177-438: Cretalamna is a genus of extinct otodontid shark that lived from the latest Early Cretaceous to Eocene epoch (about 103 to 46 million years ago). It is considered by many to be the ancestor of the largest sharks to have ever lived, such as Otodus angustidens , Otodus chubutensis , and Otodus megalodon . Cretalamna was first described by Swiss naturalist Louis Agassiz using five teeth previously identified as
354-468: A Bettrechies quarry available to Siversson et al. (2015) and for his work on Belgian Cenozoic sharks; C. ewelli , which honors paleontologist Keith Ewell who collected most of the C. ewelli type specimens in 2004; and C. hattini , which honors the late geologist Donald E. Hattin "for his work on the stratigraphy of the Niobrara Formation, western Kansas". The specific epithet of C. gertericorum
531-662: A French colonial empire rise. Internally, France was dominated by the conflict with the House of Habsburg and the French Wars of Religion between Catholics and Huguenots . France was successful in the Thirty Years' War and further increased its influence during the reign of Louis XIV . The French Revolution of 1789 overthrew the Ancien Régime and produced the Declaration of
708-438: A food chain who lose their prey. "Species coextinction is a manifestation of one of the interconnectednesses of organisms in complex ecosystems ... While coextinction may not be the most important cause of species extinctions, it is certainly an insidious one." Coextinction is especially common when a keystone species goes extinct. Models suggest that coextinction is the most common form of biodiversity loss . There may be
885-657: A nautilus to the Royal Society that was more than two feet in diameter, and morphologically distinct from any known living species. Hooke theorized that this was simply because the species lived in the deep ocean and no one had discovered them yet. While he contended that it was possible a species could be "lost", he thought this highly unlikely. Similarly, in 1695, Sir Thomas Molyneux published an account of enormous antlers found in Ireland that did not belong to any extant taxa in that area. Molyneux reasoned that they came from
1062-415: A species or a population is the variety of genetic information in its living members. A large gene pool (extensive genetic diversity ) is associated with robust populations that can survive bouts of intense selection . Meanwhile, low genetic diversity (see inbreeding and population bottlenecks ) reduces the range of adaptions possible. Replacing native with alien genes narrows genetic diversity within
1239-436: A viable population for species preservation and possible future reintroduction to the wild, through use of carefully planned breeding programs . The extinction of one species' wild population can have knock-on effects, causing further extinctions. These are also called "chains of extinction". This is especially common with extinction of keystone species . A 2018 study indicated that the sixth mass extinction started in
1416-436: A wastebasket taxon culminating to an interpretation of C. appendiculata as a variable cosmopolitan species with a 50 million year range. This changed when paleontologist Mikael Siversson found that the twenty-five syntypes actually represented a mix of at least six or more different species including three additional genera Dwardius , Cretoxyrhina , and Cretodus . To remedy the taxonomic issue, he redesignated one of
1593-399: A 2015 paper that the provision cannot be worked retroactively, and that the continued prevailing usage of 'Cretalamna' since the provision's establishment ironically secures its priority rather than threaten it. 'Cretalamna' currently remains as the most prevalent spelling and paleontologists have expressed the unlikeliness of a return to the usage of 'Cretolamna' . Cretalamna was
1770-814: A Christian in the event of victory against the Visigothic Kingdom , which was said to have guaranteed the battle. Clovis regained the southwest from the Visigoths and was baptised in 508. Clovis I was the first Germanic conqueror after the Fall of the Western Roman Empire to convert to Catholic Christianity; thus France was given the title "Eldest daughter of the Church" by the papacy, and French kings called "the Most Christian Kings of France". The Franks embraced
1947-425: A European counterweight between American and Soviet spheres of influence. However, he opposed any development of a supranational Europe , favouring sovereign nations . The revolt of May 1968 had an enormous social impact; it was a watershed moment when a conservative moral ideal (religion, patriotism, respect for authority) shifted to a more liberal moral ideal (secularism, individualism, sexual revolution). Although
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#17328691140512124-474: A Lazarus species when extant individuals were described in 2019. Attenborough's long-beaked echidna ( Zaglossus attenboroughi ) is an example of a Lazarus species from Papua New Guinea that had last been sighted in 1962 and believed to be possibly extinct, until it was recorded again in November 2023. Some species currently thought to be extinct have had continued speculation that they may still exist, and in
2301-495: A cascade of coextinction across the trophic levels . Such effects are most severe in mutualistic and parasitic relationships. An example of coextinction is the Haast's eagle and the moa : the Haast's eagle was a predator that became extinct because its food source became extinct. The moa were several species of flightless birds that were a food source for the Haast's eagle. Extinction as
2478-674: A combination of social, political and economic factors, which the Ancien Régime proved unable to manage. A financial crisis and social distress led in May 1789 to the convocation of the Estates General , which was converted into a National Assembly in June. The Storming of the Bastille on 14 July led to a series of radical measures by the Assembly, among them the abolition of feudalism , state control over
2655-429: A common ancestor with modern horses. Pseudoextinction is much easier to demonstrate for larger taxonomic groups. A Lazarus taxon or Lazarus species refers to instances where a species or taxon was thought to be extinct, but was later rediscovered. It can also refer to instances where large gaps in the fossil record of a taxon result in fossils reappearing much later, although the taxon may have ultimately become extinct at
2832-559: A coup and civil war. During the May 1958 crisis , the weak Fourth Republic gave way to the Fifth Republic , which included a strengthened presidency. The war concluded with the Évian Accords in 1962 which led to Algerian independence , at a high price: between half a million and one million deaths and over 2 million internally-displaced Algerians. Around one million Pied-Noirs and Harkis fled from Algeria to France. A vestige of empire
3009-399: A fact that was accepted by most scientists. The primary debate focused on whether this turnover caused by extinction was gradual or abrupt in nature. Cuvier understood extinction to be the result of cataclysmic events that wipe out huge numbers of species, as opposed to the gradual decline of a species over time. His catastrophic view of the nature of extinction garnered him many opponents in
3186-415: A global centre of art , science , and philosophy . It hosts the fourth-largest number of UNESCO World Heritage Sites and is the world's leading tourist destination , receiving 100 million foreign visitors in 2023 . A developed country , France has a high nominal per capita income globally , and its advanced economy ranks among the largest in the world . It is a great power , being one of
3363-708: A higher risk of extinction and die out faster than less sexually dimorphic species, the least sexually dimorphic species surviving for millions of years while the most sexually dimorphic species die out within mere thousands of years. Earlier studies based on counting the number of currently living species in modern taxa have shown a higher number of species in more sexually dimorphic taxa which have been interpreted as higher survival in taxa with more sexual selection, but such studies of modern species only measure indirect effects of extinction and are subject to error sources such as dying and doomed taxa speciating more due to splitting of habitat ranges into more small isolated groups during
3540-405: A large range, a lack of individuals of both sexes (in sexually reproducing species), or other reasons. Pinpointing the extinction (or pseudoextinction ) of a species requires a clear definition of that species . If it is to be declared extinct, the species in question must be uniquely distinguishable from any ancestor or daughter species, and from any other closely related species. Extinction of
3717-592: A later point. The coelacanth , a fish related to lungfish and tetrapods , is an example of a Lazarus taxon that was known only from the fossil record and was considered to have been extinct since the end of the Cretaceous Period . In 1938, however, a living specimen was found off the Chalumna River (now Tyolomnqa) on the east coast of South Africa. Calliostoma bullatum , a species of deepwater sea snail originally described from fossils in 1844 proved to be
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#17328691140513894-744: A maritime border with the United Kingdom to the northwest. Its metropolitan area extends from the Rhine to the Atlantic Ocean and from the Mediterranean Sea to the English Channel and the North Sea . Its eighteen integral regions (five of which are overseas) span a combined area of 643,801 km (248,573 sq mi) and have a total population of 68.4 million as of January 2024 . France
