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110-461: The 656-square-kilometre (253 sq mi) municipality is the 172nd largest by area out of the 357 municipalities in Norway. Andøy is the 191st most populous municipality in Norway with a population of 4,553. The municipality's population density is 6.9 inhabitants per square kilometre (18/sq mi) and its population has decreased by 8.8% over the previous 10-year period. The municipality of Andøy

220-607: A gazetteer compiled by Chang Qu  ( 常璩 ) during the Western Jin Dynasty (265–316), reported the discovery of dragon bones at Wucheng in Sichuan Province. Villagers in central China have long unearthed fossilized "dragon bones" for use in traditional medicines . In Europe , dinosaur fossils were generally believed to be the remains of giants and other biblical creatures. Scholarly descriptions of what would now be recognized as dinosaur bones first appeared in

330-510: A sacrum composed of three or more fused vertebrae (three are found in some other archosaurs, but only two are found in Herrerasaurus ); and a perforate acetabulum , or hip socket, with a hole at the center of its inside surface (closed in Saturnalia tupiniquim , for example). Another difficulty of determining distinctly dinosaurian features is that early dinosaurs and other archosaurs from

440-421: A "pillar-erect" configuration of the hip joint, where instead of having a projection from the femur insert on a socket on the hip, the upper pelvic bone was rotated to form an overhanging shelf. Dinosaur fossils have been known for millennia, although their true nature was not recognized. The Chinese considered them to be dragon bones and documented them as such. For example, Huayang Guo Zhi  ( 華陽國志 ),

550-488: A controversial taxon that was recently confirmed to exist after archived photos were uncovered. Bruhathkayosaurus was a titanosaur and would have most likely weighed more than even Marrapunisaurus . Recent size estimates in 2023 have placed this sauropod reaching lengths of up to 44 m (144 ft) long and a colossal weight range of around 110 000 – 170 000  kg ( 240 000 – 370 000  lb), if these upper estimates up true, Bruhathkayosaurus would have rivaled

660-448: A corresponding inwardly facing distinct head on the femur. Their erect posture enabled early dinosaurs to breathe easily while moving, which likely permitted stamina and activity levels that surpassed those of "sprawling" reptiles . Erect limbs probably also helped support the evolution of large size by reducing bending stresses on limbs. Some non-dinosaurian archosaurs, including rauisuchians , also had erect limbs but achieved this by

770-632: A dinosaur. In it he described and named a sauropod tooth , " Rutellum impicatum ", that had been found in Caswell, near Witney , Oxfordshire. Between 1815 and 1824, the Rev William Buckland , the first Reader of Geology at the University of Oxford, collected more fossilized bones of Megalosaurus and became the first person to describe a non-avian dinosaur in a scientific journal . The second non-avian dinosaur genus to be identified, Iguanodon ,

880-537: A femur of Tyrannosaurus preserved soft, flexible tissue within, including blood vessels , bone matrix , and connective tissue (bone fibers) that had retained their microscopic structure. This discovery suggested that original soft tissues could be preserved over geological time, with multiple mechanisms having been proposed. Later, in 2009, Schweitzer and colleagues reported that a Brachylophosaurus femur preserved similar microstructures, and immunohistochemical techniques (based on antibody binding) demonstrated

990-422: A group known as archosaurs, which also includes modern crocodilians. Within the archosaur group, dinosaurs are differentiated most noticeably by their gait. Dinosaur legs extend directly beneath the body, whereas the legs of lizards and crocodilians sprawl out to either side. Collectively, dinosaurs as a clade are divided into two primary branches, Saurischia and Ornithischia. Saurischia includes those taxa sharing

1100-433: A little less than the land area of Puerto Rico , 8,868 square kilometres (3,424 sq mi). Although the arithmetic density is the most common way of measuring population density, several other methods have been developed to provide alternative measures of population density over a specific area. Dinosaur Dinosaurs are a diverse group of reptiles of the clade Dinosauria . They first appeared during

1210-482: A more recent common ancestor with birds than with Ornithischia, while Ornithischia includes all taxa sharing a more recent common ancestor with Triceratops than with Saurischia. Anatomically, these two groups can be distinguished most noticeably by their pelvic structure. Early saurischians—"lizard-hipped", from the Greek sauros ( σαῦρος ) meaning "lizard" and ischion ( ἰσχίον ) meaning "hip joint"—retained

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1320-583: A now lost partial vertebral neural arch described in 1878. Extrapolating from the illustration of this bone, the animal may have been 58 meters (190 ft) long and weighed 122 400  kg ( 269 800  lb). However, recent research have placed Amphicoelias from the long, gracile diplodocid to the shorter but much stockier rebbachisaurid. Now renamed as Maraapunisaurus , this sauropod now stands as much as 40 meters (130 ft) long and weigh as much as 120 000  kg ( 260 000  lb). Another contender of this title includes Bruhathkayosaurus ,

