81-429: The chlorarachniophytes are a small group of exclusively marine algae widely distributed in tropical and temperate waters. They are typically mixotrophic , ingesting bacteria and smaller protists as well as conducting photosynthesis . Normally they have the form of small amoebae , with branching cytoplasmic extensions that capture prey and connect the cells together, forming a net. These extensions are dependent on
162-565: A West Germanic language that originated from the Anglo-Frisian dialects brought to Britain by Germanic settlers from various parts of what is now northwest Germany and the northern Netherlands. The resident population at this time was generally speaking Common Brittonic —the insular variety of Continental Celtic , which was influenced by the Roman occupation. This group of languages ( Welsh , Cornish , Cumbric ) cohabited alongside English into
243-571: A and b . Their chloroplasts are surrounded by four and three membranes, respectively, and were probably retained from ingested green algae. Chlorarachniophytes , which belong to the phylum Cercozoa , contain a small nucleomorph , which is a relict of the algae's nucleus . Euglenids , which belong to the phylum Euglenozoa , live primarily in fresh water and have chloroplasts with only three membranes. The endosymbiotic green algae may have been acquired through myzocytosis rather than phagocytosis . (Another group with green algae endosymbionts
324-408: A and c , and phycobilins. The shape can vary; they may be of discoid, plate-like, reticulate, cup-shaped, spiral, or ribbon shaped. They have one or more pyrenoids to preserve protein and starch. The latter chlorophyll type is not known from any prokaryotes or primary chloroplasts, but genetic similarities with red algae suggest a relationship there. In the first three of these groups ( Chromista ),
405-528: A polyphyletic group since they do not include a common ancestor , and although their chlorophyll -bearing plastids seem to have a single origin (from symbiogenesis with cyanobacteria ), they were acquired in different ways. Green algae are a prominent examples of algae that have primary chloroplasts derived from endosymbiont cyanobacteria. Diatoms and brown algae are examples of algae with secondary chloroplasts derived from endosymbiotic red algae , which they acquired via phagocytosis . Algae exhibit
486-549: A century as Received Pronunciation (RP). However, due to language evolution and changing social trends, some linguists argue that RP is losing prestige or has been replaced by another accent, one that the linguist Geoff Lindsey for instance calls Standard Southern British English. Others suggest that more regionally-oriented standard accents are emerging in England. Even in Scotland and Northern Ireland, RP exerts little influence in
567-526: A few species of dinoflagellates , which also have plastids originating from green algae, and the cryptomonads , which acquired their chloroplasts from a red alga . The chlorarachniophytes only include five genera, which show some variation in their life-cycles and may lack one or two of the stages described above. Genetic studies place them among the Cercozoa , a diverse group of amoeboid and amoeboid-like protozoa. The chlorarachniophytes were placed before in
648-539: A form and capabilities not possessed by the symbiont species alone (they can be experimentally isolated). The photobiont possibly triggers otherwise latent genes in the mycobiont. Trentepohlia is an example of a common green alga genus worldwide that can grow on its own or be lichenised. Lichen thus share some of the habitat and often similar appearance with specialized species of algae ( aerophytes ) growing on exposed surfaces such as tree trunks and rocks and sometimes discoloring them. Coral reefs are accumulated from
729-501: A greater movement, normally [əʊ], [əʉ] or [əɨ]. Dropping a morphological grammatical number , in collective nouns , is stronger in British English than North American English. This is to treat them as plural when once grammatically singular, a perceived natural number prevails, especially when applying to institutional nouns and groups of people. The noun 'police', for example, undergoes this treatment: Police are investigating
810-406: A lesser class or social status and often discounted or considered of a low intelligence. Another contribution to the standardisation of British English was the introduction of the printing press to England in the mid-15th century. In doing so, William Caxton enabled a common language and spelling to be dispersed among the entirety of England at a much faster rate. Samuel Johnson's A Dictionary of
