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88-586: During pre-Columbian times, the area of Panama which today includes Coclé province had a number of identifiable native cultures. Archaeologists have loosely designated these cultures by pottery style. The poorly studied La Mula period ranged from 150 BC to AD 300. It was followed by the Tonosi period, from AD 300 to AD 550, and by the Cubita period, from AD 550 to AD 700. A unified Indigenous culture appears to have flourished in this area from approximately 1200 BC until

176-527: A national park . Coclé is also home to APROVACA orchid conservation center, which conducts orchid conservation activities, including orchid reintroduction into the wild and a sponsorship program for the Panamanian national flower Peristeria elata . Folklorist Rosita Liao Gonzales was awarded the Order of Manuel José Hurtado for her work categorizing the cultures of Coclé Province. Pre-Columbian In

264-434: A single-nucleotide polymorphism mutation . More specifically, a haplotype is a combination of alleles at different chromosomal regions that are closely linked and that tend to be inherited together. As a haplogroup consists of similar haplotypes, it is usually possible to predict a haplogroup from haplotypes. Haplogroups pertain to a single line of descent . As such, membership of a haplogroup, by any individual, relies on

352-591: A different social structure. Until the accurate dating of Watson Brake and similar sites, the oldest mound complex was thought to be Poverty Point , also located in the Lower Mississippi Valley . Built about 1500 BCE, it is the centerpiece of a culture extending over 100 sites on both sides of the Mississippi . The Poverty Point site has earthworks in the form of six concentric half-circles, divided by radial aisles, together with some mounds. The entire complex

440-495: A direct female line of descent. Mutations are changes in the nitrogen bases of the DNA sequence . Single changes from the original sequence are called single nucleotide polymorphisms (SNPs). Human Y chromosomes are male-specific sex chromosomes ; nearly all humans that possess a Y chromosome will be morphologically male. Although Y chromosomes are situated in the cell nucleus and paired with X chromosomes , they only recombine with

528-439: A more recent haplogroup (which is a subgroup or subclade ) of its own to which humans carrying only mutation A do not belong. Both mtDNA and Y chromosomes are grouped into lineages and haplogroups; these are often presented as tree-like diagrams. Human Y chromosome DNA (Y-DNA) haplogroups are named from A to T, and are further subdivided using numbers and lower case letters. Y chromosome haplogroup designations are established by

616-730: A population of over 20,000. Other chiefdoms were constructed throughout the Southeast, and its trade networks reached to the Great Lakes and the Gulf of Mexico. At its peak, between the 12th and 13th centuries, Cahokia was the most populous city in North America. (Larger cities did exist in Mesoamerica and the Andes.) Monks Mound , the major ceremonial center of Cahokia, remains the largest earthen construction of

704-543: A relatively small proportion of the genetic material possessed by that individual. Each haplogroup originates from, and remains part of, a preceding single haplogroup (or paragroup ). As such, any related group of haplogroups may be precisely modelled as a nested hierarchy , in which each set (haplogroup) is also a subset of a single broader set (as opposed, that is, to biparental models, such as human family trees). Haplogroups can be further divided into subclades. Haplogroups are normally identified by an initial letter of

792-436: A set of ten Y chromosomes (derived from ten different individuals) contains a mutation, A, but only five of these chromosomes contain a second mutation, B, then it must be the case that mutation B occurred after mutation A. Furthermore, all ten individuals who carry the chromosome with mutation A are the direct male line descendants of the same man who was the first person to carry this mutation. The first man to carry mutation B

880-570: A single unified empire. The Mixtecs would eventually be conquered by the Aztecs until the Spanish conquest. The Mixtecs saw the Spanish conquest as an opportunity for liberation and established agreements with the conquistadors that allowed them to preserve their cultural traditions, though relatively few sections resisted Spanish rule. The Totonac civilization was concentrated in the present-day states of Veracruz and Puebla . The Totonacs were responsible for

968-408: Is contained in the chromosomes in the nucleus of the cell, but mtDNA is an exception. An individual inherits their cytoplasm and the organelles contained by that cytoplasm exclusively from the maternal ovum (egg cell); sperm only pass on the chromosomal DNA, all paternal mitochondria are digested in the oocyte . When a mutation arises in a mtDNA molecule, the mutation is therefore passed down in

