Misplaced Pages

Eurasian blackcap

Article snapshot taken from Wikipedia with creative commons attribution-sharealike license. Give it a read and then ask your questions in the chat. We can research this topic together.
#768231

113-421: The Eurasian blackcap ( Sylvia atricapilla ), usually known simply as the blackcap , is a common and widespread typical warbler . It has mainly olive-grey upperparts and pale grey underparts, and differences across the five subspecies are small. Both sexes have a neat coloured cap to the head, black in the male and reddish-brown in the female. The male's typical song is a rich musical warbling, often ending in

226-552: A type species but this was designated as the Eurasian blackcap ( Sylvia atricapilla ) by Charles Lucien Bonaparte in 1828. The genus name is from Modern Latin silvia , a woodland sprite, related to silva meaning "a wood". The typical warblers are now known to form a major lineage in a clade containing also the parrotbills and some taxa formerly considered to be Old World babblers . The other "Old World warblers" have been moved to their own families, entirely redelimiting

339-527: A Christmas song Sylvian Joululaulu (compare with the Latin name Sylvia atricapilla ), and is said to represent Finland before it claimed independence from Russia in 1917. Folk names for the blackcap often refer to its most obvious plumage feature (black-headed peggy, King Harry black cap and coal hoodie) or to its song, as in the "nightingale" names above. Other old names are based on its choice of nesting material (Jack Straw, hay bird, hay chat and hay Jack). There

452-556: A buckling of the lithosphere . The ten tallest peaks in Madeira exemplify the island's diverse topography. Pico Ruivo is the highest at 1,862 metres. Madeira's mountaintops offer panoramic vistas of rugged terrain and the Atlantic Ocean , attracting hikers and nature enthusiasts seeking stunning views and challenging trails. Madeira island is at the top of a massive shield volcano that rises about 6 km (20,000 ft) from

565-594: A chase, often leading to a fight. The typical territory size in a French study was 1.12 hectares (2.8 acres), but in insect-rich tall maquis in Gibraltar, the average was only 0.16 hectares (0.40 acres). Females feed within a home range which may overlap other blackcap territories, and covers up to six times the area of the defended zone. Sylvia warblers are unusual in that they vigorously defend their territories against other members of their genus as well as conspecifics . Blackcaps and garden warblers use identical habits in

678-565: A complete moult in their breeding areas in August and September prior to migration. Some birds, typically those migrating the greatest distances, have a further partial moult between December and March. Juveniles replace their loosely structured body feathers with adult plumage, starting earlier, but taking longer to complete, than the adults. Blackcaps breeding in the north of the range have an earlier and shorter post-juvenile moult than those further south, and cross-breeding of captive birds shows that

791-420: A distinct subspecies, S. a. obscura . The blackcap is a mainly grey warbler with distinct male and female plumages . The nominate subspecies is about 13 cm (5.1 in) long with a 7–8 cm (2.8–3.1 in) wing length. The weight is typically 16–25 g (0.56–0.88 oz), but can be up to 31 g (1.1 oz) for birds preparing to migrate. The adult male has olive-grey upperparts, other than

904-597: A few, and in several species has white sides. Many of the species show some sexual dimorphism , with distinctive male and female plumages , with the males in many having black or bright grey on the heads, replaced by brown, brownish-grey or similar dusky colours in females; about a third of the species also have a conspicuous red eye ring in males. Species breeding in cool temperate regions are strongly migratory , while most of those in warmer regions are partially migratory or resident. They are active warblers usually associated with open woodland, scrub, hedges or shrubs. Their diet

1017-453: A loud high-pitched crescendo, but a simpler song is given in some isolated areas, such as valleys in the Alps . The blackcap's closest relative is the garden warbler , which looks quite different but has a similar song. The blackcap breeds in much of Europe, western Asia and northwestern Africa, and its preferred habitat is mature deciduous woodland. The male holds a territory when breeding, which

1130-471: A minor increase in phylogenetic information content." The genus as currently circumscribed includes the following species: Madeira Madeira ( / m ə ˈ d ɪər ə / mə- DEER -ə or / m ə ˈ d ɛər ə / mə- DAIR -ə ; European Portuguese: [mɐˈðɐjɾɐ] ), officially the Autonomous Region of Madeira ( Portuguese : Região Autónoma da Madeira ),

1243-490: A more generalist diet than that of birds in traditional winter sites. Blackcaps did not exclusively feed in gardens; visits were linked to harsher weather. Individuals generally stayed at garden sites until immediately before spring departure, and supplemental feeding may have benefits for winter survival, When preparing for migration, abundant supplemental food may allow blackcaps to attain better body condition and may facilitate earlier and more successful breeding attempts. In

SECTION 10

#1732870154769

1356-506: A network of walking paths. Paths can provide easy and relaxing walks through the countryside, while others are narrow, crumbling ledges where a slip could result in injury or death. Some improvements have been made to these pathways following the 2010 Madeira floods and mudslides . Such improvements involved the continuous maintenance of streams, paving trails, and establishing safety fences on dangerous stretches. Due to its distinct geography, economy, social and cultural situation, as well as

1469-474: A north-northeast to south-southwest axis that extends for 1,000 kilometres (540 nmi). This structure consists of long geomorphological relief that extends from the abyssal plain to 3,500 m (11,500 ft); its highest submersed point reaches a depth of about 150 m (490 ft) (around latitude 36°N). The origins of the Tore-Madeira Ridge are not clearly established, but may have resulted from

