15162-410: The only harvestmen to lack a genital operculum, and instead exhibit a completely open gonopore. Mating in Cyphophthalmi is not well studied, but in some species at least, males and females do engage in direct contact. They further differ from most harvestmen in that the first 8 opisthosomal tergites and the prosomal carapace are fused together in what is known as a scutum completum , although this feature

15295-566: The origin of the order at about 473 million years ago (Mya), during the Ordovician. No fossils of the Cyphophthalmi or Laniatores much older than 50 million years are known, despite the former presenting a basal clade , and the latter having probably diverged from the Dyspnoi more than 300 Mya. Naturally, most finds are from comparatively recent times. More than 20 fossil species are known from

15428-587: The party system has been dominated by the Christian Democratic Union and the Social Democratic Party of Germany . So far every chancellor has been a member of one of these parties. However, the smaller liberal Free Democratic Party and the Alliance 90/The Greens have also been junior partners in coalition governments . Since 2007, the democratic socialist party The Left has been a staple in

15561-455: The people' (from diot or diota 'people'), originally used to distinguish the language of the common people from Latin and its Romance descendants . This in turn descends from Proto-Germanic * þiudiskaz 'of the people' (see also the Latinised form Theodiscus ), derived from * þeudō , descended from Proto-Indo-European * tewtéh₂- 'people', from which

15694-472: The permanent president reflected the Congress's rejection of Prussia 's rising influence. Disagreement within restoration politics partly led to the rise of liberal movements, followed by new measures of repression by Austrian statesman Klemens von Metternich . The Zollverein , a tariff union, furthered economic unity. In light of revolutionary movements in Europe , intellectuals and commoners started

15827-746: The result of sexual selection. Depending on circumstances such as temperature, the eggs may hatch at any time after the first 20 days, up to about half a year after being laid. Harvestmen variously pass through four to eight nymphal instars to reach maturity, with most known species having six instars. Most species are nocturnal and colored in hues of brown, although a number of diurnal species are known, some of which have vivid patterns in yellow, green, and black with varied reddish and blackish mottling and reticulation. Many species of harvestmen easily tolerate members of their own species, with aggregations of many individuals often found at protected sites near water. These aggregations may number 200 individuals in

15960-465: The six currently recognized families has a distinct distribution: Colombia shows the largest diversity of cyphophthalmids among countries in South America. This could reflect the large number of ecosystems found there, but could also be due to a high rate of sampling in this country. A fossil cyphophthalmid assigned to a modern genus, Siro platypedibus Dunlop & Giribet, 2003 (Sironidae),

16093-601: The south to the shores of the North Sea ( Nordsee ) in the northwest and the Baltic Sea ( Ostsee ) in the northeast. The forested uplands of central Germany and the lowlands of northern Germany (lowest point: in the municipality Neuendorf-Sachsenbande , Wilstermarsch at 3.54 metres or 11.6 feet below sea level ) are traversed by such major rivers as the Rhine, Danube and Elbe. Significant natural resources include iron ore, coal, potash , timber, lignite , uranium , copper, natural gas, salt, and nickel. Most of Germany has

16226-955: The southeast regions have more extreme temperatures. From February 2019 – 2020, average monthly temperatures in Germany ranged from a low of 3.3 °C (37.9 °F) in January 2020 to a high of 19.8 °C (67.6 °F) in June 2019. Average monthly precipitation ranged from 30 litres per square metre in February and April 2019 to 125 litres per square metre in February 2020. Average monthly hours of sunshine ranged from 45 in November 2019 to 300 in June 2019. The territory of Germany can be divided into five terrestrial ecoregions : Atlantic mixed forests , Baltic mixed forests , Central European mixed forests , Western European broadleaf forests , and Alps conifer and mixed forests . As of 2016 , 51% of Germany's land area

16359-556: The standardization of the language; the 1555 Peace of Augsburg tolerated the "Evangelical" faith ( Lutheranism ), but also decreed that the faith of the prince was to be the faith of his subjects ( cuius regio, eius religio ). From the Cologne War through the Thirty Years' Wars (1618–1648), religious conflict devastated German lands and significantly reduced the population. The Peace of Westphalia ended religious warfare among

