The Carnian pluvial episode (CPE), often called the Carnian pluvial event , was a period of major change in global climate that coincided with significant changes in Earth's biota both in the sea and on land. It occurred during the latter part of the Carnian Stage, a subdivision of the late Triassic period, and lasted for perhaps 1–2 million years (around 234–232 million years ago).
204-427: The CPE corresponds to a significant episode in the evolution and diversification of many taxa that are important today, among them some of the earliest dinosaurs (which include the ancestors of birds), lepidosaurs (the ancestors of modern-day snakes and lizards) and mammaliaforms (ancestors of mammals). In the marine realm it saw the first appearance among the microplankton of coccoliths and dinoflagellates , with
408-607: A gazetteer compiled by Chang Qu ( 常璩 ) during the Western Jin Dynasty (265–316), reported the discovery of dragon bones at Wucheng in Sichuan Province. Villagers in central China have long unearthed fossilized "dragon bones" for use in traditional medicines . In Europe , dinosaur fossils were generally believed to be the remains of giants and other biblical creatures. Scholarly descriptions of what would now be recognized as dinosaur bones first appeared in
612-513: A platycopid tolerant of shallow siliciclastic seas. Further shallowing across the Julian-Tuvalian boundary left only a few aberrant limnocytherids ( Renngartenella , Simeonella ) and cytherurids ( Kerocythere ) which could manage severe salinity fluctuations in the restricted coastal basins. Bairdiids returned in force at the end of the crisis when the basins deepened, reacquiring carbonate and better ventilation. The recent discovery of
816-510: A sacrum composed of three or more fused vertebrae (three are found in some other archosaurs, but only two are found in Herrerasaurus ); and a perforate acetabulum , or hip socket, with a hole at the center of its inside surface (closed in Saturnalia tupiniquim , for example). Another difficulty of determining distinctly dinosaurian features is that early dinosaurs and other archosaurs from
1020-421: A "pillar-erect" configuration of the hip joint, where instead of having a projection from the femur insert on a socket on the hip, the upper pelvic bone was rotated to form an overhanging shelf. Dinosaur fossils have been known for millennia, although their true nature was not recognized. The Chinese considered them to be dragon bones and documented them as such. For example, Huayang Guo Zhi ( 華陽國志 ),
1224-618: A branch of the Bourbons ascended to the throne of Sicily and Naples. During the Napoleonic Wars , north and central Italy were reorganised as Sister Republics of France and, later, as a Kingdom of Italy . The south was administered by Joachim Murat , Napoleon's brother-in-law. 1814's Congress of Vienna restored the situation of the late 18th century, but the ideals of the French Revolution could not be eradicated, and re-surfaced during
1428-460: A consequence of redox fluctuations. The ratio of mercury to organic carbon is stronger and occurs earlier in areas corresponding to open marine environments. Although the mercury spikes do not correlate with any indicators of terrestrial runoff, runoff could help maintain high mercury concentrations in the ocean through the CPE. The most parsimonious explanation is that the mercury was initially derived from
1632-488: A controversial taxon that was recently confirmed to exist after archived photos were uncovered. Bruhathkayosaurus was a titanosaur and would have most likely weighed more than even Marrapunisaurus . Recent size estimates in 2023 have placed this sauropod reaching lengths of up to 44 m (144 ft) long and a colossal weight range of around 110 000 – 170 000 kg ( 240 000 – 370 000 lb), if these upper estimates up true, Bruhathkayosaurus would have rivaled
1836-448: A corresponding inwardly facing distinct head on the femur. Their erect posture enabled early dinosaurs to breathe easily while moving, which likely permitted stamina and activity levels that surpassed those of "sprawling" reptiles . Erect limbs probably also helped support the evolution of large size by reducing bending stresses on limbs. Some non-dinosaurian archosaurs, including rauisuchians , also had erect limbs but achieved this by
2040-514: A crucially innovative role in financial development by devising banking practices, and enabling new forms of social organisation. In coastal and southern areas, maritime republics dominated the Mediterranean and monopolised trade to the Orient. They were independent thalassocratic city-states, in which merchants had considerable power. Although oligarchical, the relative political freedom they afforded
2244-632: A dinosaur. In it he described and named a sauropod tooth , " Rutellum impicatum ", that had been found in Caswell, near Witney , Oxfordshire. Between 1815 and 1824, the Rev William Buckland , the first Reader of Geology at the University of Oxford, collected more fossilized bones of Megalosaurus and became the first person to describe a non-avian dinosaur in a scientific journal . The second non-avian dinosaur genus to be identified, Iguanodon ,
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#17330862510842448-523: A famous meeting between Garibaldi and Victor Emmanuel II , the last king of Sardinia, during which Garibaldi shook Victor Emanuel's hand and hailed him as King of Italy . Cavour agreed to include Garibaldi's southern Italy in a union with the Kingdom of Sardinia in 1860. This allowed the Sardinian government to declare a united Italian kingdom on 17 March 1861, with Victor Emmanuel II as its first king. In 1865,
2652-537: A femur of Tyrannosaurus preserved soft, flexible tissue within, including blood vessels , bone matrix , and connective tissue (bone fibers) that had retained their microscopic structure. This discovery suggested that original soft tissues could be preserved over geological time, with multiple mechanisms having been proposed. Later, in 2009, Schweitzer and colleagues reported that a Brachylophosaurus femur preserved similar microstructures, and immunohistochemical techniques (based on antibody binding) demonstrated
2856-422: A group known as archosaurs, which also includes modern crocodilians. Within the archosaur group, dinosaurs are differentiated most noticeably by their gait. Dinosaur legs extend directly beneath the body, whereas the legs of lizards and crocodilians sprawl out to either side. Collectively, dinosaurs as a clade are divided into two primary branches, Saurischia and Ornithischia. Saurischia includes those taxa sharing
3060-728: A member of NATO . The Marshall Plan revived the economy, which, until the late 1960s, enjoyed a period called the Economic Miracle . In the 1950s, Italy became a founding country of the European Communities , a forerunner of the European Union. From the late 1960s until the early 1980s, the country experienced the Years of Lead , characterised by economic difficulties, especially after the 1973 oil crisis ; social conflicts; and terrorist massacres. The economy recovered and Italy became
3264-482: A more recent common ancestor with birds than with Ornithischia, while Ornithischia includes all taxa sharing a more recent common ancestor with Triceratops than with Saurischia. Anatomically, these two groups can be distinguished most noticeably by their pelvic structure. Early saurischians—"lizard-hipped", from the Greek sauros ( σαῦρος ) meaning "lizard" and ischion ( ἰσχίον ) meaning "hip joint"—retained
3468-583: A now lost partial vertebral neural arch described in 1878. Extrapolating from the illustration of this bone, the animal may have been 58 meters (190 ft) long and weighed 122 400 kg ( 269 800 lb). However, recent research have placed Amphicoelias from the long, gracile diplodocid to the shorter but much stockier rebbachisaurid. Now renamed as Maraapunisaurus , this sauropod now stands as much as 40 meters (130 ft) long and weigh as much as 120 000 kg ( 260 000 lb). Another contender of this title includes Bruhathkayosaurus ,
3672-433: A period of time, with estimates ranging from 5–10 million years to 21 million years. When dinosaurs appeared, they were not the dominant terrestrial animals. The terrestrial habitats were occupied by various types of archosauromorphs and therapsids , like cynodonts and rhynchosaurs . Their main competitors were the pseudosuchians , such as aetosaurs , ornithosuchids and rauisuchians, which were more successful than
3876-624: A permanent seat in the League of Nations 's executive council. The socialist agitations that followed the devastation of the Great War, inspired by the Russian Revolution , led to counter-revolution and repression throughout Italy. The liberal establishment, fearing a Soviet-style revolution, started to endorse the small National Fascist Party , led by Mussolini. In October 1922, the Blackshirts of
4080-652: A potential food source, radiated in the Late Triassic. Early sauropodomorphs did not have sophisticated mechanisms for processing food in the mouth, and so must have employed other means of breaking down food farther along the digestive tract. The general homogeneity of dinosaurian faunas continued into the Middle and Late Jurassic, where most localities had predators consisting of ceratosaurians , megalosauroids , and allosauroids , and herbivores consisting of stegosaurian ornithischians and large sauropods. Examples of this include
4284-656: A presentation about fossil reptiles to the British Association for the Advancement of Science in 1841, but reports of the time show that Owen did not mention the word "dinosaur", nor recognize dinosaurs as a distinct group of reptiles in his address. He introduced the Dinosauria only in the revised text version of his talk published in April 1842. With the backing of Prince Albert , the husband of Queen Victoria , Owen established
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#17330862510844488-619: A prominent δ C negative shift in higher plants' n-alkanes suggests a massive CO 2 injection in the atmosphere - ocean system at the base of the CPE. The minimum radiometric age of the CPE (≈230.9 Ma) is similar in age to the basalts of the Wrangellia large igneous province (LIP). In the geological record, LIP volcanism is often correlated to episodes of major climate changes and extinctions, which may be caused by pollution of ecosystems with massive release of volcanic gases such as CO 2 and SO 2 . Large release of CO 2 in
4692-675: A pulse of volcanic activity, particularly the Wrangellia LIP. This further supports a volcanic cause of the Carnian pluvial episode. Mercury spikes are also found alongside carbon cycle disruptions in both marine and lake sediments in China. These mercury spikes have no trace of mass-independent fractionation , meaning that their isotope distribution is most consistent with fallout from volcanic eruptions. Conodonts , ammonoids , crinoids , bryozoa and green algae experienced high extinction rates during
4896-538: A result of increased continental weathering amidst the warmth and humidity of the CPE. The first planktonic calcifiers occurred just after the CPE and might have been calcareous dinocysts, i.e., calcareous cysts of dinoflagellates . Coastal ostracod communities in Hungary experienced major changes across the CPE. Through Julian 2, land-based sediments isolated and filled in marine basins, replacing carbonate-specialists such as bairdiids and healdiids with Bektasia ,
5100-599: A review paper by Paul Sereno in 1998, were accompanied by increases in the number of published phylogenetic trees for dinosaurs. Dinosaur fossils are not limited to bones, but also include imprints or mineralized remains of skin coverings, organs, and other tissues. Of these, skin coverings based on keratin proteins are most easily preserved because of their cross-linked , hydrophobic molecular structure. Fossils of keratin-based skin coverings or bony skin coverings are known from most major groups of dinosaurs. Dinosaur fossils with scaly skin impressions have been found since