1430-433: A period of time, with estimates ranging from 5–10 million years to 21 million years. When dinosaurs appeared, they were not the dominant terrestrial animals. The terrestrial habitats were occupied by various types of archosauromorphs and therapsids , like cynodonts and rhynchosaurs . Their main competitors were the pseudosuchians , such as aetosaurs , ornithosuchids and rauisuchians, which were more successful than

1540-652: A potential food source, radiated in the Late Triassic. Early sauropodomorphs did not have sophisticated mechanisms for processing food in the mouth, and so must have employed other means of breaking down food farther along the digestive tract. The general homogeneity of dinosaurian faunas continued into the Middle and Late Jurassic, where most localities had predators consisting of ceratosaurians , megalosauroids , and allosauroids , and herbivores consisting of stegosaurian ornithischians and large sauropods. Examples of this include

1650-656: A presentation about fossil reptiles to the British Association for the Advancement of Science in 1841, but reports of the time show that Owen did not mention the word "dinosaur", nor recognize dinosaurs as a distinct group of reptiles in his address. He introduced the Dinosauria only in the revised text version of his talk published in April 1842. With the backing of Prince Albert , the husband of Queen Victoria , Owen established

1760-599: A review paper by Paul Sereno in 1998, were accompanied by increases in the number of published phylogenetic trees for dinosaurs. Dinosaur fossils are not limited to bones, but also include imprints or mineralized remains of skin coverings, organs, and other tissues. Of these, skin coverings based on keratin proteins are most easily preserved because of their cross-linked , hydrophobic molecular structure. Fossils of keratin-based skin coverings or bony skin coverings are known from most major groups of dinosaurs. Dinosaur fossils with scaly skin impressions have been found since

1870-549: A series of studies that likewise argued for active lifestyles in dinosaurs based on anatomical and ecological evidence (see § Physiology ), which were subsequently summarized in his 1986 book The Dinosaur Heresies . New revelations were supported by an increase in dinosaur discoveries. Major new dinosaur discoveries have been made by paleontologists working in previously unexplored regions, including India, South America, Madagascar, Antarctica, and most significantly China. Across theropods, sauropodomorphs, and ornithischians,

1980-603: A sister group to Dinosauria, including a large anterior trochanter, metatarsals II and IV of subequal length, reduced contact between ischium and pubis, the presence of a cnemial crest on the tibia and of an ascending process on the astragalus, and many others. A variety of other skeletal features are shared by dinosaurs. However, because they either are common to other groups of archosaurs or were not present in all early dinosaurs, these features are not considered to be synapomorphies. For example, as diapsids , dinosaurs ancestrally had two pairs of Infratemporal fenestrae (openings in

2090-600: A symbol for the fact that the municipality is situated at the sea and that fishing is important economically to the municipality. The arms were designed by Hallvard Trætteberg after an idea of Henry Oddlo Erichsen. The Church of Norway has three parishes ( sokn ) within Andøy Municipality. It is part of the Vesterålen prosti ( deanery ) in the Diocese of Sør-Hålogaland . Andøya Space , formerly Andøya Rocket Range,

2200-490: A universally agreed-upon list of their distinguishing features, nearly all dinosaurs discovered so far share certain modifications to the ancestral archosaurian skeleton, or are clearly descendants of older dinosaurs showing these modifications. Although some later groups of dinosaurs featured further modified versions of these traits, they are considered typical for Dinosauria; the earliest dinosaurs had them and passed them on to their descendants. Such modifications, originating in

2310-463: A variety of waterbirds , diversified rapidly at the beginning of the Paleogene period, entering ecological niches left vacant by the extinction of Mesozoic dinosaur groups such as the arboreal enantiornithines , aquatic hesperornithines , and even the larger terrestrial theropods (in the form of Gastornis , eogruiids , bathornithids , ratites, geranoidids , mihirungs , and " terror birds "). It

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2420-430: Is 39.7 meters (130 ft) long. There were larger dinosaurs, but knowledge of them is based entirely on a small number of fragmentary fossils. Most of the largest herbivorous specimens on record were discovered in the 1970s or later, and include the massive Argentinosaurus , which may have weighed 80 000 to 100 000 kilograms (88 to 110 short tons) and reached lengths of 30 to 40 meters (98 to 131 ft); some of

2530-499: Is 53/km (140/sq mi). This includes all continental and island land area, including Antarctica . However, if Antarctica is excluded, then population density rises to over 58 per square kilometre (150/sq mi). The European Commission's Joint Research Centre (JRC) has developed a suite of (open and free) data and tools named the Global Human Settlement Layer (GHSL) to improve the science for policy support to

