130-628: An ant colony is a population of ants , typically from a single species, capable of maintaining their complete lifecycle. Ant colonies are eusocial , communal, and efficiently organized and are very much like those found in other social Hymenoptera , though the various groups of these developed sociality independently through convergent evolution . The typical colony consists of one or more egg -laying queens, numerous sterile females (workers, soldiers) and, seasonally, many winged sexual males and females. In order to establish new colonies, ants undertake flights that occur at species-characteristic times of
260-412: A Proto-Indo-European word *morwi- was the root for Sanskrit vamrah , Greek μύρμηξ mýrmēx , Old Church Slavonic mraviji , Old Irish moirb , Old Norse maurr , Dutch mier , Swedish myra , Danish myre , Middle Dutch miere , and Crimean Gothic miera . The family Formicidae belongs to the order Hymenoptera , which also includes sawflies , bees , and wasps . Ants evolved from
390-480: A cosmopolitan distribution . They are found on all continents except Antarctica , and only a few large islands, such as Greenland , Iceland , parts of Polynesia and the Hawaiian Islands lack native ant species. Ants occupy a wide range of ecological niches and exploit many different food resources as direct or indirect herbivores, predators and scavengers. Most ant species are omnivorous generalists , but
520-500: A Hansom Cab (1886), the fastest-selling crime novel of the era, is set in Melbourne, as is Australia's best-selling book of poetry, The Songs of a Sentimental Bloke (1915) by C. J. Dennis . Contemporary Melbourne authors who have set novels in the city include Peter Carey , Helen Garner and Gerald Murnane . Melbourne has Australia's widest range of bookstores, as well as the nation's largest publishing sector. The city also hosts
650-647: A campaign to revive the economy with an aggressive development campaign of public works coupled with the promotion of the city as a tourist destination with a focus on major events and sports tourism. During this period the Australian Grand Prix moved to Melbourne from Adelaide . Major projects included the construction of a new facility for the Melbourne Museum , Federation Square , the Melbourne Convention & Exhibition Centre , Crown Casino and
780-400: A conservative estimate of the ants at about 20 × 10 (20 quadrillion). Ants range in size from 0.75 to 52 millimetres (0.030–2.0 in), the largest species being the fossil Titanomyrma giganteum , the queen of which was 6 cm ( 2 + 1 ⁄ 2 in) long with a wingspan of 15 cm (6 in). Ants vary in colour; most ants are yellow to red or brown to black, but
910-496: A contributing factor to the Australian economic depression of the 1890s and the Australian banking crisis of 1893 . The effects of the depression on the city were profound, with virtually no significant construction until the late 1890s. At the time of Australia's federation on 1 January 1901 Melbourne became the seat of government of the federated Commonwealth of Australia . The first federal parliament convened on 9 May 1901 in
1040-499: A few are specialist feeders. There is considerable variation in ant abundance across habitats, peaking in the moist tropics to nearly six times that found in less suitable habitats. Their ecological dominance has been examined primarily using estimates of their biomass : myrmecologist E. O. Wilson had estimated in 2009 that at any one time the total number of ants was between one and ten quadrillion ( short scale ) (i.e., between 10 and 10 ) and using this estimate he had suggested that
1170-406: A few species are green and some tropical species have a metallic lustre . More than 13,800 species are currently known (with upper estimates of the potential existence of about 22,000; see the article List of ant genera ), with the greatest diversity in the tropics. Taxonomic studies continue to resolve the classification and systematics of ants. Online databases of ant species, including AntWeb and
1300-505: A great deal of digging. The casts are often used for research and education purposes but many are simply given or sold to natural history museums or sold as folk art or as souvenirs. Usually, the hills are chosen after the ants have abandoned so as to not kill any ants; however in the Southeast United States, pouring into an active colony of invasive fire ants is a novel way to eliminate them. An ant-bed , in its simplest form,
1430-409: A high resolution image. They also have three small ocelli (simple eyes) on the top of the head that detect light levels and polarization . Compared to vertebrates , ants tend to have blurrier eyesight, particularly in smaller species, and a few subterranean taxa are completely blind . However, some ants, such as Australia's bulldog ant , have excellent vision and are capable of discriminating
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#17328699479941560-547: A large area unite. They still continue to recognize genetic differences in order to mate, but the different colonies within the super colony avoid aggression. Until 2000, the largest known ant supercolony was on the Ishikari coast of Hokkaidō , Japan. The colony was estimated to contain 306 million worker ants and one million queen ants living in 45,000 nests interconnected by underground passages over an area of 2.7 km (670 acres). In 2000, an enormous supercolony of Argentine ants
1690-508: A lineage within the stinging wasps , and a 2013 study suggests that they are a sister group of the Apoidea . However, since Apoidea is a superfamily, ants must be upgraded to the same rank. A more detailed basic taxonomy was proposed in 2020. Three species of the extinct mid- Cretaceous genera Camelomecia and Camelosphecia were placed outside of the Formicidae, in a separate clade within
1820-460: A major renewal of the CBD and St Kilda Road which significantly modernised the city. New fire regulations and redevelopment saw most of the taller pre-war CBD buildings either demolished or partially retained through a policy of facadism . Many of the larger suburban mansions from the boom era were also either demolished or subdivided. To counter the trend towards low-density suburban residential growth,
1950-446: A node-like petiole . The head, mesosoma , and metasoma are the three distinct body segments (formally tagmata ). The petiole forms a narrow waist between their mesosoma ( thorax plus the first abdominal segment, which is fused to it) and gaster (abdomen less the abdominal segments in the petiole). The petiole may be formed by one or two nodes (the second alone, or the second and third abdominal segments). Tergosternal fusion, when
2080-610: A pastoral estate in Bulleen , now the Heide Museum of Modern Art . The city is also home to the Australian Centre for Contemporary Art , as well as numerous independent galleries and artist-run spaces. In the 2000s, street art proliferated in Melbourne , with Banksy saying its graffiti scene "leads the world", and "laneway galleries" becoming major tourist sites; Hosier Lane for example attracts more Instagram hashtags than some of
2210-700: A prominent example of the latter style in the CBD. The 1926 Nicholas Building is the city's grandest example of the Chicago School style, while the influence of Art Deco is apparent in the Manchester Unity Building , completed in 1932. The city also features the Shrine of Remembrance , which was built as a memorial to the men and women of Victoria who served in World War I and is now a memorial to all Australians who have served in war. Residential architecture