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1056-673: Is currently divided into two general approaches. The first is the short chronology theory with the first movement beyond Alaska into the Americas occurring no earlier than 14,000–17,000 years ago, followed by successive waves of immigrants. The second belief is the long chronology theory , which proposes that the first group of people entered the hemisphere at a much earlier date, possibly 50,000–40,000 years ago or earlier. Artifacts have been found in both North and South America which have been dated to 14,000 years ago, and accordingly humans have been proposed to have reached Cape Horn at

1144-411: Is nearly a mile across. Mound building was continued by succeeding cultures, who built numerous sites in the middle Mississippi and Ohio River valleys as well, adding effigy mounds , conical and ridge mounds, and other shapes. The Woodland period of North American pre-Columbian cultures lasted from roughly 1000 BCE to 1000 CE. The term was coined in the 1930s and refers to prehistoric sites between

1232-449: Is passed solely along the patrilineal line, from father to son, while mtDNA is passed down the matrilineal line, from mother to offspring of both sexes. Neither recombines , and thus Y-DNA and mtDNA change only by chance mutation at each generation with no intermixture between parents' genetic material. Mitochondria are small organelles that lie in the cytoplasm of eukaryotic cells , such as those of humans. Their primary function

1320-522: Is reflected in the oral histories of the indigenous peoples, described by a wide range of traditional creation stories which often say that a given people have been living in a certain territory since the creation of the world. Throughout thousands of years, paleo-Indian people domesticated, bred, and cultivated many plant species, including crops that now constitute 50–60% of worldwide agriculture. In general, Arctic, Subarctic, and coastal peoples continued to live as hunters and gatherers, while agriculture

1408-578: Is the Olmec. This civilization established the cultural blueprint by which all succeeding indigenous civilizations would follow in Mexico. Pre-Olmec civilization began with the production of pottery in abundance, around 2300 BCE in the Grijalva River delta. Between 1600 and 1500 BCE, the Olmec civilization had begun, with the consolidation of power at their capital, a site today known as San Lorenzo Tenochtitlán near

1496-617: Is the determinant factor for the number of gene lineages and founding haplotypes present in today's Indigenous populations . Human settlement of the Americas occurred in stages from the Bering Sea coastline , with an initial 20,000-year layover on Beringia for the founding population . The microsatellite diversity and distributions of the Y lineage specific to South America indicate that certain Amerindian populations have been isolated since

1584-412: Is to provide energy to the cell. Mitochondria are thought to be reduced descendants of symbiotic bacteria that were once free living. One indication that mitochondria were once free living is that each contains a circular DNA , called mitochondrial DNA (mtDNA), whose structure is more similar to bacteria than eukaryotic organisms (see endosymbiotic theory ). The overwhelming majority of a human's DNA

1672-678: The Americas via the Bering Land Bridge (Beringia), now the Bering Strait , and possibly along the coast. Genetic evidence found in Indigenous peoples ' maternally inherited mitochondrial DNA (mtDNA) supports the theory of multiple genetic populations migrating from Asia. After crossing the land bridge, they moved southward along the Pacific coast and through an interior ice-free corridor. Throughout millennia, Paleo-Indians spread throughout

1760-594: The Archaic Period , numerous archaeological cultures have been identified. The unstable climate led to widespread migration, with early Paleo-Indians soon spreading throughout the Americas, diversifying into many hundreds of culturally distinct tribes. The Paleo-Indians were hunter-gatherers , likely characterized by small, mobile bands consisting of approximately 20 to 50 members of an extended family. These groups moved from place to place as preferred resources were depleted and new supplies were sought. During much of

1848-658: The Archaic period and the Mississippian cultures . The Adena culture and the ensuing Hopewell tradition during this period built monumental earthwork architecture and established continent-spanning trade and exchange networks. This period is considered a developmental stage without any massive changes in a short period but instead has a continuous development in stone and bone tools, leatherworking, textile manufacture, tool production, cultivation, and shelter construction. Some Woodland people continued to use spears and atlatls until

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1936-672: The Cliff Palace of Mesa Verde National Park in Colorado and the Great Houses in Chaco Canyon , New Mexico . The Puebloans also constructed a road system that stretched from Chaco Canyon to Kutz Canyon in the San Juan Basin . The Ancestral Puebloans are also known as "Anasazi", though the term is controversial, as the present-day Pueblo peoples consider the term to be derogatory, due to

2024-670: The Maya script . Other accounts also suggest that the Huastecs migrated as a result of the Classic Maya collapse around the year 900 CE. The Zapotecs were a civilization that thrived in the Oaxaca Valley from the late 6th century BCE until their downfall at the hands of the Spanish conquistadors. The city of Monte Albán was an important religious center for the Zapotecs and served as the capital of