2640-699: Is a synapsid ). None of them had the erect hind limb posture characteristic of true dinosaurs. Dinosaurs were the dominant terrestrial vertebrates of the Mesozoic Era , especially the Jurassic and Cretaceous periods. Other groups of animals were restricted in size and niches; mammals , for example, rarely exceeded the size of a domestic cat and were generally rodent-sized carnivores of small prey. Dinosaurs have always been recognized as an extremely varied group: over 900 non-avian dinosaur genera have been confidently identified (2018) with 1124 species (2016). Estimates put

2750-464: Is a civilian rocket range located a few kilometres south of Andenes. As of 2022, construction is in progress for a spaceport capable of launching small satellites. Andøya Space supports research as well as providing educational programming and military work. It also operates another launch site in Ny-Ålesund , Svalbard. Andøy Municipality is spread across the island of Andøya (the northernmost island in

2860-450: Is a measurement of population per unit land area. It is mostly applied to humans , but sometimes to other living organisms too. It is a key geographical term. Population density is population divided by total land area , sometimes including seas and oceans, as appropriate. Low densities may cause an extinction vortex and further reduce fertility. This is called the Allee effect after

2970-465: Is an inexact art, and reconstructing the muscles and other organs of the living animal is, at best, a process of educated guesswork. The tallest and heaviest dinosaur known from good skeletons is Giraffatitan brancai (previously classified as a species of Brachiosaurus ). Its remains were discovered in Tanzania between 1907 and 1912. Bones from several similar-sized individuals were incorporated into

3080-499: Is around 8,000,000,000 and the Earth 's total area (including land and water) is 510,000,000 km (200,000,000 sq mi). Therefore, the worldwide human population density is approximately 8,000,000,000 ÷ 510,000,000 = 16/km (41/sq mi). However, if only the Earth's land area of 150,000,000 km (58,000,000 sq mi) is taken into account, then human population density

3190-755: Is based in part on preservation bias , as large, sturdy bones are more likely to last until they are fossilized. Many dinosaurs were quite small, some measuring about 50 centimeters (20 inches) in length. The first dinosaur fossils were recognized in the early 19th century, with the name "dinosaur" (meaning "terrible lizard") being coined by Sir Richard Owen in 1842 to refer to these "great fossil lizards". Since then, mounted fossil dinosaur skeletons have been major attractions at museums worldwide, and dinosaurs have become an enduring part of popular culture . The large sizes of some dinosaurs, as well as their seemingly monstrous and fantastic nature, have ensured their regular appearance in best-selling books and films, such as

3300-459: Is between 1 and 10 metric tons (1.1 and 11.0 short tons). This contrasts sharply with the average size of Cenozoic mammals, estimated by the National Museum of Natural History as about 2 to 5 kg (4.4 to 11.0 lb). The sauropods were the largest and heaviest dinosaurs. For much of the dinosaur era, the smallest sauropods were larger than anything else in their habitat, and the largest

3410-696: Is connected to Gryllefjord on the island of Senja by the Andenes–Gryllefjord Ferry . By car, you reach Andøy on Norwegian County Road 82 , northbound from Sortland . Andøya Air Station is located next to Andenes. It is the home of the 333rd Squadron of the Royal Norwegian Air Force and houses all P-3 Orion maritime patrol aircraft in the Norwegian Armed Forces . Population density Population density (in agriculture : standing stock or plant density )

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3520-570: Is derived from a variety of fossil and non-fossil records, including fossilized bones, feces , trackways , gastroliths , feathers , impressions of skin, internal organs and other soft tissues . Many fields of study contribute to our understanding of dinosaurs, including physics (especially biomechanics ), chemistry , biology , and the Earth sciences (of which paleontology is a sub-discipline). Two topics of particular interest and study have been dinosaur size and behavior. Current evidence suggests that dinosaur average size varied through

3630-549: Is governed by a municipal council of directly elected representatives. The mayor is indirectly elected by a vote of the municipal council. The municipality is under the jurisdiction of the Midtre Hålogaland District Court and the Hålogaland Court of Appeal . The municipal council ( Kommunestyre ) of Andøy is made up of 23 representatives that are elected to four year terms. The tables below show

3740-486: Is named after the island on which it is located, Andøya ( Old Norse : Amdarøy ). The first element is the genitive case of the old uncompounded name of the island Ǫmd which has an unknown meaning. The last element is øy which means " island ". The coat of arms was granted on 7 January 1983. The official blazon is " Azure , a schnecke argent from base sinister to sinister" ( Norwegian : Delt av blått og sølv ved virvelsnitt mot venstre ). This means