2340-403: A protective casing around the body and a point of attachment for muscles, in contrast to the internal skeletons of humans and other vertebrates . Insects do not have lungs ; oxygen and other gases, such as carbon dioxide , pass through their exoskeleton via tiny valves called spiracles . Insects also lack closed blood vessels ; instead, they have a long, thin, perforated tube along the top of
2470-456: A rapid clearing trend to sunny and relatively calm weather and the temperature rising back to what it was before the shower. This can occur in the space of minutes and can be repeated many times a day, giving Melbourne a reputation for having "four seasons in one day", a phrase that is part of local popular culture . The lowest temperature on record is −2.8 °C (27.0 °F), on 21 July 1869. The highest temperature recorded in Melbourne city
2600-430: A single gene of Solenopsis invicta can decide whether the colony will have single or multiple queens. The Australian jack jumper ant ( Myrmecia pilosula ) has only a single pair of chromosomes (with the males having just one chromosome as they are haploid ), the lowest number known for any animal, making it an interesting subject for studies in the genetics and developmental biology of social insects. Genome size
2730-432: Is Flinders Street station and its main regional rail and road coach terminus is Southern Cross station . It also has Australia's most extensive freeway network and the largest urban tram network in the world . Aboriginal Australians have lived in the Melbourne area for at least 40,000 years. When European colonisers arrived in the 19th century, at least 20,000 Kulin people from three distinct language groups –
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#17328699479942860-492: Is paraphyletic In 1966, E. O. Wilson and his colleagues identified the fossil remains of an ant ( Sphecomyrma ) that lived in the Cretaceous period. The specimen, trapped in amber dating back to around 92 million years ago, has features found in some wasps, but not found in modern ants. The oldest fossils of ants date to the mid-Cretaceous, around 100 million years ago, which belong to extinct stem-groups such as
2990-566: Is a fundamental characteristic of an organism. Ants have been found to have tiny genomes, with the evolution of genome size suggested to occur through loss and accumulation of non-coding regions, mainly transposable elements , and occasionally by whole genome duplication. This may be related to colonisation processes, but further studies are needed to verify this. Melbourne, Australia Melbourne ( / ˈ m ɛ l b ər n / MEL -bərn , locally [ˈmæɫbən] ; Boonwurrung / Woiwurrung : Narrm or Naarm )
3120-408: Is a pile of soil , sand , pine needles, manure , urine , or clay or a composite of these and other materials that build up at the entrances of the subterranean dwellings of ant colonies as they are excavated. A colony is built and maintained by legions of worker ants , who carry tiny bits of dirt and pebbles in their mandibles and deposit them near the exit of the colony. They normally deposit
3250-604: Is becoming unaffordable for some. Public housing is managed and provided by the Victorian Government's Department of Families, Fairness and Housing , and operates within the framework of the Commonwealth-State Housing Agreement, by which both federal and state governments provide funding for housing. Melbourne is experiencing high population growth, generating high demand for housing. This housing boom has increased house prices and rents, as well as
3380-598: Is home to more skyscrapers than any other Australian city , the tallest being Australia 108 , situated in Southbank. Melbourne's newest planned skyscraper, Southbank By Beulah (also known as "Green Spine"), has recently been approved for construction and will be the tallest structure in Australia by 2025. The CBD and surrounds also contain many significant historic buildings such as the Royal Exhibition Building ,
3510-546: Is known for its music, theatre and arts scenes, as well as its diverse range of cultural events and festivals, including the Melbourne International Arts Festival , Melbourne Fringe Festival and Moomba , Australia's largest free community festival. For much of the 2010s, Melbourne topped The Economist Intelligence Unit 's list of the world's most liveable cities , partly due to its cultural attributes. State Library Victoria , founded in 1854,
3640-437: Is largely protected from the ocean, resulting in greater temperature variation across seasons. Melbourne's urban area is approximately 2,704 km , the largest in Australia and the 33rd largest in the world. The Hoddle Grid , a grid of streets measuring approximately 1 by 1 ⁄ 2 mi (1.61 by 0.80 km), forms the nucleus of Melbourne's central business district (CBD). The grid's southern edge fronts onto
3770-693: Is not defined by a single architectural style, but rather an eclectic mix of large McMansion -style houses (particularly in areas of urban sprawl), apartment buildings, condominiums, and townhouses which generally characterise the medium-density inner-city neighbourhoods. Freestanding dwellings with relatively large gardens are perhaps the most common type of housing outside inner city Melbourne. Victorian terrace housing, townhouses and historic Italianate , Tudor revival and Neo-Georgian mansions are all common in inner-city neighbourhoods such as Carlton, Fitzroy and further into suburban enclaves like Toorak . Often referred to as Australia's cultural capital, Melbourne
3900-554: Is noted for its street art , live music and theatre scenes. It hosts major annual sporting events, such as the Australian Grand Prix and the Australian Open , and also hosted the 1956 Summer Olympics . Melbourne ranked as the world's most livable city for much of the 2010s. Melbourne Airport is the second-busiest airport in Australia and the Port of Melbourne is the nation's busiest seaport. Its main metropolitan rail terminus
4030-481: Is often referred to as Australia's garden city, and the state of Victoria is known as the garden state . There is an abundance of parks and gardens in Melbourne , many close to the CBD with a variety of common and rare plant species amid landscaped vistas, pedestrian pathways and tree-lined avenues. Melbourne's parks are often considered the best public parks in all of Australia's major cities. There are also many parks in
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4160-424: Is often seen as the most important factor in shaping the natural history of non-colonial organisms; similarly, colony size is key in influencing how colonial organisms are collectively organized. Colonies have a significant range of sizes: some are just several ants living in a twig, while others are super-colonies with many millions of workers. Within a single ant colony, seasonal variation may be huge. For example, in
4290-543: Is one of the world's oldest free public libraries and, as of 2018, the fourth most-visited library globally. During the 19th-century boom period, Melbourne-based authors and poets Marcus Clarke , Adam Lindsay Gordon and Rolf Boldrewood produced classic visions of colonial life, and many visiting writers recorded literary responses to the city: for Henry Kendall , it was a "wild bleak Bohemia", while Henry Kingsley stated that, in its rapid growth, Melbourne "surpasses all human experience". Fergus Hume 's The Mystery of