891-592: A lineage that eventually led to the higher land plants. The innovation that defines these nonalgal plants is the presence of female reproductive organs with protective cell layers that protect the zygote and developing embryo. Hence, the land plants are referred to as the Embryophytes . The term algal turf is commonly used but poorly defined. Algal turfs are thick, carpet-like beds of seaweed that retain sediment and compete with foundation species like corals and kelps , and they are usually less than 15 cm tall. Such
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#1733085283128972-648: A process called T-glottalisation . National media, being based in London, have seen the glottal stop spreading more widely than it once was in word endings, not being heard as "no [ʔ] " and bottle of water being heard as "bo [ʔ] le of wa [ʔ] er". It is still stigmatised when used at the beginning and central positions, such as later , while often has all but regained /t/ . Other consonants subject to this usage in Cockney English are p , as in pa [ʔ] er and k as in ba [ʔ] er. In most areas of England and Wales, outside
1053-478: A red dye derived from it. The Latinization, fūcus , meant primarily the cosmetic rouge. The etymology is uncertain, but a strong candidate has long been some word related to the Biblical פוך ( pūk ), 'paint' (if not that word itself), a cosmetic eye-shadow used by the ancient Egyptians and other inhabitants of the eastern Mediterranean. It could be any color: black, red, green, or blue. The study of algae
1134-511: A regional accent or dialect. However, about 2% of Britons speak with an accent called Received Pronunciation (also called "the King's English", "Oxford English" and " BBC English" ), that is essentially region-less. It derives from a mixture of the Midlands and Southern dialects spoken in London in the early modern period. It is frequently used as a model for teaching English to foreign learners. In
1215-463: A turf may consist of one or more species, and will generally cover an area in the order of a square metre or more. Some common characteristics are listed: Many algae, particularly species of the Characeae , have served as model experimental organisms to understand the mechanisms of the water permeability of membranes, osmoregulation , turgor regulation , salt tolerance , cytoplasmic streaming , and
1296-989: A wide range of reproductive strategies, from simple asexual cell division to complex forms of sexual reproduction via spores . Algae lack the various structures that characterize plants (which evolved from freshwater green algae), such as the phyllids (leaf-like structures) and rhizoids of bryophytes ( non-vascular plants ), and the roots , leaves and other xylemic / phloemic organs found in tracheophytes ( vascular plants ). Most algae are autotrophic , although some are mixotrophic , deriving energy both from photosynthesis and uptake of organic carbon either by osmotrophy , myzotrophy or phagotrophy . Some unicellular species of green algae, many golden algae , euglenids , dinoflagellates , and other algae have become heterotrophs (also called colorless or apochlorotic algae), sometimes parasitic , relying entirely on external energy sources and have limited or no photosynthetic apparatus. Some other heterotrophic organisms, such as
1377-725: Is also due to London-centric influences. Examples of R-dropping are car and sugar , where the R is not pronounced. British dialects differ on the extent of diphthongisation of long vowels, with southern varieties extensively turning them into diphthongs, and with northern dialects normally preserving many of them. As a comparison, North American varieties could be said to be in-between. Long vowels /iː/ and /uː/ are usually preserved, and in several areas also /oː/ and /eː/, as in go and say (unlike other varieties of English, that change them to [oʊ] and [eɪ] respectively). Some areas go as far as not diphthongising medieval /iː/ and /uː/, that give rise to modern /aɪ/ and /aʊ/; that is, for example, in
1458-454: Is an informal term for any organisms of a large and diverse group of photosynthetic eukaryotes , which include species from multiple distinct clades . Such organisms range from unicellular microalgae such as Chlorella , Prototheca and the diatoms , to multicellular macroalgae such as the giant kelp , a large brown alga which may grow up to 50 metres (160 ft) in length. Most algae are aquatic organisms and lack many of
1539-769: Is included in style guides issued by various publishers including The Times newspaper, the Oxford University Press and the Cambridge University Press . The Oxford University Press guidelines were originally drafted as a single broadsheet page by Horace Henry Hart, and were at the time (1893) the first guide of their type in English; they were gradually expanded and eventually published, first as Hart's Rules , and in 2002 as part of The Oxford Manual of Style . Comparable in authority and stature to The Chicago Manual of Style for published American English ,