2112-651: The Olmec , Teotihuacan , Mayas , Zapotecs , Mixtecs , Huastecs , Purepecha , Toltecs , and Mexica / Aztecs . The Mexica civilization is also known as the Aztec Triple Alliance since they were three smaller kingdoms loosely united together. These Indigenous civilizations are credited with many inventions: building pyramid temples, mathematics , astronomy , medicine, writing, highly accurate calendars , fine arts , intensive agriculture, engineering , an abacus calculator, and complex theology . They also invented

2200-681: The Sami people . Here is a list of Y-chromosome and MtDNA geographic haplogroup assignment proposed by Bekada et al. 2013. According to SNPS haplogroups which are the age of the first extinction event tend to be around 45–50 kya. Haplogroups of the second extinction event seemed to diverge 32–35 kya according to Mal'ta . The ground zero extinction event appears to be Toba during which haplogroup CDEF* appeared to diverge into C, DE and F. C and F have almost nothing in common while D and E have plenty in common. Extinction event #1 according to current estimates occurred after Toba, although older ancient DNA could push

2288-516: The Spanish conquest of El Salvador , Cuzcatlan was forced to surrender to conquistador Pedro de Alvarado in 1528. Haplogroup A haplotype is a group of alleles in an organism that are inherited together from a single parent, and a haplogroup ( haploid from the Greek : ἁπλοῦς , haploûs , "onefold, simple" and English: group ) is a group of similar haplotypes that share a common ancestor with

2376-458: The Tlingit , Haida , Chumash , Mandan , Hidatsa , and others, and some established large settlements, even cities, such as Cahokia , in what is now Illinois . Mesoamerica is the region extending from central Mexico south to the northwestern border of Costa Rica that gave rise to a group of stratified, culturally related agrarian civilizations spanning an approximately 3,000-year period before

2464-632: The history of the Americas , the pre-Columbian era , also known as the pre-contact era , or as the pre-Cabraline era specifically in Brazil , spans from the initial peopling of the Americas in the Upper Paleolithic to the onset of European colonization , which began with Christopher Columbus 's voyage in 1492. This era encompasses the history of Indigenous cultures prior to significant European influence, which in some cases did not occur until decades or even centuries after Columbus's arrival. During

2552-407: The 16th century. Coclé Province is divided into 6 distritos (districts) and subdivided into 44 corregimientos . Estimate 2010 Estimate 2019 Parque Nacional General de División Omar Torrijos Herrera , established around the crash site of former Panamanian leader Omar Torrijos , is located in the northeastern portion of Coclé. Parque Omar, as it is known, enjoys some legal protection as

2640-625: The Americas and oral histories. Other civilizations, contemporaneous with the colonial period, were documented in European accounts of the time. For instance, the Maya civilization maintained written records, which were often destroyed by Christian Europeans such as Diego de Landa , who viewed them as pagan but sought to preserve native histories. Despite the destruction, a few original documents have survived, and others were transcribed or translated into Spanish, providing modern historians with valuable insights into ancient cultures and knowledge. Before

2728-561: The Americas, and Melanesia, as well as in parts of the Horn of Africa and North Africa; almost none have been found in Europe. The N haplogroup may represent another macrolineage that evolved outside of Africa, heading northward instead of eastward. Shortly after the migration, the large R group split off from the N. Haplogroup R consists of two subgroups defined on the basis of their geographical distributions, one found in southeastern Asia and Oceania and

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2816-472: The Europeans arrived, Indigenous peoples of North America had a wide range of lifeways from sedentary, agrarian societies to semi-nomadic hunter-gatherer societies. Many formed new tribes or confederations in response to European colonization. These are often classified by cultural regions , loosely based on geography. These can include the following: Numerous pre-Columbian societies were sedentary, such as

2904-648: The Hohokam, they constructed kivas and great houses as well as ballcourts . Several of the Sinagua ruins include Montezuma Castle , Wupatki , and Tuzigoot . The Salado resided in the Tonto Basin in southeastern Arizona from 1150 CE to the 15th century. Archaeological evidence suggests that they traded with far-away cultures, as evidenced by the presence of seashells from the Gulf of California and macaw feathers from Mexico. Most of