5304-532: A revival of Italian republicanism. Italy became a republic after the 1946 Italian institutional referendum held on 2 June, a day celebrated since as Festa della Repubblica . This was the first time women voted nationally. Victor Emmanuel III's son, Umberto II , was forced to abdicate. The Republican Constitution was approved in 1948. Under the Treaty of Paris between Italy and the Allied Powers , areas next to
5508-549: A series of studies that likewise argued for active lifestyles in dinosaurs based on anatomical and ecological evidence (see § Physiology ), which were subsequently summarized in his 1986 book The Dinosaur Heresies . New revelations were supported by an increase in dinosaur discoveries. Major new dinosaur discoveries have been made by paleontologists working in previously unexplored regions, including India, South America, Madagascar, Antarctica, and most significantly China. Across theropods, sauropodomorphs, and ornithischians,
5712-702: A series of wars in the peninsula. During the High Renaissance , popes such as Julius II (1503–1513) fought for control of Italy against foreign monarchs; Paul III (1534–1549) preferred to mediate between the European powers to secure peace. In the middle of such conflicts, the Medici popes Leo X (1513–1521) and Clement VII (1523–1534) faced the Protestant Reformation in Germany, England and elsewhere. In 1559, at
5916-615: A significant role in regional and global economic, military, cultural, and diplomatic affairs. Italy is a founding and leading member of the European Union, and is part of numerous international institutions, including NATO , the G7 and G20 , the Latin Union and the Union for the Mediterranean . As a cultural superpower , Italy has long been a renowned global centre of art , music , literature , cuisine , fashion , science and technology , and
6120-603: A sister group to Dinosauria, including a large anterior trochanter, metatarsals II and IV of subequal length, reduced contact between ischium and pubis, the presence of a cnemial crest on the tibia and of an ascending process on the astragalus, and many others. A variety of other skeletal features are shared by dinosaurs. However, because they either are common to other groups of archosaurs or were not present in all early dinosaurs, these features are not considered to be synapomorphies. For example, as diapsids , dinosaurs ancestrally had two pairs of Infratemporal fenestrae (openings in
6324-490: A universally agreed-upon list of their distinguishing features, nearly all dinosaurs discovered so far share certain modifications to the ancestral archosaurian skeleton, or are clearly descendants of older dinosaurs showing these modifications. Although some later groups of dinosaurs featured further modified versions of these traits, they are considered typical for Dinosauria; the earliest dinosaurs had them and passed them on to their descendants. Such modifications, originating in
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6528-463: A variety of waterbirds , diversified rapidly at the beginning of the Paleogene period, entering ecological niches left vacant by the extinction of Mesozoic dinosaur groups such as the arboreal enantiornithines , aquatic hesperornithines , and even the larger terrestrial theropods (in the form of Gastornis , eogruiids , bathornithids , ratites, geranoidids , mihirungs , and " terror birds "). It
6732-430: Is 39.7 meters (130 ft) long. There were larger dinosaurs, but knowledge of them is based entirely on a small number of fragmentary fossils. Most of the largest herbivorous specimens on record were discovered in the 1970s or later, and include the massive Argentinosaurus , which may have weighed 80 000 to 100 000 kilograms (88 to 110 short tons) and reached lengths of 30 to 40 meters (98 to 131 ft); some of
6936-699: Is a synapsid ). None of them had the erect hind limb posture characteristic of true dinosaurs. Dinosaurs were the dominant terrestrial vertebrates of the Mesozoic Era , especially the Jurassic and Cretaceous periods. Other groups of animals were restricted in size and niches; mammals , for example, rarely exceeded the size of a domestic cat and were generally rodent-sized carnivores of small prey. Dinosaurs have always been recognized as an extremely varied group: over 900 non-avian dinosaur genera have been confidently identified (2018) with 1124 species (2016). Estimates put
7140-465: Is an inexact art, and reconstructing the muscles and other organs of the living animal is, at best, a process of educated guesswork. The tallest and heaviest dinosaur known from good skeletons is Giraffatitan brancai (previously classified as a species of Brachiosaurus ). Its remains were discovered in Tanzania between 1907 and 1912. Bones from several similar-sized individuals were incorporated into
7344-755: Is based in part on preservation bias , as large, sturdy bones are more likely to last until they are fossilized. Many dinosaurs were quite small, some measuring about 50 centimeters (20 inches) in length. The first dinosaur fossils were recognized in the early 19th century, with the name "dinosaur" (meaning "terrible lizard") being coined by Sir Richard Owen in 1842 to refer to these "great fossil lizards". Since then, mounted fossil dinosaur skeletons have been major attractions at museums worldwide, and dinosaurs have become an enduring part of popular culture . The large sizes of some dinosaurs, as well as their seemingly monstrous and fantastic nature, have ensured their regular appearance in best-selling books and films, such as
7548-459: Is between 1 and 10 metric tons (1.1 and 11.0 short tons). This contrasts sharply with the average size of Cenozoic mammals, estimated by the National Museum of Natural History as about 2 to 5 kg (4.4 to 11.0 lb). The sauropods were the largest and heaviest dinosaurs. For much of the dinosaur era, the smallest sauropods were larger than anything else in their habitat, and the largest
7752-658: Is correlated with major ammonoid and conodont extinctions. Norwegian shale and Japanese chert from the Ladinian-Carnian boundary show a marked change in the ratio of seawater osmium isotopes. The relative abundance of osmium-187 over osmium-188 declines strongly through most of the Julian before rebounding and stabilizing in the Tuvalian. The decline is attributed to early phases of the Wrangellia large igneous province enriching
7956-617: Is derived from a variety of fossil and non-fossil records, including fossilized bones, feces , trackways , gastroliths , feathers , impressions of skin, internal organs and other soft tissues . Many fields of study contribute to our understanding of dinosaurs, including physics (especially biomechanics ), chemistry , biology , and the Earth sciences (of which paleontology is a sub-discipline). Two topics of particular interest and study have been dinosaur size and behavior. Current evidence suggests that dinosaur average size varied through
8160-478: Is influenced by the seas that surround Italy on every side except the north, which constitute a reservoir of heat and humidity. Within the southern temperate zone, they determine a Mediterranean climate with local differences. Because of the length of the peninsula and the mostly mountainous hinterland, the climate is highly diverse. In most inland northern and central regions, the climate ranges from humid subtropical to humid continental and oceanic . The Po Valley
8364-459: Is marked by disruptions to geochemical cycles, most notably the carbon cycle . Sediments corresponding to the base of the episode show a prominent –2 to –4‰ δ C excursion, indicating the release of a lightweight carbon isotope, carbon-12 , into the atmosphere. This excursion was first mentioned regarding carbonates in Israel, and was later reported in more detail from fragments of carbonized wood in
Carnian pluvial episode - Misplaced Pages Continue
8568-595: Is mostly humid subtropical, with cool winters and hot summers. The coastal areas of Liguria , Tuscany, and most of the south generally fit the Mediterranean climate stereotype, as in the Köppen climate classification . Conditions on the coast are different from those in the interior, particularly during winter when the higher altitudes tend to be cold, wet, and often snowy. The coastal regions have mild winters, and hot and generally dry summers; lowland valleys are hot in summer. Winter temperatures vary from 0 °C (32 °F) in
8772-570: Is now at the American Museum of Natural History in New York City, while Marsh's is at the Peabody Museum of Natural History at Yale University . World War II caused a pause in palaeontological research; after the war, research attention was also diverted increasingly to fossil mammals rather than dinosaurs, which were seen as sluggish and cold-blooded. At the end of the 1960s, however,
8976-644: Is often stated that mammals out-competed the neornithines for dominance of most terrestrial niches but many of these groups co-existed with rich mammalian faunas for most of the Cenozoic Era. Terror birds and bathornithids occupied carnivorous guilds alongside predatory mammals, and ratites are still fairly successful as midsized herbivores; eogruiids similarly lasted from the Eocene to Pliocene , becoming extinct only very recently after over 20 million years of co-existence with many mammal groups. Dinosaurs belong to
9180-482: Is very similar to the minimum age calculated for the CPE (≈230.9 million years ago). Ichnofossil comparisons of various tetrapods from before, during and after the CPE suggest an explosive radiation of dinosaurs due to the Carnian humid phase. However, while avemetatarsalian diversity, diversification rate, and size disparity does increase through the Carnian, it increases faster in the Ladinian and Norian, suggesting that
9384-672: Is water. Including the islands, Italy has a coastline of 7,600 kilometres (4,722 miles) on the Mediterranean Sea , the Ligurian and Tyrrhenian seas, the Ionian Sea , and the Adriatic Sea . Its border with France runs for 488 km (303 mi); Switzerland, 740 km (460 mi); Austria, 430 km (267 mi); and Slovenia, 232 km (144 mi). The sovereign states of San Marino and Vatican City (the smallest country in
9588-528: The Jurassic Park franchise. Persistent public enthusiasm for the animals has resulted in significant funding for dinosaur science, and new discoveries are regularly covered by the media. Under phylogenetic nomenclature , dinosaurs are usually defined as the group consisting of the most recent common ancestor (MRCA) of Triceratops and modern birds (Neornithes), and all its descendants. It has also been suggested that Dinosauria be defined with respect to
9792-645: The Oration on the Dignity of Man , considered the manifesto of the Renaissance. In the arts, the Italian Renaissance exercised a dominant influence on European art for centuries, with artists such as Leonardo da Vinci , Botticelli , Michelangelo , Raphael , Giotto , Donatello , and Titian , and architects such as Filippo Brunelleschi , Andrea Palladio , and Donato Bramante . Italian explorers and navigators from
9996-437: The blue whale and Perucetus colossus as one of the largest animals to have ever existed. The largest carnivorous dinosaur was Spinosaurus , reaching a length of 12.6 to 18 meters (41 to 59 ft) and weighing 7 to 20.9 metric tons (7.7 to 23.0 short tons). Other large carnivorous theropods included Giganotosaurus , Carcharodontosaurus , and Tyrannosaurus . Therizinosaurus and Deinocheirus were among