4071-587: A mathematical model that falls in all positions. By contrast, conservation biology uses the extinction vortex model to classify extinctions by cause. When concerns about human extinction have been raised, for example in Sir Martin Rees ' 2003 book Our Final Hour , those concerns lie with the effects of climate change or technological disaster. Human-driven extinction started as humans migrated out of Africa more than 60,000 years ago. Currently, environmental groups and some governments are concerned with
4248-425: A medium to large-sized shark. Based on vertebral comparisons with various extant lamniforms and Cretoxyrhina , a 2007 study by Kenshu Shimada estimated a total length of 2.3–3.0 metres (8–10 ft) for the most complete skeleton of a large individual (LACM 128126; C. hattini holotype). Shimada previously discovered that the total length of lamniform sharks is positively correlated with the size of their teeth in
4425-456: A natural part of the evolutionary process. Only recently have extinctions been recorded and scientists have become alarmed at the current high rate of extinctions . Most species that become extinct are never scientifically documented. Some scientists estimate that up to half of presently existing plant and animal species may become extinct by 2100. A 2018 report indicated that the phylogenetic diversity of 300 mammalian species erased during
4602-441: A new mega-predator or by transporting animals and plants from one part of the world to another. Such introductions have been occurring for thousands of years, sometimes intentionally (e.g. livestock released by sailors on islands as a future source of food) and sometimes accidentally (e.g. rats escaping from boats). In most cases, the introductions are unsuccessful, but when an invasive alien species does become established,
4779-610: A population a higher chance in the short term of surviving an adverse change in conditions. Effects that cause or reward a loss in genetic diversity can increase the chances of extinction of a species. Population bottlenecks can dramatically reduce genetic diversity by severely limiting the number of reproducing individuals and make inbreeding more frequent. Extinction sometimes results for species evolved to specific ecologies that are subjected to genetic pollution —i.e., uncontrolled hybridization , introgression and genetic swamping that lead to homogenization or out-competition from
4956-407: A race of animals to become extinct. A series of fossils were discovered in the late 17th century that appeared unlike any living species. As a result, the scientific community embarked on a voyage of creative rationalization, seeking to understand what had happened to these species within a framework that did not account for total extinction. In October 1686, Robert Hooke presented an impression of
5133-492: A reasonably linear relationship; thus, Shimada (2007)'s estimates enabled size estimations for Cretalamna based on teeth alone. Subsequently, in 2019, the teeth of C. appendiculata from Himedo Park, Kugushima and Wadanohana which are larger than those in LACM 128126 yielded maximum length estimates of up to 3.4 metres (11 ft), 4.5 metres (15 ft) and 5 meters (16 ft), respectively. In 2020, Shimada and colleagues estimated
5310-502: A reduction in agricultural productivity. Furthermore, increased erosion contributes to poorer water quality by elevating the levels of sediment and pollutants in rivers and streams. Habitat degradation through toxicity can kill off a species very rapidly, by killing all living members through contamination or sterilizing them. It can also occur over longer periods at lower toxicity levels by affecting life span, reproductive capacity, or competitiveness. Habitat degradation can also take
5487-486: A result of climate change has been confirmed by fossil studies. Particularly, the extinction of amphibians during the Carboniferous Rainforest Collapse , 305 million years ago. A 2003 review across 14 biodiversity research centers predicted that, because of climate change, 15–37% of land species would be "committed to extinction" by 2050. The ecologically rich areas that would potentially suffer
Cretalamna - Misplaced Pages Continue
5664-790: A similar word structure that has a person's last name prefixed onto the Latin suffix - i (from). These species are C. arambourgi , which honors paleontologist Camille Arambourg for his discovery of the C. arambourgi type specimens and his contributions to North African paleontology; C. bryanti , which honors the Bryant family who helped enhance the reputation and missions of the University of Alabama , Alabama Museum of Natural History , and McWane Science Center through their commitment to education and support; C. deschutteri , which honors paleontologist Pieter De Scutter for his efforts to make Cretalamna teeth from
5841-479: A species (or replacement by a daughter species) plays a key role in the punctuated equilibrium hypothesis of Stephen Jay Gould and Niles Eldredge . In ecology , extinction is sometimes used informally to refer to local extinction , in which a species ceases to exist in the chosen area of study, despite still existing elsewhere. Local extinctions may be made good by the reintroduction of individuals of that species taken from other locations; wolf reintroduction
6018-439: A species may come suddenly when an otherwise healthy species is wiped out completely, as when toxic pollution renders its entire habitat unliveable; or may occur gradually over thousands or millions of years, such as when a species gradually loses out in competition for food to better adapted competitors. Extinction may occur a long time after the events that set it in motion, a phenomenon known as extinction debt . Assessing
6195-404: A species or group of species. "Just as each species is unique", write Beverly and Stephen C. Stearns , "so is each extinction ... the causes for each are varied—some subtle and complex, others obvious and simple". Most simply, any species that cannot survive and reproduce in its environment and cannot move to a new environment where it can do so, dies out and becomes extinct. Extinction of
6372-518: A subsequent report, IPBES listed unsustainable fishing, hunting and logging as being some of the primary drivers of the global extinction crisis. In June 2019, one million species of plants and animals were at risk of extinction. At least 571 plant species have been lost since 1750, but likely many more. The main cause of the extinctions is the destruction of natural habitats by human activities, such as cutting down forests and converting land into fields for farming. A dagger symbol (†) placed next to
6549-668: Is a semi-presidential republic with its capital in Paris , the country's largest city and main cultural and commercial centre. Metropolitan France was settled during the Iron Age by Celtic tribes known as Gauls before Rome annexed the area in 51 BC, leading to a distinct Gallo-Roman culture . In the Early Middle Ages , the Franks formed the Kingdom of Francia , which became the heartland of
6726-451: Is a portmanteau derived from the Latin prefix bi- (two) prefixed onto the Latin auriculātā (eared), together meaning "having two ears". This is a reference to the species' large lateral cusplets, which somewhat resemble a pair of ears. The species name nigeriana is derived from the country name Niger prefixed to the suffix - iana , a feminine variation of the Latin suffix - ānus (pertaining to), together meaning "pertaining to Niger". This
6903-509: Is a reference to the species' type locality in Niger. The species name sarcoportheta is derived from the Ancient Greek σαρκός ( sarkos , meaning "flesh") prefixed to the Ancient Greek πορθητής ( porthitís , meaning "destroyer"), together meaning "destroyer of flesh". The species name catoxodon is derived from the Ancient Greek κατοξυς ( katoxys , meaning "very sharp") prefixed to
7080-517: Is also evidence to suggest that this event was preceded by another mass extinction, known as Olson's Extinction . The Cretaceous–Paleogene extinction event (K–Pg) occurred 66 million years ago, at the end of the Cretaceous period; it is best known for having wiped out non-avian dinosaurs , among many other species. According to a 1998 survey of 400 biologists conducted by New York 's American Museum of Natural History , nearly 70% believed that
7257-444: Is an example of this. Species that are not globally extinct are termed extant . Those species that are extant, yet are threatened with extinction, are referred to as threatened or endangered species . Currently, an important aspect of extinction is human attempts to preserve critically endangered species. These are reflected by the creation of the conservation status "extinct in the wild" (EW) . Species listed under this status by
Cretalamna - Misplaced Pages Continue
7434-563: Is derived from the feminine form of the Latin lātus (wide); a reference to the notably wide teeth of the species. The species name borealis is derived from the Latin boreālis (northern); this is a reference to its discovery from fossil deposits in Sweden, a boreal locality. The specific epithet of C. maroccana is a feminine form of the Latin word maroccānus ( Moroccan ), a reference to its type locality in Morocco. C. biauriculata' s specific epithet
7611-585: Is difficult to demonstrate unless one has a strong chain of evidence linking a living species to members of a pre-existing species. For example, it is sometimes claimed that the extinct Hyracotherium , which was an early horse that shares a common ancestor with the modern horse , is pseudoextinct, rather than extinct, because there are several extant species of Equus , including zebra and donkey ; however, as fossil species typically leave no genetic material behind, one cannot say whether Hyracotherium evolved into more modern horse species or merely evolved from
7788-417: Is estimated as 100 to 1,000 times "background" rates (the average extinction rates in the evolutionary time scale of planet Earth), faster than at any other time in human history, while future rates are likely 10,000 times higher. However, some groups are going extinct much faster. Biologists Paul R. Ehrlich and Stuart Pimm , among others, contend that human population growth and overconsumption are