1582-606: A number of houses and Santa Clara church were hit. The last Austrian Emperor , Charles I , was exiled to Madeira after the war. Determined to prevent an attempt to restore Charles to the throne, the Council of Allied Powers agreed he could go into exile on Madeira because it was isolated in the Atlantic and easily guarded. He died there on 1 April 1922 and his coffin lies in a chapel of the Church of Our Lady of Monte . The Madeira archipelago

1695-429: A paler grey nape and a neat black cap on the head. The underparts are light grey, becoming silvery white on the chin, throat and upper breast. The tail is dark grey, with an olive tint to the outer edge of each feather. The bill and long legs are grey, and the iris is reddish brown. The female resembles the male, but has a reddish-brown cap and a slightly browner tone to the grey of the upperparts. Juveniles are similar to

1808-489: A quiet subsong, a muted version of the full song, which is even harder to separate. The blackcap occasionally mimics the song of other birds, the most frequently copied including the garden warbler and the common nightingale . The main call is a hard tac-tac , like stones knocking together, and other vocalisations include a squeaking sweet alarm, and a low-pitched trill similar to that of a garden warbler. Male blackcaps will sometimes sing even when incubating, especially with

1921-619: A series of seven sketches of the island. In 1856, British troops recovering from cholera , and widows and orphans of soldiers fallen in the Crimean War , were stationed in Funchal, Madeira. During the Great War on 3 December 1916, a German U-boat , SM  U-38 , captained by Max Valentiner , entered Funchal harbour on Madeira. U-38 torpedoed and sank three ships, bringing the war to Portugal by extension. The ships sunk were: After attacking

2034-422: A settlement. The new settlers observed "a heavy black cloud suspended to the southwest" and upon investigation discovered the larger island they called Madeira ( Portuguese : madeira , lit.   'wood'). The first Portuguese settlers began colonizing the islands around 1420 or 1425. The first settlers were the three captain- donees and their respective families, a small group of members of

2147-617: A very narrow strait and lie 10,000 furlongs [2,000 km] from Africa. They are called the Isles of the Blessed ." Archaeological evidence suggests that the islands may have been visited by the Vikings sometime between 900 and 1030. Accounts by Muhammad al-Idrisi state that the Mugharrarin ("the adventurers" – seafarers from Lisbon) came across an island where they found "a huge quantity of sheep,

2260-624: A violent storm, their ship ran aground along the coast of an island that may have been Madeira. Later this legend was the basis of the naming of the city of Machico on the island, in memory of the young lovers. Madeira appears in several medieval manuscripts, including the Book of Knowledge of All Kingdoms from the early 14th century, the Medici-Laurentian Atlas from 1351, the Soleri Portolani from 1380 and 1385 and Corbitis Atlas from

2373-460: A way that it was not enough for domestic needs, so that sugar was imported to the island from other Portuguese colonies. Sugar mills were gradually abandoned, with few remaining, which gave way to other markets in Madeira. In the 17th century, as Portuguese sugar production was shifted to Brazil, São Tomé and Príncipe and elsewhere, Madeira's most important commodity product became its wine. Sugar plantations were replaced by vineyards, originating in

SECTION 20

#1732870154769

2486-457: A year. Madeira hosts lush laurel forests , while Porto Santo , a much flatter island, has a semiarid climate ( BSh ). In most winters snowfall occurs in the mountains. In the south, little is left of the indigenous subtropical rainforest that once covered the island (the original settlers set fires to clear the land for farming) and named it ( madeira means "wood" in Portuguese). However, in

2599-712: Is 43% (males 46%, females 29%), and 36% of juveniles live through their first year. The typical life expectancy is two years, but the record is 13 years and 10 months for a bird in the Czech Republic. The blackcap feeds mainly on insects during the breeding season, then switches to fruit in late summer, the change being triggered by an internal biological rhythm . When migrants arrive on their territories they initially take berries, pollen and nectar if there are insufficient insects available, then soon switch to their preferred diet. They mainly pick prey off foliage and twigs, but may occasionally hover, flycatch or feed on

2712-602: Is a tradition of the Royal Navy 's Fleet Air Arm bases being named for birds. A former base near Stretton in Cheshire was called HMS Blackcap . The blackcap has a very large range, and its population in Europe is estimated at 41–65 million breeding pairs. Allowing for birds breeding in Africa and Asia, the total population estimate is between 101 and 161 million individuals. It

2825-636: Is an autonomous region of Portugal . It is an archipelago situated in the North Atlantic Ocean , in the region of Macaronesia , just under 400 kilometres (250 mi) north of the Canary Islands — Spain —, 520 kilometres (320 mi) west of the Kingdom of Morocco and 805 kilometres (500 mi) southwest of mainland Portugal . Madeira sits on the African Tectonic Plate , although it

2938-429: Is approximately 22 km (14 mi) at its widest point (from Ponta da Cruz to Ponta de São Jorge), with a coastline of 150 km (90 mi). It has a mountain ridge that extends along the centre of the island, reaching 1,862 metres (6,109 feet) at its highest point ( Pico Ruivo ), staying below 200 metres along its eastern extent. The primitive volcanic foci responsible for the central mountainous area, consisted of

3051-532: Is culturally, politically and ethnically associated with Europe, with its population predominantly descended from Portuguese settlers. Its population was 251,060 in 2021. The capital of Madeira is Funchal , on the main island's south coast. The archipelago includes the islands of Madeira , Porto Santo , and the Desertas , administered together with the separate archipelago of the Savage Islands . Roughly half of