3850-570: Is now at the American Museum of Natural History in New York City, while Marsh's is at the Peabody Museum of Natural History at Yale University . World War II caused a pause in palaeontological research; after the war, research attention was also diverted increasingly to fossil mammals rather than dinosaurs, which were seen as sluggish and cold-blooded. At the end of the 1960s, however,

3960-644: Is often stated that mammals out-competed the neornithines for dominance of most terrestrial niches but many of these groups co-existed with rich mammalian faunas for most of the Cenozoic Era. Terror birds and bathornithids occupied carnivorous guilds alongside predatory mammals, and ratites are still fairly successful as midsized herbivores; eogruiids similarly lasted from the Eocene to Pliocene , becoming extinct only very recently after over 20 million years of co-existence with many mammal groups. Dinosaurs belong to

4070-528: The Jurassic Park franchise. Persistent public enthusiasm for the animals has resulted in significant funding for dinosaur science, and new discoveries are regularly covered by the media. Under phylogenetic nomenclature , dinosaurs are usually defined as the group consisting of the most recent common ancestor (MRCA) of Triceratops and modern birds (Neornithes), and all its descendants. It has also been suggested that Dinosauria be defined with respect to

4180-437: The blue whale and Perucetus colossus as one of the largest animals to have ever existed. The largest carnivorous dinosaur was Spinosaurus , reaching a length of 12.6 to 18 meters (41 to 59 ft) and weighing 7 to 20.9 metric tons (7.7 to 23.0 short tons). Other large carnivorous theropods included Giganotosaurus , Carcharodontosaurus , and Tyrannosaurus . Therizinosaurus and Deinocheirus were among

4290-666: The Anisian epoch of the Triassic, approximately 243 million years ago, which is the age of Nyasasaurus from the Manda Formation of Tanzania. However, its known fossils are too fragmentary to identify it as a dinosaur or only a close relative. The referral of the Manda Formation to the Anisian is also uncertain. Regardless, dinosaurs existed alongside non-dinosaurian ornithodirans for

4400-457: The Carnian pluvial event . Dinosaur evolution after the Triassic followed changes in vegetation and the location of continents. In the Late Triassic and Early Jurassic, the continents were connected as the single landmass Pangaea , and there was a worldwide dinosaur fauna mostly composed of coelophysoid carnivores and early sauropodomorph herbivores. Gymnosperm plants (particularly conifers ),

4510-631: The European Commission Directorate Generals and Services and as support to the United Nations system. Several of the most densely populated territories in the world are city-states , microstates and urban dependencies . In fact, 95% of the world's population is concentrated on just 10% of the world's land. These territories have a relatively small area and a high urbanization level, with an economically specialized city population drawing also on rural resources outside

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4620-613: The Ischigualasto and Santa Maria Formations of Argentina and Brazil, and the Pebbly Arkose Formation of Zimbabwe. The Ischigualasto Formation ( radiometrically dated at 231–230 million years old ) has produced the early saurischian Eoraptor , originally considered a member of the Herrerasauridae but now considered to be an early sauropodomorph, along with the herrerasaurids Herrerasaurus and Sanjuansaurus , and

4730-555: The Late Jurassic epoch , and are the only dinosaur lineage known to have survived the Cretaceous–Paleogene extinction event approximately 66 mya. Dinosaurs can therefore be divided into avian dinosaurs —birds—and the extinct non-avian dinosaurs , which are all dinosaurs other than birds. Dinosaurs are varied from taxonomic , morphological and ecological standpoints. Birds, at over 11,000 living species , are among

4840-484: The Late Triassic epoch are often poorly known and were similar in many ways; these animals have sometimes been misidentified in the literature. Dinosaurs stand with their hind limbs erect in a manner similar to most modern mammals , but distinct from most other reptiles, whose limbs sprawl out to either side. This posture is due to the development of a laterally facing recess in the pelvis (usually an open socket) and

4950-645: The Morrison Formation of North America and Tendaguru Beds of Tanzania. Dinosaurs in China show some differences, with specialized metriacanthosaurid theropods and unusual, long-necked sauropods like Mamenchisaurus . Ankylosaurians and ornithopods were also becoming more common, but primitive sauropodomorphs had become extinct. Conifers and pteridophytes were the most common plants. Sauropods, like earlier sauropodomorphs, were not oral processors, but ornithischians were evolving various means of dealing with food in

5060-487: The Natural History Museum, London , to display the national collection of dinosaur fossils and other biological and geological exhibits. In 1858, William Parker Foulke discovered the first known American dinosaur, in marl pits in the small town of Haddonfield, New Jersey . (Although fossils had been found before, their nature had not been correctly discerned.) The creature was named Hadrosaurus foulkii . It