16492-437: The suborder Cyphophthalmi, where the males deposit a spermatophore , mating involves direct copulation. The females store the sperm, which is aflagellate and immobile, at the tip of her ovipositor. The eggs are fertilized during oviposition. The males of some species offer a secretion (nuptial gift) from their chelicerae to the female before copulation. Sometimes, the male guards the female after copulation, and in many species,

16625-722: The subsequent decisive Prussian victory in the Austro-Prussian War of 1866 enabled him to create the North German Confederation which excluded Austria . After the defeat of France in the Franco-Prussian War , the German princes proclaimed the founding of the German Empire in 1871. Prussia was the dominant constituent state of the new empire; the King of Prussia ruled as its Kaiser, and Berlin became its capital. In

16758-803: The superfamily Ogoveoidea, which is the sister group to the family Troglosironidae. The systematics of the Boreophthalmi, however, is not yet fully resolved. In one recent analysis, the Boreophthalmi were recovered as paraphyletic with respect to the Sternophthalmi. Even within the Boreophthalmi, the Sironidae has been recovered either as sister family to Stylocellidae, or as paraphyletic with respect to Stylocellidae. In addition, there are 3 genera of Cyphophthalmi with uncertain affinities that have not been placed in any family or infraorder. Pettalidae Sironidae Stylocellidae Neogoveidae Ogoveidae Troglosironidae Pettalidae Stylocellidae Sironidae Neogoveidae Ogoveidae Troglosironidae Originally, Cyphophthalmi

16891-643: The total number of extant species may exceed 10,000. The order Opiliones includes five suborders: Cyphophthalmi , Eupnoi , Dyspnoi , Laniatores , and Tetrophthalmi , which were named in 2014. Representatives of each extant suborder can be found on all continents except Antarctica . Well-preserved fossils have been found in the 400-million-year-old Rhynie cherts of Scotland, and 305-million-year-old rocks in France. These fossils look surprisingly modern, indicating that their basic body shape developed very early on, and, at least in some taxa, has changed little since that time. Their phylogenetic position within

17024-406: The war and a period of Allied occupation , Germany was organized into two separate polities with limited sovereignty: the Federal Republic of Germany, or West Germany , and the German Democratic Republic, or East Germany . Berlin continued its de jure Four Power status . The Federal Republic of Germany was a founding member of the European Economic Community and the European Union , while

17157-621: The western Rhineland had been incorporated into Roman provinces . Around 260, Germanic peoples broke into Roman-controlled lands. After the invasion of the Huns in 375, and with the decline of Rome from 395, Germanic tribes moved farther southwest: the Franks established the Frankish Kingdom and pushed east to subjugate Saxony and Bavaria . Areas of what is today eastern Germany were inhabited by Western Slavic tribes. Charlemagne founded

17290-597: The word Teutons also originates. Pre-human ancestors, the Danuvius guggenmosi , who were present in Germany over 11 million years ago, are theorized to be among the earliest ones to walk on two legs. Ancient humans were present in Germany at least 600,000 years ago. The first non-modern human fossil (the Neanderthal ) was discovered in the Neander Valley . Similarly dated evidence of modern humans has been found in

17423-490: Was described from Paleogene aged Bitterfeld amber of eastern Germany . Another of this genus was described from Eocene Baltic amber as Siro balticus Dunlop & Mitov, 2011 . A third fossil, Palaeosiro burmanicum Poinar, 2008 , has been described from the Cretaceous ( Cenomanian ) aged Burmese amber . It was also assigned to Sironidae but is now thought to belong to Stylocellidae (a more typical Asian family). As

17556-484: Was recognized as a single family consisting of two subfamilies, Sironini and Stylocellini. This classification scheme existed until 1980, when a new taxonomy was proposed, separating 5 families into the now-defunct infraorders Temperophthalmi (consisting of Petallidae, Sironidae and, later, Troglosironidae) and Tropicophthalmi (consisting of Stylocellidae, Ogoveidae and Neogoveidae). These two infraorders were never recovered as monophyletic in subsequent studies, and, following

17689-491: Was the seventh-highest in the world . In response to the 2022 Russian invasion of Ukraine , Chancellor Olaf Scholz announced that German military expenditure would be increased past the NATO target of 2%, along with a one-time 2022 infusion of 100 billion euros, representing almost double the 53 billion euro military budget for 2021. In 2023, military spending according to NATO criteria amounted to $ 73.1 billion, or 1.64% of

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