1620-550: Is most commonly called phycology (from Greek phykos 'seaweed'); the term algology is falling out of use. One definition of algae is that they "have chlorophyll as their primary photosynthetic pigment and lack a sterile covering of cells around their reproductive cells ". On the other hand, the colorless Prototheca under Chlorophyta are all devoid of any chlorophyll. Although cyanobacteria are often referred to as "blue-green algae", most authorities exclude all prokaryotes , including cyanobacteria, from
1701-536: Is quite different from those of reds and browns, because they have distinct nodes, separated by internode 'stems'; whorls of branches reminiscent of the horsetails occur at the nodes. Conceptacles are another polyphyletic trait; they appear in the coralline algae and the Hildenbrandiales , as well as the browns. Most of the simpler algae are unicellular flagellates or amoeboids , but colonial and nonmotile forms have developed independently among several of
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#17330852831281782-469: Is the Latin word for 'seaweed' and retains that meaning in English. The etymology is obscure. Although some speculate that it is related to Latin algēre , 'be cold', no reason is known to associate seaweed with temperature. A more likely source is alliga , 'binding, entwining'. The Ancient Greek word for 'seaweed' was φῦκος ( phŷkos ), which could mean either the seaweed (probably red algae) or
1863-423: Is the dinoflagellate genus Lepidodinium , which has replaced its original endosymbiont of red algal origin with one of green algal origin. A nucleomorph is present, and the host genome still have several red algal genes acquired through endosymbiotic gene transfer. Also the euglenid and chlorarachniophyte genome contain genes of apparent red algal ancestry) These groups have chloroplasts containing chlorophylls
1944-506: Is thus protected from predators; the sponge is provided with oxygen and sugars which can account for 50 to 80% of sponge growth in some species. British English British English (abbreviations: BrE , en-GB , and BE ) is the set of varieties of the English language native to the United Kingdom of Great Britain and Northern Ireland . More narrowly, it can refer specifically to
2025-530: The Chambers Dictionary , and the Collins Dictionary record actual usage rather than attempting to prescribe it. In addition, vocabulary and usage change with time; words are freely borrowed from other languages and other varieties of English, and neologisms are frequent. For historical reasons dating back to the rise of London in the ninth century, the form of language spoken in London and
2106-732: The Calymmian period , early in Boring Billion , but it is difficult to track the key events because of so much time gap. Primary symbiogenesis gave rise to three divisions of archaeplastids , namely the Viridiplantae ( green algae and later plants ), Rhodophyta ( red algae ) and Glaucophyta ("grey algae"), whose plastids further spread into other protist lineages through eukaryote-eukaryote predation , engulfments and subsequent endosymbioses (secondary and tertiary symbiogenesis). This process of serial cell "capture" and "enslavement" explains
2187-637: The East Midlands became standard English within the Court, and ultimately became the basis for generally accepted use in the law, government, literature and education in Britain. The standardisation of British English is thought to be from both dialect levelling and a thought of social superiority. Speaking in the Standard dialect created class distinctions; those who did not speak the standard English would be considered of
2268-509: The English language in England , or, more broadly, to the collective dialects of English throughout the British Isles taken as a single umbrella variety, for instance additionally incorporating Scottish English , Welsh English , and Northern Irish English . Tom McArthur in the Oxford Guide to World English acknowledges that British English shares "all the ambiguities and tensions [with]
2349-609: The Infusoria (microscopic organisms). Unlike macroalgae , which were clearly viewed as plants, microalgae were frequently considered animals because they are often motile. Even the nonmotile (coccoid) microalgae were sometimes merely seen as stages of the lifecycle of plants, macroalgae, or animals. Although used as a taxonomic category in some pre-Darwinian classifications, e.g., Linnaeus (1753), de Jussieu (1789), Lamouroux (1813), Harvey (1836), Horaninow (1843), Agassiz (1859), Wilson & Cassin (1864), in further classifications,
2430-588: The International Association for Lichenology to be "an association of a fungus and a photosynthetic symbiont resulting in a stable vegetative body having a specific structure". The fungi, or mycobionts, are mainly from the Ascomycota with a few from the Basidiomycota . In nature, they do not occur separate from lichens. It is unknown when they began to associate. One or more mycobiont associates with