3164-453: Is defended against garden warblers as well as other blackcaps. The nest is a neat cup, built low in brambles or scrub, and the clutch is typically 4–6 mainly buff eggs, which hatch in about 11 days. The chicks fledge in 11–12 days, but are cared for by both adults for some time after leaving the nest. The blackcap is a partial migrant ; birds from the colder areas of its range winter in scrub or trees in northwestern Europe, around

3277-501: Is due to subtle habitat preferences rather than interspecies aggression. Blackcaps first breed when they are one year old, and are mainly monogamous , although both sexes may sometimes deviate from this. A male attracts a female to his territory through song and a display involving raising the black crown feathers, fluffing the tail, slow wingbeats, and a short flapping flight. He also builds one or more simple nests (cock nests), usually near his songpost. The final nest, which may be one of

3390-571: Is entitled to derogation from some EU policies. According to the Treaty on the Functioning of the European Union , both primary and secondary European Union law applies to Madeira, with possible derogations to take account of its "structural social and economic situation (...) which is compounded by their remoteness, insularity, small size, difficult topography and climate, economic dependence on

3503-480: Is known from the Selvagens , suggesting this seabird visited at least sporadically. Madeira is home to six species of brown mice, believed to be descendants of common European brown mice brought to the island by Vikings in the 9th century (or conceivably by 15th century Portuguese settlers), but diversified to the point where they cannot interbreed with their ancestral species or with one another. They have essentially

Eurasian blackcap - Misplaced Pages Continue

3616-454: Is largely insectivorous, though several species also eat fruit extensively, mainly small berries such as elder and ivy , particularly from late summer to late winter; one species (blackcap) also frequently takes a wide variety of human-provided foods on birdtables in winter. The genus Sylvia was introduced in 1769 by the Italian naturalist Giovanni Antonio Scopoli . Scopoli did not specify

3729-509: Is less beneficial since it tends to crush the seeds. Although any suitable fruit may be eaten, some have seasonal or local importance; elder makes up a large proportion of the diet of northern birds preparing for migration, and energy-rich olives and lentisc are favoured by blackcaps wintering in the Mediterranean. The continental birds wintering in British gardens rely on provided food, and

3842-666: Is located 520 km (280 nmi) from the African coast, 805 km (430 nmi) from the closest point in the European coast (the Portuguese town of Sagres , in Algarve ) and 1,000 km (540 nmi) from the capital of Portugal, Lisbon (approximately a one-and-a-half-hour flight). Madeira inhabits the extreme south of the Tore-Madeira Ridge , a bathymetric structure oriented along

3955-451: Is low because the cuckoo's eggs are often rejected. Blackcaps have evolved adaptations which make it difficult for the parasitic species to succeed, despite the cuckoo's tendency to lay eggs which resemble those of their host. Blackcaps are good at spotting alien eggs, and their own eggs are very alike within a clutch, making it easier to spot the intruder. There is, however, considerable variation between different clutches, making it harder for

4068-881: Is mature deciduous woodland, with good scrub cover below the trees. Other habitats, such as parks, large gardens and overgrown hedges, are used as long as they meet the essential requirements of tall trees for songposts and an established understory . Where other Sylvia warblers also breed, blackcaps tend to use taller trees than their relatives, preferably those with a good canopy, such as pedunculate oak . In prime habitat, breeding densities reach 100–200 pairs per square kilometre (250–500 pairs per square mile) in northern Europe, and 500–900 pairs per square kilometre (1,250–2,250 pairs per square mile) in Italy. Densities are much lower in poorer habitats such as conifer forests. Breeding occurs in Europe at altitudes up to 2,200 m (7,200 ft). The preferred winter habitat around

4181-546: Is more typical. The clutch is typically 4–6 eggs (range 2–7), which are usually buff with grey and brown blotches and a few dark brown spots. The average size of the egg is 19.7 mm × 14.7 mm (0.78 in × 0.58 in). The eggs are incubated for an average of 11 days (range 10–16). Both adults incubate, although only the female stays on the nest at night. The chicks are altricial , hatching naked and with closed eyes, and are fed by both parents. They fledge about 11–12 days after hatching, leaving

4294-451: Is noted for its Madeira wine , flora , and fauna , with its pre-historic laurel forest , classified as a UNESCO World Heritage Site . The destination is certified by EarthCheck . The main harbour in Funchal has long been the leading Portuguese port in cruise ship dockings, an important stopover for Atlantic passenger cruises between Europe, the Caribbean and North Africa . In addition,

4407-545: Is still open to this day. The British first amicably occupied the island in 1801 whereafter Colonel William Henry Clinton became governor. A detachment of the 85th Regiment of Foot under Lieutenant-colonel James Willoughby Gordon garrisoned the island. After the Peace of Amiens , British troops withdrew in 1802, only to reoccupy Madeira in 1807 until the end of the Peninsular War in 1814. In 1846 James Julius Wood wrote

4520-513: Is therefore classified by the International Union for Conservation of Nature as being of least concern . Blackcaps and other small birds are illegally trapped and hunted in large numbers in Mediterranean countries, particularly in Lebanon, Syria, Palestine, Malta, Libya, Egypt and Cyprus, where they are considered as a delicacy . Despite hunting and natural hazards, the European population of

