A biplane is a fixed-wing aircraft with two main wings stacked one above the other. The first powered, controlled aeroplane to fly, the Wright Flyer , used a biplane wing arrangement, as did many aircraft in the early years of aviation . While a biplane wing structure has a structural advantage over a monoplane , it produces more drag than a monoplane wing. Improved structural techniques, better materials and higher speeds made the biplane configuration obsolete for most purposes by the late 1930s.
144-427: The Bristol Berkeley was built to a British government specification for a single-engine day or night bomber. Three of these two-seat biplanes were built, but no contract for further production was awarded. In August 1923, British aircraft manufacturers were invited to submit designs to Air Ministry Specification 26/23 , which called for a single Rolls-Royce Condor -engined two-seat day or night bomber. The Berkeley
288-525: A last universal common ancestor that lived about 3.5 billion years ago . Geologists have developed a geologic time scale that divides the history of the Earth into major divisions, starting with four eons ( Hadean , Archean , Proterozoic , and Phanerozoic ), the first three of which are collectively known as the Precambrian , which lasted approximately 4 billion years. Each eon can be divided into eras, with
432-440: A lipid bilayer , including cholesterols that sit between phospholipids to maintain their fluidity at various temperatures. Cell membranes are semipermeable , allowing small molecules such as oxygen, carbon dioxide, and water to pass through while restricting the movement of larger molecules and charged particles such as ions . Cell membranes also contain membrane proteins , including integral membrane proteins that go across
576-443: A nucleus , and prokaryotic cells, which do not. Prokaryotes are single-celled organisms such as bacteria , whereas eukaryotes can be single-celled or multicellular. In multicellular organisms , every cell in the organism's body is derived ultimately from a single cell in a fertilized egg . Every cell is enclosed within a cell membrane that separates its cytoplasm from the extracellular space . A cell membrane consists of
720-504: A number of shapes , ranging from spheres to rods and spirals . Bacteria were among the first life forms to appear on Earth, and are present in most of its habitats . Bacteria inhabit soil, water, acidic hot springs , radioactive waste , and the deep biosphere of the Earth's crust . Bacteria also live in symbiotic and parasitic relationships with plants and animals. Most bacteria have not been characterised, and only about 27 percent of
864-646: A W shape cabane, however as it does not connect the wings to each other, it does not add to the number of bays. Large transport and bombing biplanes often needed still more bays to provide sufficient strength. These are often referred to as multi-bay biplanes . A small number of biplanes, such as the Zeppelin-Lindau D.I have no interplane struts and are referred to as being strutless . Because most biplanes do not have cantilever structures, they require rigging wires to maintain their rigidity. Early aircraft used simple wire (either braided or plain), however during
1008-412: A biplane aircraft, two wings are placed one above the other. Each provides part of the lift, although they are not able to produce twice as much lift as a single wing of similar size and shape because the upper and the lower are working on nearly the same portion of the atmosphere and thus interfere with each other's behaviour. In a biplane configuration with no stagger from the upper wing to the lower wing,
1152-471: A cell's size, shape, membrane potential , metabolic activity , and responsiveness to signals, which are largely due to highly controlled modifications in gene expression and epigenetics . With a few exceptions, cellular differentiation almost never involves a change in the DNA sequence itself. Thus, different cells can have very different physical characteristics despite having the same genome . Morphogenesis, or
1296-408: A chemical (e.g., nitrous acid , benzopyrene ) or radiation (e.g., x-ray , gamma ray , ultraviolet radiation , particles emitted by unstable isotopes). Mutations can lead to phenotypic effects such as loss-of-function, gain-of-function , and conditional mutations. Some mutations are beneficial, as they are a source of genetic variation for evolution. Others are harmful if they were to result in
1440-608: A documented jet-kill, as one Lockheed F-94 Starfire was lost while slowing down to 161 km/h (100 mph) – below its stall speed – during an intercept in order to engage the low flying Po-2. Later biplane trainers included the de Havilland Tiger Moth in the Royal Air Force (RAF), Royal Canadian Air Force (RCAF) and others and the Stampe SV.4 , which saw service postwar in the French and Belgian Air Forces. The Stearman PT-13
1584-509: A faster and more comfortable successor to the Dragon. As the available engine power and speed increased, the drag penalty of external bracing increasingly limited aircraft performance. To fly faster, it would be necessary to reduce external bracing to create an aerodynamically clean design; however, early cantilever designs were either too weak or too heavy. The 1917 Junkers J.I sesquiplane utilized corrugated aluminum for all flying surfaces, with
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#17328690850951728-1227: A few archaea have very different shapes, such as the flat and square cells of Haloquadratum walsbyi . Despite this morphological similarity to bacteria, archaea possess genes and several metabolic pathways that are more closely related to those of eukaryotes, notably for the enzymes involved in transcription and translation . Other aspects of archaeal biochemistry are unique, such as their reliance on ether lipids in their cell membranes , including archaeols . Archaea use more energy sources than eukaryotes: these range from organic compounds , such as sugars, to ammonia , metal ions or even hydrogen gas . Salt-tolerant archaea (the Haloarchaea ) use sunlight as an energy source, and other species of archaea fix carbon , but unlike plants and cyanobacteria , no known species of archaea does both. Archaea reproduce asexually by binary fission , fragmentation , or budding ; unlike bacteria, no known species of Archaea form endospores . The first observed archaea were extremophiles , living in extreme environments, such as hot springs and salt lakes with no other organisms. Improved molecular detection tools led to
1872-403: A final electron acceptor, which is usually the oxidized form of NADP , which is reduced to NADPH, a process that takes place in a protein complex called photosystem I (PSI). The transport of electrons is coupled to the movement of protons (or hydrogen) from the stroma to the thylakoid membrane, which forms a pH gradient across the membrane as hydrogen becomes more concentrated in the lumen than in
2016-422: A given wing area. However, interference between the airflow over each wing increases drag substantially, and biplanes generally need extensive bracing, which causes additional drag. Biplanes are distinguished from tandem wing arrangements, where the wings are placed forward and aft, instead of above and below. The term is also occasionally used in biology , to describe the wings of some flying animals . In
2160-759: A loss of function of genes needed for survival. Gene expression is the molecular process by which a genotype encoded in DNA gives rise to an observable phenotype in the proteins of an organism's body. This process is summarized by the central dogma of molecular biology , which was formulated by Francis Crick in 1958. According to the Central Dogma, genetic information flows from DNA to RNA to protein. There are two gene expression processes: transcription (DNA to RNA) and translation (RNA to protein). The regulation of gene expression by environmental factors and during different stages of development can occur at each step of
2304-507: A minimum of struts; however, it was relatively easy to damage the thin metal skin and required careful handling by ground crews. The 1918 Zeppelin-Lindau D.I fighter was an all-metal stressed-skin monocoque fully cantilevered biplane, but its arrival had come too late to see combat use in the conflict. By the 1930s, biplanes had reached their performance limits, and monoplanes become increasingly predominant, particularly in continental Europe where monoplanes had been increasingly common from