2992-1304: The Horn of Africa and the Arabian Peninsula (Saudi Arabia, Yemen, Oman), separated only by a narrow strait between the Red Sea and the Gulf of Aden. CZ – Many Siberians; branch C – Some Amerindian; branch Z – Many Saami, some Korean, some North Chinese, some Central Asian populations. D – Some Amerindians, many Siberians and northern East Asians E – Malay, Borneo, Philippines, Taiwanese aborigines , Papua New Guinea G – Many Northeast Siberians, northern East Asians, and Central Asians Q – Melanesian, Polynesian, New Guinean populations The N type consists of: A – Found in many Amerindians and some East Asians and Siberians I – 10% frequency in Northern, Eastern Europe S – Some Indigenous Australian (First Nations People of Australia) W – Some Eastern Europeans, South Asians, and southern East Asians X – Some Amerindians, Southern Siberians, Southwest Asians, and Southern Europeans Y – Most Nivkhs and people of Nias ; many Ainus, Tungusic people , and Austronesians ; also found with low frequency in some other populations of Siberia, East Asia, and Central Asia R – Large group found within

3080-663: The Levant (modern Lebanon area), found in 25% frequency in Bedouin populations; branch JT (branch J; branch T) – North, Eastern Europe, Indus, Mediterranean U – High frequency in West Eurasia, Indian sub-continent, and Algeria, found from India to the Mediterranean and to the rest of Europe; U5 in particular shows high frequency in Scandinavia and Baltic countries with the highest frequency in

3168-707: The Maya city of Chichen Itza . The Toltecs established vast trading relations with other Mesoamerican civilizations in Central America and the Puebloans in present-day New Mexico . During the Post-Classic era, the Toltecs suffered a subsequent collapse in the early 12th century, due to famine and civil war. The Toltec civilization was so influential to the point where many groups such as the Aztecs claimed to be descended from. With

3256-450: The Maya civilization cannot be overstated: it transformed political power, artistic depictions, and the nature of economics. Within the city of Teotihuacan was a diverse and cosmopolitan population. Most of the regional ethnicities of Mexico were represented in the city, such as Zapotecs from the Oaxaca region. They lived in apartment communities where they worked their trades and contributed to

3344-802: The Mississippian groups had vanished, and vast swaths of their territory were virtually uninhabited. The Ancestral Puebloans thrived in what is now the Four Corners region in the United States. It is commonly suggested that the culture of the Ancestral Puebloans emerged during the Early Basketmaker II Era during the 12th century BCE. The Ancestral Puebloans were a complex Oasisamerican society that constructed kivas , multi-story houses, and apartment blocks made from stone and adobe, such as

3432-841: The N type. Populations contained therein can be divided geographically into West Eurasia and East Eurasia. Almost all European populations and a large number of Middle-Eastern population today are contained within this branch. A smaller percentage is contained in other N type groups (See above). Below are subclades of R : B – Some Chinese, Tibetans, Mongolians, Central Asians, Koreans, Amerindians, South Siberians, Japanese, Austronesians F – Mainly found in southeastern Asia, especially Vietnam ; 8.3% in Hvar Island in Croatia. R0 – Found in Arabia and among Ethiopians and Somalis; branch HV (branch H; branch V) – Europe, Western Asia, North Africa; Pre-JT – Arose in

3520-483: The Olmec resulted in a power vacuum in Mexico. Emerging from that vacuum was Teotihuacan, first settled in 300 BCE. By 150 CE, Teotihuacan had risen to become the first true metropolis of what is now called North America. Teotihuacan established a new economic and political order never before seen in Mexico. Its influence stretched across Mexico into Central America, founding new dynasties in the Maya cities of Tikal , Copan , and Kaminaljuyú . Teotihuacan's influence over

3608-732: The Olmecs, Teotihuacan, the Toltecs, the Mexica, and the Mayas. These civilizations (except for the politically fragmented Maya) extended their reach across Mesoamerica—and beyond—like no others. They consolidated power and distributed influence in matters of trade, art, politics, technology, and theology. Other regional power players made economic and political alliances with these civilizations over 4,000 years. Many made war with them, but almost all peoples found themselves within one of their spheres of influence. Regional communications in ancient Mesoamerica have been

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3696-662: The Paleo-Indian period, bands are thought to have subsisted primarily through hunting now-extinct giant land animals such as mastodon and ancient bison . Paleo-Indian groups carried a variety of tools, including distinctive projectile points and knives, as well as less distinctive butchering and hide-scraping implements. The vastness of the North American continent, and the variety of its climates, ecology , vegetation , fauna , and landforms, led ancient peoples to coalesce into many distinct linguistic and cultural groups. This