4633-479: Is unclear, since birds from northwest Africa may be of this form. About 2% of male blackcaps on Madeira and the Azores are melanistic , with black plumage on the whole head and upper breast. Melanistic females are rarer, and are characterised by grey-brown underparts. This dark morph has also been recorded from the Canary Islands , but not from Cape Verde . The melanistic birds were formerly sometimes considered to be

Eurasian blackcap - Misplaced Pages Continue

4746-642: Is wet in the northwest, but dry in the southeast. In the 16th century the Portuguese started building levadas ( aqueducts ) to carry water to the agricultural regions in the south. Due to its mountainous landscape, building levadas promised to be a difficult venture, often tasked to criminals and convicts from continental Portugal to build these water canals. Many are cut into the mountainsides, accompanied by 40 km (25 mi) of tunnels, some of which are still accessible. Today levadas supply water and also provide hydro-electric power . Over 2,170 km (1,350 mi) of levadas were built and later provided

4859-583: The Gulf Stream and Canary Current , giving it mild to warm year-round temperatures. According to the Instituto de Meteorologia (IPMA), the average annual temperature at Funchal weather station is 19.6 °C (67.3 °F) for the 1981–2010 period. Relief is a determinant factor on precipitation levels; areas such as the Madeira Natural Park can get as much as 2,800 mm (110 in) of precipitation

4972-582: The International Business Centre of Madeira , also known as the Madeira Free Trade Zone, was established in the 1980s. It includes (mainly tax-related) incentives. Plutarch in his Parallel Lives ( Sertorius , 75 AD) referring to the military commander Quintus Sertorius (d. 72 BC), relates that after his return to Cádiz , he met sailors who spoke of idyllic Atlantic islands: "The islands are said to be two in number separated by

5085-546: The Leiern (drawling) song by the German ornithologists who first described it. The song's introduction is like that of other blackcaps, but the final warbling part is a simple alternation between two notes, as in a great tit 's call but more fluting. The main song is confusable with that of the garden warbler, but it is slightly higher pitched than in that species, more broken into discrete song segments, and less mellow. Both species have

5198-709: The Portuguese constitution and other laws, Madeira possesses its own political and administrative statute and has its own government . The branches of Government are the Regional Government and the Legislative Assembly , the latter elected by universal suffrage , using the D'Hondt method of proportional representation . The president of the Regional Government is appointed by the Representative of

5311-552: The Sylviidae . A molecular phylogenetic study using mitochondrial DNA sequence data published in 2011 found that the species in the genus Sylvia formed two distinct clades. Based on these results, the ornithologists Edward Dickinson and Leslie Christidis in the fourth edition of Howard and Moore Complete Checklist of the Birds of the World , chose to split the genus and moved most of

5424-613: The Trocaz pigeon , the Madeira Chaffinch and the Madeira firecrest . In addition extinct species include the Madeiran scops owl , two rail species, Rallus adolfocaesaris and R. lowei , and two quail species, Coturnix lignorum and C. alabrevis , and the Madeiran wood pigeon , a subspecies of the common wood pigeon and which was last seen in the early 20th century. A Great Auk bone

5537-711: The West Indies and England itself. The Madeira wine became very popular in the markets and it is also said to have been used in a toast during the Declaration of Independence by the Founding Fathers of the United States . In the eighteenth and nineteenth centuries, Madeira stands out for its climate and therapeutic effects. In the nineteenth century, visitors to the island integrated four major groups: patients, travellers, tourists and scientists. Most visitors belonged to

5650-475: The conquest of Ceuta and West Africans after further exploration of the African coast. Barbary corsairs from North Africa, who enslaved Europeans from ships and coastal communities throughout the Mediterranean region, captured 1,200 people in Porto Santo in 1617. Until the first half of the sixteenth century, Madeira was one of the major sugar markets of the Atlantic. Apparently, it is in Madeira that, in

5763-403: The garden warbler eventually metamorphosed into a blackcap. The blackcap's song has led to it being described as the "mock nightingale" or "country nightingale", and John Clare , in "The March Nightingale" describes the listener as believing that the rarer species has arrived prematurely. "He stops his own and thinks the nightingale/Hath of her monthly reckoning counted wrong". The song is also

SECTION 50

#1732870154769

5876-495: The 1980s. Birds of both species are longer-winged and lighter than in the past, suggesting a longer migration as the breeding range expands northwards. In recent decades, substantial numbers of continental European birds have taken to wintering in gardens in Great Britain, and, to a lesser extent, Ireland, where the blackcap was formerly just a summer visitor. Although the British climate is sub-optimal, compensatory factors include

5989-541: The African coast in the service of Prince Henry the Navigator , were driven off course by a storm to an island which they named Porto Santo (English: "holy harbour") in gratitude for divine deliverance from a shipwreck. The following year, Zarco and Vaz organised an expedition with Bartolomeu Perestrello . The trio travelled to the island of Porto Santo, claimed it on behalf of the Portuguese Crown, and established

6102-496: The Iberian Peninsula, migratory blackcaps (and European robins ) track similar climatic conditions over the season, which sedentary individuals must cope with great variation in climate over the year. This suggests a trade-off between the cost of travelling long distances of migrants, and the flexibility required by sedentary individuals to tolerate a wide variety of environmental conditions. The blackcap's main breeding habitat