4420-504: Is the capital and most populous city of the Australian state of Victoria , and the second-most populous city in Australia, after Sydney . Its name generally refers to a 9,993 km (3,858 sq mi) metropolitan area also known as Greater Melbourne , comprising an urban agglomeration of 31 local municipalities , although the name is also used specifically for the local municipality of City of Melbourne based around its central business area . The metropolis occupies much of
4550-530: The Australian Dream . This, coupled with the popularity of the private automobile after 1945, led to the auto-centric urban structure now present today in the middle and outer suburbs. Much of metropolitan Melbourne is accordingly characterised by low-density sprawl, whilst its inner-city areas feature predominantly medium-density, transit-oriented urban forms. The city centre, Docklands, St. Kilda Road and Southbank areas feature high-density forms. Melbourne
4680-449: The CityLink tollway . Other strategies included the privatisation of some of Melbourne's services, including power and public transport, and a reduction in funding to public services such as health, education and public transport infrastructure. Since the mid-1990s, Melbourne has maintained significant population and employment growth. There has been substantial international investment in
4810-538: The Cretaceous period. More than 13,800 of an estimated total of 22,000 species have been classified. They are easily identified by their geniculate (elbowed) antennae and the distinctive node-like structure that forms their slender waists. Ants form colonies that range in size from a few dozen individuals often living in small natural cavities to highly organised colonies that may occupy large territories with sizeable nest that consist of millions of individuals or into
4940-573: The Empire of Japan , and the government requisitioned the Melbourne Cricket Ground for military use. In the immediate years after World War II , Melbourne expanded rapidly, its growth boosted by post-war immigration to Australia , primarily from Southern Europe and the Mediterranean . While the "Paris End" of Collins Street began Melbourne's boutique shopping and open air cafe cultures,
5070-673: The Haidomyrmecinae , Sphecomyrminae and Zigrasimeciinae , with modern ant subfamilies appearing towards the end of the Cretaceous around 80–70 million years ago. Ants diversified extensively during the Angiosperm Terrestrial Revolution and assumed ecological dominance around 60 million years ago. Some groups, such as the Leptanillinae and Martialinae , are suggested to have diversified from early primitive ants that were likely to have been predators underneath
5200-584: The Heidelberg School of impressionists, named after a suburb where they camped to paint en plein air in the 1880s. The Australian tonalists followed in the 1910s, some of whom founded Montsalvat in Eltham , Australia's oldest surviving art colony. Mid-century Melbourne became a stronghold of figurative modernism through the paintings of the Antipodeans and Angry Penguins ; the latter group often met at
5330-593: The Melbourne Bounce genre and the Melbourne Shuffle dance style, both of which emerged from the city's underground rave scene. Established in 1861, the National Gallery of Victoria is Australia's oldest and largest art museum, and houses its collection across two sites: NGV International in Southbank and NGV Australia at Federation Square. Several art movements originated in Melbourne, most famously
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5460-535: The Melbourne Town Hall and Parliament House . Although the area is described as the centre , it is not actually the demographic centre of Melbourne at all, due to an urban sprawl to the southeast, the demographic centre being located at Camberwell . Melbourne is typical of Australian capital cities in that after the turn of the 20th century, it expanded with the underlying notion of a 'quarter acre home and garden' for every family, often referred to locally as
5590-426: The Melbourne Town Hall , St Patrick's cathedral, though many remained incomplete for decades. The layout of the inner suburbs on a largely one-mile grid pattern, cut through by wide radial boulevards and parklands surrounding the central city, was largely established in the 1850s and 1860s. These areas rapidly filled with the ubiquitous terrace houses, as well as with detached houses and grand mansions, while some of
5720-750: The Melbourne Writers Festival and the Victorian Premier's Literary Awards . In 2008, it became the second UNESCO City of Literature . Melbourne is home to many theatres , eight of which are concentrated in the East End Theatre District , including the Victorian era Athenaeum , Her Majesty's and Princess theatres, as well as the Forum and the Regent . Other heritage-listed theatres include
5850-466: The Otway and Macedon Ranges , which block much of the rainfall arriving from the north and west. Port Phillip is often warmer than the surrounding oceans or the land mass, particularly in spring and autumn; this can set up a " bay effect rain ", similar to the " lake effect " seen in colder climates, where showers are intensified leeward of the bay. Relatively narrow streams of heavy showers can often affect
5980-787: The Paleogene period . By the Oligocene and Miocene , ants had come to represent 20–40% of all insects found in major fossil deposits. Of the species that lived in the Eocene epoch, around one in 10 genera survive to the present. Genera surviving today comprise 56% of the genera in Baltic amber fossils (early Oligocene), and 92% of the genera in Dominican amber fossils (apparently early Miocene). Termites live in colonies and are sometimes called "white ants", but termites are only distantly related to ants. They are
6110-472: The Royal Exhibition Building , subsequently moving to the Victorian Parliament House, where it sat until it moved to Canberra in 1927. The Governor-General of Australia resided at Government House in Melbourne until 1930, and many major national institutions remained in Melbourne well into the twentieth century. During World War II the city hosted American military forces who were fighting
6240-815: The Selwyn fault , which transects Mount Martha and Cranbourne . The western portion of the metropolitan area lies within the Victorian Volcanic Plain grasslands vegetation community, and the southeast falls in the Gippsland Plains Grassy Woodland zone. Melbourne extends northward through the undulating bushland valleys of the Yarra Valley 's tributaries— Moonee Ponds Creek (toward Melbourne Airport ), Merri Creek , Darebin Creek and Plenty River . The city reaches southeast through Dandenong to
6370-575: The World Heritage -listed Royal Exhibition Building (1880), the General Post Office (1867), Hotel Windsor (1884) and the Block Arcade (1891). Comparatively little remains of Melbourne's pre-gold rush architecture; St James Old Cathedral (1839) and St Francis' Church (1845) are among the few examples left in the CBD. Many of the CBD's Victorian boom-time landmarks were also demolished in
6500-645: The Wurundjeri , Bunurong and Wathaurong – resided in the area. It was an important meeting place for the clans of the Kulin nation alliance and a vital source of food and water. In June 2021, the boundaries between the land of two of the traditional owner groups, the Wurundjeri and Bunurong, were agreed after being drawn up by the Victorian Aboriginal Heritage Council . The borderline runs across