7965-417: Is estimated that there are currently around 8.7 million species of eukaryotes globally, and possibly many times more if microorganisms , such as bacteria , are included. Notable extinct animal species include non-avian dinosaurs , saber-toothed cats , dodos , mammoths , ground sloths , thylacines , trilobites , golden toads , and passenger pigeons . Through evolution , species arise through
8142-522: Is given in the dental formula S2?.A2.I1.L10(+?) s?.a2.i1.l4 , constructed in a 2007 study of LACM 128126 by paleontologist Kenshu Shimada. It is possible that Cretalamna contained more than two symphysial tooth rows, as the related Cretoxyrhina mantelli possessed four upper symphysial tooth rows. In C. hattini , the upper and lower jaws are similar to that of Cretoxyrhina mantelli . The jaws also resemble those of modern alopiids (thresher sharks) and lamnids . Limited fossil evidence suggests that
8319-462: Is indicative of an active fast-swimming pelagic shark likely partially warm-blooded through regional endothermy . Cretalamna teeth are distinguished by a broad triangular cusp and two lateral cusplets. The cutting edges of the teeth are razor-like, while the sides have a smooth surface. Teeth symmetry is variable; some have exact bilateral symmetry whereas others have high asymmetry. Adjacent teeth do not overlap. The exact dentition of Cretalamna
8496-614: Is structured differently; it is derived from the names "Gert", "Eric", and the Latin suffix - orum (a masculine plural declension). The derived names "Gert" and "Eric" refer to fossil collectors Gert De Bie and Eric Collier, both of whom collected the majority of Cretalamna teeth examined in Siversson et al. (2015) that were from the Bettrechies quarry. The valid spelling of Cretalamna , specifically between it and 'Cretolamna' , has been subject to controversy. Originally, Glickman described
8673-558: Is the French overseas departments and territories . During the Cold War , de Gaulle pursued a policy of "national independence" towards the Western and Eastern blocs . He withdrew from NATO's military-integrated command (while remaining within the alliance), launched a nuclear development programme and made France the fourth nuclear power . He restored cordial Franco-German relations to create
8850-592: Is the destruction of ocean floors by bottom trawling . Diminished resources or introduction of new competitor species also often accompany habitat degradation. Global warming has allowed some species to expand their range, bringing competition to other species that previously occupied that area. Sometimes these new competitors are predators and directly affect prey species, while at other times they may merely outcompete vulnerable species for limited resources. Vital resources including water and food can also be limited during habitat degradation, leading to extinction. In
9027-525: Is the most important determinant of genus extinction at background rates but becomes increasingly irrelevant as mass extinction arises. Limited geographic range is a cause both of small population size and of greater vulnerability to local environmental catastrophes. Extinction rates can be affected not just by population size, but by any factor that affects evolvability , including balancing selection , cryptic genetic variation , phenotypic plasticity , and robustness . A diverse or deep gene pool gives
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#17328691140519204-562: Is uncertain due to poor fossil representation. Traditionally, most reconstructions of its dentition were constructed from individual shed teeth. Based on a specimen of C. hattini (LACM 128126), the dentition of the shark follows a lamnoid pattern with at least fifteen upper tooth rows and eight lower tooth rows on each side of the jaw. The upper tooth rows contain, from front to back: two symphysial, two anterior, one intermediate, and ten lateral tooth rows. The lower tooth rows contain: two anterior, one intermediate, and five lateral tooth rows. This
9381-512: The 1973 oil crisis , which now accounts for 75 per cent of its electricity production and results in less pollution. According to the 2020 Environmental Performance Index conducted by Yale and Columbia , France was the fifth most environmentally conscious country in the world. Like all European Union state members, France agreed to cut carbon emissions by at least 20% of 1990 levels by 2020. As of 2009 , French carbon dioxide emissions per capita were lower than that of China. The country
9558-609: The 1995 public transport bombings , France has been targeted by Islamist organisations, notably the Charlie Hebdo attack in 2015 which provoked the largest public rallies in French history, gathering 4.4 million people, the November 2015 Paris attacks which resulted in 130 deaths, the deadliest attack on French soil since World War II and the deadliest in the European Union since
9735-700: The 3rd Earl of Enniskillen from the Speeton Clay in Yorkshire ; one tooth of the collection of a Strasbourg Museum from an unspecified location; one tooth of the collection of the Hancock Museum from the Marly Chalk near Cambridge ; and six teeth of the collection of German paleontologist Heinrich Georg Bronn from chalk around Aachen ), he described a species whose teeth had thick bulged roots, lateral cusplets, and extreme variability. Agassiz remarked that some of
9912-642: The Battle of Hastings in 1066, William the Conqueror added "King of England" to his titles, becoming vassal and the equal of the king of France, creating recurring tensions. The Carolingian dynasty ruled France until 987, when Hugh Capet was crowned king of the Franks . His descendants unified the country through wars and inheritance. From 1190, the Capetian rulers began to be referred as "kings of France" rather than "kings of
10089-574: The British Isles and Switzerland . The wars were ended by Henry IV 's Edict of Nantes , which granted some freedom of religion to the Huguenots. Spanish troops, assisted the Catholics from 1589 to 1594 and invaded France in 1597. Spain and France returned to all-out war between 1635 and 1659. The war cost France 300,000 casualties. Under Louis XIII , Cardinal Richelieu promoted centralisation of
10266-719: The Carolingian Empire . The Treaty of Verdun of 843 partitioned the empire, with West Francia evolving into the Kingdom of France . In the High Middle Ages , France was a powerful but decentralized feudal kingdom, but from the mid-14th to the mid-15th centuries, France was plunged into a dynastic conflict with England known as the Hundred Years' War . In the 16th century, the French Renaissance saw culture flourish and
10443-753: The Catholic Church in France , and a declaration of rights . The next three years were dominated by struggle for political control, exacerbated by economic depression . Military defeats following the outbreak of the French Revolutionary Wars in April 1792 resulted in the insurrection of 10 August 1792 . The monarchy was abolished and replaced by the French First Republic in September, while Louis XVI
10620-592: The Consulate seized power in a coup led by Napoleon . Napoleon became First Consul in 1799 and later Emperor of the French Empire (1804–1814; 1815). Changing sets of European coalitions declared wars on Napoleon's empire . His armies conquered most of continental Europe with swift victories such as the battles of Jena-Auerstadt and Austerlitz . Members of the Bonaparte family were appointed monarchs in some of
10797-1049: The Hercynian uplift in the Paleozoic Era , the Armorican Massif , the Massif Central , the Morvan , the Vosges and Ardennes ranges and the island of Corsica were formed. These massifs delineate several sedimentary basins such as the Aquitaine Basin in the southwest and the Paris Basin in the north. Various routes of natural passage, such as the Rhône Valley , allow easy communication. The Alpine, Pyrenean and Jura mountains are much younger and have less eroded forms. At 4,810.45 metres (15,782 ft) above sea level, Mont Blanc , located in
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#173286911405110974-556: The House of Plantagenet , rulers of the County of Anjou , established its dominion over the surrounding provinces of Maine and Touraine , then built an "empire" from England to the Pyrenees , covering half of modern France. Tensions between France and the Plantagenet empire would last a hundred years, until Philip II of France conquered, between 1202 and 1214, most continental possessions of
11151-474: The International Union for Conservation of Nature (IUCN) are not known to have any living specimens in the wild and are maintained only in zoos or other artificial environments. Some of these species are functionally extinct, as they are no longer part of their natural habitat and it is unlikely the species will ever be restored to the wild. When possible, modern zoological institutions try to maintain
11328-477: The Late Pleistocene could take up to 5 to 7 million years to restore mammal diversity to what it was before the human era. Extinction of a parent species where daughter species or subspecies are still extant is called pseudoextinction or phyletic extinction. Effectively, the old taxon vanishes, transformed ( anagenesis ) into a successor, or split into more than one ( cladogenesis ). Pseudoextinction
11505-557: The Madrid train bombings in 2004 . Opération Chammal , France's military efforts to contain ISIS , killed over 1,000 ISIS troops between 2014 and 2015. The vast majority of France's territory and population is situated in Western Europe and is called Metropolitan France . It is bordered by the North Sea in the north, the English Channel in the northwest, the Atlantic Ocean in the west and
11682-402: The Mediterranean Sea in the southeast. Its land borders consist of Belgium and Luxembourg in the northeast, Germany and Switzerland in the east, Italy and Monaco in the southeast, and Andorra and Spain in the south and southwest. Except for the northeast, most of France's land borders are roughly delineated by natural boundaries and geographic features: to the south and southeast,