6215-513: The Madeira sugar trade, with the refining and distribution concentrated in Antwerp. By the 1490s Madeira had overtaken Cyprus as a producer of sugar." Sugarcane production was the primary engine of the island's economy, which quickly afforded the Funchal metropolis economic prosperity. The production of sugar cane attracted adventurers and merchants from all parts of Europe, especially Italians , Basques , Catalans , and Flemish . This meant that, in

6328-766: The Mediterranean and Atlantic islands and in the milder west and south of the main Eurasian distribution often winter within the nesting range, but populations elsewhere are migratory . The blackcap is a leap-frog migrant; birds from the north of the breeding range travel furthest south, whereas Mediterranean breeders move much shorter distances. The wintering areas overlap with the breeding range, but also include extensive areas in West Africa , East Africa south to Lake Malawi , and further north in Ethiopia, South Sudan and Eritrea. The large majority of birds wintering in eastern Africa are of

6441-415: The Mediterranean and in tropical Africa. Some birds from Germany and western continental Europe have adapted to spending the winter in gardens in Great Britain and Ireland. Insects are the main food in the breeding season, but, for the rest of the year, blackcaps survive primarily on small fruit. Garden birds also eat bread, fat and peanuts in winter. Despite extensive hunting in Mediterranean countries and

6554-416: The Mediterranean is scrub and olive orchards, where densities approach the levels found in the best breeding areas. The British wintering population is atypical, with 95% found in gardens, mostly in towns at altitudes below 100 m (330 ft). In Africa, habitats include cultivated land, acacia scrub, mangroves and forest, and these warblers are found at altitudes up to 3,600 m (11,800 ft) in

6667-465: The Mediterranean or western Africa. This is because the British migrants arrive back on the breeding grounds earlier than blackcaps wintering around the Mediterranean, and form pairs before the southern birds arrive. Mixed pairings are also selected against because the hybrid young would migrate in an intermediate direction, which would take them into the Bay of Biscay . It now appears that the wintering birds in

6780-673: The Republic according to the results of the election to the legislative assemblies. The sovereignty of the Portuguese Republic is represented in Madeira by the Representative of the Republic, appointed by the President of the Republic on the advice of the Government of the Republic . The tasks of the Representative of the Republic are to sign and order the publication of regional legislative decrees and regional regulatory decrees or to exercise

6893-778: The UK come from a much wider area than previously thought. The majority come from France, and some individuals come from as far away as Spain and Poland. The steady supply of winter food in gardens gives even Spanish-breeding birds an opportunity to put on weight quicker than in their home range. A 2021 paper showed that blackcaps, particularly adults, wintering in Britain and Ireland showed high site fidelity and low movement between wintering sites, in contrast to blackcaps wintering in their traditional winter ranges. Adults that frequented gardens had better body condition, smaller fat stores, longer bills, and rounder wing tips. The bill and wing tip shapes reflected

SECTION 60

#1732870154769

7006-595: The blackcap has been rising for several decades as the range extends northwards, for example to Scotland and Denmark. There are occasional nesting records from outside the main range, such as in northern Israel and the Faroes , and wandering birds may appear further afield in Iceland or on the islands of Arctic Russia. In the Baltic, the spread of the blackcap appears to have been helped by the availability of territories formerly occupied by

7119-678: The blackcap in subgenus Sylvia and the garden warbler in Epilais . These sister species have a breeding range which extends farther northeast than all other Sylvia species except the lesser whitethroat and common whitethroat . The nearest relatives of the garden warbler outside the sister group are believed to be the African hill babbler and Dohrn's thrush-babbler , both of which should probably be placed in Sylvia rather than their current genera, Pseudoalcippe and Horizorhinus respectively. The blackcap

7232-457: The blackcap's song. The genus Sylvia , the typical warblers , forms part of a large family of Old World warblers, the Sylviidae . The blackcap and its nearest relative, the garden warbler , are an ancient species pair which diverged very early from the rest of the genus at between 12 and 16 million years ago. In the course of time, these two species have become sufficiently distinctive that they have been placed in separate subgenera, with

7345-438: The chicks to grow is replaced by fruit sugar which helps the birds to fatten for migration. Aphids are still taken while they are available, since they often contain sugars from the plant sap on which they feed. Blackcaps eat a wide range of small fruit, and squeeze out any seeds on a branch before consuming the pulp. This technique makes them an important propagator of mistletoe . The mistle thrush , which also favours that plant,

7458-407: The cock nests or built from scratch, is a neat cup of roots, stems and grasses lined with fine material such as hair. The nest is typically 5.5 cm (2.2 in) deep and 10 cm (3.9 in) across, and is built in the cover of bramble , scrubs or trees. It is constructed mainly by the female, and may be up to 4.5 m (15 ft) above the ground, although lower than 1 m (3.3 ft)

7571-414: The context of sugar production, slave labour was applied for the first time. The colonial system of sugar production was put into practice on the island of Madeira, on a much smaller scale, and later transferred, on a large scale, to other overseas production areas. Later on, this small scale of production was outmatched by Brazilian and São Tomean plantations. Madeiran sugar production declined in such

7684-454: The cuckoo to convincingly mimic a blackcap egg. The open habitat and cup nest of the warbler make it a potential target for the cuckoo; it may have experienced much higher levels of parasitism in the past, and countermeasures would have spread rapidly once they evolved. The only blood parasites found in a study of blackcaps trapped on migration were protozoans from the genera Haemoproteus and Plasmodium . The study concluded that 45.5% of