2448-417: A more successful evolutionary theory based on natural selection ; similar reasoning and evidence led Alfred Russel Wallace to independently reach the same conclusions. The basis for modern genetics began with the work of Gregor Mendel in 1865. This outlined the principles of biological inheritance. However, the significance of his work was not realized until the early 20th century when evolution became
2592-457: A new cycle. In contrast to mitosis, meiosis results in four haploid daughter cells by undergoing one round of DNA replication followed by two divisions. Homologous chromosomes are separated in the first division ( meiosis I ), and sister chromatids are separated in the second division ( meiosis II ). Both of these cell division cycles are used in the process of sexual reproduction at some point in their life cycle. Both are believed to be present in
2736-409: A process known as allopatric speciation . A phylogeny is an evolutionary history of a specific group of organisms or their genes. It can be represented using a phylogenetic tree , a diagram showing lines of descent among organisms or their genes. Each line drawn on the time axis of a tree represents a lineage of descendants of a particular species or population. When a lineage divides into two, it
2880-437: A rectangular cross section. The pilot sat forward of the leading edge of the wing in an open cockpit and the gunner/observer in a cockpit well aft, fitted with a ring-mounted .303 in (7.7 mm) Lewis Gun . He could also access a bomb aimer's position, when he lay prone on the aircraft floor. The horizontal tail was positioned at the top of the fuselage and braced below, carrying elevators whose balances protruded beyond
3024-408: A region of deoxyribonucleic acid (DNA) that carries genetic information that controls form or function of an organism. DNA is composed of two polynucleotide chains that coil around each other to form a double helix . It is found as linear chromosomes in eukaryotes , and circular chromosomes in prokaryotes . The set of chromosomes in a cell is collectively known as its genome . In eukaryotes, DNA
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#17328690850953168-541: A role in the synthesis and packaging of proteins, respectively. Biomolecules such as proteins can be engulfed by lysosomes , another specialized organelle. Plant cells have additional organelles that distinguish them from animal cells such as a cell wall that provides support for the plant cell, chloroplasts that harvest sunlight energy to produce sugar, and vacuoles that provide storage and structural support as well as being involved in reproduction and breakdown of plant seeds. Eukaryotic cells also have cytoskeleton that
3312-436: A separate clade as some protists may be more closely related to plants, fungi, or animals than they are to other protists. Like groupings such as algae , invertebrates , or protozoans , the protist grouping is not a formal taxonomic group but is used for convenience. Most protists are unicellular; these are called microbial eukaryotes. Plants are mainly multicellular organisms , predominantly photosynthetic eukaryotes of
3456-423: A single carbon atom can form four single covalent bonds such as in methane , two double covalent bonds such as in carbon dioxide (CO 2 ), or a triple covalent bond such as in carbon monoxide (CO). Moreover, carbon can form very long chains of interconnecting carbon–carbon bonds such as octane or ring-like structures such as glucose . The simplest form of an organic molecule is the hydrocarbon , which
3600-463: A small degree, but more often was used to improve access to the cockpit. Many biplanes have staggered wings. Common examples include the de Havilland Tiger Moth , Bücker Bü 131 Jungmann and Travel Air 2000 . Alternatively, the lower wing can instead be moved ahead of the upper wing, giving negative stagger, and similar benefits. This is usually done in a given design for structural reasons, or to improve visibility. Examples of negative stagger include
3744-473: A somewhat unusual sesquiplane arrangement, possessing a more substantial lower wing with two spars that eliminated the flutter problems encountered by single-spar sesquiplanes. The stacking of wing planes was suggested by Sir George Cayley in 1843. Hiram Maxim adopted the idea for his steam-powered test rig, which lifted off but was held down by safety rails, in 1894. Otto Lilienthal designed and flew two different biplane hang gliders in 1895, though he
3888-446: A specific enzyme. Enzymes are crucial to metabolism because they allow organisms to drive desirable reactions that require energy that will not occur by themselves, by coupling them to spontaneous reactions that release energy. Enzymes act as catalysts —they allow a reaction to proceed more rapidly without being consumed by it—by reducing the amount of activation energy needed to convert reactants into products . Enzymes also allow
4032-399: A unified theory as the modern synthesis reconciled Darwinian evolution with classical genetics . In the 1940s and early 1950s, a series of experiments by Alfred Hershey and Martha Chase pointed to DNA as the component of chromosomes that held the trait-carrying units that had become known as genes . A focus on new kinds of model organisms such as viruses and bacteria, along with
4176-421: A varied mix of traits, and reproduction is able to increase any population, Darwin argued that in the natural world, it was nature that played the role of humans in selecting for specific traits. Darwin inferred that individuals who possessed heritable traits better adapted to their environments are more likely to survive and produce more offspring than other individuals. He further inferred that this would lead to
4320-492: A water molecule again. In pure water , the number of hydrogen ions balances (or equals) the number of hydroxyl ions, resulting in a pH that is neutral. Organic compounds are molecules that contain carbon bonded to another element such as hydrogen. With the exception of water, nearly all the molecules that make up each organism contain carbon. Carbon can form covalent bonds with up to four other atoms, enabling it to form diverse, large, and complex molecules. For example,
4464-536: Is evolution , which explains the unity and diversity of life. Energy processing is also important to life as it allows organisms to move , grow, and reproduce . Finally, all organisms are able to regulate their own internal environments . Biologists are able to study life at multiple levels of organization , from the molecular biology of a cell to the anatomy and physiology of plants and animals, and evolution of populations. Hence, there are multiple subdisciplines within biology , each defined by
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4608-444: Is a central feature of sexual reproduction in eukaryotes, and the most fundamental function of meiosis appears to be conservation of the integrity of the genome that is passed on to progeny by parents. Two aspects of sexual reproduction , meiotic recombination and outcrossing , are likely maintained respectively by the adaptive advantages of recombinational repair of genomic DNA damage and genetic complementation which masks
4752-641: Is a large family of organic compounds that are composed of hydrogen atoms bonded to a chain of carbon atoms. A hydrocarbon backbone can be substituted by other elements such as oxygen (O), hydrogen (H), phosphorus (P), and sulfur (S), which can change the chemical behavior of that compound. Groups of atoms that contain these elements (O-, H-, P-, and S-) and are bonded to a central carbon atom or skeleton are called functional groups . There are six prominent functional groups that can be found in organisms: amino group , carboxyl group , carbonyl group , hydroxyl group , phosphate group , and sulfhydryl group . In 1953,
4896-480: Is a small polar molecule with a bent shape formed by the polar covalent bonds of two hydrogen (H) atoms to one oxygen (O) atom (H 2 O). Because the O–H bonds are polar, the oxygen atom has a slight negative charge and the two hydrogen atoms have a slight positive charge. This polar property of water allows it to attract other water molecules via hydrogen bonds, which makes water cohesive . Surface tension results from