11859-514: The Popular Front government (e.g., annual leave , eight-hour workdays , women in government ). In 1940, France was invaded and quickly defeated by Nazi Germany . France was divided into a German occupation zone in the north, an Italian occupation zone and an unoccupied territory, the rest of France, which consisted of the southern France and the French empire. The Vichy government , an authoritarian regime collaborating with Germany, ruled
12036-488: The Seven Years' War (1756–1763). Its European territory kept growing, however, with acquisitions such as Lorraine and Corsica . Louis XV's weak rule, including the decadence of his court, discredited the monarchy, which in part paved the way for the French Revolution . Louis XVI (r. 1774–1793) supported America with money, fleets and armies , helping them win independence from Great Britain . France gained revenge, but verged on bankruptcy—a factor that contributed to
12213-589: The common smooth-hound and collected by English paleontologist Gideon Mantell from the Southerham Grey Pit near Lewes , East Sussex . In his 1835 publication Rapport sur les poissons fossiles découverts en Angleterre , he reidentified them as a new species of porbeagle shark under the taxon Lamna appendiculata . In 1843, Agassiz published Recherches sur les poissons fossiles , which reexamined Mantell's five teeth. Using them, eight additional teeth collected by Mantell, and twenty more teeth collected by various paleontologists in various locations (One tooth found by
12390-470: The official language of France and Europe's aristocracy. France became rivals of the House of Habsburg during the Italian Wars , which would dictate much of their later foreign policy until the mid-18th century. French explorers claimed lands in the Americas, paving expansion of the French colonial empire . The rise of Protestantism led France to a civil war known as the French Wars of Religion . This forced Huguenots to flee to Protestant regions such as
12567-405: The porbeagle and salmon sharks in build, with a compact fusiform body, large pectoral and first dorsal fins and tail, and small second dorsal, pelvic, and anal fins. The first dorsal fin was positioned directly above the pectoral fins unlike its analog species, where the first dorsal fin is usually positioned behind it. The tail fin was semi- lunate , similar to the whale shark . Such a body plan
12744-410: The slender-billed curlew ( Numenius tenuirostris ), not seen since 2007. As long as species have been evolving, species have been going extinct. It is estimated that over 99.9% of all species that ever lived are extinct. The average lifespan of a species is 1–10 million years, although this varies widely between taxa. A variety of causes can contribute directly or indirectly to the extinction of
12921-484: The strata of the Paris basin. They saw alternating saltwater and freshwater deposits, as well as patterns of the appearance and disappearance of fossils throughout the record. From these patterns, Cuvier inferred historic cycles of catastrophic flooding, extinction, and repopulation of the earth with new species. Cuvier's fossil evidence showed that very different life forms existed in the past than those that exist today,
13098-463: The trap-bath split such as Received Pronunciation , though it can be also heard in some other dialects such as Cardiff English . The oldest traces of archaic humans in what is now France date from approximately 1.8 million years ago. Neanderthals occupied the region into the Upper Paleolithic era but were slowly replaced by Homo sapiens around 35,000 BC. This period witnessed
13275-508: The 1970s France faced an economic crisis and allowed new immigrants (mostly from the Maghreb , in northwest Africa) to permanently settle in France with their families and acquire citizenship. It resulted in hundreds of thousands of Muslims living in subsidised public housing and suffering from high unemployment rates. The government had a policy of assimilation of immigrants, where they were expected to adhere to French values and norms. Since
13452-619: The 20 biodiversity goals laid out by the Aichi Biodiversity Targets in 2010, only 6 were "partially achieved" by the deadline of 2020. The report warned that biodiversity will continue to decline if the status quo is not changed, in particular the "currently unsustainable patterns of production and consumption, population growth and technological developments". In a 2021 report published in the journal Frontiers in Conservation Science , some top scientists asserted that even if
13629-599: The 200 years of the Crusades, in such a fashion that the Arabs referred to crusaders as Franj . French Crusaders imported French into the Levant , making Old French the base of the lingua franca ("Frankish language") of the Crusader states . The Albigensian Crusade was launched in 1209 to eliminate the heretical Cathars in the southwest of modern-day France. From the 11th century,
13806-534: The 2000 Edition of the Code in order to address this situation in the future, which states that "when an unjustified emendation is in prevailing usage and is attributed to the original author and date it is deemed to be a justified emendation". While Cappetta argued in a 2012 handbook that this new provision justifies the priority of 'Cretolamna' due to the spelling's overwhelmingly prevailing usage prior to its replacement by Siversson in 1999, Siversson himself pointed out in
13983-646: The 4th century, a period of revival and prosperity. In 312, Emperor Constantine I converted to Christianity . Christians, who had been persecuted, increased. But from the 5th century, the Barbarian Invasions resumed. Teutonic tribes invaded the region, the Visigoths settling in the southwest, the Burgundians along the Rhine River Valley, and the Franks in the north. In Late antiquity , ancient Gaul
14160-451: The Aichi Biodiversity Targets set for 2020 had been achieved, it would not have resulted in a significant mitigation of biodiversity loss. They added that failure of the global community to reach these targets is hardly surprising given that biodiversity loss is "nowhere close to the top of any country's priorities, trailing far behind other concerns such as employment, healthcare, economic growth, or currency stability." For much of history,
14337-748: The Alps on the France–Italy border , is the highest point in Western Europe. Although 60% of municipalities are classified as having seismic risks (though moderate). The coastlines offer contrasting landscapes: mountain ranges along the French Riviera , coastal cliffs such as the Côte d'Albâtre , and wide sandy plains in the Languedoc . Corsica lies off the Mediterranean coast. France has an extensive river system consisting of
14514-459: The Ancient Greek ὀδών ( odon , meaning "tooth". Together they mean "very sharp tooth", referring to the unusually sharp cutting edges of some C. catoxodon teeth. Six of the Cretalamna species have specific epithets that are named in honor of specific people, either for their contributions to the research of their associated species or for notable work they undertook. Of these six, five share
14691-562: The Atlantic Ocean. Other water courses drain towards the Meuse and Rhine along the northeastern borders. France has 11,000,000 km (4,200,000 sq mi) of marine waters within three oceans under its jurisdiction, of which 97% are overseas. France was one of the first countries to create an environment ministry, in 1971. France is ranked 19th by carbon dioxide emissions due to the country's heavy investment in nuclear power following
14868-621: The Christian Gallo-Roman culture , and ancient Gaul was renamed Francia ("Land of the Franks"). The Germanic Franks adopted Romanic languages . Clovis made Paris his capital and established the Merovingian dynasty , but his kingdom would not survive his death. The Franks treated land as a private possession and divided it among their heirs, so four kingdoms emerged from that of Clovis: Paris, Orléans , Soissons , and Rheims . The last Merovingian kings lost power to their mayors of
15045-605: The Earth is currently in the early stages of a human-caused mass extinction, known as the Holocene extinction . In that survey, the same proportion of respondents agreed with the prediction that up to 20% of all living populations could become extinct within 30 years (by 2028). A 2014 special edition of Science declared there is widespread consensus on the issue of human-driven mass species extinctions. A 2020 study published in PNAS stated that
15222-538: The Franks was known as the francisca ), although these weapons may have been named because of their use by the Franks, not the other way around. In English, 'France' is pronounced / f r æ n s / FRANSS in American English and / f r ɑː n s / FRAHNSS or / f r æ n s / FRANSS in British English. The pronunciation with / ɑː / is mostly confined to accents with
15399-521: The Franks". Later kings expanded their directly possessed domaine royal to cover over half of modern France by the 15th century. Royal authority became more assertive, centred on a hierarchically conceived society distinguishing nobility , clergy, and commoners . The nobility played a prominent role in Crusades to restore Christian access to the Holy Land . French knights made up most reinforcements in
15576-464: The French government's longtime historical association with the Catholic Church , Charlemagne tried to revive the Western Roman Empire and its cultural grandeur. Charlemagne's son, Louis I kept the empire united, however in 843, it was divided between Louis' three sons, into East Francia , Middle Francia and West Francia . West Francia approximated the area occupied by modern France and