6630-499: The Yarra River . More recent office, commercial and public developments in the adjoining districts of Southbank and Docklands have made these areas into extensions of the CBD in all but name. A byproduct of the CBD's layout is its network of lanes and arcades , such as Block Arcade and Royal Arcade . Melbourne's CBD has become Australia's most densely populated area, with approximately 19,500 residents per square kilometre, and
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#17328699479946760-401: The genetic bottleneck of the imported ants. In 2009, it was demonstrated that the largest Japanese, Californian and European Argentine ant supercolonies were in fact part of a single global "megacolony". This intercontinental megacolony represents the most populous recorded animal society on earth, other than humans. Another supercolony, measuring approximately 100 km (62 mi) wide,
6890-438: The haplodiploid sex-determination system in ants, workers of a number of species can lay unfertilised eggs that become fully fertile, haploid males. The role of workers may change with their age and in some species, such as honeypot ants , young workers are fed until their gasters are distended, and act as living food storage vessels. These food storage workers are called repletes . For instance, these replete workers develop in
7020-485: The mandibles , used to carry food, manipulate objects, construct nests, and for defence. In some species, a small pocket (infrabuccal chamber) inside the mouth stores food, so it may be passed to other ants or their larvae. Both the legs and wings of the ant are attached to the mesosoma ("thorax"). The legs terminate in a hooked claw which allows them to hook on and climb surfaces. Only reproductive ants ( queens and males) have wings. Queens shed their wings after
7150-536: The market town of Melbourne , Derbyshire . That year, the settlement's general post office officially opened with that name. Between 1836 and 1842, Victorian Aboriginal groups were largely dispossessed of their land by British colonists. In 1840, the Superintendent of the Port Phillip District , Charles La Trobe issued a directive to banish Aborigines from the immediate vicinity of Melbourne. This
7280-457: The nuptial flight , leaving visible stubs, a distinguishing feature of queens. In a few species, wingless queens ( ergatoids ) and males occur. The metasoma (the "abdomen") of the ant houses important internal organs, including those of the reproductive, respiratory (tracheae), and excretory systems. Workers of many species have their egg-laying structures modified into stings that are used for subduing prey and defending their nests. In
7410-605: The red imported fire ant ( Solenopsis invicta ) of South America, are regarded as invasive species in other parts of the world, establishing themselves in areas where they have been introduced accidentally. The word ant and the archaic word emmet are derived from ante , emete of Middle English , which come from ǣmette of Old English ; these are all related to Low Saxon e(e)mt , empe and varieties ( Old Saxon emeta ) and to German Ameise ( Old High German āmeiza ). All of these words come from West Germanic * ǣmaitjōn , and
7540-725: The 2010s the State Government of Victoria initiated a number of major infrastructure projects designed to reduce congestion in Melbourne and encourage economic growth, including the Metro Tunnel , the West Gate Tunnel , the Level Crossing Removal Project and the Suburban Rail Loop . New urban renewal zones were initiated in inner-city areas like Fisherman's Bend and Arden , while suburban growth continued on
7670-462: The 2024 Global Financial Centres Index . The city's eclectic architecture blends Victorian era structures, such as the World Heritage -listed Royal Exhibition Building , with one of the world's tallest skylines . Additional landmarks include the Melbourne Cricket Ground and the National Gallery of Victoria . Noted for its cultural heritage , the city gave rise to Australian rules football , Australian impressionism and Australian cinema , and
7800-595: The Bay was filled with teatree scrub where boorrimul (emu) and marram (kangaroo) were hunted. The first British settlement in Victoria , then part of the penal colony of New South Wales , was established by Colonel David Collins in October 1803, at Sullivan Bay , near present-day Sorrento . The following year, due to a perceived lack of resources, these settlers relocated to Van Diemen's Land (present-day Tasmania ) and founded
7930-589: The Colony of Victoria, with Melbourne as its capital. The discovery of gold in Victoria in mid-1851 sparked a gold rush , and Melbourne, the colony's major port, experienced rapid growth. Within months, the city's population had nearly doubled from 25,000 to 40,000 inhabitants. Exponential growth ensued, and by 1865 Melbourne had overtaken Sydney as Australia's most populous city. An influx of intercolonial and international migrants, particularly from Europe and China, saw
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#17328699479948060-499: The Hymenoptera Name Server, help to keep track of the known and newly described species. The relative ease with which ants may be sampled and studied in ecosystems has made them useful as indicator species in biodiversity studies. Ants are distinct in their morphology from other insects in having geniculate (elbowed) antennae , metapleural glands , and a strong constriction of their second abdominal segment into
8190-469: The Melbourne CBD in the back-beaches of Rye , Sorrento and Portsea . Melbourne has a temperate oceanic climate ( Köppen climate classification Cfb ), with warm summers and cool winters. Melbourne is well known for its changeable weather conditions , mainly due to it being located on the boundary of hot inland areas and the cool southern ocean. This temperature differential is most pronounced in
8320-745: The North American honeypot ant Myrmecocystus mexicanus . Usually the largest workers in the colony develop into repletes; and, if repletes are removed from the colony, other workers become repletes, demonstrating the flexibility of this particular polymorphism . This polymorphism in morphology and behaviour of workers initially was thought to be determined by environmental factors such as nutrition and hormones that led to different developmental paths ; however, genetic differences between worker castes have been noted in Acromyrmex sp. These polymorphisms are caused by relatively small genetic changes; differences in
8450-561: The ant Dolichoderus mariae , one colony can shift from around 300 workers in the summer to over 2,000 workers per queen in the winter. Genetics and environmental factors can cause the variation among different colonies of a single species to be even bigger. Different ant species, even those in the same genus, may have enormous colony size disparities: Formica yessensis has colony sizes that are reported to be 306 million workers while Formica fusca colonies sometimes comprise only 500 workers. A supercolony occurs when many ant colonies over