5040-918: Is a term of convenience as not all algae are closely related. Algae comprise several distinct clades such as glaucophytes , which are microscopic freshwater algae that may have resembled in form to the early unicellular ancestor of Plantae. Unlike glaucophytes, the other algal clades such as red and green algae are multicellular. Green algae comprise three major clades: chlorophytes , coleochaetophytes , and stoneworts . Fungi are eukaryotes that digest foods outside their bodies, secreting digestive enzymes that break down large food molecules before absorbing them through their cell membranes. Many fungi are also saprobes , feeding on dead organic matter, making them important decomposers in ecological systems. Animals are multicellular eukaryotes. With few exceptions, animals consume organic material , breathe oxygen , are able to move , can reproduce sexually , and grow from
5184-547: Is better known for his monoplanes. By 1896 a group of young men in the United States, led by Octave Chanute , were flying hang gliders including biplanes and concluded that the externally braced biplane offered better prospects for powered flight than the monoplane. In 1903, the Wright Flyer biplane became the first successful powered aeroplane. Throughout the pioneer years, both biplanes and monoplanes were common, but by
5328-403: Is catalyzed by lactate dehydrogenase in a reversible reaction. Lactate can also be used as an indirect precursor for liver glycogen. During recovery, when oxygen becomes available, NAD attaches to hydrogen from lactate to form ATP. In yeast, the waste products are ethanol and carbon dioxide. This type of fermentation is known as alcoholic or ethanol fermentation . The ATP generated in this process
5472-441: Is made by substrate-level phosphorylation , which does not require oxygen. Photosynthesis is a process used by plants and other organisms to convert light energy into chemical energy that can later be released to fuel the organism's metabolic activities via cellular respiration. This chemical energy is stored in carbohydrate molecules, such as sugars, which are synthesized from carbon dioxide and water. In most cases, oxygen
5616-535: Is made up of microtubules , intermediate filaments , and microfilaments , all of which provide support for the cell and are involved in the movement of the cell and its organelles. In terms of their structural composition, the microtubules are made up of tubulin (e.g., α-tubulin and β-tubulin ) whereas intermediate filaments are made up of fibrous proteins. Microfilaments are made up of actin molecules that interact with other strands of proteins. All cells require energy to sustain cellular processes. Metabolism
5760-526: Is mainly in the cell nucleus . In prokaryotes, the DNA is held within the nucleoid . The genetic information is held within genes, and the complete assemblage in an organism is called its genotype . DNA replication is a semiconservative process whereby each strand serves as a template for a new strand of DNA. Mutations are heritable changes in DNA. They can arise spontaneously as a result of replication errors that were not corrected by proofreading or can be induced by an environmental mutagen such as
5904-399: Is one of the key ways a cell releases chemical energy to fuel cellular activity. The overall reaction occurs in a series of biochemical steps, some of which are redox reactions. Although cellular respiration is technically a combustion reaction , it clearly does not resemble one when it occurs in a cell because of the slow, controlled release of energy from the series of reactions. Sugar in
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6048-407: Is pulled to the separate poles of the bacterium as it increases the size to prepare for splitting. Growth of a new cell wall begins to separate the bacterium (triggered by FtsZ polymerization and "Z-ring" formation). The new cell wall ( septum ) fully develops, resulting in the complete split of the bacterium. The new daughter cells have tightly coiled DNA rods, ribosomes , and plasmids . Meiosis
6192-399: Is released as a waste product. Most plants, algae , and cyanobacteria perform photosynthesis, which is largely responsible for producing and maintaining the oxygen content of the Earth's atmosphere, and supplies most of the energy necessary for life on Earth. Photosynthesis has four stages: Light absorption , electron transport, ATP synthesis, and carbon fixation . Light absorption is
6336-412: Is represented as a fork or split on the phylogenetic tree. Phylogenetic trees are the basis for comparing and grouping different species. Different species that share a feature inherited from a common ancestor are described as having homologous features (or synapomorphy ). Phylogeny provides the basis of biological classification. This classification system is rank-based, with the highest rank being
6480-468: Is the process by which genes and traits are passed on from parents to offspring. It has several principles. The first is that genetic characteristics, alleles , are discrete and have alternate forms (e.g., purple vs. white or tall vs. dwarf), each inherited from one of two parents. Based on the law of dominance and uniformity , which states that some alleles are dominant while others are recessive ; an organism with at least one dominant allele will display
6624-485: Is the set of chemical reactions in an organism. The three main purposes of metabolism are: the conversion of food to energy to run cellular processes; the conversion of food/fuel to monomer building blocks; and the elimination of metabolic wastes . These enzyme-catalyzed reactions allow organisms to grow and reproduce, maintain their structures, and respond to their environments. Metabolic reactions may be categorized as catabolic —the breaking down of compounds (for example,
6768-410: Is the study of chemical processes within and relating to living organisms . Molecular biology is the branch of biology that seeks to understand the molecular basis of biological activity in and between cells, including molecular synthesis, modification, mechanisms, and interactions. Life arose from the Earth's first ocean, which formed some 3.8 billion years ago. Since then, water continues to be
6912-493: Is to store, transmit, and express hereditary information. Cell theory states that cells are the fundamental units of life, that all living things are composed of one or more cells, and that all cells arise from preexisting cells through cell division . Most cells are very small, with diameters ranging from 1 to 100 micrometers and are therefore only visible under a light or electron microscope . There are generally two types of cells: eukaryotic cells, which contain
7056-524: The Bristol M.1 , that caused even those with relatively high performance attributes to be overlooked in favour of 'orthodox' biplanes, and there was an allegedly widespread belief held at that time that monoplane aircraft were inherently unsafe during combat. Between the years of 1914 and 1925, a clear majority of new aircraft introduced were biplanes; however, during the latter years of the First World War,
7200-629: The Cambrian explosion . During the Permian period, synapsids , including the ancestors of mammals , dominated the land, but most of this group became extinct in the Permian–Triassic extinction event 252 million years ago. During the recovery from this catastrophe, archosaurs became the most abundant land vertebrates; one archosaur group, the dinosaurs, dominated the Jurassic and Cretaceous periods. After
7344-495: The Cretaceous–Paleogene extinction event 66 million years ago killed off the non-avian dinosaurs, mammals increased rapidly in size and diversity . Such mass extinctions may have accelerated evolution by providing opportunities for new groups of organisms to diversify. Bacteria are a type of cell that constitute a large domain of prokaryotic microorganisms . Typically a few micrometers in length, bacteria have
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#17328690850957488-638: The First World War -era Fokker D.VII fighter and the Second World War de Havilland Tiger Moth basic trainer. The larger two-seat Curtiss JN-4 Jenny is a two bay biplane , the extra bay being necessary as overlong bays are prone to flexing and can fail. The SPAD S.XIII fighter, while appearing to be a two bay biplane, has only one bay, but has the midpoints of the rigging braced with additional struts; however, these are not structurally contiguous from top to bottom wing. The Sopwith 1½ Strutter has