15753-503: The Mentès well in Tahoua , Niger , which he assigned to the subspecies Lamna biauriculata nigeriana . This subspecies would also be elevated to its own species in 1991. In 1975, Cappetta and American paleontologist Gerard Case examined Cretalamna teeth described by Arambourg in 1952 from Danian deposits in Morocco and proposed that it represents a new subspecies of the type species and assigned it
15930-628: The Neolithic, including the Carnac stones site (approximately 3,300 BC). In 600 BC, Ionian Greeks from Phocaea founded the colony of Massalia (present-day Marseille ). Celtic tribes penetrated parts of eastern and northern France, spreading through the rest of the country between the 5th and 3rd century BC. Around 390 BC, the Gallic chieftain Brennus and his troops made their way to Roman Italy , defeated
16107-515: The North American moose and that the animal had once been common on the British Isles . Rather than suggest that this indicated the possibility of species going extinct, he argued that although organisms could become locally extinct, they could never be entirely lost and would continue to exist in some unknown region of the globe. The antlers were later confirmed to be from the extinct deer Megaloceros . Hooke and Molyneux's line of thinking
16284-620: The North Atlantic, the French West Indies , and many islands in Oceania and the Indian Ocean , giving it one of the largest discontiguous exclusive economic zones in the world . Metropolitan France shares borders with Belgium and Luxembourg to the north, Germany to the northeast, Switzerland to the east, Italy and Monaco to the southeast, Andorra and Spain to the south, and
16461-518: The Paris basin, could be formed by a slow rise and fall of sea levels . The concept of extinction was integral to Charles Darwin 's On the Origin of Species , with less fit lineages disappearing over time. For Darwin, extinction was a constant side effect of competition . Because of the wide reach of On the Origin of Species , it was widely accepted that extinction occurred gradually and evenly (a concept now referred to as background extinction ). It
16638-688: The Pyrenees and the Alps and the Jura, respectively, and to the east, the Rhine river. Metropolitan France includes various coastal islands, of which the largest is Corsica . Metropolitan France is situated mostly between latitudes 41° and 51° N , and longitudes 6° W and 10° E , on the western edge of Europe, and thus lies within the northern temperate zone. Its continental part covers about 1000 km from north to south and from east to west. Metropolitan France covers 551,500 square kilometres (212,935 sq mi),
16815-516: The Revolution. Some of the Enlightenment occurred in French intellectual circles, and scientific breakthroughs, such as the naming of oxygen (1778) and the first hot air balloon carrying passengers (1783), were achieved by French scientists. French explorers took part in the voyages of scientific exploration through maritime expeditions. Enlightenment philosophy, in which reason is advocated as
16992-663: The Rights of Man , which expresses the nation's ideals to this day. France reached its political and military zenith in the early 19th century under Napoleon Bonaparte , subjugating part of continental Europe and establishing the First French Empire . The collapse of the empire initiated a period of relative decline, in which France endured the Bourbon Restoration until the founding of the French Second Republic which
17169-606: The Romans in the Battle of the Allia , and besieged and ransomed Rome. This left Rome weakened, and the Gauls continued to harass the region until 345 BC when they entered into a peace treaty. But the Romans and the Gauls remained adversaries for centuries. Around 125 BC, the south of Gaul was conquered by the Romans, who called this region Provincia Nostra ("Our Province"), which evolved into Provence in French. Julius Caesar conquered
17346-503: The U.S. are of Cretaceous and Paleocene age; and deposits in Mali are of Cretaceous ( Maastrichtian ) age. C. maroccana is more prevalent in Morocco and Jordan, while C. appendiculata is more prevalent in the United States. Both species overlapped at one point in time. Fossil evidence of Cretalamna is found in deposits representing a diverse set of marine environments, indicating that it
17523-720: The United States government, to force the removal of Native Americans , many of whom relied on the bison for food. France – in Europe (green & dark grey) – in the European Union (green) France , officially the French Republic , is a country located primarily in Western Europe . Its overseas regions and territories include French Guiana in South America , Saint Pierre and Miquelon in
17700-429: The accumulation of slightly deleterious mutations , then a population will go extinct. Smaller populations have fewer beneficial mutations entering the population each generation, slowing adaptation. It is also easier for slightly deleterious mutations to fix in small populations; the resulting positive feedback loop between small population size and low fitness can cause mutational meltdown . Limited geographic range
17877-600: The consequences can be catastrophic. Invasive alien species can affect native species directly by eating them, competing with them, and introducing pathogens or parasites that sicken or kill them; or indirectly by destroying or degrading their habitat. Human populations may themselves act as invasive predators. According to the "overkill hypothesis", the swift extinction of the megafauna in areas such as Australia (40,000 years before present), North and South America (12,000 years before present), Madagascar , Hawaii (AD 300–1000), and New Zealand (AD 1300–1500), resulted from
18054-789: The constitutional July Monarchy ; French troops began the conquest of Algeria . Unrest led to the French Revolution of 1848 and the end of the July Monarchy. The abolition of slavery and introduction of male universal suffrage was re-enacted in 1848. In 1852, president of the French Republic, Louis-Napoléon Bonaparte , Napoleon I's nephew, was proclaimed emperor of the Second Empire , as Napoleon III. He multiplied French interventions abroad, especially in Crimea , Mexico and Italy . Napoleon III
18231-418: The contemporary extinction crisis "may be the most serious environmental threat to the persistence of civilization, because it is irreversible." Biologist E. O. Wilson estimated in 2002 that if current rates of human destruction of the biosphere continue, one-half of all plant and animal species of life on earth will be extinct in 100 years. More significantly, the current rate of global species extinctions
18408-541: The death of its last member if it loses the capacity to reproduce and recover. Because a species' potential range may be very large, determining this moment is difficult, and is usually done retrospectively. This difficulty leads to phenomena such as Lazarus taxa , where a species presumed extinct abruptly "reappears" (typically in the fossil record ) after a period of apparent absence. More than 99% of all species that ever lived on Earth , amounting to over five billion species, are estimated to have died out. It
18585-471: The deliberate destruction of some species, such as dangerous viruses , and the total destruction of other problematic species has been suggested. Other species were deliberately driven to extinction, or nearly so, due to poaching or because they were "undesirable", or to push for other human agendas. One example was the near extinction of the American bison , which was nearly wiped out by mass hunts sanctioned by
18762-718: The emergence of cave painting in the Dordogne and Pyrenees , including at Lascaux , dated to c. 18,000 BC. At the end of the Last Glacial Period (10,000 BC), the climate became milder; from approximately 7,000 BC, this part of Western Europe entered the Neolithic era, and its inhabitants became sedentary . After demographic and agricultural development between the 4th and 3rd millennia BC, metallurgy appeared , initially working gold, copper and bronze , then later iron . France has numerous megalithic sites from
18939-516: The empire, leaving England and Aquitaine to the Plantagenets. Charles IV the Fair died without an heir in 1328. The crown passed to Philip of Valois , rather than Edward of Plantagenet, who became Edward III of England . During the reign of Philip, the monarchy reached the height of its medieval power. However Philip's seat on the throne was contested by Edward in 1337, and England and France entered
19116-403: The endangered wild water buffalo is most threatened with extinction by genetic pollution from the abundant domestic water buffalo ). Such extinctions are not always apparent from morphological (non-genetic) observations. Some degree of gene flow is a normal evolutionary process; nevertheless, hybridization (with or without introgression) threatens rare species' existence. The gene pool of
19293-594: The ensuing uprising of European monarchies against his rule, Napoleon was defeated. About a million Frenchmen died during the Napoleonic Wars . After his brief return from exile, Napoleon was finally defeated in 1815 at the Battle of Waterloo , and the Bourbon monarchy was restored with new constitutional limitations. The discredited Bourbon dynasty was overthrown by the July Revolution of 1830, which established
19470-575: The event of rediscovery would be considered Lazarus species. Examples include the thylacine , or Tasmanian tiger ( Thylacinus cynocephalus ), the last known example of which died in Hobart Zoo in Tasmania in 1936; the Japanese wolf ( Canis lupus hodophilax ), last sighted over 100 years ago; the American ivory-billed woodpecker ( Campephilus principalis ), with the last universally accepted sighting in 1944; and
19647-418: The examined teeth may be variable enough to belong to a separate species, but ultimately unified them under a new taxon Otodus appendiculatus . The species would later be found in 1958 by Soviet paleontologist Leonid Glickman to belong to a distinct new genus- Cretalamna . Despite Agassiz's remarks on variability, his ultimately broad interpretation of O. appendiculatus subsequently led the species to become