8580-1167: The ants appear to operate as a unified entity, collectively working together to support the colony. Ants have colonised almost every landmass on Earth . The only places lacking indigenous ants are Antarctica and a few remote or inhospitable islands. Ants thrive in moist tropical ecosystems and may exceed the combined biomass of wild birds and mammals. Their success in so many environments has been attributed to their social organisation and their ability to modify habitats, tap resources, and defend themselves. Their long co-evolution with other species has led to mimetic , commensal , parasitic , and mutualistic relationships. Ant societies have division of labour , communication between individuals, and an ability to solve complex problems . These parallels with human societies have long been an inspiration and subject of study. Many human cultures make use of ants in cuisine, medication, and rites. Some species are valued in their role as biological pest control agents. Their ability to exploit resources may bring ants into conflict with humans, however, as they can damage crops and invade buildings. Some species, such as
8710-573: The area and established a settlement at the site of the current Melbourne Immigration Museum . Batman and his group arrived the following month and the two groups ultimately agreed to share the settlement, initially known by the native name of Dootigala. Batman's Treaty with the Aboriginal elders was annulled by Richard Bourke , the Governor of New South Wales (who at the time governed all of eastern mainland Australia), with compensation paid to members of
8840-535: The association. In 1836, Bourke declared the city the administrative capital of the Port Phillip District of New South Wales , and commissioned the first plan for its urban layout, the Hoddle Grid , in 1837. Known briefly as Batmania, the settlement was named Melbourne on 10 April 1837 by Bourke after the British Prime Minister , William Lamb, 2nd Viscount Melbourne , whose seat was Melbourne Hall in
8970-499: The availability of all types of housing. Subdivision regularly occurs in the outer areas of Melbourne, with numerous developers offering house and land packages. However, since the release of Melbourne 2030 in 2002, planning policies have encouraged medium-density and high-density development in existing areas with good access to public transport and other services. As a result of this, Melbourne's middle and outer-ring suburbs have seen significant brownfields redevelopment. On
9100-732: The avant-garde picture palace The Capitol and St Kilda's Palais Theatre , Australia's largest seated theatre with a capacity of 3,000 people. The Arts Precinct in Southbank is home to Arts Centre Melbourne (which includes the State Theatre and Hamer Hall ), as well as the Melbourne Recital Centre , Malthouse Theatre and Southbank Theatre , home of the Melbourne Theatre Company , Australia's oldest professional theatre company. The Australian Ballet , Opera Australia and Melbourne Symphony Orchestra are also based in
9230-436: The back of the 1850s gold rush and 1880s land boom, Melbourne became renowned as one of the world's great Victorian-era cities, a reputation that persists due to its diverse range of Victorian architecture . High concentrations of well-preserved Victorian-era buildings can be found in the inner suburbs, such as Carlton , East Melbourne and South Melbourne . Outstanding examples of Melbourne's built Victorian heritage include
9360-434: The blocks. The Aboriginal population continued to decline, with an estimated 80% total decrease by 1863, due primarily to introduced diseases (particularly smallpox ), frontier violence and dispossession of their lands. The 1880s saw extraordinary growth: consumer confidence, easy access to credit, and steep increases in land prices led to an enormous amount of construction. During this "land boom", Melbourne reputedly became
9490-556: The body (called the "dorsal aorta") that functions like a heart, and pumps haemolymph toward the head, thus driving the circulation of the internal fluids. The nervous system consists of a ventral nerve cord that runs the length of the body, with several ganglia and branches along the way reaching into the extremities of the appendages. An ant's head contains many sensory organs . Like most insects, ants have compound eyes made from numerous tiny lenses attached together. Ant eyes are good for acute movement detection, but do not offer
9620-551: The city centre was seen by many as stale—the dreary domain of office workers—something expressed by John Brack in his famous painting Collins St., 5 pm (1955). Up until the 21st century, Melbourne was considered Australia's "industrial heartland". Height limits in the CBD were lifted in 1958, after the construction of ICI House , transforming the city's skyline with the introduction of skyscrapers. Suburban expansion then intensified, served by new indoor malls beginning with Chadstone Shopping Centre . The post-war period also saw
9750-399: The city from west to east, with the CBD , Richmond and Hawthorn included in Wurundjeri land, and Albert Park , St Kilda and Caulfield on Bunurong land. However, this change in boundaries is still disputed by people on both sides of the dispute including N'arweet Carolyn Briggs . The name Narrm is commonly used by the broader Aboriginal community to refer to the city, stemming from
9880-530: The city of Hobart . It would be 30 years before another settlement was attempted. In May and June 1835, John Batman , a leading member of the Port Phillip Association in Van Diemen's Land, explored the Melbourne area, and later claimed to have negotiated a purchase of 2,400 km (600,000 acres) with eight Wurundjeri elders. However, the nature of the treaty has been heavily disputed, as none of
10010-430: The city's industries and property market. Major inner-city urban renewal has occurred in areas such as Southbank , Port Melbourne , Melbourne Docklands and South Wharf . Melbourne sustained the highest population increase and economic growth rate of any Australian capital city from 2001 to 2004. From 2006, the growth of the city extended into "green wedges" and beyond the city's urban growth boundary . Predictions of
10140-653: The city's population reaching 5 million people pushed the state government to review the growth boundary in 2008 as part of its Melbourne @ Five Million strategy. In 2009, Melbourne was less affected by the Great Recession in comparison to other Australian cities. At this time, more new jobs were created in Melbourne than any other Australian city—almost as many as the next two fastest growing cities, Brisbane and Perth, combined, and Melbourne's property market remained highly priced, resulting in historically high property prices and widespread rent increases. Beginning in
10270-441: The city. Nauru 's then booming economy resulted in several ambitious investments in Melbourne, such as Nauru House . Melbourne remained Australia's main business and financial centre until the late 1970s, when it began to lose this primacy to Sydney. Melbourne experienced an economic downturn between 1989 and 1992, following the collapse of several local financial institutions. In 1992, the newly elected Kennett government began
10400-519: The city. The establishment of the Melbourne Hydraulic Power Company in 1886 led to the availability of high-pressure piped water, allowing for the installation of hydraulically powered elevators , which led to the construction of the first high-rise buildings in the city. The period also saw the huge expansion of a significant radial rail-based transport network throughout the city and suburbs. Melbourne's land-boom peaked in 1888,