7797-490: The declining barred warbler . Typical warbler See text Parisoma Swainson in Richardson , 1832 The typical warblers are small birds belonging to the genus Sylvia in the "Old World warbler" (or sylviid warbler) family Sylviidae . There are seven species in the genus. Typical warblers occur in the temperate to tropical regions of Europe , western and central Asia , and Africa , with

7910-410: The development of agriculture on the island, the settlers had to chop down part of the dense forest and build a large number of water channels, called “levadas”, to carry the abundant waters on the north coast to the south coast of the island. Initially, the settlers produced wheat for their own sustenance but later began to export wheat to mainland Portugal. In earlier times, fish and vegetables were

8023-402: The disease wanes. The novel was adapted as films of the same name in 1917, 1943 and 1993. The last version was directed by Franco Zeffirelli , and its English-language version was retitled as Sparrow . In Saint François d'Assise , an opera by Messiaen , the orchestration is based on bird song. St Francis himself is represented by the blackcap. In Finland, a caged blackcap is featured in

8136-399: The east of the continent. Wintering birds wander in search of good fruit supplies, but often stay in good feeding areas, and return in subsequent winters. Migrants may occur in a wide variety of habitats, such as reed bed and fen , but show a preference for shrubland. When male blackcaps return to their breeding areas, they establish a territory . Adults that have previously bred return to

8249-413: The female, but their upperparts have a slight rufous tinge, and the breast and flanks have a more olive tone; young males have a darker brown cap than their female counterparts. This species is unmistakable; other dark-headed Sylvia species, such Sardinian and Orphean warblers have extensive black on the head instead of a small cap. They are also larger and have white edges on the tail. Blackcaps have

8362-486: The first water-driven sugar mill on Madeira, sugar production increased to over 6,000 arrobas (an arroba was equal to 11 to 12 kilograms or 24 to 26 pounds) by 1455, using advisers from Sicily and financed by Genoese capital (Genoa acted as an integral part of the island economy until the 17th century). The accessibility of Madeira attracted Genoese and Flemish traders, who were keen to bypass Venetian monopolies. "By 1480 Antwerp had some seventy ships engaged in

8475-672: The floor of the Atlantic Ocean, on the Tore underwater mountain range. The volcano formed atop an east–west rift in the oceanic crust along the African Plate , beginning during the Miocene epoch (5 million years ago), continuing into the Pleistocene (700,000 years ago). This was followed by extensive erosion , producing two large amphitheatres opening southward in the central part of

8588-463: The gentry, people of modest conditions and some former inmates of the kingdom . The majority of settlers were fishermen and peasant farmers who willingly left Portugal for a new life on the islands, a better one, they hoped, than was possible in a Portugal which had been ravaged by the Black Death , and where the best farmlands were strictly controlled by the nobility. To have minimum conditions for

8701-476: The ground. Blackcaps eat a wide range of invertebrate prey, although aphids are particularly important early in the season, and flies , beetles and caterpillars are also taken in large numbers. Small snails are swallowed whole, since the shell is a source of calcium for the bird's eggs. Chicks are mainly fed soft-bodied insects, fruit only if invertebrates are scarce. In July, the diet switches increasingly to fruit. The protein needed for egg-laying and for

8814-432: The hand. Blackcaps are caught by Eurasian sparrowhawks in the breeding range, and by Eleonora's falcons on migration. Eurasian jays and Eurasian magpies take eggs and young, as do mammals such as stoats , weasels and squirrels . Domestic cats are the most important predator, possibly killing up to 10% of blackcaps. Blackcaps are occasionally hosts of the common cuckoo , a brood parasite . The level of parasitism

8927-497: The head)". Fossils and subfossils of the blackcap have been found in a number of European countries; the oldest, dated to 1.2–1.0 million years ago, are from the Early Pleistocene of Bulgaria. Fossils from France show that the genus Sylvia dates back at least 20 million years. The differences between subspecies are small, making subspecific boundaries hard to define, and the exact distribution of S. a. heineken

9040-478: The highest species diversity centred on the Mediterranean . They are strongly built, with stouter legs and a slightly thicker bill than many other warblers . The plumage is in varying shades of grey and brown, usually darker above and paler below, with bluish or pinkish tones in several species; several also have orange-brown or rufous fringed wing feathers. The tail is square-ended in most, slightly rounded in

9153-462: The historical autonomic aspirations of the population, the Autonomous Regions of Madeira was established in 1976. Although it is a politico-administrative autonomous region, the Portuguese constitution specifies both a regional and national connection, obliging their administrations to maintain democratic principles and promote regional interests, while reinforcing national unity. As defined by

9266-429: The interior generally inaccessible. Daily life is concentrated in the many villages at the mouths of the ravines, through which the heavy autumn and winter rains travel to the sea. Madeira has many different bioclimates. Based on differences in sun exposure, humidity, and annual mean temperature, clear variations distinguish north- and south-facing regions, as well as some islands. The islands are strongly influenced by

9379-436: The island. Volcanic activity later resumed, producing scoria cones and lava flows atop the eroded shield. The most recent volcanic eruptions were on the west-central part of the island 6,500 years ago, creating more cinder cones and lava flows. It is the largest island of the group with an area of 741 km (286 sq mi), a length of 57 km (35 mi) (from Ponte de São Lourenço to Ponta do Pargo). It