5170-611: The Pietraroja Plattenkalk of Italy. It preserves portions of the intestines, colon, liver, muscles, and windpipe. Concurrently, a line of work led by Mary Higby Schweitzer , Jack Horner , and colleagues reported various occurrences of preserved soft tissues and proteins within dinosaur bone fossils. Various mineralized structures that likely represented red blood cells and collagen fibres had been found by Schweitzer and others in tyrannosaurid bones as early as 1991. However, in 2005, Schweitzer and colleagues reported that

5280-650: The Triassic period , between 243 and 233.23  million years ago (mya), although the exact origin and timing of the evolution of dinosaurs is a subject of active research. They became the dominant terrestrial vertebrates after the Triassic–Jurassic extinction event 201.3 mya and their dominance continued throughout the Jurassic and Cretaceous periods. The fossil record shows that birds are feathered dinosaurs , having evolved from earlier theropods during

5390-676: The United States and displayed in Pittsburgh 's Carnegie Museum of Natural History in 1907. The longest dinosaur known from good fossil material is Patagotitan : the skeleton mount in the American Museum of Natural History in New York is 37 meters (121 ft) long. The Museo Municipal Carmen Funes in Plaza Huincul , Argentina, has an Argentinosaurus reconstructed skeleton mount that

5500-688: The Vesterålen archipelago ) and the northeastern part of the island of Hinnøya (the largest and most populous island in Norway). The municipality also consists of smaller skerries including Bleiksøya, Vomma, and Stavaøyan. Andøy is located between the Gavlfjorden and the Andfjorden , and the Risøysundet strait separates the two main islands in Andøy. The Andøy Bridge connects the two islands. The highest point in

5610-427: The largest and smallest dinosaurs to have ever existed. This is because only a tiny percentage of animals were ever fossilized and most of these remain buried in the earth. Few non-avian dinosaur specimens that are recovered are complete skeletons, and impressions of skin and other soft tissues are rare. Rebuilding a complete skeleton by comparing the size and morphology of bones to those of similar, better-known species

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5720-530: The "lizard-hipped" dinosaurs—birds evolved from earlier dinosaurs with "lizard hips". The following is a simplified classification of dinosaur groups based on their evolutionary relationships, and those of the main dinosaur groups Theropoda, Sauropodomorpha and Ornithischia, compiled by Justin Tweet. Further details and other hypotheses of classification may be found on individual articles. Timeline of major dinosaur groups per Holtz (2007) . Knowledge about dinosaurs

5830-677: The 1970s , however, has indicated that dinosaurs were active animals with elevated metabolisms and numerous adaptations for social interaction. Some were herbivorous , others carnivorous . Evidence suggests that all dinosaurs were egg-laying , and that nest -building was a trait shared by many dinosaurs, both avian and non-avian. While dinosaurs were ancestrally bipedal , many extinct groups included quadrupedal species, and some were able to shift between these stances. Elaborate display structures such as horns or crests are common to all dinosaur groups, and some extinct groups developed skeletal modifications such as bony armor and spines . While

5940-409: The 19th century. Samuel Beckles discovered a sauropod forelimb with preserved skin in 1852 that was incorrectly attributed to a crocodile; it was correctly attributed by Marsh in 1888 and subject to further study by Reginald Hooley in 1917. Among ornithischians, in 1884 Jacob Wortman found skin impressions on the first known specimen of Edmontosaurus annectens , which were largely destroyed during

6050-501: The English paleontologist Sir Richard Owen coined the term "dinosaur", using it to refer to the "distinct tribe or sub-order of Saurian Reptiles" that were then being recognized in England and around the world. The term is derived from Ancient Greek δεινός (deinos)  'terrible, potent or fearfully great' and σαῦρος (sauros)  'lizard or reptile'. Though

6160-577: The Greek ornitheios (ὀρνίθειος) meaning "of a bird" and ischion (ἰσχίον) meaning "hip joint"—had a pelvis that superficially resembled a bird's pelvis: the pubic bone was oriented caudally (rear-pointing). Unlike birds, the ornithischian pubis also usually had an additional forward-pointing process. Ornithischia includes a variety of species that were primarily herbivores. Despite the terms "bird hip" (Ornithischia) and "lizard hip" (Saurischia), birds are not part of Ornithischia. Birds instead belong to Saurischia,

6270-501: The Late Cretaceous, was the evolution of flowering plants . At the same time, several groups of dinosaurian herbivores evolved more sophisticated ways to orally process food. Ceratopsians developed a method of slicing with teeth stacked on each other in batteries, and iguanodontians refined a method of grinding with dental batteries , taken to its extreme in hadrosaurids. Some sauropods also evolved tooth batteries, best exemplified by