10530-515: The colonies of a few ant species, there are physical castes—workers in distinct size-classes, called minor (micrergates), median, and major ergates (macrergates). Often, the larger ants have disproportionately larger heads, and correspondingly stronger mandibles . Although formally known as dinergates, such individuals are sometimes called "soldier" ants because their stronger mandibles make them more effective in fighting, although they still are workers and their "duties" typically do not vary greatly from
10660-447: The colony to find the best nesting site or to find food. Some social-parasitic species of ants, known as the slave-making ant , raid and steal larvae from neighboring colonies. Ant hill art is a growing collecting hobby. It involves pouring molten metal (typically non-toxic zinc or aluminum), plaster or cement down an ant colony mound acting as a mold and upon hardening, one excavates the resulting structure. In some cases, this involves
10790-548: The countryside from the late 1850s. Construction started on further major public buildings in the 1860s and 1870s, such as the Supreme Court , Government House , and the Queen Victoria Market . The central city filled up with shops and offices, workshops, and warehouses. Large banks and hotels faced the main streets, with fine townhouses in the east end of Collins Street, contrasting with tiny cottages down laneways within
10920-417: The day. Swarms of the winged sexuals (known as alates ) depart the nest in search of other nests. The males die shortly thereafter, along with most of the females. A small percentage of the females survive to initiate new nests. The term "ant colony" refers to a population of workers, reproductive individuals, and brood that live together, cooperate, and treat one another non-aggressively. Often this comprises
11050-645: The decades after World War II, including the Federal Coffee Palace (1888) and the APA Building (1889), one of the tallest early skyscrapers upon completion. Heritage listings and heritage overlays have since been introduced in an effort to prevent further losses of the city's historic fabric. In line with the city's expansion during the early 20th century, suburbs such as Hawthorn and Camberwell are defined largely by Federation and Edwardian architectural styles. The City Baths , built in 1903, are
11180-426: The dirt or vegetation at the top of the hill to prevent it from sliding back into the colony, but in some species, they actively sculpt the materials into specific shapes and may create nest chambers within the mound. Ant Ants are eusocial insects of the family Formicidae and, along with the related wasps and bees , belong to the order Hymenoptera . Ants evolved from vespoid wasp ancestors in
11310-532: The distance and size of objects moving nearly a meter away. Based on experiments conducted to test their ability to differentiate between selected wavelengths of light, some ant species such as Camponotus blandus, Solenopsis invicta, and Formica cunicularia are thought to possess a degree of colour vision. Two antennae ("feelers") are attached to the head; these organs detect chemicals, air currents , and vibrations ; they also are used to transmit and receive signals through touch. The head has two strong jaws,
11440-471: The eastern fringe at Gembrook . Melbourne receives 48.6 clear days annually. Dewpoint temperatures in the summer range from 9.5 to 11.7 °C (49.1 to 53.1 °F). Melbourne is also prone to isolated convective showers forming when a cold pool crosses the state, especially if there is considerable daytime heating. These showers are often heavy and can include hail, squalls, and significant drops in temperature, but they often pass through very quickly with
11570-468: The establishment of slums, including Chinatown and a temporary "tent city" on the southern banks of the Yarra. In the aftermath of the 1854 Eureka Rebellion , mass public support for the plight of the miners resulted in major political changes to the colony, including improvements in working conditions across mining, agriculture, manufacturing and other local industries. At least twenty nationalities took part in
11700-561: The five peoples of the Kulin nation, the traditional custodians of the land encompassing Melbourne are the Boonwurrung , Woiwurrung and the Wurundjeri peoples. In 1803, a short-lived British penal settlement was established at Port Phillip, then part of the Colony of New South Wales . Melbourne was founded in 1835 with the arrival of free settlers from Van Diemen's Land (modern-day Tasmania ). It
11830-862: The general superfamily Formicoidea , which, together with Apoidea, forms the higher-ranking group Formicapoidina . Fernández et al. (2021) suggest that the common ancestors of ants and apoids within the Formicapoidina probably existed as early as in the end of the Jurassic period, before divergence in the Cretaceous. Chrysidoidea Vespidae Rhopalosomatidae Pompilidae Mutillidae Tiphiidae Chyphotidae Scolioidea Apoidea Formicidae Myrmicinae Ectatomminae Heteroponerinae Formicinae Dolichoderinae Aneuretinae Pseudomyrmecinae Myrmeciinae Dorylinae‡ Ponerinae Agroecomyrmecinae Paraponerinae Proceratiinae Amblyoponinae Apomyrminae Leptanillinae Martialinae *Cerapachyinae
11960-512: The genetically related progeny from a single queen, although this is not universal across ants. The name "ant farm" is commonly given to ant nests that are kept in formicaria , isolated from their natural habitat . These formicaria are formed so scientists can study by rearing or temporarily maintaining them. Another name is "formicary", which derives from the Medieval Latin word formīcārium . The word also derives from formica . "Ant nests" are
12090-539: The government began a series of controversial public housing projects in the inner city by the Housing Commission of Victoria , which resulted in the demolition of many neighbourhoods and a proliferation of high-rise towers. In later years, with the rapid rise of motor vehicle ownership, the investment in freeway and highway developments greatly accelerated the outward suburban sprawl and declining inner-city population. The Bolte government sought to rapidly accelerate
12220-549: The growth corridor of Pakenham towards West Gippsland . In the west, it extends along the Maribyrnong River and its tributaries north towards Sunbury . Melbourne's major bayside beaches are in the various suburbs along the shores of Port Phillip Bay, in areas like Port Melbourne , Albert Park , St Kilda , Elwood , Brighton , Sandringham , Mentone , Frankston , Altona , Williamstown and Werribee South. The nearest surf beaches are 85 km (53 mi) south of
12350-403: The hundreds of millions in super colonies . Typical colonies consist of various castes of sterile, wingless females, most of which are workers (ergates), as well as soldiers (dinergates) and other specialised groups. Nearly all ant colonies also have some fertile males called "drones" and one or more fertile females called " queens " ( gynes ). The colonies are described as superorganisms because
12480-562: The late 1970s and early 1980s, including the Little Band scene and St Kilda 's Crystal Ballroom scene, which gave rise to Dead Can Dance and Nick Cave and the Bad Seeds . More recent independent acts from Melbourne to achieve global recognition include The Avalanches , Gotye and King Gizzard and the Lizard Wizard . Melbourne is also regarded as a centre of EDM , and lends its name to