19824-443: The extinction crisis. According to the International Union for Conservation of Nature (IUCN), 784 extinctions have been recorded since the year 1500, the arbitrary date selected to define "recent" extinctions, up to the year 2004; with many more likely to have gone unnoticed. Several species have also been listed as extinct since 2004. If adaptation increasing population fitness is slower than environmental degradation plus
20001-471: The extinction of Cretalamna is increased competition with newer generalist sharks during the Cenozoic. It is widely believed that Otodus (and thus Carcharocles ) is derived from Cretalamna due to its strong similarity to certain species within the genus. Extinction Extinction is the termination of a taxon by the death of its last member . A taxon may become functionally extinct before
20178-475: The extinction of species caused by humanity, and they try to prevent further extinctions through a variety of conservation programs. Humans can cause extinction of a species through overharvesting , pollution , habitat destruction , introduction of invasive species (such as new predators and food competitors ), overhunting, and other influences. Explosive, unsustainable human population growth and increasing per capita consumption are essential drivers of
20355-533: The field of zoology , and biology in general, and has also become an area of concern outside the scientific community. A number of organizations, such as the Worldwide Fund for Nature , have been created with the goal of preserving species from extinction. Governments have attempted, through enacting laws, to avoid habitat destruction, agricultural over-harvesting, and pollution . While many human-caused extinctions have been accidental, humans have also engaged in
20532-545: The five permanent members of the United Nations Security Council and an official nuclear-weapon state . France is a founding and leading member of the European Union and the eurozone , as well as a member of the Group of Seven , North Atlantic Treaty Organization (NATO), Organisation for Economic Co-operation and Development (OECD), and Francophonie . Originally applied to the whole Frankish Empire ,
20709-582: The forefront of the development of a supranational European Union , notably by signing the Maastricht Treaty in 1992, establishing the eurozone in 1999 and signing the Treaty of Lisbon in 2007. France has fully reintegrated into NATO and since participated in most NATO-sponsored wars. Since the 19th century, France has received many immigrants , often male foreign workers from European Catholic countries who generally returned home when not employed. During
20886-435: The form of a physical destruction of niche habitats. The widespread destruction of tropical rainforests and replacement with open pastureland is widely cited as an example of this; elimination of the dense forest eliminated the infrastructure needed by many species to survive. For example, a fern that depends on dense shade for protection from direct sunlight can no longer survive without forest to shelter it. Another example
21063-823: The four major rivers Seine , the Loire , the Garonne , the Rhône and their tributaries, whose combined catchment includes over 62% of the metropolitan territory. The Rhône divides the Massif Central from the Alps and flows into the Mediterranean Sea at the Camargue . The Garonne meets the Dordogne just after Bordeaux, forming the Gironde estuary , the largest estuary in Western Europe which after approximately 100 kilometres (62 mi) empties into
21240-453: The genus Lamna , which is a romanization of the Ancient Greek λάμνα (lámna, meaning "kind of fierce shark"). When put together they mean "chalk-shark", which refers to chalk deposits from which the species' type specimens were found in. The type species name appendiculata is a feminine form of the Latin word appendiculātus (having an appendage), a reference to the thick bulged roots found in C. appendiculata teeth. The species name lata
21417-473: The genus with the intention of naming it as 'Cretolamna' , but during publication of the corresponding 1958 paper a typographical error occurred, with the print misspelling it as 'Cretalamna' . Glickman pointed out the spelling as an error and continued to use his intended spelling 'Cretolamna' in later works. This spelling was universally adopted until 1999 when Siversson remarked that this violates ICZN Articles 32 and 33, reinstating 'Cretalamna' as
21594-580: The habitat retreat of taxa approaching extinction. Possible causes of the higher extinction risk in species with more sexual selection shown by the comprehensive fossil studies that rule out such error sources include expensive sexually selected ornaments having negative effects on the ability to survive natural selection , as well as sexual selection removing a diversity of genes that under current ecological conditions are neutral for natural selection but some of which may be important for surviving climate change. There have been at least five mass extinctions in
21771-660: The heaviest losses include the Cape Floristic Region and the Caribbean Basin . These areas might see a doubling of present carbon dioxide levels and rising temperatures that could eliminate 56,000 plant and 3,700 animal species. Climate change has also been found to be a factor in habitat loss and desertification . Studies of fossils following species from the time they evolved to their extinction show that species with high sexual dimorphism , especially characteristics in males that are used to compete for mating, are at
21948-447: The history of life on earth, and four in the last 350 million years in which many species have disappeared in a relatively short period of geological time. A massive eruptive event that released large quantities of tephra particles into the atmosphere is considered to be one likely cause of the " Permian–Triassic extinction event " about 250 million years ago, which is estimated to have killed 90% of species then existing. There
22125-475: The human era since the Late Pleistocene would require 5 to 7 million years to recover. According to the 2019 Global Assessment Report on Biodiversity and Ecosystem Services by IPBES , the biomass of wild mammals has fallen by 82%, natural ecosystems have lost about half their area and a million species are at risk of extinction—all largely as a result of human actions. Twenty-five percent of plant and animal species are threatened with extinction. In
22302-431: The introduced ( or hybrid ) species. Endemic populations can face such extinctions when new populations are imported or selectively bred by people, or when habitat modification brings previously isolated species into contact. Extinction is likeliest for rare species coming into contact with more abundant ones; interbreeding can swamp the rarer gene pool and create hybrids, depleting the purebred gene pool (for example,
22479-475: The largest among European Union members. France's total land area, with its overseas departments and territories (excluding Adélie Land ), is 643,801 km (248,573 sq mi), 0.45% of the total land area on Earth. France possesses a wide variety of landscapes, from coastal plains in the north and west to mountain ranges of the Alps in the southeast, the Massif Central in the south-central and Pyrenees in
22656-485: The main drivers of the modern extinction crisis. In January 2020, the UN's Convention on Biological Diversity drafted a plan to mitigate the contemporary extinction crisis by establishing a deadline of 2030 to protect 30% of the Earth's land and oceans and reduce pollution by 50%, with the goal of allowing for the restoration of ecosystems by 2050. The 2020 United Nations ' Global Biodiversity Outlook report stated that of
22833-514: The maximum possible length of C. borealis up to 3.5 meters (11 ft) based on an upper jaw tooth specimen (LO 11350t) from Åsen locality. The body plan of Cretalamna is almost completely known, informed by near-complete fossil impressions with soft tissue preserving the shark's outline from the Hjoula lagerstätte in Lebanon as documented by Pfiel (2021) and Greenfield (2022). It was most similar to
23010-421: The modern understanding of extinction as the end of a species was incompatible with the prevailing worldview. Prior to the 19th century, much of Western society adhered to the belief that the world was created by God and as such was complete and perfect. This concept reached its heyday in the 1700s with the peak popularity of a theological concept called the great chain of being , in which all life on earth, from
23187-484: The most-used language in diplomacy, science, and literature until the 20th century. France took control of territories in the Americas, Africa and Asia. In 1685, Louis XIV revoked the Edict of Nantes , forcing thousands of Huguenots into exile and published the Code Noir providing the legal framework for slavery and expelling Jews from French colonies. Under the wars of Louis XV (r. 1715–1774), France lost New France and most Indian possessions after its defeat in
23364-418: The name France comes from the Latin Francia , or "realm of the Franks ". The name of the Franks is related to the English word frank ("free"): the latter stems from the Old French franc ("free, noble, sincere"), and ultimately from the Medieval Latin word francus ("free, exempt from service; freeman, Frank"), a generalisation of the tribal name that emerged as a Late Latin borrowing of
23541-490: The name of a species or other taxon normally indicates its status as extinct. Examples of species and subspecies that are extinct include: A species is extinct when the last existing member dies. Extinction therefore becomes a certainty when there are no surviving individuals that can reproduce and create a new generation. A species may become functionally extinct when only a handful of individuals survive, which cannot reproduce due to poor health, age, sparse distribution over