9492-433: The late 14th century. These texts refer to Madeira as Lecmane , Lolegname, Legnami (the isle of wood), Puerto or Porto Santo, deserte or deserta, and desierta . It is widely accepted that knowledge of these Atlantic islands existed before their better-documented discovery and successful settlement by the Kingdom of Portugal . In 1418, two captains, João Gonçalves Zarco and Tristão Vaz Teixeira , while exploring

9605-517: The major items are bread and fat, each making up around 20% of the diet; one bird survived the whole winter eating only Christmas cake . Fruit is also eaten, notably cotoneaster (41% of the fruit consumed), ivy and honeysuckle , and apple if available. Some birds have learned to take peanuts from feeders. Blackcaps defend good winter food sources in the wild, and at garden feeding stations they repel competitors as large as starlings and blackbirds . Birds occasionally become tame enough to feed from

9718-405: The males and 22.7% of the females were affected, but the number of parasites was small, and the ability to store fat for the migration flight was unimpaired. Seventeen strains of H. parabelopolskyi are found only in the blackcap, and form a monophyletic group; three further members of that group are found only in the garden warbler, and another three occur in the African hill babbler, supporting

9831-452: The meat of which was bitter and inedible" before going to the more inhabited Canary Islands , in Spain . This island, possibly Madeira or Hierro , must have been inhabited or previously visited by people for livestock to be present. During the reign of King Edward III of England , lovers Robert Machim and Anna d'Arfet were said to have fled from England to France in 1346. Driven off course by

9944-424: The milder climate of the Mediterranean and the Atlantic islands; triple brooding has been observed once, the female laying a total of 23 eggs in the season. Of eggs laid, 65–93% hatch successfully, and 75–92% of the chicks go on to fledge. The productivity (young fledged per nest) varies with location, level of predation and quality of habitat, but the national figure for the UK was 2.5. The adult annual survival rate

10057-535: The moneyed aristocracy . As a result of a high demand for the season, there was a need to prepare guides for visitors. The first tourist guide of Madeira appeared in 1850 and focused on elements of history, geology, flora , fauna and customs of the island. Regarding hotel infrastructures, the British and the Germans were the first to launch the Madeiran hotel chain. The historic Belmond Reid's Palace opened in 1891 and

10170-432: The natural hazards of predation and disease, the blackcap has been extending its range for several decades, and is classified by the International Union for Conservation of Nature as least concern . Its rich and varied song has led to it being described as the "mock nightingale" and it has featured in literature, films and music. In Messiaen's opera Saint François d'Assise , the saint is represented by themes based on

10283-463: The nest shortly before they are able to fly. They are assisted with feeding for a further two or three weeks. If the nest is threatened, the non-incubating bird gives an alarm call so that the sitting parent and chicks stay still and quiet. A male blackcap may mob a potential predator, or try to lure it away with disjointed runs and flaps on the ground. The blackcap normally raises just one brood, but second nestings are sometimes recorded, particularly in

10396-437: The north, the valleys harbor native trees. These laurissilva forests , notably those on the northern slopes, are designated as a UNESCO World Heritage Site . Madeira's paleobotanical record reveals that laurissilva forest has existed for at least 1.8 million years. Critically endangered species such as the vine Jasminum azoricum and the rowan Sorbus maderensis are endemic. The Madeiran large white butterfly

10509-457: The peaks: Ruivo (1,862 m), Torres (1,851 m), Arieiro (1,818 m), Cidrão (1,802 m), Cedro (1,759 m), Casado (1,725 m), Grande (1,657 m), Ferreiro (1,582 m). At the end of this eruptive phase, reefs encircled the island, its marine vestiges evident in a calcareous layer in the area of Lameiros, in São Vicente. Sea cliffs, such as Cabo Girão , valleys and ravines extend from this central spine, leaving

10622-751: The population lives in Funchal. The region has political and administrative autonomy through the Administrative Political Statute of the Autonomous Region of Madeira provided for in the Portuguese Constitution . The region is an integral part of the European Union as an outermost region . Madeira generally has a mild/moderate subtropical climate with mediterranean summer droughts and winter rain. Many microclimates are found at different elevations. Madeira, uninhabited at

10735-559: The ready availability of food, (particularly from bird tables ), a shorter migration distance, and the avoidance of the Alps and the Sahara Desert . It was originally thought that most of these wintering birds come from Germany, and isotope analysis (which enables the wintering location to be determined) showed that the continental birds wintering in Britain tend to mate only among themselves, and do not usually interbreed with those wintering in

10848-516: The right of veto over regional laws, should these laws be unconstitutional. Before the sixth amendment to the Portuguese Constitution passed in 2006, this responsibility was held by a more-powerful Minister of the Republic, who was proposed by the Government and appointed by the President. Madeira is an Outermost Region (OMR) of the European Union , meaning that due to its geographical situation, it

10961-467: The same genes, but rearranged to give different chromosome numbers: the ancestral species has 40 chromosomes, whereas the Madeira species have from 22 to 30. The deep valleys of Madeira are separated by high ground, and the different species of mice do not encounter each other. A levada is an irrigation channel or aqueduct specific to the Portuguese Atlantic region of Madeira. Madeira island

11074-410: The same wood, yet aggressive interactions mean that their territories never overlap. Similar songs are a feature of the Sylvia warblers as a group, and it has been suggested that this promotes interspecific competition and helps to segregate territories between related species. It appears more likely from later studies that segregation of sympatric species, other than the blackcap and garden warbler,