3784-459: The Spanish colonists. The Wichita people were a loose confederation that consisted of sedentary agriculturalists and hunter-gatherers who resided in the eastern Great Plains . They lived in permanent settlements and even established a city called Etzanoa , which had a population of 20,000 people. The city was eventually abandoned around the 18th century after it was encountered by Spanish conquistadors Jusepe Gutierrez and Juan de Oñate . When

3872-602: The Tlaxcalans for preserving their culture and for their assistance in defeating the Aztecs. The Tlaxcalans would once again assist to the Spaniards during the Mixtón War and the conquest of Guatemala . Cuzcatlan was a Pipil confederacy of kingdoms and city-states located in present-day El Salvador . According to legend, Cuzcatlan was established by Toltec migrants during the Classic Maya collapse in approximately 1200 CE. During

3960-467: The X chromosome at the ends of the Y chromosome ; the remaining 95% of the Y chromosome does not recombine. Therefore, the Y chromosome and any mutations that arise in it are passed down in a direct male line of descent. Other chromosomes, autosomes and X chromosomes (when another X chromosome is available to pair with it), share their genetic material during meiosis , the process of cell division which produces gametes . Effectively this means that

4048-715: The Y Chromosome Consortium. Y-chromosomal Adam is the name given by researchers to the male who is the most recent common patrilineal (male-lineage) ancestor of all living humans. Major Y-chromosome haplogroups, and their geographical regions of occurrence (prior to the recent European colonization), include: (mutation M168 occurred ~50,000 bp) (mutation M89 occurred ~45,000 bp) (mutation M9 occurred ~40,000 bp ) Human mtDNA haplogroups are lettered: A , B , C , CZ , D , E , F , G , H , HV , I , J , pre- JT , JT , K , L0 , L1 , L2 , L3 , L4 , L5 , L6 , M , N , O , P , Q , R , R0 , S , T , U , V , W , X , Y , and Z . The most up-to-date version of

4136-418: The alphabet, and refinements consist of additional number and letter combinations, such as (for example) A → A1 → A1a . The alphabetical nomenclature was published in 2002 by the Y Chromosome Consortium . In human genetics , the haplogroups most commonly studied are Y-chromosome (Y-DNA) haplogroups and mitochondrial DNA (mtDNA) haplogroups , each of which can be used to define genetic populations . Y-DNA

4224-547: The calendar, were bequest from the former inhabitants of Tula, the Toltecs. The Mexica-Aztecs were the rulers of much of central Mexico by about 1400 (while Yaquis , Coras, and Apaches commanded sizable regions of northern desert), having subjugated most of the other regional states by the 1470s. At their peak, the Valley of Mexico where the Aztec Empire presided, saw a population growth that included nearly one million people during

4312-507: The city's economic and cultural prowess. Teotihuacan's economic pull impacted areas in northern Mexico as well. It was a city whose monumental architecture reflected a monumental new era in Mexican civilization, declining in political power about 650 CE—but lasting in cultural influence for the better part of a millennium, to around 950 CE. Contemporary to Teotihuacan's greatness was that of the Maya civilization. The period between 250 CE and 650 CE

4400-516: The civilizations in its area, the Tarascan Empire was the most prominent in metallurgy, harnessing copper, silver, and gold to create items such as tools, decorations, and even weapons and armor. Bronze was also used. The great victories over the Aztecs by the Tarascans cannot be understated. Nearly every war they fought in resulted in a Tarascan victory. Because the Tarascan Empire had little links to

4488-712: The cliff dwellings constructed by the Salado are primarily located in Tonto National Monument . The Iroquois League of Nations or "People of the Long House" was a politically advanced, democratic society, which is thought by some historians to have influenced the United States Constitution , with the Senate passing a resolution to this effect in 1988. Other historians have contested this interpretation and believe

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4576-404: The coast in southeast Veracruz . The Olmec influence extended across Mexico, into Central America , and along the Gulf of Mexico . They transformed many peoples' thinking toward a new way of government, pyramid temples, writing, astronomy, art, mathematics, economics, and religion. Their achievements paved the way for the Maya civilization and the civilizations in central Mexico. The decline of

4664-541: The de Soto expedition wandered the American Southeast for four years, becoming more bedraggled, losing more men and equipment, and eventually arriving in Mexico as a fraction of its original size. The local people fared much worse though, as the fatalities of diseases introduced by the expedition devastated the populations and produced much social disruption. By the time Europeans returned a hundred years later, nearly all of

4752-656: The decline of the Toltec civilization came political fragmentation in the Valley of Mexico . Into this new political game of contenders to the Toltec throne stepped outsiders: the Mexica . They were also a desert people, one of seven groups who formerly called themselves "Azteca", in memory of Aztlán , but they changed their name after years of migrating. Since they were not from the Valley of Mexico , they were initially seen as crude and unrefined in