10200-657: The Achaeans founded various cities. Greek colonisation placed the Italic peoples in contact with democratic forms of government and high artistic and cultural expressions. Ancient Rome, a settlement on the river Tiber in central Italy, founded in 753 BC, was ruled for 244 years by a monarchical system. In 509 BC, the Romans, favouring a government of the Senate and the People ( SPQR ), expelled
10404-598: The Adriatic Sea were annexed by Yugoslavia , resulting in the Istrian-Dalmatian exodus , which involved the emigration of around 300,000 Istrian and Dalmatian Italians . Italy lost all colonial possessions, ending the Italian Empire . Fears of a Communist takeover proved crucial in 1948 , when the Christian Democrats , under Alcide De Gasperi , won a landslide victory. Consequently, in 1949 Italy became
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#173308625108410608-666: The Anisian epoch of the Triassic, approximately 243 million years ago, which is the age of Nyasasaurus from the Manda Formation of Tanzania. However, its known fossils are too fragmentary to identify it as a dinosaur or only a close relative. The referral of the Manda Formation to the Anisian is also uncertain. Regardless, dinosaurs existed alongside non-dinosaurian ornithodirans for
10812-474: The Carnian pluvial episode , was tentatively considered to be a result of oceanic and/or volcanic instability related to the early rifting of Pangea, but at that time direct evidence of this was lacking. Simms and Ruffell published several more papers in the coming years, but their hypothesis was not widely accepted. A strong critique by Visscher et al. (1994) argued that aridity-adapted pollen stayed abundant through
11016-408: The Carnian pluvial event . Dinosaur evolution after the Triassic followed changes in vegetation and the location of continents. In the Late Triassic and Early Jurassic, the continents were connected as the single landmass Pangaea , and there was a worldwide dinosaur fauna mostly composed of coelophysoid carnivores and early sauropodomorph herbivores. Gymnosperm plants (particularly conifers ),
11220-483: The Clean Hands investigation—demanded radical reform. The Christian Democrats, who had ruled for almost 50 years, underwent a crisis and disbanded, splitting into factions. The Communists reorganised as a social-democratic force. During the 1990s and 2000s, centre-right (dominated by media magnate Silvio Berlusconi ) and centre-left coalitions (led by professor Romano Prodi ) alternately governed. In 2011, amidst
11424-788: The Crimean War . Sardinia fought the Austrian Empire in the Second Italian War of Independence of 1859, with the aid of France, resulting in liberating Lombardy . On the basis of the Plombières Agreement , the Sardinia ceded Savoy and Nice to France, an event that caused the Niçard exodus . In 1860–1861, Garibaldi led the drive for unification in Naples and Sicily. Teano was the site of
11628-645: The Crusades , providing support, transport, but mostly taking political and trading opportunities. Italy first felt the economic changes which led to the commercial revolution : Venice was able to sack Byzantine's capital and finance Marco Polo 's voyages to Asia; the first universities were formed in Italian cities, and scholars such as Aquinas obtained international fame; capitalism and banking families emerged in Florence, where Dante and Giotto were active around 1300. In
11832-809: The Empire of Japan , and strongly supported Francisco Franco in the Spanish Civil War . In 1939, Italy annexed Albania . Italy entered World War II on 10 June 1940. At different times, Italians advanced in British Somaliland , Egypt , the Balkans , and eastern fronts. They were, however, defeated on the Eastern Front as well as in the East African and North African campaigns, losing their territories in Africa and
12036-429: The Great Recession , Berlusconi resigned and was replaced by the technocratic cabinet of Mario Monti . In 2014, Matteo Renzi became prime minister and the government started constitutional reform. This was rejected in a 2016 referendum and Paolo Gentiloni became prime minister. During the European migrant crisis of the 2010s, Italy was the entry point and leading destination for most asylum seekers entering
12240-440: The Gregorian calendar ; the Jesuit China mission ; the French Wars of Religion ; end of the Thirty Years' War (1618–1648); and the Great Turkish War . The Italian economy declined in the 1600s and 1700s. During the war of the Spanish succession (1700–1714), Austria acquired most of the Spanish domains in Italy, namely Milan, Naples and Sardinia; the latter was given to the House of Savoy in exchange for Sicily in 1720. Later,
12444-401: The House of Savoy to establish a united kingdom encompassing the entire Italian Peninsula . By the mid-19th century, rising Italian nationalism led to revolution. Following the Congress of Vienna in 1815, the political and social Italian unification movement, or Risorgimento , emerged to unite Italy by consolidating the states and liberating them from foreign control. A radical figure was
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#173308625108412648-436: The Ischigualasto and Santa Maria Formations of Argentina and Brazil, and the Pebbly Arkose Formation of Zimbabwe. The Ischigualasto Formation ( radiometrically dated at 231–230 million years old ) has produced the early saurischian Eoraptor , originally considered a member of the Herrerasauridae but now considered to be an early sauropodomorph, along with the herrerasaurids Herrerasaurus and Sanjuansaurus , and
12852-428: The Italian Peninsula , Sicily and Sardinia (the biggest islands of the Mediterranean), and many smaller islands . Some of Italy's territory extends beyond the Alpine basin, and some islands are located outside the Eurasian continental shelf. The country's area is 301,230 square kilometres (116,306 sq mi), of which 294,020 km (113,522 sq mi) is land and 7,210 km (2,784 sq mi)
13056-525: The Italian Republic , is a country in Southern and Western Europe . It consists of a peninsula that extends into the Mediterranean Sea , with the Alps on its northern land border, as well as nearly 800 islands, notably Sicily and Sardinia . Italy shares its borders with France, Switzerland, Austria, Slovenia, and two enclaves: Vatican City and San Marino . It is the tenth-largest country in Europe by area, covering 301,340 km (116,350 sq mi), and third-most populous member state of
13260-418: The Italic peoples . The main historic peoples of possible non-Indo-European or pre-Indo-European heritage include the Etruscans , the Elymians and Sicani of Sicily, and the prehistoric Sardinians , who gave birth to the Nuragic civilisation . Other ancient populations include the Rhaetian people and Camunni , known for their rock drawings in Valcamonica . A natural mummy, Ötzi , dated 3400–3100 BC,
13464-594: The Jurassic and Cretaceous periods. The fossil record shows that birds are feathered dinosaurs , having evolved from earlier theropods during the Late Jurassic epoch , and are the only dinosaur lineage known to have survived the Cretaceous–Paleogene extinction event approximately 66 mya. Dinosaurs can therefore be divided into avian dinosaurs —birds—and the extinct non-avian dinosaurs , which are all dinosaurs other than birds. Dinosaurs are varied from taxonomic , morphological and ecological standpoints. Birds, at over 11,000 living species , are among
13668-461: The Kvarner Gulf , and the Dalmatian city of Zara . The subsequent Treaty of Rome (1924) led to annexation of Fiume by Italy. Italy did not receive other territories promised by the Treaty of London, so this outcome was denounced as a " mutilated victory ", by Benito Mussolini , which helped lead to the rise of Italian fascism . Historians regard "mutilated victory" as a "political myth", used by fascists to fuel Italian imperialism . Italy gained
13872-429: The Late Triassic epoch are often poorly known and were similar in many ways; these animals have sometimes been misidentified in the literature. Dinosaurs stand with their hind limbs erect in a manner similar to most modern mammals , but distinct from most other reptiles, whose limbs sprawl out to either side. This posture is due to the development of a laterally facing recess in the pelvis (usually an open socket) and
14076-408: The Middle East . In the wake of Julius Caesar 's assassination in 44 BC, Rome grew into a massive empire stretching from Britain to the borders of Persia , engulfing the whole Mediterranean basin, in which Greek, Roman, and other cultures merged into a powerful civilisation. The long reign of the first emperor, Augustus , began an age of peace and prosperity. Roman Italy remained the metropole of
14280-537: The Morrison Formation of North America and Tendaguru Beds of Tanzania. Dinosaurs in China show some differences, with specialized metriacanthosaurid theropods and unusual, long-necked sauropods like Mamenchisaurus . Ankylosaurians and ornithopods were also becoming more common, but primitive sauropodomorphs had become extinct. Conifers and pteridophytes were the most common plants. Sauropods, like earlier sauropodomorphs, were not oral processors, but ornithischians were evolving various means of dealing with food in
14484-479: The National Republican Army . German troops, with RSI collaboration, committed massacres and deported thousands of Jews to death camps. The post-armistice period saw the emergence of the Italian Resistance , who fought a guerrilla war against the Nazi German occupiers and collaborators. This has been described as an Italian civil war due to fighting between partisans and fascist RSI forces. In April 1945, with defeat looming, Mussolini attempted to escape north, but
14688-487: The Natural History Museum, London , to display the national collection of dinosaur fossils and other biological and geological exhibits. In 1858, William Parker Foulke discovered the first known American dinosaur, in marl pits in the small town of Haddonfield, New Jersey . (Although fossils had been found before, their nature had not been correctly discerned.) The creature was named Hadrosaurus foulkii . It
14892-747: The Norse ; and Amerigo Vespucci , for whom the continent of America is named. A defensive alliance known as the Italic League was formed between Venice, Naples, Florence, Milan, and the Papacy. Lorenzo the Magnificent de Medici was the Renaissance's greatest patron, his support allowed the League to abort invasion by the Turks. The alliance, however, collapsed in the 1490s; the invasion of Charles VIII of France initiated
15096-611: The Pietraroja Plattenkalk of Italy. It preserves portions of the intestines, colon, liver, muscles, and windpipe. Concurrently, a line of work led by Mary Higby Schweitzer , Jack Horner , and colleagues reported various occurrences of preserved soft tissues and proteins within dinosaur bone fossils. Various mineralized structures that likely represented red blood cells and collagen fibres had been found by Schweitzer and others in tyrannosaurid bones as early as 1991. However, in 2005, Schweitzer and colleagues reported that
15300-763: The Roman Empire . With the spread of Christianity, Rome became the seat of the Catholic Church and the Papacy . Between late antiquity and the Early Middle Ages , Italy experienced the arrival of Germanic tribes and the fall of the Western Roman Empire . By the 11th century, Italian city-states and maritime republics expanded, bringing renewed prosperity through commerce and laying the groundwork for modern capitalism. The Italian Renaissance flourished during
15504-625: The Schilfsandstein of Germany and various members of the United Kingdom 's Mercia Mudstone Group ) are intervals of river sediments enriched with kaolinitic clay and plant debris, despite having been deposited between more arid strata. Humidity-adapted palynomorphs in New Brunswick , karst topography in the U.K., and a carbon isotope excursion in Israel were all reported for the middle of