2511-582: The Late Cambrian / Early Ordovician period, from sessile shallow freshwater charophyte algae much like Chara , which likely got stranded ashore when riverine / lacustrine water levels dropped during dry seasons . These charophyte algae probably already developed filamentous thalli and holdfasts that superficially resembled plant stems and roots , and probably had an isomorphic alternation of generations . They perhaps evolved some 850 mya and might even be as early as 1 Gya during
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2592-484: The Royal Spanish Academy with Spanish. Standard British English differs notably in certain vocabulary, grammar, and pronunciation features from standard American English and certain other standard English varieties around the world. British and American spelling also differ in minor ways. The accent, or pronunciation system, of standard British English, based in southeastern England, has been known for over
2673-486: The Scots language or Scottish Gaelic ). Each group includes a range of dialects, some markedly different from others. The various British dialects also differ in the words that they have borrowed from other languages. Around the middle of the 15th century, there were points where within the 5 major dialects there were almost 500 ways to spell the word though . Following its last major survey of English Dialects (1949–1950),
2754-560: The University of Leeds has started work on a new project. In May 2007 the Arts and Humanities Research Council awarded a grant to Leeds to study British regional dialects. The team are sifting through a large collection of examples of regional slang words and phrases turned up by the "Voices project" run by the BBC , in which they invited the public to send in examples of English still spoken throughout
2835-976: The Vindhya basin have been dated to 1.6 to 1.7 billion years ago. Because of the wide range of algae types, they have increasingly different industrial and traditional applications in human society. Traditional seaweed farming practices have existed for thousands of years and have strong traditions in East Asia food cultures. More modern algaculture applications extend the food traditions for other applications, including cattle feed, using algae for bioremediation or pollution control, transforming sunlight into algae fuels or other chemicals used in industrial processes, and in medical and scientific applications. A 2020 review found that these applications of algae could play an important role in carbon sequestration to mitigate climate change while providing lucrative value-added products for global economies. The singular alga
2916-601: The West Country and other near-by counties of the UK, the consonant R is not pronounced if not followed by a vowel, lengthening the preceding vowel instead. This phenomenon is known as non-rhoticity . In these same areas, a tendency exists to insert an R between a word ending in a vowel and a next word beginning with a vowel. This is called the intrusive R . It could be understood as a merger, in that words that once ended in an R and words that did not are no longer treated differently. This
2997-601: The apicomplexans are also parasites derived from ancestors that possessed plastids, but are not included in any group traditionally seen as algae. Algae are polyphyletic thus their origin cannot be traced back to single hypothetical common ancestor . It is thought that they came into existence when photosynthetic coccoid cyanobacteria got phagocytized by a unicellular heterotrophic eukaryote (a protist ), giving rise to double-membranous primary plastids . Such symbiogenic events (primary symbiogenesis) are believed to have occurred more than 1.5 billion years ago during
3078-435: The apicomplexans , are also derived from cells whose ancestors possessed chlorophyllic plastids, but are not traditionally considered as algae. Algae have photosynthetic machinery ultimately derived from cyanobacteria that produce oxygen as a byproduct of splitting water molecules , unlike other organisms that conduct anoxygenic photosynthesis such as purple and green sulfur bacteria . Fossilized filamentous algae from
3159-545: The calcareous exoskeletons of marine invertebrates of the order Scleractinia (stony corals ). These animals metabolize sugar and oxygen to obtain energy for their cell-building processes, including secretion of the exoskeleton, with water and carbon dioxide as byproducts. Dinoflagellates (algal protists) are often endosymbionts in the cells of the coral-forming marine invertebrates, where they accelerate host-cell metabolism by generating sugar and oxygen immediately available through photosynthesis using incident light and
3240-432: The endoplasmic reticulum , and contain a small nucleomorph between the middle two, which is a remnant of the alga's nucleus. This contains a small amount of DNA and divides without forming a mitotic spindle. The origin of the chloroplasts from green algae is supported by their pigmentation, which includes chlorophylls a and b , and by genetic similarities. The only other groups of algae that contain nucleomorphs are