6380-781: The MRCA of Megalosaurus and Iguanodon , because these were two of the three genera cited by Richard Owen when he recognized the Dinosauria. Both definitions cover the same known genera: Dinosauria = Ornithischia + Saurischia . This includes major groups such as ankylosaurians (armored herbivorous quadrupeds), stegosaurians (plated herbivorous quadrupeds), ceratopsians (bipedal or quadrupedal herbivores with neck frills ), pachycephalosaurians (bipedal herbivores with thick skulls), ornithopods (bipedal or quadrupedal herbivores including " duck-bills "), theropods (mostly bipedal carnivores and birds), and sauropodomorphs (mostly large herbivorous quadrupeds with long necks and tails). Birds are

6490-401: The Middle to Late Triassic epochs, roughly 20 million years after the devastating Permian–Triassic extinction event wiped out an estimated 96% of all marine species and 70% of terrestrial vertebrate species approximately 252 million years ago. The oldest dinosaur fossils known from substantial remains date to the Carnian epoch of the Triassic period and have been found primarily in

6600-519: The Triassic, Early Jurassic, Late Jurassic and Cretaceous. Predatory theropod dinosaurs, which occupied most terrestrial carnivore niches during the Mesozoic, most often fall into the 100-to-1,000 kg (220-to-2,200 lb) category when sorted by estimated weight into categories based on order of magnitude , whereas recent predatory carnivoran mammals peak in the 10-to-100 kg (22-to-220 lb) category. The mode of Mesozoic dinosaur body masses

6710-583: The United States, known as dinosaur mania. Dinosaur mania was exemplified by the fierce rivalry between Edward Drinker Cope and Othniel Charles Marsh , both of whom raced to be the first to find new dinosaurs in what came to be known as the Bone Wars . This fight between the two scientists lasted for over 30 years, ending in 1897 when Cope died after spending his entire fortune on the dinosaur hunt. Many valuable dinosaur specimens were damaged or destroyed due to

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6820-988: The area, illustrating the difference between high population density and overpopulation . Deserts have very limited potential for growing crops as there is not enough rain to support them. Thus, their population density is generally low. However, some cities in the Middle East, such as Dubai , have been increasing in population and infrastructure growth at a fast pace. Cities with high population densities are, by some, considered to be overpopulated, though this will depend on factors like quality of housing and infrastructure and access to resources. Very densely populated cities are mostly in Asia (particularly Southeast Asia ); Africa's Lagos , Kinshasa , and Cairo ; South America's Bogotá , Lima , and São Paulo ; and Mexico City and Saint Petersburg also fall into this category. City population and especially area are, however, heavily dependent on

6930-401: The arms have a field (background) that is divided by a line called a schnecke (a swirling counter-clockwise spiral design that is looks like a wave ). The field located below the line has a tincture of argent which means it is commonly colored white, but if it is made out of metal, then silver is used. The tincture above the line is blue. The colors and design of the arms were chosen as

7040-666: The best-known genera are remarkable for their large size, many Mesozoic dinosaurs were human-sized or smaller, and modern birds are generally small in size. Dinosaurs today inhabit every continent, and fossils show that they had achieved global distribution by the Early Jurassic epoch at latest. Modern birds inhabit most available habitats, from terrestrial to marine, and there is evidence that some non-avian dinosaurs (such as Microraptor ) could fly or at least glide, and others, such as spinosaurids , had semiaquatic habits. While recent discoveries have made it more difficult to present

7150-455: The body . Other prehistoric animals, including pterosaurs , mosasaurs , ichthyosaurs , plesiosaurs , and Dimetrodon , while often popularly conceived of as dinosaurs, are not taxonomically classified as dinosaurs. Pterosaurs are distantly related to dinosaurs, being members of the clade Ornithodira . The other groups mentioned are, like dinosaurs and pterosaurs, members of Sauropsida (the reptile and bird clade), except Dimetrodon (which

7260-429: The bone as the lower extremity of the femur of a large animal, and recognized that it was too large to belong to any known species. He therefore concluded it to be the femur of a huge human, perhaps a Titan or another type of giant featured in legends. Edward Lhuyd , a friend of Sir Isaac Newton , published Lithophylacii Britannici ichnographia (1699), the first scientific treatment of what would now be recognized as

7370-467: The common theropods, and ankylosaurids and early ceratopsians like Psittacosaurus became important herbivores. Meanwhile, Australia was home to a fauna of basal ankylosaurians, hypsilophodonts , and iguanodontians. The stegosaurians appear to have gone extinct at some point in the late Early Cretaceous or early Late Cretaceous . A major change in the Early Cretaceous, which would be amplified in

7480-414: The current and historical composition of the council by political party . The mayor ( Norwegian : ordfører ) of Andøy is the political leader of the municipality and the chairperson of the municipal council. Here is a list of people who have held this position: The Hurtigruten boat stops at Risøyhamn in Andøy. Andøya Airport at Andenes is served by Widerøe airlines. During the summer, Andenes