15708-676: The United States and displayed in Pittsburgh 's Carnegie Museum of Natural History in 1907. The longest dinosaur known from good fossil material is Patagotitan : the skeleton mount in the American Museum of Natural History in New York is 37 meters (121 ft) long. The Museo Municipal Carmen Funes in Plaza Huincul , Argentina, has an Argentinosaurus reconstructed skeleton mount that
15912-555: The collapse of the Fascist regime on 25 July. Mussolini was deposed and arrested by order of King Victor Emmanuel III. On 8 September, Italy signed the Armistice of Cassibile , ending its war with the Allies. The Germans, with the assistance of Italian fascists, succeeded in taking control of north and central Italy. The country remained a battlefield, with the Allies moving up from the south. In
16116-625: The eponymous geographical region , is located in Southern Europe (and is also considered part of Western Europe ) between latitudes 35° and 47° N , and longitudes 6° and 19° E . To the north, from west to east, Italy borders France, Switzerland, Austria, and Slovenia, and is roughly delimited by the Alpine watershed , enclosing the Po Valley and the Venetian Plain . It consists of the entirety of
16320-427: The largest and smallest dinosaurs to have ever existed. This is because only a tiny percentage of animals were ever fossilized and most of these remain buried in the earth. Few non-avian dinosaur specimens that are recovered are complete skeletons, and impressions of skin and other soft tissues are rare. Rebuilding a complete skeleton by comparing the size and morphology of bones to those of similar, better-known species
16524-606: The political upheavals that characterised the early 19th century. The first adoption of the Italian tricolour by an Italian state, the Cispadane Republic , occurred during Napoleonic Italy , following the French Revolution, which advocated national self-determination . This event is celebrated by Tricolour Day . The birth of the Kingdom of Italy was the result of efforts of Italian nationalists and monarchists loyal to
16728-530: The "lizard-hipped" dinosaurs—birds evolved from earlier dinosaurs with "lizard hips". The following is a simplified classification of dinosaur groups based on their evolutionary relationships, and those of the main dinosaur groups Theropoda, Sauropodomorpha and Ornithischia, compiled by Justin Tweet. Further details and other hypotheses of classification may be found on individual articles. Timeline of major dinosaur groups per Holtz (2007) . Knowledge about dinosaurs
16932-454: The 1400s and 1500s, Italy was the birthplace and heart of the Renaissance . This era marked the transition from the medieval period to the modern age and was fostered by the wealth accumulated by merchant cities and the patronage of dominant families. Italian polities were now regional states effectively ruled by princes, in control of trade and administration, and their courts became centres of
17136-723: The 15th and 16th centuries and spread to the rest of Europe. Italian explorers discovered new routes to the Far East and the New World , contributing significantly to the European Age of Discovery . After centuries of political and territorial divisions, Italy was almost entirely unified in 1861, following wars of independence and the Expedition of the Thousand , establishing the Kingdom of Italy . From
17340-677: The 1970s , however, has indicated that dinosaurs were active animals with elevated metabolisms and numerous adaptations for social interaction. Some were herbivorous , others carnivorous . Evidence suggests that all dinosaurs were egg-laying , and that nest -building was a trait shared by many dinosaurs, both avian and non-avian. While dinosaurs were ancestrally bipedal , many extinct groups included quadrupedal species, and some were able to shift between these stances. Elaborate display structures such as horns or crests are common to all dinosaur groups, and some extinct groups developed skeletal modifications such as bony armor and spines . While
17544-409: The 19th century. Samuel Beckles discovered a sauropod forelimb with preserved skin in 1852 that was incorrectly attributed to a crocodile; it was correctly attributed by Marsh in 1888 and subject to further study by Reginald Hooley in 1917. Among ornithischians, in 1884 Jacob Wortman found skin impressions on the first known specimen of Edmontosaurus annectens , which were largely destroyed during
17748-689: The Adriatic or Tyrrhenian Sea. The longest is the Po , which flows from the Alps on the western border, and crosses the Padan plain to the Adriatic. The Po Valley is the largest plain, with 46,000 km (18,000 sq mi), and contains over 70% of the country's lowlands . The largest lakes are, in descending size: Garda (367.94 km or 142 sq mi), Maggiore (212.51 km or 82 sq mi), and Como (145.9 km or 56 sq mi). The climate
17952-768: The Alps form most of its northern boundary, where Italy's highest point is located on the summit of Mont Blanc ( Monte Bianco ) at 4,810 m (15,780 ft). Other well-known mountains include the Matterhorn ( Monte Cervino ) in the western Alps, and the Dolomites in the eastern Alps. Many parts of Italy are of volcanic origin . Most small islands and archipelagos in the south are volcanic islands . There are active volcanoes: Mount Etna in Sicily (the largest in Europe), Vulcano , Stromboli , and Vesuvius . Most rivers of Italy drain into
18156-500: The Balkans. Italian war crimes included extrajudicial killings and ethnic cleansing by deportation of about 25,000 people—mainly Yugoslavs—to Italian concentration camps and elsewhere. Yugoslav Partisans perpetrated their own crimes against the ethnic Italian population during and after the war, including the foibe massacres . An Allied invasion of Sicily began in July 1943, leading to
18360-471: The CPE was not a major influence on the rise of dinosaurs. The CPE had a profound effect on the diversity and morphological disparity of herbivorous tetrapods. This is exemplified in rhynchosaurs , a group of reptiles with strong shearing and grinding jaws. Rhynchosaur lineages which were common in the Middle Triassic went extinct, leaving only the specialized hyperodapedontines as representatives of
18564-409: The CPE, as well as its relevance for understanding modern climate change. It also attempted to standardize the nomenclature of the CPE; rejecting descriptors such as "event" (typically applied to geological processes under a million years in duration) or "middle Carnian" (a nebulous term with no equivalent geological substage). The Carnian pluvial episode introduced markedly more humid conditions across
18768-496: The CPE, which show an approximately 1.5 ‰ negative shift in the stable isotope δ O , suggesting global warming of 3–4 °C during the CPE and/or a change in seawater salinity . This warming was probably related to extensive volcanic activity at the time, evidenced by carbon isotope trends across the CPE. This volcanic activity was in turn probably related to the formation of the Wrangellia Large igneous province around
18972-707: The CPE. Though tiny amber traces can be found in rocks as old as the Carboniferous , the earliest widespread amber deposits date back to the CPE. Carnian amber droplets from Italian paleosols are the oldest amber deposits known to preserve arthropods and microorganisms . Amber would not reappear in the fossil record until the Late Jurassic , though it would take until the Early Cretaceous for amber to occur in concentrations equivalent to or exceeding Carnian amber. Radiolarians increased in their diversity, likely as
19176-484: The CPE. Other organisms radiated and diversified during the interval, such as dinosaurs , calcareous nannofossils , corals and conifers . The CPE has been interpreted as a key geobiological event facilitating the radiation of dinosaurs. The oldest dinosaur-bearing fossil assemblage, the Ischigualasto Formation of Argentina , has been radiometrically dated back to 230.3 to 231.4 million years ago. This age
19380-434: The Carnian prior to 1989. The Julian-Tuvalian boundary experienced high extinction rates among many marine invertebrates, while an extinction among land vertebrates was suggested to occur in the late Carnian. In 1989, a paper by Michael J. Simms and Alastair H. Ruffell combined these disparate observations into a new hypothesis, pointing to an episode of increased rainfall synchronous with significant ecological turnover in
19584-630: The Carnian stage long before a global climate perturbation was proposed. Schlager & Schöllnberger (1974) drew attention to a dark siliciclastic layer which abruptly interrupted a long period of carbonate deposition in the Northern Limestone Alps . They termed this stratigraphic "wende" (turning point) the Reingrabener Wende, and it has also been called the Reingraben event or Raibl event . Several Carnian terrestrial formations (namely
19788-601: The Church and new political dynasties. In 1453, despite activity by Pope Nicholas V to support the Byzantines, the city of Constantinople fell to the Ottomans . This led to the migration of Greek scholars and texts to Italy, fuelling the rediscovery of Greek humanism . Humanist rulers such as Federico da Montefeltro and Pope Pius II worked to establish ideal cities , founding Urbino and Pienza . Pico della Mirandola wrote
19992-464: The Dolomites. It has been confirmed in various carbon-based sediments throughout Europe and Asia. More precise stratigraphic evaluation of European outcrops has resolved this excursion into three or possibly four major pulses, spanning the late Julian and early Tuvalian. Each pulse can be equated with an interval of abnormal sedimentation on land and sea. The third excursion, at the Julian-Tuvalian boundary,
20196-529: The EU. Between 2013 and 2018, it took in over 700,000 migrants, mainly from sub-Saharan Africa, which put a strain on the public purse and led to a surge in support for far-right or euro-sceptic parties. After the 2018 general election , Giuseppe Conte became prime minister of a populist coalition . With more than 155,000 victims, Italy was one of the countries with the most deaths in the COVID-19 pandemic and one of
20400-501: The English paleontologist Sir Richard Owen coined the term "dinosaur", using it to refer to the "distinct tribe or sub-order of Saurian Reptiles" that were then being recognized in England and around the world. The term is derived from Ancient Greek δεινός (deinos) 'terrible, potent or fearfully great' and σαῦρος (sauros) 'lizard or reptile'. Though
20604-483: The European Union , with a population of nearly 60 million. Italy's capital and largest city is Rome ; other major urban areas include Milan , Naples , Turin , Palermo , Bologna , Florence , Genoa , and Venice . The history of Italy goes back to numerous Italic peoples , notably including the ancient Romans , who conquered the Mediterranean world during the Roman Republic and ruled it for centuries during
20808-743: The First World War in 1915 with the aim of completing national unity, so it is also considered the Fourth Italian War of Independence, from a historiographical perspective, as the conclusion of the unification of Italy . Italy, nominally allied with German and the Austro-Hungarian empires in the Triple Alliance , in 1915 joined the Allies , entering World War I with a promise of substantial territorial gains that included west Inner Carniola ,
21012-577: The Greek ornitheios (ὀρνίθειος) meaning "of a bird" and ischion (ἰσχίον) meaning "hip joint"—had a pelvis that superficially resembled a bird's pelvis: the pubic bone was oriented caudally (rear-pointing). Unlike birds, the ornithischian pubis also usually had an additional forward-pointing process. Ornithischia includes a variety of species that were primarily herbivores. Despite the terms "bird hip" (Ornithischia) and "lizard hip" (Saurischia), birds are not part of Ornithischia. Birds instead belong to Saurischia,