3321-720: The "algae" are seen as an artificial, polyphyletic group. Throughout the 20th century, most classifications treated the following groups as divisions or classes of algae: cyanophytes , rhodophytes , chrysophytes , xanthophytes , bacillariophytes , phaeophytes , pyrrhophytes ( cryptophytes and dinophytes ), euglenophytes , and chlorophytes . Later, many new groups were discovered (e.g., Bolidophyceae ), and others were splintered from older groups: charophytes and glaucophytes (from chlorophytes), many heterokontophytes (e.g., synurophytes from chrysophytes, or eustigmatophytes from xanthophytes), haptophytes (from chrysophytes), and chlorarachniophytes (from xanthophytes). With
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3402-603: The 21st century. RP, while long established as the standard English accent around the globe due to the spread of the British Empire , is distinct from the standard English pronunciation in some parts of the world; most prominently, RP notably contrasts with standard North American accents. In the 21st century, dictionaries like the Oxford English Dictionary , the Longman Dictionary of Contemporary English ,
3483-816: The English Language (1755) was a large step in the English-language spelling reform , where the purification of language focused on standardising both speech and spelling. By the early 20th century, British authors had produced numerous books intended as guides to English grammar and usage, a few of which achieved sufficient acclaim to have remained in print for long periods and to have been reissued in new editions after some decades. These include, most notably of all, Fowler's Modern English Usage and The Complete Plain Words by Sir Ernest Gowers . Detailed guidance on many aspects of writing British English for publication
3564-644: The Germanic schwein ) is the animal in the field bred by the occupied Anglo-Saxons and pork (like the French porc ) is the animal at the table eaten by the occupying Normans. Another example is the Anglo-Saxon cu meaning cow, and the French bœuf meaning beef. Cohabitation with the Scandinavians resulted in a significant grammatical simplification and lexical enrichment of the Anglo-Frisian core of English;
3645-891: The Oxford Manual is a fairly exhaustive standard for published British English that writers can turn to in the absence of specific guidance from their publishing house. British English is the basis of, and very similar to, Commonwealth English . Commonwealth English is English as spoken and written in the Commonwealth countries , though often with some local variation. This includes English spoken in Australia , Malta , New Zealand , Nigeria , and South Africa . It also includes South Asian English used in South Asia, in English varieties in Southeast Asia , and in parts of Africa. Canadian English
3726-701: The South East, there are significantly different accents; the Cockney accent spoken by some East Londoners is strikingly different from Received Pronunciation (RP). Cockney rhyming slang can be (and was initially intended to be) difficult for outsiders to understand, although the extent of its use is often somewhat exaggerated. Londoners speak with a mixture of accents, depending on ethnicity, neighbourhood, class, age, upbringing, and sundry other factors. Estuary English has been gaining prominence in recent decades: it has some features of RP and some of Cockney. Immigrants to
3807-537: The UK in recent decades have brought many more languages to the country and particularly to London. Surveys started in 1979 by the Inner London Education Authority discovered over 125 languages being spoken domestically by the families of the inner city's schoolchildren. Notably Multicultural London English , a sociolect that emerged in the late 20th century spoken mainly by young, working-class people in multicultural parts of London . Since
3888-621: The United Kingdom , as well as within the countries themselves. The major divisions are normally classified as English English (or English as spoken in England (which is itself broadly grouped into Southern English , West Country , East and West Midlands English and Northern English ), Northern Irish English (in Northern Ireland), Welsh English (not to be confused with the Welsh language ), and Scottish English (not to be confused with
3969-458: The West Scottish accent. Phonological features characteristic of British English revolve around the pronunciation of the letter R, as well as the dental plosive T and some diphthongs specific to this dialect. Once regarded as a Cockney feature, in a number of forms of spoken British English, /t/ has become commonly realised as a glottal stop [ʔ] when it is in the intervocalic position, in