7632-684: The Grumman Ag Cat are available in upgraded versions with turboprop engines. The two most produced biplane designs were the 1913 British Avro 504 of which 11,303 were built, and the 1928 Soviet Polikarpov Po-2 of which over 20,000 were built, with the Po-2 being the direct replacement for the Soviet copy of the Avro 504. Both were widely used as trainers. The Antonov An-2 was very successful too, with more than 18,000 built. Although most ultralights are monoplanes,
7776-595: The Lite Flyer Biplane, the Sherwood Ranger , and the Murphy Renegade . The feathered dinosaur Microraptor gui glided, and perhaps even flew, on four wings, which may have been configured in a staggered sesquiplane arrangement. This was made possible by the presence of flight feathers on both forelimbs and hindlimbs, with the feathers on the forelimbs opening to a greater span. It has been suggested that
7920-535: The Miller–Urey experiment showed that organic compounds could be synthesized abiotically within a closed system mimicking the conditions of early Earth , thus suggesting that complex organic molecules could have arisen spontaneously in early Earth (see abiogenesis ). Macromolecules are large molecules made up of smaller subunits or monomers . Monomers include sugars, amino acids, and nucleotides. Carbohydrates include monomers and polymers of sugars. Lipids are
8064-539: The Nieuport-Delage NiD 42 / 52 / 62 series, Fokker C.Vd & e, and Potez 25 , all serving across a large number of air forces. In the general aviation sector, aircraft such as the Waco Custom Cabin series proved to be relatively popular. The Saro Windhover was a sesquiplane with the upper wing smaller than the lower, which was a much rarer configuration than the reverse. The Pfalz D.III also featured
8208-492: The Precambrian about 1.5 billion years ago and can be classified into eight major clades : alveolates , excavates , stramenopiles , plants, rhizarians , amoebozoans , fungi , and animals. Five of these clades are collectively known as protists , which are mostly microscopic eukaryotic organisms that are not plants, fungi, or animals. While it is likely that protists share a common ancestor (the last eukaryotic common ancestor ), protists by themselves do not constitute
8352-494: The Sopwith Dolphin , Breguet 14 and Beechcraft Staggerwing . However, positive (forward) stagger is much more common. The space enclosed by a set of interplane struts is called a bay (much as the architectural form is used), hence a biplane or triplane with one set of such struts connecting the wings on each side of the aircraft is a single-bay biplane . This provided sufficient strength for smaller aircraft such as
8496-727: The activator is the transcription factor that stimulates transcription when it binds to the sequence near or at the promoter. Negative regulation occurs when another transcription factor called a repressor binds to a DNA sequence called an operator , which is part of an operon, to prevent transcription. Repressors can be inhibited by compounds called inducers (e.g., allolactose ), thereby allowing transcription to occur. Specific genes that can be activated by inducers are called inducible genes , in contrast to constitutive genes that are almost constantly active. In contrast to both, structural genes encode proteins that are not involved in gene regulation. In addition to regulatory events involving
8640-585: The bacterial phyla have species that can be grown in the laboratory. Archaea constitute the other domain of prokaryotic cells and were initially classified as bacteria, receiving the name archaebacteria (in the Archaebacteria kingdom ), a term that has fallen out of use. Archaeal cells have unique properties separating them from the other two domains , Bacteria and Eukaryota . Archaea are further divided into multiple recognized phyla . Archaea and bacteria are generally similar in size and shape, although
8784-404: The cell . In 1838, Schleiden and Schwann began promoting the now universal ideas that (1) the basic unit of organisms is the cell and (2) that individual cells have all the characteristics of life, although they opposed the idea that (3) all cells come from the division of other cells, continuing to support spontaneous generation . However, Robert Remak and Rudolf Virchow were able to reify
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#17328690850958928-403: The cell membrane of another cell or located deep inside a cell. There are generally four types of chemical signals: autocrine , paracrine , juxtacrine , and hormones . In autocrine signaling, the ligand affects the same cell that releases it. Tumor cells, for example, can reproduce uncontrollably because they release signals that initiate their own self-division. In paracrine signaling,
9072-416: The circulatory systems of animals or vascular systems of plants to reach their target cells. Once a ligand binds with a receptor, it can influence the behavior of another cell, depending on the type of receptor. For instance, neurotransmitters that bind with an inotropic receptor can alter the excitability of a target cell. Other types of receptors include protein kinase receptors (e.g., receptor for
9216-528: The domain followed by kingdom , phylum , class , order , family , genus , and species . All organisms can be classified as belonging to one of three domains : Archaea (originally Archaebacteria), bacteria (originally eubacteria), or eukarya (includes the fungi, plant, and animal kingdoms). The history of life on Earth traces how organisms have evolved from the earliest emergence of life to present day. Earth formed about 4.5 billion years ago and all life on Earth, both living and extinct, descended from
9360-481: The duplication of its DNA and some of its organelles , and the subsequent partitioning of its cytoplasm into two daughter cells in a process called cell division . In eukaryotes (i.e., animal, plant, fungal , and protist cells), there are two distinct types of cell division: mitosis and meiosis . Mitosis is part of the cell cycle, in which replicated chromosomes are separated into two new nuclei. Cell division gives rise to genetically identical cells in which
9504-403: The kingdom Plantae, which would exclude fungi and some algae . Plant cells were derived by endosymbiosis of a cyanobacterium into an early eukaryote about one billion years ago, which gave rise to chloroplasts. The first several clades that emerged following primary endosymbiosis were aquatic and most of the aquatic photosynthetic eukaryotic organisms are collectively described as algae, which
9648-419: The lift coefficient is reduced by 10 to 15 percent compared to that of a monoplane using the same airfoil and aspect ratio . The lower wing is usually attached to the fuselage , while the upper wing is raised above the fuselage with an arrangement of cabane struts , although other arrangements have been used. Either or both of the main wings can support ailerons , while flaps are more usually positioned on
9792-423: The microscope . It was then that scholars discovered spermatozoa , bacteria, infusoria and the diversity of microscopic life. Investigations by Jan Swammerdam led to new interest in entomology and helped to develop techniques of microscopic dissection and staining . Advances in microscopy had a profound impact on biological thinking. In the early 19th century, biologists pointed to the central importance of
9936-412: The mitochondrial cristae . Oxidative phosphorylation comprises the electron transport chain, which is a series of four protein complexes that transfer electrons from one complex to another, thereby releasing energy from NADH and FADH 2 that is coupled to the pumping of protons (hydrogen ions) across the inner mitochondrial membrane ( chemiosmosis ), which generates a proton motive force . Energy from
10080-546: The phenotype of that dominant allele. During gamete formation, the alleles for each gene segregate, so that each gamete carries only one allele for each gene. Heterozygotic individuals produce gametes with an equal frequency of two alleles. Finally, the law of independent assortment , states that genes of different traits can segregate independently during the formation of gametes, i.e., genes are unlinked. An exception to this rule would include traits that are sex-linked . Test crosses can be performed to experimentally determine