21216-599: The Italian Front, secured dissolution of the Austro-Hungarian Empire, and was instrumental in ending the war less than two weeks later. During the war, more than 650,000 Italian soldiers and as many civilians died, and the kingdom was on the brink of bankruptcy. The Treaty of Saint-Germain-en-Laye (1919) and Treaty of Rapallo (1920) allowed for annexation of Trentino Alto-Adige , the Julian March , Istria ,
21420-501: The Late Cretaceous, was the evolution of flowering plants . At the same time, several groups of dinosaurian herbivores evolved more sophisticated ways to orally process food. Ceratopsians developed a method of slicing with teeth stacked on each other in batteries, and iguanodontians refined a method of grinding with dental batteries , taken to its extreme in hadrosaurids. Some sauropods also evolved tooth batteries, best exemplified by
21624-781: The MRCA of Megalosaurus and Iguanodon , because these were two of the three genera cited by Richard Owen when he recognized the Dinosauria. Both definitions cover the same known genera: Dinosauria = Ornithischia + Saurischia . This includes major groups such as ankylosaurians (armored herbivorous quadrupeds), stegosaurians (plated herbivorous quadrupeds), ceratopsians (bipedal or quadrupedal herbivores with neck frills ), pachycephalosaurians (bipedal herbivores with thick skulls), ornithopods (bipedal or quadrupedal herbivores including " duck-bills "), theropods (mostly bipedal carnivores and birds), and sauropodomorphs (mostly large herbivorous quadrupeds with long necks and tails). Birds are
21828-401: The Middle to Late Triassic epochs, roughly 20 million years after the devastating Permian–Triassic extinction event wiped out an estimated 96% of all marine species and 70% of terrestrial vertebrate species approximately 252 million years ago. The oldest dinosaur fossils known from substantial remains date to the Carnian epoch of the Triassic period and have been found primarily in
22032-571: The National Fascist Party organised a mass demonstration and the " March on Rome " coup . King Victor Emmanuel III appointed Mussolini as prime minister, transferring power to the fascists without armed conflict. Mussolini banned political parties and curtailed personal liberties, establishing a dictatorship. These actions attracted international attention and inspired similar dictatorships in Nazi Germany and Francoist Spain . Fascism
22236-686: The Papacy, with city-states siding with the former ( Ghibellines ) or with the latter ( Guelphs ) for momentary advantage. The Germanic emperor and Roman pontiff became the universal powers of medieval Europe. However, conflict over the Investiture Controversy and between Guelphs and Ghibellines ended the imperial-feudal system in the north, where cities gained independence. In 1176, the Lombard League of city-states, defeated Holy Roman Emperor Frederick Barbarossa , ensuring their independence. City-states—e.g. Milan , Florence, Venice —played
22440-567: The Roman Republic's expansion, the name was used by Greeks for the land between the strait of Messina and the line connecting the gulfs of Salerno and Taranto , corresponding to Calabria. The Greeks came to apply "Italia" to a larger region. In addition to the " Greek Italy " in the south, historians have suggested the existence of an "Etruscan Italy", which consisted of areas of central Italy. The borders of Roman Italy , Italia , are better established. Cato's Origines describes Italy as
22644-519: The Triassic, Early Jurassic, Late Jurassic and Cretaceous. Predatory theropod dinosaurs, which occupied most terrestrial carnivore niches during the Mesozoic, most often fall into the 100-to-1,000 kg (220-to-2,200 lb) category when sorted by estimated weight into categories based on order of magnitude , whereas recent predatory carnivoran mammals peak in the 10-to-100 kg (22-to-220 lb) category. The mode of Mesozoic dinosaur body masses
22848-525: The Triassic. Spores of ferns and freshwater algae are frequently abundant in palynological samples. The Carnian saw the reestablishment of large inland lakes and peat swamps , ending the Early-Middle Triassic " coal gap " caused by the Permian-Triassic mass extinction. Increased plant growth and coal burial probably helped to draw down CO2, returning the atmosphere to a more normal state after
23052-439: The Triassic. Conversely, more versatile and generalist herbivores such as aetosaurs and sauropodomorph dinosaurs would diversify after the CPE. Conifers , ferns , and the now-extinct bennettitaleans all diversified greatly during and after the CPE, establishing themselves as mainstays of Mesozoic flora. Most areas show a higher proportion of hygrophytic (moisture-loving) plants during the episode compared to earlier parts of
23256-583: The United States, known as dinosaur mania. Dinosaur mania was exemplified by the fierce rivalry between Edward Drinker Cope and Othniel Charles Marsh , both of whom raced to be the first to find new dinosaurs in what came to be known as the Bone Wars . This fight between the two scientists lasted for over 30 years, ending in 1897 when Cope died after spending his entire fortune on the dinosaur hunt. Many valuable dinosaur specimens were damaged or destroyed due to
23460-560: The arts and sciences. These princedoms were led by political dynasties and merchant families, such as the Medici of Florence. After the end of the Western Schism , newly elected Pope Martin V returned to the Papal States and restored Italy as the sole centre of Western Christianity. The Medici Bank was made the credit institution of the Papacy, and significant ties were established between
23664-448: The atmosphere-ocean system by Wrangellia can explain the increased supply of siliciclastic material into basins, as observed during the CPE. The increase of CO 2 in the atmosphere could have resulted in global warming and consequent acceleration of the hydrological cycle, thus strongly enhancing the continental weathering . Moreover, if rapid enough, a sudden rise of pCO 2 levels could have resulted in acidification of seawater with
23868-666: The best-known genera are remarkable for their large size, many Mesozoic dinosaurs were human-sized or smaller, and modern birds are generally small in size. Dinosaurs today inhabit every continent, and fossils show that they had achieved global distribution by the Early Jurassic epoch at latest. Modern birds inhabit most available habitats, from terrestrial to marine, and there is evidence that some non-avian dinosaurs (such as Microraptor ) could fly or at least glide, and others, such as spinosaurids , had semiaquatic habits. While recent discoveries have made it more difficult to present
24072-455: The body . Other prehistoric animals, including pterosaurs , mosasaurs , ichthyosaurs , plesiosaurs , and Dimetrodon , while often popularly conceived of as dinosaurs, are not taxonomically classified as dinosaurs. Pterosaurs are distantly related to dinosaurs, being members of the clade Ornithodira . The other groups mentioned are, like dinosaurs and pterosaurs, members of Sauropsida (the reptile and bird clade), except Dimetrodon (which
24276-429: The bone as the lower extremity of the femur of a large animal, and recognized that it was too large to belong to any known species. He therefore concluded it to be the femur of a huge human, perhaps a Titan or another type of giant featured in legends. Edward Lhuyd , a friend of Sir Isaac Newton , published Lithophylacii Britannici ichnographia (1699), the first scientific treatment of what would now be recognized as
24480-462: The calf ( Latin : vitulus ; Umbrian : vitlo ; Oscan : Víteliú ). Several ancient authors said it was named after a local ruler Italus . The ancient Greek term for Italy initially referred only to the south of the Bruttium peninsula and parts of Catanzaro and Vibo Valentia . The larger concept of Oenotria and "Italy" became synonymous, and the name applied to most of Lucania as well. Before
24684-530: The coming years supported a strong link between the Carnian climate disturbances and the Wrangellia large igneous province, but many questions remain unanswered. A geological workshop focusing on the CPE met in 2018 at the Hanse-Wissenschaftskolleg (HWK) Institute for Advanced Study in Delmenhorst, Germany . The workshop was intended to spur further research on the mechanisms, impact and stratigraphy of
24888-467: The common theropods, and ankylosaurids and early ceratopsians like Psittacosaurus became important herbivores. Meanwhile, Australia was home to a fauna of basal ankylosaurians, hypsilophodonts , and iguanodontians. The stegosaurians appear to have gone extinct at some point in the late Early Cretaceous or early Late Cretaceous . A major change in the Early Cretaceous, which would be amplified in
25092-427: The consequent rise of the carbonate compensation depth (CCD) and a crisis of carbonate precipitation (e.g. demise of carbonate platforms in the western Tethys ). On top of all that, the global warming brought on by the flood basalt event was likely exacerbated by the release of methane clathrates. According to an alternative hypothesis, the Carnian pluvial episode was a regional climatic perturbation mostly visible in
25296-713: The dinosaur subgroup Maniraptora , which are coelurosaurs , which are theropods, which are saurischians. Research by Matthew G. Baron, David B. Norman , and Paul M. Barrett in 2017 suggested a radical revision of dinosaurian systematics. Phylogenetic analysis by Baron et al. recovered the Ornithischia as being closer to the Theropoda than the Sauropodomorpha, as opposed to the traditional union of theropods with sauropodomorphs. This would cause sauropods and kin to fall outside traditional dinosaurs, so they re-defined Dinosauria as
25500-427: The dinosaurs' modern-day surviving avian lineage (birds) are generally small due to the constraints of flight, many prehistoric dinosaurs (non-avian and avian) were large-bodied—the largest sauropod dinosaurs are estimated to have reached lengths of 39.7 meters (130 feet) and heights of 18 m (59 ft) and were the largest land animals of all time. The misconception that non-avian dinosaurs were uniformly gigantic
25704-626: The dinosaurs. Most of these other animals became extinct in the Triassic, in one of two events. First, at about 215 million years ago, a variety of basal archosauromorphs, including the protorosaurs , became extinct. This was followed by the Triassic–Jurassic extinction event (about 201 million years ago), that saw the end of most of the other groups of early archosaurs, like aetosaurs, ornithosuchids, phytosaurs , and rauisuchians. Rhynchosaurs and dicynodonts survived (at least in some areas) at least as late as early –mid Norian and late Norian or earliest Rhaetian stages , respectively, and
25908-406: The early Carnian, were replaced by low-relief carbonate platforms featuring low-angle slopes (i.e., ramps). This turnover is related to a major change in the biological community responsible for calcium carbonate precipitation (i.e. carbonate factory). The highly-productive, mainly bacterial-dominated biological community (M-factory) whose action led to the carbonate production on high-relief platforms
26112-469: The early saurischian Alwalkeria , are known from the Upper Maleri and Lower Maleri Formations of India. The Carnian-aged Chañares Formation of Argentina preserves primitive, dinosaur-like ornithodirans such as Lagosuchus and Lagerpeton in Argentina , making it another important site for understanding dinosaur evolution. These ornithodirans support the model of early dinosaurs as small, bipedal predators. Dinosaurs may have appeared as early as