23718-469: The natural course of events, species become extinct for a number of reasons, including but not limited to: extinction of a necessary host, prey or pollinator, interspecific competition , inability to deal with evolving diseases and changing environmental conditions (particularly sudden changes) which can act to introduce novel predators, or to remove prey. Recently in geological time, humans have become an additional cause of extinction of some species, either as
23895-410: The newly emerging school of uniformitarianism . Jean-Baptiste Lamarck , a gradualist and colleague of Cuvier, saw the fossils of different life forms as evidence of the mutable character of species. While Lamarck did not deny the possibility of extinction, he believed that it was exceptional and rare and that most of the change in species over time was due to gradual change. Unlike Cuvier, Lamarck
24072-405: The newly established kingdoms. These victories led to the worldwide expansion of French revolutionary ideals and reforms, such as the metric system , Napoleonic Code and Declaration of the Rights of Man. In 1812 Napoleon attacked Russia , reaching Moscow. Thereafter his army disintegrated through supply problems, disease, Russian attacks, and finally winter. After this catastrophic campaign and
24249-448: The off-and-on Hundred Years' War . Boundaries changed, but landholdings inside France by English Kings remained extensive for decades. With charismatic leaders, such as Joan of Arc , French counterattacks won back most English continental territories. France was struck by the Black Death , from which half of the 17 million population died. The French Renaissance saw cultural development and standardisation of French, which became
24426-399: The original population, thereby increasing the chance of extinction. Habitat degradation is currently the main anthropogenic cause of species extinctions. The main cause of habitat degradation worldwide is agriculture, with urban sprawl , logging, mining, and some fishing practices close behind. The degradation of a species' habitat may alter the fitness landscape to such an extent that
24603-464: The palace (head of household). One mayor of the palace, Charles Martel , defeated an Umayyad invasion of Gaul at the Battle of Tours (732). His son, Pepin the Short , seized the crown of Francia from the weakened Merovingians and founded the Carolingian dynasty . Pepin's son, Charlemagne , reunited the Frankish kingdoms and built an empire across Western and Central Europe . Proclaimed Holy Roman Emperor by Pope Leo III and thus establishing
24780-506: The primary source of legitimacy , undermined the power of and support for the monarchy and was a factor in the Revolution. The French Revolution was a period of political and societal change that began with the Estates General of 1789 , and ended with the coup of 18 Brumaire in 1799 and the formation of the French Consulate . Many of its ideas are fundamental principles of liberal democracy , while its values and institutions remain central to modern political discourse. Its causes were
24957-702: The process of speciation —where new varieties of organisms arise and thrive when they are able to find and exploit an ecological niche —and species become extinct when they are no longer able to survive in changing conditions or against superior competition . The relationship between animals and their ecological niches has been firmly established. A typical species becomes extinct within 10 million years of its first appearance, although some species, called living fossils , survive with little to no morphological change for hundreds of millions of years. Mass extinctions are relatively rare events; however, isolated extinctions of species and clades are quite common, and are
25134-465: The reconstructed Frankish endonym * Frank . It has been suggested that the meaning "free" was adopted because, after the conquest of Gaul , only Franks were free of taxation, or more generally because they had the status of freemen in contrast to servants or slaves. The etymology of *Frank is uncertain. It is traditionally derived from the Proto-Germanic word * frankōn , which translates as "javelin" or "lance" (the throwing axe of
25311-410: The relative importance of genetic factors compared to environmental ones as the causes of extinction has been compared to the debate on nature and nurture . The question of whether more extinctions in the fossil record have been caused by evolution or by competition or by predation or by disease or by catastrophe is a subject of discussion; Mark Newman, the author of Modeling Extinction , argues for
25488-413: The remainder of Gaul and overcame a revolt by Gallic chieftain Vercingetorix in 52 BC. Gaul was divided by Augustus into provinces and many cities were founded during the Gallo-Roman period , including Lugdunum (present-day Lyon ), the capital of the Gauls. In 250–290 AD, Roman Gaul suffered a crisis with its fortified borders attacked by barbarians . The situation improved in the first half of
25665-399: The revolt was a political failure (the Gaullist party emerged stronger than before) it announced a split between the French and de Gaulle, who resigned. In the post-Gaullist era, France remained one of the most developed economies in the world but faced crises that resulted in high unemployment rates and increasing public debt. In the late 20th and early 21st centuries, France has been at
25842-436: The second-largest behind the British Empire . Including metropolitan France, the total area reached almost 13 million square kilometres in the 1920s and 1930s, 9% of the world's land. Known as the Belle Époque , the turn of the century was characterised by optimism, regional peace, economic prosperity and technological, scientific and cultural innovations. In 1905, state secularism was officially established . France
26019-421: The southwest. Due to its numerous overseas departments and territories scattered across the planet, France possesses the second-largest exclusive economic zone (EEZ) in the world, covering 11,035,000 km (4,261,000 sq mi). Its EEZ covers approximately 8% of the total surface of all the EEZs of the world. Metropolitan France has a wide variety of topographical sets and natural landscapes. During
26196-460: The species is no longer able to survive and becomes extinct. This may occur by direct effects, such as the environment becoming toxic , or indirectly, by limiting a species' ability to compete effectively for diminished resources or against new competitor species. Habitat destruction, particularly the removal of vegetation that stabilizes soil, enhances erosion and diminishes nutrient availability in terrestrial ecosystems. This degradation can lead to
26373-488: The state and reinforced royal power. He destroyed castles of defiant lords and denounced the use of private armies. By the end of the 1620s, Richelieu established "the royal monopoly of force". France fought in the Thirty Years' War , supporting the Protestant side against the Habsburgs. From the 16th to the 19th century, France was responsible for about 10% of the transatlantic slave trade . During Louis XIV 's minority, trouble known as The Fronde occurred. This rebellion
26550-430: The sudden introduction of human beings to environments full of animals that had never seen them before and were therefore completely unadapted to their predation techniques. Coextinction refers to the loss of a species due to the extinction of another; for example, the extinction of parasitic insects following the loss of their hosts. Coextinction can also occur when a species loses its pollinator , or to predators in
26727-465: The syntypes as the sole lectotype of C. appendiculata in 1999. In 2015, he led a study which revisited the taxonomic situation and established a renewed description of the species, which led to the erection of six additional Cretalamna species- C. catoxodon , C. deschutteri , C. ewelli , C. gertericorum , C. hattini , and C. sarcoporthea . Before Siversson, other Cretalamna species have been described. Another species described by Agassiz under
26904-421: The taxon Cretolamna appendiculata arambourgi , which Siversson et al. (2015) elevated into its own species. In 2018, American paleontologists Jun Ebersole and Dana Ehret described a new species of Cretalamna from various teeth from the Eutaw Formation and Mooreville Chalk in Alabama , which they named C. bryanti . The genus Cretalamna is a portmanteau of creta , the Latin word for "chalk", prefixed to
27081-491: The taxon Lamna borealis . This would be revised to 'Cretolamna borealis' by Glickman in a 1980 paper. In 1902, German paleontologist Johannes Wanner described teeth from Egyptian Cretaceous deposits near the Dakhla Oasis and Farafra . He noted that the teeth are almost identical with that of the Otodus appendiculatus teeth, except that the Egyptian teeth also contained two clear pairs of lateral cusplets (a feature not seen in Otodus appendiculatus ). Wanner concluded that
27258-399: The taxon Otodus latus was demoted to a variation of C. appendiculata in 1908, promoted into a subspecies in 1977 by Belgian paleontologist Jaques Herman, and finally elevated to the species level as Cretolamna lata by Herman and paleontologist Van Waes Hilde in 2012. In 1897, French paleontologist Fernand Priem described a single tooth from the Köpinge Sandstone in Scania , Sweden under
27435-435: The teeth were of a closely related new species and placed it under the taxon Otodus biauriculatus . In 1935, French Paleontologist Camille Arambourg described a new subspecies of C. biauriculata from teeth found in Moroccan phosphates under the taxon Lamna biauriculata maroccana , which was elevated into its own species in 1997. In 1972, French ichthyologist Henri Cappetta described teeth from Maastrichtian deposits near