10224-497: The regulation of the rate of a metabolic reaction, for example in response to changes in the cell's environment or to signals from other cells. Cellular respiration is a set of metabolic reactions and processes that take place in cells to convert chemical energy from nutrients into adenosine triphosphate (ATP), and then release waste products. The reactions involved in respiration are catabolic reactions , which break large molecules into smaller ones, releasing energy. Respiration
10368-503: The CR.42 was able to achieve success in the defensive night fighter role against RAF bombers that were striking industrial targets throughout northern Italy. The British Fleet Air Arm operated the Fairey Swordfish torpedo bomber from its aircraft carriers, and used the type in the anti-submarine warfare role until the end of the conflict, largely due to their ability to operate from
10512-518: The Caribou , performed the first non-stop flight between the Canadian mainland and Britain in 30 hours 55 minutes, although the intended target for this long distance flight had originally been Baghdad , Iraq . Despite its relative success, British production of the Dragon was quickly ended when in favour of the more powerful and elegant de Havilland Dragon Rapide , which had been specifically designed to be
10656-503: The First World War, the British Royal Aircraft Factory developed airfoil section wire named RAFwire in an effort to both increase the strength and reduce the drag. Four types of wires are used in the biplane wing structure. Drag wires inside the wings prevent the wings from being folded back against the fuselage, running inside a wing bay from the forward inboard corner to the rear outboard corner. Anti-drag wires prevent
10800-565: The French Nieuport 17 and German Albatros D.III , offered lower drag than a conventional biplane while being stronger than a monoplane. During the Interwar period , numerous biplane airliners were introduced. The British de Havilland Dragon was a particularly successful aircraft, using straightforward design to could carry six passengers on busy routes, such as London-Paris services. During early August 1934, one such aircraft, named Trail of
10944-644: The Germans had been experimenting with a new generation of monoplanes, such as the Fokker D.VIII , that might have ended the biplane's advantages earlier had the conflict not ended when it had. The French were also introducing the Morane-Saulnier AI , a strut-braced parasol monoplane , although the type was quickly relegated to the advanced trainer role following the resolution of structural issues. Sesquiplane types, which were biplanes with abbreviated lower wings such as
11088-475: The Phanerozoic eon that began 539 million years ago being subdivided into Paleozoic , Mesozoic , and Cenozoic eras. These three eras together comprise eleven periods ( Cambrian , Ordovician , Silurian , Devonian , Carboniferous , Permian , Triassic , Jurassic , Cretaceous , Tertiary , and Quaternary ). The similarities among all known present-day species indicate that they have diverged through
11232-696: The accumulation of favorable traits over successive generations, thereby increasing the match between the organisms and their environment. A species is a group of organisms that mate with one another and speciation is the process by which one lineage splits into two lineages as a result of having evolved independently from each other. For speciation to occur, there has to be reproductive isolation . Reproductive isolation can result from incompatibilities between genes as described by Bateson–Dobzhansky–Muller model . Reproductive isolation also tends to increase with genetic divergence . Speciation can occur when there are physical barriers that divide an ancestral species,
11376-736: The all-metal third one in the following June. All three went to the Royal Aircraft Establishment (RAE) for experimental flight. The second aircraft undertook comparative trials of a four-blade wooden airscrew against its original two-blade steel one. One of the three Berkeleys was still flying with the RAE at the end of 1930. Data from Barnes 1964 , p. 188 General characteristics Performance Armament Biplane Biplanes offer several advantages over conventional cantilever monoplane designs: they permit lighter wing structures, low wing loading and smaller span for
11520-407: The biplane naturally has a deep structure and is therefore easier to make both light and strong. Rigging wires on non-cantilevered monoplanes are at a much sharper angle, thus providing less tension to ensure stiffness of the outer wing. On a biplane, since the angles are closer to the ideal of being in direct line with the forces being opposed, the overall structure can then be made stiffer. Because of
11664-586: The body plan and the number, identity, and pattern of body parts. Among the most important toolkit genes are the Hox genes . Hox genes determine where repeating parts, such as the many vertebrae of snakes, will grow in a developing embryo or larva. Evolution is a central organizing concept in biology. It is the change in heritable characteristics of populations over successive generations . In artificial selection , animals were selectively bred for specific traits. Given that traits are inherited, populations contain
11808-450: The breaking down of glucose to pyruvate by cellular respiration ); or anabolic —the building up ( synthesis ) of compounds (such as proteins, carbohydrates, lipids, and nucleic acids). Usually, catabolism releases energy, and anabolism consumes energy. The chemical reactions of metabolism are organized into metabolic pathways , in which one chemical is transformed through a series of steps into another chemical, each step being facilitated by
11952-410: The cohesive force due to the attraction between molecules at the surface of the liquid. Water is also adhesive as it is able to adhere to the surface of any polar or charged non-water molecules. Water is denser as a liquid than it is as a solid (or ice). This unique property of water allows ice to float above liquid water such as ponds, lakes, and oceans, thereby insulating the liquid below from
12096-433: The cold air above. Water has the capacity to absorb energy, giving it a higher specific heat capacity than other solvents such as ethanol . Thus, a large amount of energy is needed to break the hydrogen bonds between water molecules to convert liquid water into water vapor . As a molecule, water is not completely stable as each water molecule continuously dissociates into hydrogen and hydroxyl ions before reforming into
12240-598: The competition aerobatics role and format for such a biplane well-defined by the mid-1930s by the Udet U 12 Flamingo and Waco Taperwing . The Pitts Special dominated aerobatics for many years after World War II and is still in production. The vast majority of biplane designs have been fitted with reciprocating engines . Exceptions include the Antonov An-3 and WSK-Mielec M-15 Belphegor , fitted with turboprop and turbofan engines respectively. Some older biplane designs, such as
12384-501: The cytoplasm of a cell, there are many biomolecules such as proteins and nucleic acids . In addition to biomolecules, eukaryotic cells have specialized structures called organelles that have their own lipid bilayers or are spatially units. These organelles include the cell nucleus , which contains most of the cell's DNA, or mitochondria , which generate adenosine triphosphate (ATP) to power cellular processes. Other organelles such as endoplasmic reticulum and Golgi apparatus play
12528-455: The cytoplasm, where it is converted to waste products that may be removed from the cell. This serves the purpose of oxidizing the electron carriers so that they can perform glycolysis again and removing the excess pyruvate. Fermentation oxidizes NADH to NAD so it can be re-used in glycolysis. In the absence of oxygen, fermentation prevents the buildup of NADH in the cytoplasm and provides NAD for glycolysis. This waste product varies depending on
12672-673: The development of biological knowledge. He explored biological causation and the diversity of life. His successor, Theophrastus , began the scientific study of plants. Scholars of the medieval Islamic world who wrote on biology included al-Jahiz (781–869), Al-Dīnawarī (828–896), who wrote on botany, and Rhazes (865–925) who wrote on anatomy and physiology . Medicine was especially well studied by Islamic scholars working in Greek philosopher traditions, while natural history drew heavily on Aristotelian thought. Biology began to quickly develop with Anton van Leeuwenhoek 's dramatic improvement of