4050-670: The abandonment of plant-animal dichotomous classification, most groups of algae (sometimes all) were included in Protista , later also abandoned in favour of Eukaryota . However, as a legacy of the older plant life scheme, some groups that were also treated as protozoans in the past still have duplicated classifications (see ambiregnal protists ). Some parasitic algae (e.g., the green algae Prototheca and Helicosporidium , parasites of metazoans, or Cephaleuros , parasites of plants) were originally classified as fungi , sporozoans , or protistans of incertae sedis , while others (e.g.,
4131-410: The adjective little is predominant elsewhere. Nevertheless, there is a meaningful degree of uniformity in written English within the United Kingdom, and this could be described by the term British English . The forms of spoken English, however, vary considerably more than in most other areas of the world where English is spoken and so a uniform concept of British English is more difficult to apply to
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#17330852831284212-481: The award of the grant in 2007, Leeds University stated: that they were "very pleased"—and indeed, "well chuffed"—at receiving their generous grant. He could, of course, have been "bostin" if he had come from the Black Country , or if he was a Scouser he would have been well "made up" over so many spondoolicks, because as a Geordie might say, £460,000 is a "canny load of chink". Most people in Britain speak with
4293-451: The carbon dioxide produced by the host. Reef-building stony corals ( hermatypic corals ) require endosymbiotic algae from the genus Symbiodinium to be in a healthy condition. The loss of Symbiodinium from the host is known as coral bleaching , a condition which leads to the deterioration of a reef. Endosymbiontic green algae live close to the surface of some sponges, for example, breadcrumb sponges ( Halichondria panicea ). The alga
4374-420: The chloroplast has four membranes, retaining a nucleomorph in cryptomonads , and they likely share a common pigmented ancestor, although other evidence casts doubt on whether the heterokonts , Haptophyta , and cryptomonads are in fact more closely related to each other than to other groups. The typical dinoflagellate chloroplast has three membranes, but considerable diversity exists in chloroplasts within
4455-624: The composition of the three major groups of algae. Their lineage relationships are shown in the figure in the upper right. Many of these groups contain some members that are no longer photosynthetic. Some retain plastids, but not chloroplasts, while others have lost plastids entirely. Phylogeny based on plastid not nucleocytoplasmic genealogy: Cyanobacteria Glaucophytes Rhodophytes Stramenopiles Cryptophytes Haptophytes Euglenophytes Chlorarachniophytes Chlorophytes Charophytes Land plants (Embryophyta) These groups have green chloroplasts containing chlorophylls
4536-613: The country. The BBC Voices project also collected hundreds of news articles about how the British speak English from swearing through to items on language schools. This information will also be collated and analysed by Johnson's team both for content and for where it was reported. "Perhaps the most remarkable finding in the Voices study is that the English language is as diverse as ever, despite our increased mobility and constant exposure to other accents and dialects through TV and radio". When discussing
4617-436: The definition of algae. The algae contain chloroplasts that are similar in structure to cyanobacteria. Chloroplasts contain circular DNA like that in cyanobacteria and are interpreted as representing reduced endosymbiotic cyanobacteria . However, the exact origin of the chloroplasts is different among separate lineages of algae, reflecting their acquisition during different endosymbiotic events. The table below describes
4698-493: The distinct cell and tissue types, such as stomata , xylem and phloem that are found in land plants . The largest and most complex marine algae are called seaweeds . In contrast, the most complex freshwater forms are the Charophyta , a division of green algae which includes, for example, Spirogyra and stoneworts . Algae that are carried passively by water are plankton , specifically phytoplankton . Algae constitute
4779-425: The diversity of photosynthetic eukaryotes. Recent genomic and phylogenomic approaches have significantly clarified plastid genome evolution , the horizontal movement of endosymbiont genes to the "host" nuclear genome , and plastid spread throughout the eukaryotic tree of life . Fossils of isolated spores suggest land plants may have been around as long as 475 million years ago (mya) during