27612-412: The tiniest microorganism to God, is linked in a continuous chain. The extinction of a species was impossible under this model, as it would create gaps or missing links in the chain and destroy the natural order. Thomas Jefferson was a firm supporter of the great chain of being and an opponent of extinction, famously denying the extinction of the woolly mammoth on the grounds that nature never allows
27789-448: The total extinction of the dodo and the extirpation of indigenous horses to the British Isles. He similarly argued against mass extinctions , believing that any extinction must be a gradual process. Lyell also showed that Cuvier's original interpretation of the Parisian strata was incorrect. Instead of the catastrophic floods inferred by Cuvier, Lyell demonstrated that patterns of saltwater and freshwater deposits , like those seen in
27966-424: The unoccupied territory. Free France , the government-in-exile led by Charles de Gaulle , was set up in London. From 1942 to 1944, about 160,000 French citizens, including around 75,000 Jews , were deported to death and concentration camps . On 6 June 1944, the Allies invaded Normandy , and in August they invaded Provence . The Allies and French Resistance emerged victorious, and French sovereignty
28143-424: The upper jaws extended over the lower jaws, giving Cretalamna a subterminal mouth. Cretalamna was a widespread genus found in North Africa ( Morocco ), the Near East (Jordan), West Africa ( Mali ), North America both on the East Coast and in the Midwest and Central America ( Tonosí , Panama ). Deposits in Morocco are usually Eocene in age; deposits in Jordan are of Cretaceous and Eocene in age; most deposits in
28320-577: The valid spelling. Since then, the reinstatement of 'Cretalamna' gained prominence and by the 2010s, was accepted by the majority of paleontologists. However, some paleontologists including Cappetta strongly opposed it. In an attempt to suppress the usage of 'Cretalamna' , Cappetta appealed to a representative of the ICZN, arguing that the original intentions of Glickman and the prevailing usage of 'Cretolamna' prior to Siversson (1999) secures its priority. The ICZN, who reportedly were impressed by Cappetta's "spirit", subsequently erected Article 33.3.1 of
28497-530: The wider scientific community of his theory. Cuvier was a well-regarded geologist, lauded for his ability to reconstruct the anatomy of an unknown species from a few fragments of bone. His primary evidence for extinction came from mammoth skulls found in the Paris basin . Cuvier recognized them as distinct from any known living species of elephant, and argued that it was highly unlikely such an enormous animal would go undiscovered. In 1812, Cuvier, along with Alexandre Brongniart and Geoffroy Saint-Hilaire , mapped
28674-401: Was invaded by Germany and defended by Great Britain at the start of World War I in August 1914. A rich industrial area in the north was occupied. France and the Allies emerged victorious against the Central Powers at tremendous human cost. It left 1.4 million French soldiers dead, 4% of its population. Interwar was marked by intense international tensions and social reforms introduced by
28851-410: Was a founding member of NATO and attempted to regain control of French Indochina , but was defeated by the Viet Minh in 1954. France faced another anti-colonialist conflict in Algeria , then part of France and home to over one million European settlers ( Pied-Noir ). The French systematically used torture and repression, including extrajudicial killings to keep control. This conflict nearly led to
29028-401: Was a generalist. It was a predator and preyed upon large bony fish, turtles , mosasaurs , squids, and other sharks. For example, multiple teeth of C. appendiculata have been found around elasmosaurid Futabasaurus , suggesting it predated or scavenged that elasmosaur. Some tooth specimens of Cretalamna exhibit heavy wear—likely the result of drastic diet changes. A possible factor to
29205-459: Was able to adapt to a wide range of habitats. This may have attributed to its ability to exist through a long temporal range. The fusiform body of Cretalamna suggests it was a pelagic shark. The Cretaceous waters inhabited by Cretalamna were also home to a diverse range of cartilaginous fishes, bony fishes, turtles, squamates, plesiosaurs, pterosaurs, birds, and even some non-avian dinosaurs. The tooth morphology of Cretalamna implies that it
29382-503: Was among the Allies of World War II , but it surrendered and was occupied in 1940. Following its liberation in 1944 , the short-lived Fourth Republic was established and later dissolved in the course of the defeat in the Algerian War . The current Fifth Republic was formed in 1958 by Charles de Gaulle . Algeria and most French colonies became independent in the 1960s, with the majority retaining close economic and military ties with France . France retains its centuries-long status as
29559-425: Was difficult to disprove. When parts of the world had not been thoroughly examined and charted, scientists could not rule out that animals found only in the fossil record were not simply "hiding" in unexplored regions of the Earth. Georges Cuvier is credited with establishing the modern conception of extinction in a 1796 lecture to the French Institute , though he would spend most of his career trying to convince
29736-451: Was divided into Germanic kingdoms and a remaining Gallo-Roman territory. Celtic Britons , fleeing the Anglo-Saxon settlement of Britain , settled in west Armorica ; the Armorican peninsula was renamed Brittany and Celtic culture was revived. The first leader to unite all Franks was Clovis I , who began his reign as king of the Salian Franks in 481, routing the last forces of the Roman governors in 486. Clovis said he would be baptised
29913-479: Was driven by feudal lords and sovereign courts as a reaction to the royal absolute power . The monarchy reached its peak during the 17th century and reign of Louis XIV. By turning lords into courtiers at the Palace of Versailles , his command of the military went unchallenged. The "Sun King" made France the leading European power. France became the most populous European country and had tremendous influence over European politics, economy, and culture. French became
30090-405: Was executed in January 1793. After another revolt in June 1793 , the constitution was suspended and power passed from the National Convention to the Committee of Public Safety . About 16,000 people were executed in a Reign of Terror , which ended in July 1794 . Weakened by external threats and internal opposition, the Republic was replaced in 1795 by the Directory . Four years later in 1799,
30267-414: Was its precursor. During the 9th and 10th centuries, threatened by Viking invasions , France became a decentralised state: the nobility's titles and lands became hereditary, and authority of the king became more religious than secular, and so was less effective and challenged by noblemen. Thus was established feudalism in France. Some king's vassals grew so powerful they posed a threat to the king. After
30444-430: Was not until 1982, when David Raup and Jack Sepkoski published their seminal paper on mass extinctions, that Cuvier was vindicated and catastrophic extinction was accepted as an important mechanism . The current understanding of extinction is a synthesis of the cataclysmic extinction events proposed by Cuvier, and the background extinction events proposed by Lyell and Darwin. Extinction is an important research topic in
30621-453: Was restored with the Provisional Government of the French Republic (GPRF). This interim government, established by de Gaulle, continued to wage war against Germany and to purge collaborators from office . It made important reforms e.g. suffrage extended to women and the creation of a social security system. A new constitution resulted in the Fourth Republic (1946–1958), which saw strong economic growth ( les Trente Glorieuses ). France
30798-608: Was set to impose a carbon tax in 2009; however, the plan was abandoned due to fears of burdening French businesses. Forests account for 31 per cent of France's land area—the fourth-highest proportion in Europe—representing an increase of 7 per cent since 1990. French forests are some of the most diverse in Europe, comprising more than 140 species of trees. France had a 2018 Forest Landscape Integrity Index mean score of 4.52/10, ranking it 123rd globally. There are nine national parks and 46 natural parks in France. A regional nature park (French: parc naturel régional or PNR)
30975-571: Was skeptical that catastrophic events of a scale large enough to cause total extinction were possible. In his geological history of the earth titled Hydrogeologie, Lamarck instead argued that the surface of the earth was shaped by gradual erosion and deposition by water, and that species changed over time in response to the changing environment. Charles Lyell , a noted geologist and founder of uniformitarianism , believed that past processes should be understood using present day processes. Like Lamarck, Lyell acknowledged that extinction could occur, noting
31152-461: Was succeeded by the Second French Empire upon Napoleon III 's takeover. His empire collapsed during the Franco-Prussian War in 1870. This led to the establishment of the Third French Republic , and subsequent decades saw a period of economic prosperity and cultural and scientific flourishing known as the Belle Époque . France was one of the major participants of World War I , from which it emerged victorious at great human and economic cost. It
31329-404: Was unseated following defeat in the Franco-Prussian War of 1870, and his regime replaced by the Third Republic . By 1875, the French conquest of Algeria was complete, with approximately 825,000 Algerians killed from famine, disease, and violence. France had colonial possessions since the beginning of the 17th century, but in the 19th and 20th centuries its empire extended greatly and became
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