12816-418: The development of body form, is the result of spatial differences in gene expression. A small fraction of the genes in an organism's genome called the developmental-genetic toolkit control the development of that organism. These toolkit genes are highly conserved among phyla , meaning that they are ancient and very similar in widely separated groups of animals. Differences in deployment of toolkit genes affect
12960-440: The developmental fate of a cell, which becomes more restrictive during development. Differentiation is the process by which specialized cells arise from less specialized cells such as stem cells . Stem cells are undifferentiated or partially differentiated cells that can differentiate into various types of cells and proliferate indefinitely to produce more of the same stem cell. Cellular differentiation dramatically changes
13104-411: The discovery of archaea in almost every habitat , including soil, oceans, and marshlands . Archaea are particularly numerous in the oceans, and the archaea in plankton may be one of the most abundant groups of organisms on the planet. Archaea are a major part of Earth's life . They are part of the microbiota of all organisms. In the human microbiome , they are important in the gut , mouth, and on
13248-420: The discovery of the double-helical structure of DNA by James Watson and Francis Crick in 1953, marked the transition to the era of molecular genetics . From the 1950s onwards, biology has been vastly extended in the molecular domain. The genetic code was cracked by Har Gobind Khorana , Robert W. Holley and Marshall Warren Nirenberg after DNA was understood to contain codons . The Human Genome Project
13392-470: The drag from the number of struts used. The structural forces acting on the spars of a biplane wing tend to be lower as they are divided between four spars rather than two, so the wing can use less material to obtain the same overall strength and is therefore lighter. A given area of wing also tends to be shorter, reducing bending moments on the spars, which then allow them to be more lightly built as well. The biplane does however need extra struts to maintain
13536-616: The earliest terrestrial ecosystems , at least 2.7 billion years ago. Microorganisms are thought to have paved the way for the inception of land plants in the Ordovician period. Land plants were so successful that they are thought to have contributed to the Late Devonian extinction event . Ediacara biota appear during the Ediacaran period, while vertebrates , along with most other modern phyla originated about 525 million years ago during
13680-542: The end of World War I . At the start of World War II , several air forces still had biplane combat aircraft in front line service but they were no longer competitive, and most were used in niche roles, such as training or shipboard operation, until shortly after the end of the war. The British Gloster Gladiator biplane, the Italian Fiat CR.42 Falco and Soviet I-153 sesquiplane fighters were all still operational after 1939. According to aviation author Gianni Cattaneo,
13824-473: The expression of deleterious recessive mutations . The beneficial effect of genetic complementation, derived from outcrossing (cross-fertilization) is also referred to as hybrid vigor or heterosis. Charles Darwin in his 1878 book The Effects of Cross and Self-Fertilization in the Vegetable Kingdom at the start of chapter XII noted “The first and most important of the conclusions which may be drawn from
13968-427: The fixed surfaces. The rudder was tall and also horn-balanced, but more elegantly than the elevators with the edge running smoothly into the fin. The undercarriage was of wide track, mounted to the wings below the centre section interplane struts and braced to the fuselage. The 650 hp (490 kW) Condor engine drove a two-blade propeller and had, after some Air Ministry input, a nose-mounted radiator under
14112-455: The form of glucose is the main nutrient used by animal and plant cells in respiration. Cellular respiration involving oxygen is called aerobic respiration, which has four stages: glycolysis , citric acid cycle (or Krebs cycle), electron transport chain , and oxidative phosphorylation . Glycolysis is a metabolic process that occurs in the cytoplasm whereby glucose is converted into two pyruvates , with two net molecules of ATP being produced at
14256-416: The gap between the wings, which add both weight and drag. The low power supplied by the engines available in the first years of aviation limited aeroplanes to fairly low speeds. This required an even lower stalling speed, which in turn required a low wing loading , combining both large wing area with light weight. Obtaining a large enough wing area without the wings being long, and thus dangerously flexible
14400-513: The hind limbs could not have opened out sideways but in flight would have hung below and slightly behind the fore limbs. Biology Biology is the scientific study of life . It is a natural science with a broad scope but has several unifying themes that tie it together as a single, coherent field. For instance, all organisms are made up of at least one cell that processes hereditary information encoded in genes , which can be transmitted to future generations. Another major theme
14544-426: The hormone insulin ) and G protein-coupled receptors . Activation of G protein-coupled receptors can initiate second messenger cascades. The process by which a chemical or physical signal is transmitted through a cell as a series of molecular events is called signal transduction . The cell cycle is a series of events that take place in a cell that cause it to divide into two daughter cells. These events include
14688-414: The initial step of photosynthesis whereby light energy is absorbed by chlorophyll pigments attached to proteins in the thylakoid membranes . The absorbed light energy is used to remove electrons from a donor (water) to a primary electron acceptor, a quinone designated as Q. In the second stage, electrons move from the quinone primary electron acceptor through a series of electron carriers until they reach
14832-406: The last eukaryotic common ancestor. Prokaryotes (i.e., archaea and bacteria) can also undergo cell division (or binary fission ). Unlike the processes of mitosis and meiosis in eukaryotes, binary fission in prokaryotes takes place without the formation of a spindle apparatus on the cell. Before binary fission, DNA in the bacterium is tightly coiled. After it has uncoiled and duplicated, it
14976-405: The ligand diffuses to nearby cells and affects them. For example, brain cells called neurons release ligands called neurotransmitters that diffuse across a synaptic cleft to bind with a receptor on an adjacent cell such as another neuron or muscle cell . In juxtacrine signaling, there is direct contact between the signaling and responding cells. Finally, hormones are ligands that travel through
15120-612: The low speeds and simple construction involved have inspired a small number of biplane ultralights, such as Larry Mauro's Easy Riser (1975–). Mauro also made a version powered with solar cells driving an electric motor called the Solar Riser . Mauro's Easy Riser was used by "Father Goose", Bill Lishman . Other biplane ultralights include the Belgian-designed Aviasud Mistral , the German FK12 Comet (1997–),
15264-519: The lower wing. Bracing is nearly always added between the upper and lower wings, in the form of interplane struts positioned symmetrically on either side of the fuselage and bracing wires to keep the structure from flexing, where the wings are not themselves cantilever structures. The primary advantage of the biplane over a monoplane is its ability to combine greater stiffness with lower weight. Stiffness requires structural depth and where early monoplanes had to have this provided with external bracing,
15408-444: The membrane serving as membrane transporters , and peripheral proteins that loosely attach to the outer side of the cell membrane, acting as enzymes shaping the cell. Cell membranes are involved in various cellular processes such as cell adhesion , storing electrical energy , and cell signalling and serve as the attachment surface for several extracellular structures such as a cell wall , glycocalyx , and cytoskeleton . Within
15552-403: The mitotic phase of an animal cell cycle—the division of the mother cell into two genetically identical daughter cells. The cell cycle is a vital process by which a single-celled fertilized egg develops into a mature organism, as well as the process by which hair, skin, blood cells , and some internal organs are renewed. After cell division, each of the daughter cells begin the interphase of