4860-432: The first to divide macroscopic algae into four divisions based on their pigmentation. This is the first use of a biochemical criterion in plant systematics. Harvey's four divisions are: red algae (Rhodospermae), brown algae (Melanospermae), green algae (Chlorospermae), and Diatomaceae. At this time, microscopic algae were discovered and reported by a different group of workers (e.g., O. F. Müller and Ehrenberg ) studying
4941-530: The genera Volvox and Corallina , and a species of Acetabularia (as Madrepora ), among the animals. In 1768, Samuel Gottlieb Gmelin (1744–1774) published the Historia Fucorum , the first work dedicated to marine algae and the first book on marine biology to use the then new binomial nomenclature of Linnaeus. It included elaborate illustrations of seaweed and marine algae on folded leaves. W. H. Harvey (1811–1866) and Lamouroux (1813) were
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#17330852831285022-446: The generation of action potentials . Plant hormones are found not only in higher plants, but in algae, too. Some species of algae form symbiotic relationships with other organisms. In these symbioses, the algae supply photosynthates (organic substances) to the host organism providing protection to the algal cells. The host organism derives some or all of its energy requirements from the algae. Examples are: Lichens are defined by
5103-538: The green algae Phyllosiphon and Rhodochytrium , parasites of plants, or the red algae Pterocladiophila and Gelidiocolax mammillatus , parasites of other red algae, or the dinoflagellates Oodinium , parasites of fish) had their relationship with algae conjectured early. In other cases, some groups were originally characterized as parasitic algae (e.g., Chlorochytrium ), but later were seen as endophytic algae. Some filamentous bacteria (e.g., Beggiatoa ) were originally seen as algae. Furthermore, groups like
5184-563: The group, and a number of endosymbiotic events apparently occurred. The Apicomplexa , a group of closely related parasites, also have plastids called apicoplasts , which are not photosynthetic, but appear to have a common origin with dinoflagellate chloroplasts. Linnaeus , in Species Plantarum (1753), the starting point for modern botanical nomenclature , recognized 14 genera of algae, of which only four are currently considered among algae. In Systema Naturae , Linnaeus described
5265-453: The groups. Some of the more common organizational levels, more than one of which may occur in the lifecycle of a species, are In three lines, even higher levels of organization have been reached, with full tissue differentiation. These are the brown algae, —some of which may reach 50 m in length ( kelps ) —the red algae, and the green algae. The most complex forms are found among the charophyte algae (see Charales and Charophyta ), in
5346-453: The idea of two different morphemes, one that causes the double negation, and one that is used for the point or the verb. Standard English in the United Kingdom, as in other English-speaking nations, is widely enforced in schools and by social norms for formal contexts but not by any singular authority; for instance, there is no institution equivalent to the Académie française with French or
5427-511: The last southern Midlands accent to use the broad "a" in words like bath or grass (i.e. barth or grarss ). Conversely crass or plastic use a slender "a". A few miles northwest in Leicestershire the slender "a" becomes more widespread generally. In the town of Corby , five miles (8 km) north, one can find Corbyite which, unlike the Kettering accent, is largely influenced by
5508-417: The late phase of the Boring Billion . A range of algal morphologies is exhibited, and convergence of features in unrelated groups is common. The only groups to exhibit three-dimensional multicellular thalli are the reds and browns , and some chlorophytes . Apical growth is constrained to subsets of these groups: the florideophyte reds, various browns, and the charophytes. The form of charophytes
5589-508: The later Norman occupation led to the grafting onto that Germanic core of a more elaborate layer of words from the Romance branch of the European languages. This Norman influence entered English largely through the courts and government. Thus, English developed into a "borrowing" language of great flexibility and with a huge vocabulary . Dialects and accents vary amongst the four countries of
5670-452: The mass internal migration to Northamptonshire in the 1940s and given its position between several major accent regions, it has become a source of various accent developments. In Northampton the older accent has been influenced by overspill Londoners. There is an accent known locally as the Kettering accent, which is a transitional accent between the East Midlands and East Anglian . It is
5751-458: The modern period, but due to their remoteness from the Germanic languages , influence on English was notably limited . However, the degree of influence remains debated, and it has recently been argued that its grammatical influence accounts for the substantial innovations noted between English and the other West Germanic languages. Initially, Old English was a diverse group of dialects, reflecting