4840-451: The development of archaeology in the 19th century, historians of the pre-Columbian period mainly interpreted the records of the European conquerors and the accounts of early European travelers and antiquaries. It was not until the nineteenth century that the work of people such as John Lloyd Stephens , Eduard Seler , and Alfred Maudslay , and institutions such as the Peabody Museum of Archaeology and Ethnology of Harvard University , led to

4928-430: The empire from 700 BCE to 700 CE. The Zapotecs resisted the expansion of the Aztecs until they were subjugated in 1502 under Aztec emperor Ahuitzotl . After the Spanish conquest of the Aztec Empire , the Zapotecs resisted Spanish rule until King Cosijopii I surrendered in 1563. Like the Zapotecs, the Mixtecs thrived in the Oaxaca Valley. The Mixtecs consisted of separate independent kingdoms and city-states, rather than

5016-409: The end of the period when they were replaced by bows and arrows . The Mississippian culture was spread across the Southeast and Midwest of what is today the United States, from the Atlantic coast to the edge of the plains, from the Gulf of Mexico to the Upper Midwest, although most intensively in the area along the Mississippi River and Ohio River . One of the distinguishing features of this culture

5104-408: The establishment of cities, such as El Tajín as important commercial trading centers. The Totonacs would later assist in the Spanish conquest of the Aztec Empire as an opportunity to liberate themselves from Aztec military imperialism. The Toltec civilization was established in the 8th century CE. The Toltec Empire expanded its political borders to as far south as the Yucatán peninsula , including

5192-403: The first Cazonci, Tariacuri, united these communities and built them into one of the most advanced civilizations in Mesoamerica. Their capital at Tzintzuntzan was just one of the many cities—there were ninety more under its control. The Tarascan Empire was among the largest in Central America, so it is no surprise that they routinely came into conflict with the neighboring Aztec Empire . Out of all

5280-559: The following are common divisions for mtDNA haplogroups: The mitochondrial haplogroups are divided into three main groups, which are designated by the sequential letters L, M, N. Humanity first split within the L group between L0 and L1-6. L1-6 gave rise to other L groups, one of which, L3, split into the M and N group. The M group comprises the first wave of human migration which is thought to have evolved outside of Africa, following an eastward route along southern coastal areas. Descendant lineages of haplogroup M are now found throughout Asia,

5368-400: The former Toltec Empire , they were also quite independent in culture from their neighbors. The Aztecs, Tlaxcaltec , Olmec, Mixtec, Maya, and others were very similar to each other, however. This is because they were all directly preceded by the Toltecs, and they therefore shared almost identical cultures. The Tarascans, however, possessed a unique religion, as well as other things. Tlaxcala

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5456-438: The genetic material from these chromosomes gets mixed up in every generation, and so any new mutations are passed down randomly from parents to offspring. The special feature that both Y chromosomes and mtDNA display is that mutations can accrue along a certain segment of both molecules and these mutations remain fixed in place on the DNA. Furthermore, the historical sequence of these mutations can also be inferred. For example, if

5544-414: The ground zero extinction event to long before Toba, and push the first extinction event here back to Toba. Haplogroups with extinction event notes by them have a dubious origin and this is because extinction events lead to severe bottlenecks, so all notes by these groups are just guesses. Note that the SNP counting of ancient DNA can be highly variable meaning that even though all these groups diverged around

5632-543: The impact was minimal or did not exist, pointing to numerous differences between the two systems and the ample precedents for the constitution in European political thought. The Calusa were a complex paramountcy/kingdom that resided in southern Florida . Instead of agriculture, the Calusa economy relied on abundant fishing. According to Spanish sources, the "king's house" at Mound Key was large enough to house 2,000 people. The Calusa ultimately collapsed into extinction at around 1750 after succumbing to diseases introduced by

5720-687: The influence that astronomical activities had upon Mesoamerican people before the arrival of Europeans. Many of the later Mesoamerican civilizations carefully built their cities and ceremonial centers according to specific astronomical events. The biggest Mesoamerican cities, such as Teotihuacan , Tenochtitlan , and Cholula , were among the largest in the world. These cities grew as centers of commerce, ideas, ceremonies, and theology, and they radiated influence outwards onto neighboring cultures in central Mexico. While many city-states, kingdoms, and empires competed with one another for power and prestige, Mesoamerica can be said to have had five major civilizations:

5808-429: The initial colonization of the region. The Na-Dené , Inuit , and Indigenous Alaskan populations exhibit haplogroup Q-M242 (Y-DNA) mutations, however, and are distinct from other Indigenous peoples with various mtDNA mutations. This suggests that the earliest migrants into the northern extremes of North America and Greenland derived from later populations. Asian nomadic Paleo-Indians are thought to have entered

5896-472: The late Aztec period (1350–1519). Their capital, Tenochtitlan , is the site of modern-day Mexico City . At its peak, it was one of the largest cities in the world with population estimates of 200,000–300,000. The market established there was the largest ever seen by the conquistadores on arrival. Initially, the lands that would someday comprise the lands of the powerful Tarascan Empire were inhabited by several independent communities. Around 1300, however,

5984-607: The late twentieth century, archeologists have studied, analyzed, and dated these sites, realizing that the earliest complexes were built by hunter-gatherer societies, whose people occupied the sites on a seasonal basis. Watson Brake , a large complex of eleven platform mounds, was constructed beginning in 3400 BCE and added to over 500 years. This has changed earlier assumptions that complex construction arose only after societies had adopted agriculture, and become sedentary, with stratified hierarchy and usually ceramics. These ancient people had organized to build complex mound projects under

6072-400: The majority of the Y-chromosome is unique and does not recombine during meiosis . This has the effect that the historical pattern of mutations can easily be studied. The pattern indicates Indigenous peoples of the Americas experienced two very distinctive genetic episodes: first with the initial peopling of the Americas and second with European colonization of the Americas . The former

6160-416: The mtDNA tree is maintained by Mannis van Oven on the PhyloTree website. Phylogenetic tree of human mitochondrial DNA (mtDNA) haplogroups Mitochondrial Eve is the name given by researchers to the woman who is the most recent common matrilineal (female-lineage) ancestor of all living humans. Haplogroups can be used to define genetic populations and are often geographically oriented. For example,

6248-457: The only true writing system native to the Americas using pictographs and syllabic elements in the form of texts and codices inscribed on stone, pottery, wood, or perishable books made from bark paper. The Huastecs were a Maya ethnic group that migrated northwards to the Gulf Coast of Mexico. The Huastecs are considered to be distinct from the Maya civilization, as they separated from the main Maya branch at around 2000 BCE and did not possess

6336-458: The other containing almost all of the modern European populations. Haplogroup N(xR), i.e. mtDNA that belongs to the N group but not to its R subgroup, is typical of Australian aboriginal populations, while also being present at low frequencies among many populations of Eurasia and the Americas. The L type consists of nearly all Africans. The M type consists of: M1 – Ethiopian, Somali and Indian populations. Likely due to much gene flow between

6424-646: The people abandoned their settlements, likely due to drought. The Mogollon resided in the present-day states of Arizona , New Mexico, and Texas as well as Sonora and Chihuahua . Like most other cultures in Oasisamerica, the Mogollon constructed sophisticated kivas and cliff dwellings. In the village of Paquimé , the Mogollon are revealed to have housed pens for scarlet macaws , which were introduced from Mesoamerica through trade. The Sinagua were hunter-gatherers and agriculturalists who lived in central Arizona. Like

6512-422: The pre-Columbian era, many civilizations developed permanent settlements, cities, agricultural practices, civic and monumental architecture, major earthworks , and complex societal hierarchies. Some of these civilizations had declined by the time of the establishment of the first permanent European colonies, around the late 16th to early 17th centuries, and are known primarily through archaeological research of

6600-416: The prehistoric Americas . The culture reached its peak in about 1200–1400 CE, and in most places, it seems to have been in decline before the arrival of Europeans. Many Mississippian peoples were encountered by the expedition of Hernando de Soto in the 1540s, mostly with disastrous results for both sides. Unlike the Spanish expeditions in Mesoamerica, which conquered vast empires with relatively few men,

6688-613: The reconsideration and criticism of the early European sources. Now, the scholarly study of pre-Columbian cultures is most often based on scientific and multidisciplinary methodologies. The haplogroup most commonly associated with Indigenous Amerindian genetics is Y-chromosome haplogroup Q1a3a . Researchers have found genetic evidence that the Q1a3a haplogroup has been in South America since at least 18,000 BCE. Y-chromosome DNA , like mtDNA , differs from other nuclear chromosomes in that