15696-437: The most abundant molecule in every organism. Water is important to life because it is an effective solvent , capable of dissolving solutes such as sodium and chloride ions or other small molecules to form an aqueous solution . Once dissolved in water, these solutes are more likely to come in contact with one another and therefore take part in chemical reactions that sustain life. In terms of its molecular structure , water
15840-561: The most famed copies was the Siemens-Schuckert D.I . The Albatros D.III and D.V , which had also copied the general layout from Nieuport, similarly provided the backbone of the German forces during the First World War. The Albatros sesquiplanes were widely acclaimed by their aircrews for their maneuverability and high rate of climb. During interwar period , the sesquiplane configuration continued to be popular, with numerous types such as
15984-403: The nature of their research questions and the tools that they use. Like other scientists, biologists use the scientific method to make observations , pose questions, generate hypotheses , perform experiments, and form conclusions about the world around them. Life on Earth, which emerged more than 3.7 billion years ago, is immensely diverse. Biologists have sought to study and classify
16128-588: The night ground attack role throughout the Second World War. In the case of the Po-2, production of the aircraft continued even after the end of the conflict, not ending until around 1952. A significant number of Po-2s were fielded by the Korean People's Air Force during the Korean War , inflicting serious damage during night raids on United Nations bases. The Po-2 is also the only biplane to be credited with
16272-612: The night bombing role; unfortunately for Handley-Page and Bristol, the Air Ministry had already decided, on the basis of experience with the Avro Aldershot that single-engine aircraft were not suitable for night-bombing. In the end, therefore the only successful contender was the Horsley, chosen for the day-bomber role. The second Berkeley was accepted by the Air Ministry in December 1925 and
16416-441: The observations given in this volume, is that generally cross-fertilisation is beneficial and self-fertilisation often injurious, at least with the plants on which I experimented.” Genetic variation , often produced as a byproduct of sexual reproduction, may provide long-term advantages to those sexual lineages that engage in outcrossing . Genetics is the scientific study of inheritance. Mendelian inheritance , specifically,
16560-517: The only class of macromolecules that are not made up of polymers. They include steroids , phospholipids , and fats, largely nonpolar and hydrophobic (water-repelling) substances. Proteins are the most diverse of the macromolecules. They include enzymes , transport proteins , large signaling molecules, antibodies , and structural proteins . The basic unit (or monomer) of a protein is an amino acid . Twenty amino acids are used in proteins. Nucleic acids are polymers of nucleotides . Their function
16704-404: The organism. In skeletal muscles, the waste product is lactic acid . This type of fermentation is called lactic acid fermentation . In strenuous exercise, when energy demands exceed energy supply, the respiratory chain cannot process all of the hydrogen atoms joined by NADH. During anaerobic glycolysis, NAD regenerates when pairs of hydrogen combine with pyruvate to form lactate. Lactate formation
16848-483: The outbreak of the First World War biplanes had gained favour after several monoplane structural failures resulted in the RFC's "Monoplane Ban" when all monoplanes in military service were grounded, while the French also withdrew most monoplanes from combat roles and relegated them to training. Figures such as aviation author Bruce observed that there was an apparent prejudice held even against newly-designed monoplanes, such as
16992-428: The possibility of common descent . Serious evolutionary thinking originated with the works of Jean-Baptiste Lamarck , who presented a coherent theory of evolution. The British naturalist Charles Darwin , combining the biogeographical approach of Humboldt , the uniformitarian geology of Lyell , Malthus's writings on population growth, and his own morphological expertise and extensive natural observations, forged
17136-907: The process of evolution from their common ancestor. Biologists regard the ubiquity of the genetic code as evidence of universal common descent for all bacteria , archaea , and eukaryotes . Microbial mats of coexisting bacteria and archaea were the dominant form of life in the early Archean eon and many of the major steps in early evolution are thought to have taken place in this environment. The earliest evidence of eukaryotes dates from 1.85 billion years ago, and while they may have been present earlier, their diversification accelerated when they started using oxygen in their metabolism . Later, around 1.7 billion years ago, multicellular organisms began to appear, with differentiated cells performing specialised functions. Algae-like multicellular land plants are dated back to about 1 billion years ago, although evidence suggests that microorganisms formed
17280-539: The process such as transcription , RNA splicing , translation , and post-translational modification of a protein. Gene expression can be influenced by positive or negative regulation, depending on which of the two types of regulatory proteins called transcription factors bind to the DNA sequence close to or at a promoter. A cluster of genes that share the same promoter is called an operon , found mainly in prokaryotes and some lower eukaryotes (e.g., Caenorhabditis elegans ). In positive regulation of gene expression,
17424-517: The promoter, gene expression can also be regulated by epigenetic changes to chromatin , which is a complex of DNA and protein found in eukaryotic cells. Development is the process by which a multicellular organism (plant or animal) goes through a series of changes, starting from a single cell, and taking on various forms that are characteristic of its life cycle. There are four key processes that underlie development: Determination , differentiation , morphogenesis , and growth. Determination sets
17568-407: The propeller shaft. The Ministry advised that the wings of the first two Berkeleys of the three specified in the contract should have wooden wings for speed of completion, with the third to be all metal. Leitner-Watts Metal airscrews were required for the second and third machine. The first Berkeley flew on 5 March 1925. The Type 90 Berkeley was the first Bristol aircraft to receive a type number at
17712-406: The proton motive force drives the enzyme ATP synthase to synthesize more ATPs by phosphorylating ADPs . The transfer of electrons terminates with molecular oxygen being the final electron acceptor . If oxygen were not present, pyruvate would not be metabolized by cellular respiration but undergoes a process of fermentation . The pyruvate is not transported into the mitochondrion but remains in
17856-424: The reduced stiffness, wire braced monoplanes often had multiple sets of flying and landing wires where a biplane could easily be built with one bay, with one set of landing and flying wires. The extra drag from the wires was not enough to offset the aerodynamic disadvantages from having two airfoils interfering with each other however. Strut braced monoplanes were tried but none of them were successful, not least due to
18000-634: The relatively compact decks of escort carriers . Its low stall speed and inherently tough design made it ideal for operations even in the often severe mid-Atlantic weather conditions. By the end of the conflict, the Swordfish held the distinction of having caused the destruction of a greater tonnage of Axis shipping than any other Allied aircraft. Both the German Heinkel He 50 and the Soviet Polikarpov Po-2 were used with relative success in
18144-449: The same time. Each pyruvate is then oxidized into acetyl-CoA by the pyruvate dehydrogenase complex , which also generates NADH and carbon dioxide. Acetyl-CoA enters the citric acid cycle, which takes places inside the mitochondrial matrix. At the end of the cycle, the total yield from 1 glucose (or 2 pyruvates) is 6 NADH, 2 FADH 2 , and 2 ATP molecules. Finally, the next stage is oxidative phosphorylation, which in eukaryotes, occurs in