5832-965: The order Rhizochloridales, class Xanthophyceae (e.g., Smith, 1938 ), as algae, or in order Rhizochloridea, class Xanthomonadina (e.g., Deflandre, 1956), as protozoa. So far sexual reproduction has only been reported in two species; Chlorarachnion reptans and Cryptochlora perforans . Based on Shiratori et al. 2024. Amorphochlora amoebiformis Viridiuvalis adhaerens Gymnochlora dimorpha Gymnochlora stellata L. globosa L. oceanica L. reticulosa L. vacuolata Partenskyella glossopodia Chlorarachnion reptans Norrisiella sphaerica Bigelowiella natans Bigelowiella longifila Minorisa minuta Minorisa fusiformis Minorisa megafusiformis Minorisa magna Rhabdamoeba marina Alga Algae ( UK : / ˈ æ l ɡ iː / AL -ghee , US : / ˈ æ l dʒ iː / AL -jee ; sg. : alga / ˈ æ l ɡ ə / AL -gə )
5913-408: The presence of light and polymerization of the actin cytoskeleton . They may also form flagellate zoospores , which characteristically have a single subapical flagellum that spirals backwards around the cell body, and walled coccoid cells. The chloroplasts were presumably acquired by ingesting some green alga . They are surrounded by four membranes, the outermost of which is continuous with
5994-405: The same phycobiont species, from the green algae, except that alternatively, the mycobiont may associate with a species of cyanobacteria (hence "photobiont" is the more accurate term). A photobiont may be associated with many different mycobionts or may live independently; accordingly, lichens are named and classified as fungal species. The association is termed a morphogenesis because the lichen has
6075-401: The spoken language. Globally, countries that are former British colonies or members of the Commonwealth tend to follow British English, as is the case for English used by European Union institutions. In China, both British English and American English are taught. The UK government actively teaches and promotes English around the world and operates in over 200 countries . English is
6156-587: The theft of work tools worth £500 from a van at the Sprucefield park and ride car park in Lisburn. A football team can be treated likewise: Arsenal have lost just one of 20 home Premier League matches against Manchester City. This tendency can be observed in texts produced already in the 19th century. For example, Jane Austen , a British author, writes in Chapter 4 of Pride and Prejudice , published in 1813: All
6237-403: The traditional accent of Newcastle upon Tyne , 'out' will sound as 'oot', and in parts of Scotland and North-West England, 'my' will be pronounced as 'me'. Long vowels /iː/ and /uː/ are diphthongised to [ɪi] and [ʊu] respectively (or, more technically, [ʏʉ], with a raised tongue), so that ee and oo in feed and food are pronounced with a movement. The diphthong [oʊ] is also pronounced with
6318-733: The varied origins of the Anglo-Saxon kingdoms of England. One of these dialects, Late West Saxon , eventually came to dominate. The original Old English was then influenced by two waves of invasion: the first was by speakers of the Scandinavian branch of the Germanic family, who settled in parts of Britain in the eighth and ninth centuries; the second was the Normans in the 11th century, who spoke Old Norman and ultimately developed an English variety of this called Anglo-Norman . These two invasions caused English to become "mixed" to some degree (though it
6399-402: The word 'British' and as a result can be used and interpreted in two ways, more broadly or more narrowly, within a range of blurring and ambiguity". Variations exist in formal (both written and spoken) English in the United Kingdom. For example, the adjective wee is almost exclusively used in parts of Scotland, north-east England, Northern Ireland, Ireland, and occasionally Yorkshire , whereas
6480-560: The world are good and agreeable in your eyes. However, in Chapter 16, the grammatical number is used. The world is blinded by his fortune and consequence. Some dialects of British English use negative concords, also known as double negatives . Rather than changing a word or using a positive, words like nobody, not, nothing, and never would be used in the same sentence. While this does not occur in Standard English, it does occur in non-standard dialects. The double negation follows
6561-422: Was never a truly mixed language in the strictest sense of the word; mixed languages arise from the cohabitation of speakers of different languages, who develop a hybrid tongue for basic communication). The more idiomatic, concrete and descriptive English is, the more it is from Anglo-Saxon origins. The more intellectual and abstract English is, the more it contains Latin and French influences, e.g. swine (like
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