6776-511: The rest of North and South America. Exactly when the first people migrated into the Americas is the subject of much debate. One of the earliest identifiable cultures was the Clovis culture , with sites dating from some 13,000 years ago. However, older sites dating back to 20,000 years ago have been claimed. Some genetic studies estimate the colonization of the Americas dates from between 40,000 and 13,000 years ago. The chronology of migration models

6864-545: The southern tip of South America by this time. In that case, the Inuit would have arrived separately and at a much later date, probably no more than 2,000 years ago, moving across the ice from Siberia into Alaska. The North American climate was unstable as the ice age receded during the Lithic stage . It finally stabilized about 10,000 years ago; climatic conditions were then very similar to today's. Within this time frame, roughly about

6952-545: The subject of considerable research. There is evidence of trade routes starting as far north as the Mexico Central Plateau , and going down to the Pacific coast. These trade routes and cultural contacts then went on as far as Central America . These networks operated with various interruptions from pre-Olmec times and up to the Late Classical Period (600–900 CE). The earliest known civilization in Mesoamerica

7040-550: The visits to the Caribbean by Christopher Columbus. Mesoamerican is the adjective generally used to refer to that group of pre-Columbian cultures. This refers to an environmental area occupied by an assortment of ancient cultures that shared religious beliefs, art, architecture, and technology in the Americas for more than three thousand years. Between 2000 and 300 BCE, complex cultures began to form in Mesoamerica. Some matured into advanced pre-Columbian Mesoamerican civilizations such as

7128-496: The ways of the Nahua civilization. Through political maneuvers and ferocious martial skills, they managed to rule Mexico as the head of the 'Triple Alliance' which included two other Aztec cities, Tetxcoco and Tlacopan . Latecomers to Mexico's central plateau , the Mexica thought of themselves, nevertheless, as heirs of the civilizations that had preceded them. For them, arts, sculpture, architecture, engraving, feather-mosaic work, and

7216-420: The wheel, but it was used solely as a toy. In addition, they used native copper , silver , and gold for metalworking. Archaic inscriptions on rocks and rock walls all over northern Mexico (especially in the state of Nuevo León ) demonstrate an early propensity for counting. Their number system was base 20 and included zero . These early count markings were associated with astronomical events and underscore

7304-654: The word tracing its origins to a Navajo word meaning "ancestor enemies". The Hohokam thrived in the Sonoran desert in what is now the U.S. state of Arizona and the Mexican state of Sonora . The Hohokam were responsible for the construction of a series of irrigation canals that led to the successful establishment of Phoenix, Arizona via the Salt River Project . The Hohokam also established complex settlements such as Snaketown , which served as an important commercial trading center. After 1375 CE, Hohokam society collapsed and

7392-487: Was a Nahua republic and confederation in central Mexico. The Tlaxcalans fiercely resisted Aztec expansion during the Flower Wars ever since the Aztecs expelled them from Lake Texcoco . The Tlaxcalans would later ally with the Spanish conquistadors under Hernán Cortés as an opportunity to liberate them from the Aztecs and managed to successfully conquer the Aztecs with the help of the conquistadors. The Spaniards would reward

7480-416: Was a time of intense flourishing of Maya civilized accomplishments. While the many Maya city-states never achieved political unity on the order of the central Mexican civilizations, they exerted tremendous intellectual influence upon Mexico and Central America. The Maya built some of the most elaborate cities on the continent and made innovations in mathematics, astronomy, and calendrics. The Maya also developed

7568-573: Was adopted in more temperate and sheltered regions, permitting a dramatic rise in population. After the migration or migrations, it was several thousand years before the first complex societies arose, the earliest emerging about seven to eight thousand years ago. As early as 5500 BCE, people in the Lower Mississippi Valley at Monte Sano and other sites in present-day Louisiana , Mississippi , and Florida were building complex earthwork mounds , probably for religious purposes. Beginning in

7656-422: Was also a direct male line descendant of this man, but is also the direct male line ancestor of all men carrying mutation B. Series of mutations such as this form molecular lineages. Furthermore, each mutation defines a set of specific Y chromosomes called a haplogroup. All humans carrying mutation A form a single haplogroup, and all humans carrying mutation B are part of this haplogroup, but mutation B also defines

7744-538: Was the construction of complexes of large earthen mounds and grand plazas, continuing the mound-building traditions of earlier cultures. They grew maize and other crops intensively, participated in an extensive trade network, and had a complex stratified society. The Mississippians first appeared around 1000 CE, following and developing out of the less agriculturally intensive and less centralized Woodland period. The largest urban site of these people, Cahokia —located near modern East St. Louis, Illinois —may have reached

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