18288-676: The second and third stages, respectively, provide the energy and electrons to drive the synthesis of glucose by fixing atmospheric carbon dioxide into existing organic carbon compounds, such as ribulose bisphosphate (RuBP) in a sequence of light-independent (or dark) reactions called the Calvin cycle . Cell signaling (or communication) is the ability of cells to receive, process, and transmit signals with its environment and with itself. Signals can be non-chemical such as light, electrical impulses , and heat, or chemical signals (or ligands ) that interact with receptors , which can be found embedded in
18432-700: The series of Nieuport military aircraft—from the Nieuport 10 through to the Nieuport 27 which formed the backbone of the Allied air forces between 1915 and 1917. The performance of the Nieuport sesquiplanes was so impressive that the Idflieg (the German Inspectorate of flying troops) requested their aircraft manufacturers to produce copies, an effort which was aided by several captured aircraft and detailed drawings; one of
18576-553: The skin. Their morphological, metabolic, and geographical diversity permits them to play multiple ecological roles: carbon fixation; nitrogen cycling; organic compound turnover; and maintaining microbial symbiotic and syntrophic communities, for example. Eukaryotes are hypothesized to have split from archaea, which was followed by their endosymbioses with bacteria (or symbiogenesis ) that gave rise to mitochondria and chloroplasts, both of which are now part of modern-day eukaryotic cells. The major lineages of eukaryotes diversified in
18720-682: The start of its design rather than retrospectively. The first Berkeley was accepted for trials at RAF Martlesham Heath in May 1925 and remained there until November. Other aircraft competing for production contracts were the Handley Page Handcross , the Hawker Horsley and the Westland Yeovil . The Handcross and the Berkeley were the two larger machines and for that reason judged more suitable for
18864-476: The stroma. This is analogous to the proton-motive force generated across the inner mitochondrial membrane in aerobic respiration. During the third stage of photosynthesis, the movement of protons down their concentration gradients from the thylakoid lumen to the stroma through the ATP synthase is coupled to the synthesis of ATP by that same ATP synthase. The NADPH and ATPs generated by the light-dependent reactions in
19008-415: The structural problems associated with monoplanes, but offered little improvement for biplanes. The default design for a biplane has the wings positioned directly one above the other. Moving the upper wing forward relative to the lower one is called positive stagger or, more often, simply stagger. It can increase lift and reduce drag by reducing the aerodynamic interference effects between the two wings by
19152-471: The third tenet, and by the 1860s most biologists accepted all three tenets which consolidated into cell theory . Meanwhile, taxonomy and classification became the focus of natural historians. Carl Linnaeus published a basic taxonomy for the natural world in 1735, and in the 1750s introduced scientific names for all his species. Georges-Louis Leclerc, Comte de Buffon , treated species as artificial categories and living forms as malleable—even suggesting
19296-439: The total number of chromosomes is maintained. In general, mitosis (division of the nucleus) is preceded by the S stage of interphase (during which the DNA is replicated) and is often followed by telophase and cytokinesis ; which divides the cytoplasm , organelles and cell membrane of one cell into two new cells containing roughly equal shares of these cellular components. The different stages of mitosis all together define
19440-665: The two planes when the high pressure air under the top wing and the low pressure air above the lower wing cancel each other out. This means that a biplane does not in practice obtain twice the lift of the similarly-sized monoplane. The farther apart the wings are spaced the less the interference, but the spacing struts must be longer, and the gap must be extremely large to reduce it appreciably. As engine power and speeds rose late in World War I , thick cantilever wings with inherently lower drag and higher wing loading became practical, which in turn made monoplanes more attractive as it helped solve
19584-431: The underlying genotype of an organism with a dominant phenotype. A Punnett square can be used to predict the results of a test cross. The chromosome theory of inheritance , which states that genes are found on chromosomes, was supported by Thomas Morgans 's experiments with fruit flies , which established the sex linkage between eye color and sex in these insects. A gene is a unit of heredity that corresponds to
19728-400: The upper and lower wings together. The sesquiplane is a type of biplane where one wing (usually the lower) is significantly smaller than the other. The word, from Latin, means "one-and-a-half wings". The arrangement can reduce drag and weight while retaining the biplane's structural advantages. The lower wing may have a significantly shorter span, or a reduced chord . Examples include
19872-656: The various forms of life, from prokaryotic organisms such as archaea and bacteria to eukaryotic organisms such as protists , fungi, plants, and animals. These various organisms contribute to the biodiversity of an ecosystem , where they play specialized roles in the cycling of nutrients and energy through their biophysical environment . The earliest of roots of science, which included medicine, can be traced to ancient Egypt and Mesopotamia in around 3000 to 1200 BCE . Their contributions shaped ancient Greek natural philosophy . Ancient Greek philosophers such as Aristotle (384–322 BCE) contributed extensively to
20016-434: The wings from folding up, and run from the underside of the outer wing to the lower wing root. Conversely, landing wires prevent the wings from sagging, and resist the forces when an aircraft is landing, and run from the upper wing centre section to outboard on the lower wings. Additional drag and anti-drag wires may be used to brace the cabane struts which connect the fuselage to the wings, and interplane struts, which connect
20160-421: The wings from moving forward when the aircraft stops and run the opposite direction to the drag wires. Both of these are usually hidden within the wings, and if the structure is sufficiently stiff otherwise, may be omitted in some designs. Indeed many early aircraft relied on the fabric covering of the wing to provide this rigidity, until higher speeds and forces made this inadequate. Externally, lift wires prevent
20304-419: Was Bristol's response, designed largely by W.T. Reid with finishing touches from Bristol's longtime chief designer, Frank Barnwell . It was a fabric-covered all-metal structured three-bay biplane, with equal span, unswept and unstaggered wings with Frise-type ailerons on the upper and lower planes. Structurally, the wings were of rolled steel and duralumin . The fuselage was built from steel tubes and had
20448-434: Was launched in 1990 to map the human genome . All organisms are made up of chemical elements ; oxygen , carbon , hydrogen , and nitrogen account for most (96%) of the mass of all organisms, with calcium , phosphorus , sulfur , sodium , chlorine , and magnesium constituting essentially all the remainder. Different elements can combine to form compounds such as water, which is fundamental to life. Biochemistry
20592-404: Was more readily accomplished with a biplane. The smaller biplane wing allows greater maneuverability . Following World War I, this helped extend the era of the biplane and, despite the performance disadvantages, most fighter aircraft were biplanes as late as the mid-1930s. Specialist sports aerobatic biplanes are still made in small numbers. Biplanes suffer aerodynamic interference between
20736-741: Was widely used by the United States Army Air Force (USAAF) while the US Navy operated the Naval Aircraft Factory N3N . In later civilian use in the US, the Stearman became particularly associated with stunt flying such as wing-walking , and with crop dusting, where its compactness worked well at low levels, where it had to dodge obstacles. Modern biplane designs still exist in specialist roles such as aerobatics and agricultural aircraft with
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