12610-633: The major roads developed as shopping streets. Melbourne quickly became a major finance centre, home to several banks, the Royal Mint , and (in 1861) Australia's first stock exchange . In 1855, the Melbourne Cricket Club secured possession of its now famous ground, the MCG . Members of the Melbourne Football Club codified Australian football in 1859, and in 1861, the first Melbourne Cup race
12740-484: The minor or median workers. In a few species, the median workers are absent, creating a sharp divide between the minors and majors. Weaver ants , for example, have a distinct bimodal size distribution. Some other species show continuous variation in the size of workers. The smallest and largest workers in Carebara diversa show nearly a 500-fold difference in their dry weights. Workers cannot mate; however, because of
12870-430: The modernisation of Melbourne. Major road projects including the remodelling of St Kilda Junction , the widening of Hoddle Street and then the extensive 1969 Melbourne Transportation Plan changed the face of the city into a car-dominated environment. Australia's financial and mining booms during 1969 and 1970 resulted in establishment of the headquarters of many major companies ( BHP and Rio Tinto , among others) in
13000-550: The most famous Melburnian of the Edwardian era. The Sidney Myer Music Bowl in Kings Domain hosted the largest crowd ever for a music concert in Australia when an estimated 200,000 attendees saw Melbourne band The Seekers in 1967. Airing between 1974 and 1987, Melbourne's Countdown helped launch the careers of local acts as diverse as AC/DC and Kylie Minogue . Several distinct post-punk scenes flourished in Melbourne during
13130-440: The nation's largest city in 2029–30 at just over 5.9 million, exceeding 6 million people the following year. Melbourne is in the southeastern part of mainland Australia, within the state of Victoria . Geologically, it is built on the confluence of Quaternary lava flows to the west, Silurian mudstones to the east, and Holocene sand accumulation to the southeast along Port Phillip . The southeastern suburbs are situated on
13260-488: The nest protecting the queen and young. Sometimes, a queen is not present and is replaced by egg-laying workers. These worker ants can only lay haploid eggs producing sterile offspring. Despite the title of queen, she doesn't delegate the tasks to the worker ants; however, the ants choose their tasks based on individual preference. Ants as a colony also work as a collective "super mind". Ants can compare areas and solve complex problems by using information gained by each member of
13390-446: The north and Dandenong Ranges National Park to the east. There are also a number of significant state parks just outside Melbourne. The extensive area covered by urban Melbourne is formally divided into hundreds of suburbs (for addressing and postal purposes), and administered as local government areas, 31 of which are located within the metropolitan area. Melbourne has minimal public housing and high demand for rental housing, which
13520-730: The northern and eastern coastlines of Port Phillip Bay and spreads into the Mornington Peninsula , part of West Gippsland , as well as the hinterlands towards the Yarra Valley , the Dandenong Ranges , and the Macedon Ranges . As of 2023, the population of the metropolitan area was 5.2 million (19% of the population of Australia ); inhabitants are referred to as "Melburnians". The area of Melbourne has been home to Aboriginal Victorians for over 40,000 years and serves as an important meeting place for local Kulin nation clans. Of
13650-544: The original meaning of the word was "the biter" (from Proto-Germanic * ai- , "off, away" + * mait- "cut"). The family name Formicidae is derived from the Latin formīca ("ant") from which the words in other Romance languages , such as the Portuguese formiga , Italian formica , Spanish hormiga , Romanian furnică , and French fourmi are derived. It has been hypothesised that
13780-483: The parties spoke the same language, and the elders likely perceived it as part of the gift exchanges which had taken place over the previous few days amounting to a tanderrum ceremony which allows temporary, not permanent, access to and use of the land. Batman selected a site on the northern bank of the Yarra River , declaring that "this will be the place for a village" before returning to Van Diemen's Land. In August 1835, another group of Vandemonian settlers arrived in
13910-483: The physical spaces in which the ants live. These can be underground, in trees, under rocks, or even inside a single acorn. The name "anthill" (or "ant hill") applies to aboveground nests where the workers pile sand or soil outside the entrance, forming a large mound. Colony size (the number of individuals that make up the colony) is very important to ants: it can affect how they forage, how they defend their nests, how they mate, and even their physical appearances. Body size
14040-563: The precinct. Many of Melbourne's theatres join the Melbourne Town Hall in hosting the annual Melbourne International Comedy Festival , one of the world's three largest comedy festivals. Melbourne has been called "the live music capital of the world"; one study found it has more music venues per capita than any other world city sampled, with 17.5 million patron visits to 553 venues in 2016. Australia's first global music star, opera singer Nellie Melba , took her stage name from her hometown. Composer Percy Grainger followed her in becoming
14170-614: The rebellion, giving some indication of immigration flows at the time. With the wealth brought in from the gold rush and the subsequent need for public buildings, a program of grand civic construction soon began. The 1850s and 1860s saw the commencement of Parliament House , the Treasury Building , the Old Melbourne Gaol , Victoria Barracks , the State Library , University of Melbourne , General Post Office , Customs House ,
14300-639: The richest city in the world, and the second-largest (after London) in the British Empire . The decade began with the Melbourne International Exhibition in 1880, held in the large purpose-built Exhibition Building . A telephone exchange was established that year, and the foundations of St Paul's were laid. In 1881, electric light was installed in the Eastern Market , and a generating station capable of supplying 2,000 incandescent lamps
14430-399: The same places (usually the eastern suburbs) for an extended period, while the rest of Melbourne and surrounds stays dry. Overall, the area around Melbourne is, owing to its rain shadow, nonetheless significantly drier than average for southern Victoria. Within the city and surrounds, rainfall varies widely, from around 425 mm (17 in) at Little River to 1,250 mm (49 in) on
14560-416: The spring and summer months and can cause strong cold fronts to form. These cold fronts can be responsible for varied forms of severe weather from gales to thunderstorms and hail , large temperature drops and heavy rain. Winters, while exceptionally dry by southern Victorian standards, are nonetheless drizzly and overcast. The lack of winter rainfall is owed to Melbourne's rain shadowed location between
14690-509: The sub-order Isoptera , and together with cockroaches , they form the order Blattodea . Blattodeans are related to mantids , crickets , and other winged insects that do not undergo complete metamorphosis . Like ants, termites are eusocial , with sterile workers, but they differ greatly in the genetics of reproduction. The similarity of their social structure to that of ants is attributed to convergent evolution . Velvet ants look like large ants, but are wingless female wasps . Ants have