26316-444: The emergence of Christianity as a world religion, are among the many legacies of Roman dominance. After the fall of the Western Roman Empire , Italy fell under the Odoacer's kingdom , and was seized by the Ostrogoths . Invasions resulted in a chaotic succession of kingdoms and the supposed " Dark Ages ". The invasion of another Germanic tribe in the 6th century, the Lombards, reduced Byzantine presence and ended political unity of
26520-446: The empire, homeland of the Romans and territory of the capital. As Roman provinces were being established throughout the Mediterranean, Italy maintained a special status which made it domina provinciarum ('ruler of the provinces'), and—especially in relation to the first centuries of imperial stability — rectrix mundi ('governor of the world') and omnium terrarum parens ('parent of all lands'). The Roman Empire
26724-424: The end of the Italian wars between France and the Habsburgs, about half of Italy (the southern Kingdoms of Naples , Sicily , Sardinia , and the Duchy of Milan ) was under Spanish rule, while the other half remained independent (many states continued to be formally part of the Holy Roman Empire). The Papacy launched the Counter-Reformation , whose key events include: the Council of Trent (1545–1563); adoption of
26928-403: The end of the Cretaceous, caused the extinction of all dinosaur groups except for the neornithine birds. Some other diapsid groups, including crocodilians , dyrosaurs , sebecosuchians , turtles, lizards , snakes , sphenodontians , and choristoderans , also survived the event. The surviving lineages of neornithine birds, including the ancestors of modern ratites , ducks and chickens , and
27132-483: The entire Carnian of Germany, suggesting that the Schilfsandstein was simply indicative of an invading river system rather than widespread climate change. Their critique also coined the term " Carnian pluvial event ", which would eventually become among the most widespread names for the climatic disturbance. The obscurity of Simms and Ruffell's hypothesis began to dissipate in the late 2000s, as further support accumulated from studies on Carnian sites in Italy . Interest in
27336-509: The entire peninsula south of the Alps. In 264 BC, Roman Italy extended from the Arno and Rubicon rivers of the centre-north to the entire south. The northern area, Cisalpine Gaul , considered geographically part of Italy, was occupied by Rome in the 220s BC, but remained politically separated. It was legally merged into the administrative unit of Italy in 42 BC. Sardinia, Corsica , Sicily, and Malta were added to Italy by Diocletian in 292 AD, which made late-ancient Italy coterminous with
27540-401: The exact date of their extinction is uncertain. These losses left behind a land fauna of crocodylomorphs , dinosaurs, mammals, pterosaurians, and turtles . The first few lines of early dinosaurs diversified through the Carnian and Norian stages of the Triassic, possibly by occupying the niches of the groups that became extinct. Also notably, there was a heightened rate of extinction during
27744-433: The field of dinosaur research experienced a surge in activity that remains ongoing. Several seminal studies led to this activity. First, John Ostrom discovered the bird-like dromaeosaurid theropod Deinonychus and described it in 1969. Its anatomy indicated that it was an active predator that was likely warm-blooded, in marked contrast to the then-prevailing image of dinosaurs. Concurrently, Robert T. Bakker published
27948-411: The former Austrian Littoral , and Dalmatia , as well as parts of the Ottoman Empire . The country's contribution to the Allied victory earned it a place as one of the " Big Four " powers. Reorganisation of the army and conscription led to Italian victories. In October 1918, the Italians launched a massive offensive, culminating in victory at the Battle of Vittorio Veneto . This marked the end of war on
28152-464: The globe, interrupting the otherwise arid climate of the Late Triassic period. This humidity was related to increased rainfall during the CPE evidence of which includes: This usually wet climate of the CPE was periodically interrupted by drier climates typical of the rest of the Late Triassic period. Global warming was also prevalent during the Carnian pluvial event. This is evidenced by oxygen isotope analyses performed on conodont apatite from
28356-545: The group. Immediately after the CPE, hyperodapedontines were widespread and abundant in the late Carnian world, suggesting that they benefited from the climate fluctuations or floral turnover. Hyperodapedontine abundance was not sustained for long, and they too would die out in the early Norian. By cutting rhynchosaurs off from a greater variety of niches, the CPE would have reduced their versatility and increased their vulnerability to extinction. Similar trends are observed in dicynodonts , though they would survive until much later in
28560-486: The hip structure of their ancestors, with a pubis bone directed cranially , or forward. This basic form was modified by rotating the pubis backward to varying degrees in several groups ( Herrerasaurus , therizinosauroids, dromaeosaurids, and birds ). Saurischia includes the theropods (exclusively bipedal and with a wide variety of diets) and sauropodomorphs (long-necked herbivores which include advanced, quadrupedal groups). By contrast, ornithischians—"bird-hipped", from
28764-533: The hypothesis was greatly enhanced by a 2008 meeting and workshop on Triassic climate at the Museum of Nature South Tyrol in Bolzano, Italy . However, even as the global nature of the CPE became increasingly accepted, its ultimate cause was still hotly debated going into the 2010s. Even its nomenclature was not agreed upon, with various authors applying names such as the middle Carnian wet intermezzo , Carnian humid episode , Carnian pluvial phase , and Carnian crisis . Carbon and Osmium isotope records published over
28968-435: The kingdom's capital was moved from Turin to Florence. In 1866, Victor Emmanuel II, allied with Prussia during the Austro-Prussian War , waged the Third Italian War of Independence , which resulted in Italy annexing Venetia . Finally, in 1870, as France abandoned Rome during the Franco-Prussian War , the Italians captured the Papal States , unification was completed, and the capital moved to Rome. Sardinia's constitution
29172-432: The last common ancestor of Triceratops horridus , Passer domesticus and Diplodocus carnegii , and all of its descendants, to ensure that sauropods and kin remain included as dinosaurs. They also resurrected the clade Ornithoscelida to refer to the group containing Ornithischia and Theropoda. Using one of the above definitions, dinosaurs can be generally described as archosaurs with hind limbs held erect beneath
29376-448: The late 17th century in England. Part of a bone, now known to have been the femur of a Megalosaurus , was recovered from a limestone quarry at Cornwell near Chipping Norton , Oxfordshire, in 1676. The fragment was sent to Robert Plot , Professor of Chemistry at the University of Oxford and first curator of the Ashmolean Museum , who published a description in his The Natural History of Oxford-shire (1677). He correctly identified
29580-421: The late 19th to the early 20th century, Italy rapidly industrialised, mainly in the north, and acquired a colonial empire , while the south remained largely impoverished, fueling a large immigrant diaspora to the Americas. From 1915 to 1918, Italy took part in World War I with the Entente against the Central Powers . In 1922, the Italian fascist dictatorship was established. During World War II , Italy
29784-410: The latter linked to the rapid diversification of scleractinian corals through the establishment of symbiotic zooxanthellae within them. The CPE also saw the extinction of many aquatic invertebrate species, especially among the ammonoids , bryozoa , and crinoids . Evidence for the CPE is observed in Carnian strata worldwide and in sediments of both terrestrial and marine environments. On land,
29988-478: The longest were the 33.5-meter (110 ft) long Diplodocus hallorum (formerly Seismosaurus ), the 33-to-34-meter (108 to 112 ft) long Supersaurus , and 37-meter (121 ft) long Patagotitan ; and the tallest, the 18-meter (59 ft) tall Sauroposeidon , which could have reached a sixth-floor window. There were a few dinosaurs that was considered either the heaviest and longest. The most famous one include Amphicoelias fragillimus , known only from
30192-480: The marine realm. Dinosaur Dinosaurs are a diverse group of reptiles of the clade Dinosauria . They first appeared during the Triassic period , between 243 and 233.23 million years ago (mya), although the exact origin and timing of the evolution of dinosaurs is a subject of active research. They became the dominant terrestrial vertebrates after the Triassic–Jurassic extinction event 201.3 mya and their dominance continued throughout
30396-407: The maritime republics, eager to find an alternative route to the Indies to bypass the Ottomans, offered their services to monarchs of Atlantic countries and played a key role in ushering the Age of Discovery and colonization of the Americas. The most notable were: Christopher Columbus , who opened the Americas for conquest by Europeans; John Cabot , the first European to explore North America since
30600-534: The mid-Carnian. The paper was inspired by a conversation between Simms and Ruffell, on 10 November 1987 at Birmingham University, that connected Ruffell's research on lithological changes in the Mercia Mudstone Group to Simms's research on crinoid extinction. A key aspect of their hypothesis was that the evidence used to demonstrate the climate change was entirely independent of the evidence for biotic change; fossils were not used in any way to infer climate change. Their hypothesized climatic disturbance, which they named
30804-413: The modern Italian geographical region . The Latin Italicus was used to describe "a man of Italy" as opposed to a provincial , or one from the Roman province . The adjective italianus , from which Italian was derived, is from medieval Latin and was used alternatively with Italicus during the early modern period . After the fall of the Western Roman Empire , the Ostrogothic Kingdom of Italy
31008-420: The monarchy and established an oligarchic republic. The Italian Peninsula, named Italia , was consolidated into a unified entity during Roman expansion, the conquest of new territories often at the expense of the other Italic tribes , Etruscans , Celts , and Greeks . A permanent association, with most of the local tribes and cities, was formed, and Rome began the conquest of Western Europe, North Africa, and
31212-447: The most affected economically . In February 2021, after a government crisis , Conte resigned. Mario Draghi , former president of the European Central Bank , formed a national unity government supported by most main parties, pledging to implement an economic stimulus to face the crisis caused by the pandemic. In 2022, Giorgia Meloni was sworn in as Italy's first female prime minister. Italy, whose territory largely coincides with
31416-479: The most complete theropod specimens, while North American localities have produced the most complete sauropodomorph specimens. Prior to the dinosaur renaissance, dinosaurs were mostly classified using the traditional rank-based system of Linnaean taxonomy . The renaissance was also accompanied by the increasingly widespread application of cladistics , a more objective method of classification based on ancestry and shared traits, which has proved tremendously useful in