11187-542: The second brood. This appears to be intended to maintain the bond with the female. Wintering birds in Africa are quiet initially, but start singing in January or February prior to their return north. The continental breeding range of the blackcap lies between the 14–30° July isotherms , and is occupied by the nominate subspecies, the other forms being restricted to islands or fringe areas in the Caucasus and eastern Iberia. Birds on

11300-438: The second half of the fifteenth century, the city of Funchal became a mandatory port of call for European trade routes. Slaves were used during the island's period of sugar trade to cultivate sugar cane alongside paid workers, though slave owners were only a small minority of the Madeiran population, and those who did own slaves owned only a few. Slaves consisted of Guanches from the nearby Canary islands, captured Berbers from

11413-400: The settlers' main means of subsistence. Grain production began to fall and the ensuing crisis forced Henry the Navigator to order other commercial crops to be planted so that the islands could be profitable. These specialised plants, and their associated industrial technology, created one of the major revolutions on the islands and fuelled Portuguese industry. Following the introduction of

11526-490: The shared ancestry of the three bird species. The protozoan Isospora ashmoonensis was first identified in a blackcap in Egypt. Blackcaps may carry parasitic worms that sometimes kill their hosts. External parasites include chewing lice and feather mites . The latter do little damage, although heavy infestations cause individual tail feathers to develop asymmetrically. Aristotle , in his History of Animals , considered that

11639-469: The ships, U-38 bombarded Funchal for two hours from a range of about 3 kilometres (2 mi). Batteries on Madeira returned fire and eventually forced U-38 to withdraw. On 12 December 1917, two German U-boats, SM U-156 and SM U-157 (captained by Max Valentiner ), again bombarded Funchal. This time the attack lasted around 30 minutes. The U-boats fired 40 120 and 150 mm (4.7 and 5.9 in) shells. There were three fatalities and 17 wounded;

11752-409: The site they have used in previous summers, whereas inexperienced birds either wander until they find a suitable area, or establish a very large initial territory which contracts under pressure from neighbours. Territorial boundaries are established initially by loud singing, performed while the male displays with his crown raised, tail fanned and slow wingbeats. This display is followed, if necessary, by

11865-531: The so-called ‘Wine Culture’, which acquired international fame and provided the rise of a new social class, the Bourgeoisie . With the increase of commercial treaties with England, important English merchants settled on the Island and, ultimately, controlled the increasingly important island wine trade. The English traders settled in the Funchal as of the seventeenth century, consolidating the markets from North America,

11978-467: The southwest Asian race, S. a. dammholzi . There is a migratory divide in Europe at longitude 10–11°E. Birds to the west of this line head southwest towards Iberia or West Africa, whereas populations to the east migrate to the eastern Mediterranean and on to East Africa. Cross-breeding in captivity of birds from the resident population on the Canary Islands with migratory blackcaps from Germany showed

12091-468: The species into a resurrected genus Curruca retaining only the Eurasian blackcap and the garden warbler in Sylvia . In an additional change they moved the African hill babbler and Dohrn's thrush-babbler into Sylvia . The split was not made by the British Ornithologists' Union on the grounds that "a split into two genera would unnecessarily destabilize nomenclature and results in only

12204-484: The time, was claimed by Portuguese sailors in the service of Prince Henry the Navigator in 1419 and settled after 1420. The archipelago is the first territorial discovery of the exploratory period of the Age of Discovery . Madeira is a year-round resort, particularly for Portuguese, but also British (148,000 visits in 2021), and Germans (113,000). It is by far the most populous and densely populated Portuguese island. The region

12317-417: The timing is genetically controlled. The male's song is a rich musical warbling, often ending in a loud high-pitched crescendo, which is given in bursts of up to 30 seconds. The song is repeated for about two-and-a-half minutes, with a short pause before each repetition. In some geographically isolated areas, such as islands, peninsulas and valleys in the Alps , a simplified fluting song occurs, named

12430-437: The topic of Italian poet Giovanni Pascoli 's "La Capinera" [The Blackcap]. Giovanni Verga 's 1871 novel Storia di una capinera , according to its author, was inspired by a story of a blackcap trapped and caged by children. The bird, silent and pining for its lost freedom, eventually dies. In the book, a nun evacuated from her convent by cholera falls in love with a family friend, only to have to return to her confinement when

12543-584: The urge to migrate is genetically controlled, the offspring showing intermediate behaviour in terms of restlessness at migration time. Similar experiments using birds from southern Germany and eastern Austria, on opposite sides of the migratory divide, demonstrated that the direction of migration is also genetically determined. Climate change appears to be affecting the migration pattern of the garden warbler and blackcap. Both are arriving in Europe earlier than previously, and blackcaps and juvenile (but not adult) garden warblers are departing nearly two weeks later than in

12656-475: Was an endemic subspecies of the large white that inhabited the laurissilva forests but has not been seen since 1977. Hogna ingens , the Deserta Grande wolf spider, is endemic to the Madeira archipelago, specifically Deserta Grande Island . It is critically endangered. It is considered the largest member of its family . Restoration efforts are underway. Three species of birds are endemic to Madeira:

12769-463: Was one of the many bird species originally described by Carl Linnaeus in his landmark 1758 10th edition of Systema Naturae , as Motacilla atricapilla . The current genus name is from Modern Latin silvia , a woodland sprite, related to silva , a wood. The species name, like the English name, refers to the male's black cap. Atricapilla is from the Latin ater , "black", and capillus , "hair (of

#768231