14820-518: The surface of the soil. During the Cretaceous period, a few species of primitive ants ranged widely on the Laurasian supercontinent (the Northern Hemisphere ). Their representation in the fossil record is poor, in comparison to the populations of other insects, representing only about 1% of fossil evidence of insects in the era. Ants became dominant after adaptive radiation at the beginning of
14950-496: The surrounding suburbs of Melbourne, such as in the municipalities of Stonnington , Boroondara and Port Phillip , southeast of the central business district. Several national parks have been designated around the urban area of Melbourne, including the Mornington Peninsula National Park , Port Phillip Heads Marine National Park and Point Nepean National Park in the southeast, Organ Pipes National Park to
15080-464: The tergite and sternite of a segment fuse together, can occur partly or fully on the second, third and fourth abdominal segment and is used in identification. Fourth abdominal tergosternal fusion was formerly used as character that defined the poneromorph subfamilies, Ponerinae and relatives within their clade, but this is no longer considered a synapomorphic character. Like other arthropods, ants have an exoskeleton , an external covering that provides
15210-400: The total biomass of all the ants in the world was approximately equal to the total biomass of the entire human race. More careful estimates made in 2022 which take into account regional variations puts the global ant contribution at 12 megatons of dry carbon, which is about 20% of the total human contribution, but greater than that of the wild birds and mammals combined. This study also puts
15340-625: The traditional name recorded for the area on which the Melbourne city centre is built. The word is closely related to Narm-narm , being the Boonwurrung word for Port Phillip Bay . Narrm means scrub in Eastern Kulin languages which reflects the Creation Story of how the Bay was filled by the creation of the Birrarung (Yarra River). Before this, the dry Melbourne region extended out into the Bay and
15470-553: The urban periphery in Melbourne's outer west and east in suburbs like Wyndham Vale and Cranbourne . Middle suburbs like Box Hill became denser as a greater proportion of Melburnians began living in apartments. A construction boom resulted in 34 new skyscrapers being built in the central business district between 2010 and 2020. In 2020, Melbourne was classified as an Alpha city by the Globalization and World Cities Research Network . Out of all major Australian cities, Melbourne
15600-476: The world's largest and wealthiest metropolises. After the federation of Australia in 1901, Melbourne served as the interim seat of government of the new nation until Canberra became the permanent capital in 1927. Today, Melbourne is culturally diverse and, among world cities, has the 4th largest foreign born population . It is a leading financial centre in the Asia-Pacific region, ranking 28th globally in
15730-464: The year it hosted the Centennial Exhibition. The brash boosterism that had typified Melbourne during that time ended in the early 1890s. The bubble supporting the local finance and property industries burst, resulting in a severe economic depression. Sixteen small land banks and building societies collapsed, and 133 limited companies went into liquidation. The Melbourne financial crisis was
15860-537: Was 46.4 °C (115.5 °F), on 7 February 2009 . While snow is occasionally seen at higher elevations in the outskirts of the city, and dustings were observed in 2020, it has not been recorded in the Central Business District since 1986. The sea temperature in Melbourne is warmer than the surrounding ocean during the summer months, and colder during the winter months. This is predominately due to Port Phillip Bay being an enclosed and shallow bay that
15990-484: Was enforced later that same year by the mass-arrest and imprisonment of hundreds of Indigenous people during the Lettsom raid . However, Aboriginal people still managed to continue living near the settlement and by January 1844 there were said to be 675 residing in squalid camps around Melbourne. The British Colonial Office had appointed five Aboriginal Protectors for the Aboriginal people of Victoria, in 1839, but their work
16120-420: Was found beneath Melbourne , Australia in 2004. The following terminology is commonly used among myrmecologists to describe the behaviors demonstrated by ants when founding and organizing colonies: Ant colonies have a complex social structure. Ants’ jobs are determined and can be changed by age. As ants grow older their jobs move them farther from the queen, or center of the colony. Younger ants work within
16250-638: Was found in Southern Europe (report published in 2002). Of 33 ant populations tested along the 6,004-kilometre (3,731 mi) stretch along the Mediterranean and Atlantic coasts in Southern Europe, 30 belonged to one supercolony with estimated millions of nests and billions of workers, interspersed with three populations of another supercolony. The researchers claim that this case of unicoloniality cannot be explained by loss of their genetic diversity due to
16380-457: Was held. Melbourne acquired its first public monument, the Burke and Wills statue, in 1864. With the gold rush largely over by 1860, Melbourne continued to grow on the back of continuing gold-mining, as the major port for exporting the agricultural products of Victoria (especially wool) and with a developing manufacturing sector protected by high tariffs. An extensive radial railway network spread into
16510-499: Was in operation by 1882. The Melbourne cable tramway system opened in 1885 and became one of the world's most extensive systems by 1890. In 1885, visiting English journalist George Augustus Henry Sala coined the phrase "Marvellous Melbourne", which stuck long into the twentieth century and has come to refer to the opulence and energy of the 1880s, during which time large commercial buildings, grand hotels, banks, coffee palaces , terrace housing and palatial mansions proliferated in
16640-483: Was incorporated as a Crown settlement in 1837, and named after the then- Prime Minister of the United Kingdom , William Lamb, 2nd Viscount Melbourne . Declared a city by Queen Victoria in 1847, it became the capital of the newly separated Colony of Victoria in 1851. During the 1850s Victorian gold rush , the city entered a lengthy boom period that, by the late 1880s, had transformed it into Australia's, and one of
16770-523: Was nullified by a land policy that favoured squatters who took possession of Aboriginal lands. By 1845, fewer than 240 wealthy Europeans held all the pastoral licences then issued in Victoria and became a powerful political and economic force in Victoria for generations to come. Letters patent of Queen Victoria , issued on 25 June 1847, declared Melbourne a city. On 1 July 1851, the Port Phillip District separated from New South Wales to become
16900-471: Was the worst affected by the COVID-19 pandemic and spent a long time under lockdown restrictions, with Melbourne experiencing six lockdowns totalling 262 days. While this contributed to a net outflow of migration causing a slight reduction in Melbourne's population over the course of 2020 to 2022, Melbourne is projected to be the fastest growing capital city in Australia from 2023–24 onwards, overtaking Sydney as
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