31620-495: The most diverse groups of vertebrates. Using fossil evidence, paleontologists have identified over 900 distinct genera and more than 1,000 different species of non-avian dinosaurs. Dinosaurs are represented on every continent by both extant species (birds) and fossil remains. Through the first half of the 20th century, before birds were recognized as dinosaurs, most of the scientific community believed dinosaurs to have been sluggish and cold-blooded . Most research conducted since
31824-509: The most important advantage was dietary. Large animals are more efficient at digestion than small animals, because food spends more time in their digestive systems. This also permits them to subsist on food with lower nutritive value than smaller animals. Sauropod remains are mostly found in rock formations interpreted as dry or seasonally dry, and the ability to eat large quantities of low-nutrient browse would have been advantageous in such environments. Scientists will probably never be certain of
32028-478: The most recent common ancestor of a certain taxonomic group, are called the synapomorphies of such a group. A detailed assessment of archosaur interrelations by Sterling Nesbitt confirmed or found the following twelve unambiguous synapomorphies, some previously known: Nesbitt found a number of further potential synapomorphies and discounted a number of synapomorphies previously suggested. Some of these are also present in silesaurids , which Nesbitt recovered as
32232-400: The mouth, including potential cheek -like organs to keep food in the mouth, and jaw motions to grind food. Another notable evolutionary event of the Jurassic was the appearance of true birds, descended from maniraptoran coelurosaurians. By the Early Cretaceous and the ongoing breakup of Pangaea, dinosaurs were becoming strongly differentiated by landmass. The earliest part of this time saw
32436-413: The north, the Germans set up the Italian Social Republic (RSI), a Nazi puppet state and collaborationist regime with Mussolini installed as leader after he was rescued by German paratroopers. What remained of the Italian troops was organised into the Italian Co-belligerent Army , which fought alongside the Allies, while other Italian forces, loyal to Mussolini, opted to fight alongside the Germans in
32640-715: The now-splitting supercontinent Gondwana , abelisaurids were the common theropods, and titanosaurian sauropods the common herbivores. Finally, in Europe, dromaeosaurids, rhabdodontid iguanodontians, nodosaurid ankylosaurians, and titanosaurian sauropods were prevalent. Flowering plants were greatly radiating, with the first grasses appearing by the end of the Cretaceous. Grinding hadrosaurids and shearing ceratopsians became very diverse across North America and Asia. Theropods were also radiating as herbivores or omnivores , with therizinosaurians and ornithomimosaurians becoming common. The Cretaceous–Paleogene extinction event, which occurred approximately 66 million years ago at
32844-756: The number of named genera began to increase exponentially in the 1990s. As of 2008, over 30 new species of dinosaurs were named each year. At least sauropodomorphs experienced a further increase in the number of named species in the 2010s, with an average of 9.3 new species having been named each year between 2009 and 2020. As a consequence, more sauropodomorphs were named between 1990 and 2020 than in all previous years combined. These new localities also led to improvements in overall specimen quality, with new species being increasingly named not on scrappy fossils but on more complete skeletons, sometimes from multiple individuals. Better specimens also led to new species being invalidated less frequently. Asian localities have produced
33048-468: The ocean and continent, and thus generating a monsoon . Summer monsoonal winds were thus intercepted by the Cimmerian mountain range and generated strong rain, thus explaining the switch to humid climate recognized in western Tethys sediments. Highstands in sea level and crises amongst marine organisms during the CPE are closely correlated to maxima in the 1.2-Myr obliquity modulation cycle, suggesting that this Milankovitch cycle regulated biotic turnovers in
33252-483: The ocean floor to form sediments. Climate change during the Carnian pluvial event is reflected in chemical changes in Carnian strata across the CPE which suggest that global warming was prevalent at the time. This climate change was probably linked to the eruption of extensive flood basalts as the Wrangellia Terrane was accreted onto the northwestern end of the North American Plate . Environmental disturbance and high extinction rates were observed for sediments of
33456-405: The ocean with osmium-188. Osmium-188 is preferentially sourced directly from the mantle, while osmium-187 is a radiogenic isotope supplied from eroded land. In the Alps , moderate to high concentrations of mercury occur alongside carbon cycle disruptions, just prior to the sediment disruption which marks the CPE. These mercury spikes occur in well-oxygenated mudstones, meaning that they are not
33660-438: The pair's rough methods: for example, their diggers often used dynamite to unearth bones. Modern paleontologists would find such methods crude and unacceptable, since blasting easily destroys fossil and stratigraphic evidence. Despite their unrefined methods, the contributions of Cope and Marsh to paleontology were vast: Marsh unearthed 86 new species of dinosaur and Cope discovered 56, a total of 142 new species. Cope's collection
33864-399: The patriotic journalist Giuseppe Mazzini , founder of the political movement Young Italy in the 1830s, who favoured a unitary republic and advocated a broad nationalist movement. 1847 saw the first public performance of " Il Canto degli Italiani ", which became the national anthem in 1946. The most famous member of Young Italy was the revolutionary and general Giuseppe Garibaldi who led
34068-406: The peninsula. The north formed the Lombard kingdom, central-south was also controlled by the Lombards, and other parts remained Byzantine. The Lombard kingdom was absorbed into Francia by Charlemagne in the late 8th century and became the Kingdom of Italy. The Franks helped form the Papal States . Until the 13th century, politics was dominated by relations between the Holy Roman Emperors and
34272-408: The presence of proteins such as collagen, elastin , and laminin . Both specimens yielded collagen protein sequences that were viable for molecular phylogenetic analyses , which grouped them with birds as would be expected. The extraction of fragmentary DNA has also been reported for both of these fossils, along with a specimen of Hypacrosaurus . In 2015, Sergio Bertazzo and colleagues reported
34476-458: The preservation of collagen fibres and red blood cells in eight Cretaceous dinosaur specimens that did not show any signs of exceptional preservation, indicating that soft tissue may be preserved more commonly than previously thought. Suggestions that these structures represent bacterial biofilms have been rejected, but cross-contamination remains a possibility that is difficult to detect. Dinosaurs diverged from their archosaur ancestors during
34680-464: The prevailing arid climate across much of the supercontinent Pangea shifted briefly to a hotter and more humid climate, with a significant increase in rainfall and runoff. In the oceans there was reduced deposition of carbonate minerals . This may reflect the extinction of many carbonate-forming organisms , but may also be due to a rise in the carbonate compensation depth , below which most carbonate shells dissolve and leave few carbonate particles on
34884-422: The rebbachisaurid Nigersaurus . There were three general dinosaur faunas in the Late Cretaceous. In the northern continents of North America and Asia, the major theropods were tyrannosaurids and various types of smaller maniraptoran theropods, with a predominantly ornithischian herbivore assemblage of hadrosaurids, ceratopsians, ankylosaurids, and pachycephalosaurians. In the southern continents that had made up
35088-469: The republican drive for unification in southern Italy. However, the Italian monarchy of the House of Savoy, in the Kingdom of Sardinia , whose government was led by Camillo Benso, Count of Cavour , also had ambitions of establishing a united Italian state. In the context of the 1848 liberal revolutions that swept Europe, an unsuccessful First Italian War of Independence was declared against Austria . In 1855, Sardinia became an ally of Britain and France in
35292-1307: The rocks that produced the Jehol (Early Cretaceous) and Yanliao (Mid-to-Late Jurassic) biotas of northeastern China, from which hundreds of dinosaur specimens bearing impressions of feather-like structures (both closely related to birds and otherwise, see § Origin of birds ) have been described by Xing Xu and colleagues. In living reptiles and mammals, pigment-storing cellular structures known as melanosomes are partially responsible for producing colouration. Both chemical traces of melanin and characteristically shaped melanosomes have been reported from feathers and scales of Jehol and Yanliao dinosaurs, including both theropods and ornithischians. This has enabled multiple full-body reconstructions of dinosaur colouration , such as for Sinosauropteryx and Psittacosaurus by Jakob Vinther and colleagues, and similar techniques have also been extended to dinosaur fossils from other localities. (However, some researchers have also suggested that fossilized melanosomes represent bacterial remains. ) Stomach contents in some Jehol and Yanliao dinosaurs closely related to birds have also provided indirect indications of diet and digestive system anatomy (e.g., crops ). More concrete evidence of internal anatomy has been reported in Scipionyx from
35496-422: The same time, which created vast quantities of igneous (volcanic) rocks that were accreted onto the northwest end of the North American Plate (now the Wrangell Mountains , Alaska , and an estimated 6km thick layer underlying most of Vancouver Island) There is some evidence for seabed euxinia (no oxygen and high toxic sulfide concentrations) during the CPE. Limestones are enriched in manganese ions near
35700-450: The sauropodomorphs Bagualosaurus , Buriolestes , Guaibasaurus , Macrocollum , Nhandumirim , Pampadromaeus , Saturnalia , and Unaysaurus . The Pebbly Arkose Formation, which is of uncertain age but was likely comparable to the other two, has produced the sauropodomorph Mbiresaurus , along with an unnamed herrerasaurid. Less well-preserved remains of the sauropodomorphs Jaklapallisaurus and Nambalia , along with
35904-416: The sauropodomorphs Chromogisaurus , Eodromaeus , and Panphagia . Eoraptor 's likely resemblance to the common ancestor of all dinosaurs suggests that the first dinosaurs would have been small, bipedal predators . The Santa Maria Formation (radiometrically dated to be older, at 233.23 million years old ) has produced the herrerasaurids Gnathovorax and Staurikosaurus , along with
36108-426: The skeleton now mounted and on display at the Museum für Naturkunde in Berlin ; this mount is 12 meters (39 ft) tall and 21.8 to 22.5 meters (72 to 74 ft) long, and would have belonged to an animal that weighed between 30 000 and 60 000 kilograms ( 70 000 and 130 000 lb). The longest complete dinosaur is the 27 meters (89 ft) long Diplodocus , which was discovered in Wyoming in
36312-406: The skull behind the eyes), and as members of the diapsid group Archosauria, had additional openings in the snout and lower jaw. Additionally, several characteristics once thought to be synapomorphies are now known to have appeared before dinosaurs, or were absent in the earliest dinosaurs and independently evolved by different dinosaur groups. These include an elongated scapula , or shoulder blade;