7590-547: The definition of "urban area" used: densities are almost invariably higher for the center only than when suburban settlements and intervening rural areas are included, as in the agglomeration or metropolitan area (the latter sometimes including neighboring cities). In comparison, based on a world population of 8 billion, the world's inhabitants, if conceptualized as a loose crowd occupying just under 1 m (10 sq ft) per person (cf. Jacobs Method ), would occupy an area of 8,000 square kilometres (3,100 sq mi)

7700-713: The dinosaur subgroup Maniraptora , which are coelurosaurs , which are theropods, which are saurischians. Research by Matthew G. Baron, David B. Norman , and Paul M. Barrett in 2017 suggested a radical revision of dinosaurian systematics. Phylogenetic analysis by Baron et al. recovered the Ornithischia as being closer to the Theropoda than the Sauropodomorpha, as opposed to the traditional union of theropods with sauropodomorphs. This would cause sauropods and kin to fall outside traditional dinosaurs, so they re-defined Dinosauria as

7810-427: The dinosaurs' modern-day surviving avian lineage (birds) are generally small due to the constraints of flight, many prehistoric dinosaurs (non-avian and avian) were large-bodied—the largest sauropod dinosaurs are estimated to have reached lengths of 39.7 meters (130 feet) and heights of 18 m (59 ft) and were the largest land animals of all time. The misconception that non-avian dinosaurs were uniformly gigantic

7920-686: The dinosaurs. Most of these other animals became extinct in the Triassic, in one of two events. First, at about 215 million years ago, a variety of basal archosauromorphs, including the protorosaurs , became extinct. This was followed by the Triassic–Jurassic extinction event (about 201 million years ago), that saw the end of most of the other groups of early archosaurs, like aetosaurs, ornithosuchids, phytosaurs , and rauisuchians. Rhynchosaurs and dicynodonts survived (at least in some areas) at least as late as early –mid Norian and late Norian or earliest Rhaetian stages , respectively, and

8030-574: The early saurischian Alwalkeria , are known from the Upper Maleri and Lower Maleri Formations of India. The Carnian-aged Chañares Formation of Argentina preserves primitive, dinosaur-like ornithodirans such as Lagosuchus and Lagerpeton in Argentina , making it another important site for understanding dinosaur evolution. These ornithodirans support the model of early dinosaurs as small, bipedal predators. Dinosaurs may have appeared as early as

8140-456: The end of the Cretaceous, caused the extinction of all dinosaur groups except for the neornithine birds. Some other diapsid groups, including crocodilians , dyrosaurs , sebecosuchians , turtles, lizards , snakes , sphenodontians , and choristoderans , also survived the event. The surviving lineages of neornithine birds, including the ancestors of modern ratites , ducks and chickens , and

8250-451: The exact date of their extinction is uncertain. These losses left behind a land fauna of crocodylomorphs , dinosaurs, mammals, pterosaurians, and turtles . The first few lines of early dinosaurs diversified through the Carnian and Norian stages of the Triassic, possibly by occupying the niches of the groups that became extinct. Also notably, there was a heightened rate of extinction during

8360-433: The field of dinosaur research experienced a surge in activity that remains ongoing. Several seminal studies led to this activity. First, John Ostrom discovered the bird-like dromaeosaurid theropod Deinonychus and described it in 1969. Its anatomy indicated that it was an active predator that was likely warm-blooded, in marked contrast to the then-prevailing image of dinosaurs. Concurrently, Robert T. Bakker published

8470-486: The hip structure of their ancestors, with a pubis bone directed cranially , or forward. This basic form was modified by rotating the pubis backward to varying degrees in several groups ( Herrerasaurus , therizinosauroids, dromaeosaurids, and birds ). Saurischia includes the theropods (exclusively bipedal and with a wide variety of diets) and sauropodomorphs (long-necked herbivores which include advanced, quadrupedal groups). By contrast, ornithischians—"bird-hipped", from

8580-432: The last common ancestor of Triceratops horridus , Passer domesticus and Diplodocus carnegii , and all of its descendants, to ensure that sauropods and kin remain included as dinosaurs. They also resurrected the clade Ornithoscelida to refer to the group containing Ornithischia and Theropoda. Using one of the above definitions, dinosaurs can be generally described as archosaurs with hind limbs held erect beneath

8690-558: The late 17th century in England. Part of a bone, now known to have been the femur of a Megalosaurus , was recovered from a limestone quarry at Cornwell near Chipping Norton , Oxfordshire, in 1676. The fragment was sent to Robert Plot , Professor of Chemistry at the University of Oxford and first curator of the Ashmolean Museum , who published a description in his The Natural History of Oxford-shire (1677). He correctly identified