36516-434: The sole surviving dinosaurs. In traditional taxonomy , birds were considered a separate class that had evolved from dinosaurs, a distinct superorder . However, most contemporary paleontologists reject the traditional style of classification based on anatomical similarity, in favor of phylogenetic taxonomy based on deduced ancestry, in which each group is defined as all descendants of a given founding genus. Birds belong to
36720-474: The source of multiple inventions and discoveries . It has the highest number of World Heritage Sites ( 60 ) and is the fifth-most visited country in the world. Hypotheses for the etymology of Italia are numerous. One theory suggests it originated from an Ancient Greek term for the land of the Italói , a tribe that resided in the region now known as Calabria . Originally thought to be named Vituli , some scholars suggest their totemic animal to be
36924-408: The south, Sicily had become an Arab Islamic emirate in the 9th century, thriving until the Italo-Normans conquered it in the late 11th century, together with most of the Lombard and Byzantine principalities of southern Italy. The region was subsequently divided between the Kingdom of Sicily and Kingdom of Naples . The Black Death of 1348 killed perhaps a third of Italy's population. During
37128-515: The specimen's excavation. Owen and Hooley subsequently described skin impressions of Hypsilophodon and Iguanodon in 1885 and 1917. Since then, scale impressions have been most frequently found among hadrosaurids, where the impressions are known from nearly the entire body across multiple specimens. Starting from the 1990s, major discoveries of exceptionally preserved fossils in deposits known as conservation Lagerstätten contributed to research on dinosaur soft tissues. Chiefly among these were
37332-423: The spread of ankylosaurians, iguanodontians , and brachiosaurids through Europe, North America, and northern Africa . These were later supplemented or replaced in Africa by large spinosaurid and carcharodontosaurid theropods, and rebbachisaurid and titanosaurian sauropods, also found in South America . In Asia , maniraptoran coelurosaurians like dromaeosaurids, troodontids , and oviraptorosaurians became
37536-468: The study of dinosaur systematics and evolution. Cladistic analysis, among other techniques, helps to compensate for an often incomplete and fragmentary fossil record. Reference books summarizing the state of dinosaur research, such as David B. Weishampel and colleagues' The Dinosauria , made knowledge more accessible and spurred further interest in dinosaur research. The release of the first and second editions of The Dinosauria in 1990 and 2004, and of
37740-438: The tallest of the theropods. The largest ornithischian dinosaur was probably the hadrosaurid Shantungosaurus giganteus which measured 16.6 meters (54 ft). The largest individuals may have weighed as much as 16 metric tons (18 short tons). Italy – in Europe (light green & dark grey) – in the European Union (light green) – [ Legend ] Italy , officially
37944-459: The taxonomic name has often been interpreted as a reference to dinosaurs' teeth, claws, and other fearsome characteristics, Owen intended it also to evoke their size and majesty. Owen recognized that the remains that had been found so far, Iguanodon , Megalosaurus and Hylaeosaurus , shared distinctive features, and so decided to present them as a distinct taxonomic group. As clarified by British geologist and historian Hugh Torrens, Owen had given
38148-519: The top of the Zhuganpo Formation of south China. Manganese ions are concentrated and soluble in deep euxinic waters, but precipitate in carbonates at the base of the oxygenated zone. Increasing manganese concentrations indicate a narrowing of the oxygenated zone and a corresponding expansion of euxinic water. At the onset of the CPE a sharp change in carbonate platform geometries is recorded in western Tethys . High-relief, mainly isolated, small carbonate platforms surrounded by steep slopes, typical of
38352-849: The total number of dinosaur genera preserved in the fossil record at 1850, nearly 75% still undiscovered, and the number that ever existed (in or out of the fossil record) at 3,400. A 2016 estimate put the number of dinosaur species living in the Mesozoic at 1,543–2,468, compared to the number of modern-day birds (avian dinosaurs) at 10,806 species. Extinct dinosaurs, as well as modern birds, include genera that are herbivorous and others carnivorous, including seed-eaters, fish-eaters, insectivores, and omnivores. While dinosaurs were ancestrally bipedal (as are all modern birds), some evolved into quadrupeds, and others, such as Anchisaurus and Iguanodon , could walk as easily on two or four legs. Cranial modifications like horns and crests are common dinosaurian traits, and some extinct species had bony armor. Although
38556-405: The traditional liberal and conservative establishment. In the last two decades of the 19th century, Italy developed into a colonial power by subjugating Eritrea , Somalia , Tripolitania , and Cyrenaica in Africa. In 1913, male universal suffrage was adopted. The pre- World War I period was dominated by Giovanni Giolitti , prime minister five times between 1892 and 1921. Italy entered into
38760-399: The western Tethys and related to the uplift of a new mountain range , the Cimmerian orogen , which resulted from the closing of a Tethyan northern branch, east of the present European continent. The new mountain range was forming on the southern side of Laurasia , and acted then as the Himalayas and Asia do today for the Indian Ocean , maintaining a strong pressure gradient between
38964-419: The world and headquarters of the worldwide Catholic Church under the governance of the Holy See ) are enclaves within Italy, while Campione d'Italia is an Italian exclave in Switzerland. The border with San Marino is 39 km (24 mi) long, that with Vatican City, 3.2 km (2.0 mi). Over 35% of Italian territory is mountainous. The Apennine Mountains form the peninsula's backbone, and
39168-400: The world's fifth-largest industrial nation after it gained entry into the G7 in the 1970s. However, national debt skyrocketed past 100% of GDP. Between 1992 and 1993, Italy faced terror attacks perpetrated by the Sicilian Mafia as a consequence of new anti-mafia measures by the government. Voters—disenchanted with political paralysis, massive public debt and extensive corruption uncovered by
39372-422: Was among the largest in history, wielding great economical, cultural, political, and military power. At its greatest extent, it had an area of 5 million square kilometres (1.9 million square miles). The Roman legacy has deeply influenced Western civilisation shaping the modern world. The widespread use of Romance languages derived from Latin, numerical system , modern Western alphabet and calendar, and
39576-401: Was an extremely important find: Hadrosaurus was one of the first nearly complete dinosaur skeletons found ( the first was in 1834, in Maidstone, England ), and it was clearly a bipedal creature. This was a revolutionary discovery as, until that point, most scientists had believed dinosaurs walked on four feet, like other lizards. Foulke's discoveries sparked a wave of interests in dinosaurs in
39780-437: Was an order of magnitude more massive than anything else that has since walked the Earth. Giant prehistoric mammals such as Paraceratherium (the largest land mammal ever) were dwarfed by the giant sauropods, and only modern whales approach or surpass them in size. There are several proposed advantages for the large size of sauropods, including protection from predation, reduction of energy use, and longevity, but it may be that
39984-458: Was based upon Italian nationalism and imperialism, seeking to expand Italian possessions via irredentist claims based on the legacy of the Roman and Venetian empires. For this reason the fascists engaged in interventionist foreign policy . In 1935, Mussolini invaded Ethiopia and founded Italian East Africa , resulting in international isolation and leading to Italy's withdrawal from the League of Nations . Italy then allied with Nazi Germany and
40188-417: Was captured and summarily executed by partisans. Hostilities ended on 29 April 1945, when the German forces in Italy surrendered . Nearly half a million Italians died in the conflict, society was divided, and the economy all but destroyed—per capita income in 1944 was at its lowest point since 1900. The aftermath left Italy angry with the monarchy for its endorsement of the Fascist regime, contributing to
40392-560: Was conducive to academic and artistic advancement. The best-known maritime republics were Venice, Genoa , Pisa , and Amalfi . Each had dominion over overseas lands, islands, lands on the Adriatic, Aegean, and Black seas, and commercial colonies in the Near East and North Africa. Venice and Genoa were Europe's gateways to the East, and producers of fine glass, while Florence was a centre of silk, wool, banking, and jewellery. The wealth generated meant large public and private artistic projects could be commissioned. The republics participated in
40596-530: Was created. After the Lombard invasions, Italia was retained as the name for their kingdom, and its successor kingdom within the Holy Roman Empire . Lower Paleolithic artefacts, dating back 850,000 years, have been recovered from Monte Poggiolo . Excavations throughout Italy revealed a Neanderthal presence in the Middle Palaeolithic period 200,000 years ago, while modern humans appeared about 40,000 years ago at Riparo Mochi . The ancient peoples of pre-Roman Italy were Indo-European , specifically
40800-499: Was discovered in the Similaun glacier in 1991. The first colonisers were the Phoenicians , who established emporiums on the coasts of Sicily and Sardinia. Some became small urban centers and developed parallel to Greek colonies . During the 8th and 7th centuries, Greek colonies were established at Pithecusae , eventually extending along the south of the Italian Peninsula and the coast of Sicily, an area later known as Magna Graecia . Ionians , Doric colonists, Syracusans , and
41004-496: Was extended to all of Italy in 1861, and provided basic freedoms for the new state; but electoral laws excluded the non-propertied classes. The new kingdom was governed by a parliamentary constitutional monarchy dominated by liberals. As northern Italy quickly industrialised, southern and northern rural areas remained underdeveloped and overpopulated, forcing millions to migrate and fuelling a large and influential diaspora . The Italian Socialist Party increased in strength, challenging
41208-448: Was first part of the Axis until its surrender to the Allied powers (1940–1943), then a co-belligerent of the Allies during the Italian resistance and the liberation of Italy (1943–1945). Following the war, the monarchy was replaced by a republic and the country enjoyed a strong recovery . A developed country , Italy has the ninth-largest nominal GDP in the world, the second-largest manufacturing industry in Europe, and plays
41412-450: Was purportedly discovered in 1822 by Mary Ann Mantell , the wife of English geologist Gideon Mantell , though this is disputed and some historians say Gideon had acquired remains years earlier. Gideon Mantell recognized similarities between his fossils and the bones of modern iguanas and published his findings in 1825. The study of these "great fossil lizards" soon became of great interest to European and American scientists, and in 1842
41616-454: Was substituted by a less productive mollusc-metazoan-dominated community (C-T factories). In the South China block, the demise of carbonate platforms is coupled with the deposition of sediments typical of anoxic environments (black shales ). Thanks to low oxygen levels, animal remains were often well-preserved in sedimentary deposits called Lagerstätten . These Lagerstätten are rich in crinoids and reptiles, such as ichthyosaurs . The CPE
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