8800-551: The latter a Ramsar site of international importance. The bird cliff at Bleiksøya, outside of Bleik , is a spectacular and photogenic landmark outside the northern part of Andøya, with many species of seabirds including puffin . Andøya has a subpolar oceanic climate (Cfc). Andøy Municipality is responsible for primary education (through 10th grade), outpatient health services , senior citizen services, welfare and other social services , zoning , economic development , and municipal roads and utilities. The municipality

8910-478: The longest were the 33.5-meter (110 ft) long Diplodocus hallorum (formerly Seismosaurus ), the 33-to-34-meter (108 to 112 ft) long Supersaurus , and 37-meter (121 ft) long Patagotitan ; and the tallest, the 18-meter (59 ft) tall Sauroposeidon , which could have reached a sixth-floor window. There were a few dinosaurs that was considered either the heaviest and longest. The most famous one include Amphicoelias fragillimus , known only from

9020-479: The most complete theropod specimens, while North American localities have produced the most complete sauropodomorph specimens. Prior to the dinosaur renaissance, dinosaurs were mostly classified using the traditional rank-based system of Linnaean taxonomy . The renaissance was also accompanied by the increasingly widespread application of cladistics , a more objective method of classification based on ancestry and shared traits, which has proved tremendously useful in

9130-495: The most diverse groups of vertebrates. Using fossil evidence, paleontologists have identified over 900 distinct genera and more than 1,000 different species of non-avian dinosaurs. Dinosaurs are represented on every continent by both extant species (birds) and fossil remains. Through the first half of the 20th century, before birds were recognized as dinosaurs, most of the scientific community believed dinosaurs to have been sluggish and cold-blooded . Most research conducted since

9240-509: The most important advantage was dietary. Large animals are more efficient at digestion than small animals, because food spends more time in their digestive systems. This also permits them to subsist on food with lower nutritive value than smaller animals. Sauropod remains are mostly found in rock formations interpreted as dry or seasonally dry, and the ability to eat large quantities of low-nutrient browse would have been advantageous in such environments. Scientists will probably never be certain of

9350-478: The most recent common ancestor of a certain taxonomic group, are called the synapomorphies of such a group. A detailed assessment of archosaur interrelations by Sterling Nesbitt confirmed or found the following twelve unambiguous synapomorphies, some previously known: Nesbitt found a number of further potential synapomorphies and discounted a number of synapomorphies previously suggested. Some of these are also present in silesaurids , which Nesbitt recovered as

9460-458: The mouth, including potential cheek -like organs to keep food in the mouth, and jaw motions to grind food. Another notable evolutionary event of the Jurassic was the appearance of true birds, descended from maniraptoran coelurosaurians. By the Early Cretaceous and the ongoing breakup of Pangaea, dinosaurs were becoming strongly differentiated by landmass. The earliest part of this time saw

9570-492: The municipality is the 889.18-metre (2,917.3 ft) tall mountain Skrivartinden. The midsection of the island consists of bogs and marshes , known for their Arctic cloudberries . There are numerous lakes on the island including Bleiksvatnet and Skogvollvatnet . The island is also the only place in continental Norway where coal and fossils from dinosaurs are found. Forfjorddalen nature reserve on Hinnøya has some of

9680-764: The now-splitting supercontinent Gondwana , abelisaurids were the common theropods, and titanosaurian sauropods the common herbivores. Finally, in Europe, dromaeosaurids, rhabdodontid iguanodontians, nodosaurid ankylosaurians, and titanosaurian sauropods were prevalent. Flowering plants were greatly radiating, with the first grasses appearing by the end of the Cretaceous. Grinding hadrosaurids and shearing ceratopsians became very diverse across North America and Asia. Theropods were also radiating as herbivores or omnivores , with therizinosaurians and ornithomimosaurians becoming common. The Cretaceous–Paleogene extinction event, which occurred approximately 66 million years ago at

9790-756: The number of named genera began to increase exponentially in the 1990s. As of 2008, over 30 new species of dinosaurs were named each year. At least sauropodomorphs experienced a further increase in the number of named species in the 2010s, with an average of 9.3 new species having been named each year between 2009 and 2020. As a consequence, more sauropodomorphs were named between 1990 and 2020 than in all previous years combined. These new localities also led to improvements in overall specimen quality, with new species being increasingly named not on scrappy fossils but on more complete skeletons, sometimes from multiple individuals. Better specimens also led to new species being invalidated less frequently. Asian localities have produced

9900-502: The oldest pine trees in Scandinavia , some more than 700 years old. Lying furthest north in the area known as Vesterålen, Andøy is a mixture of vast areas of marshland fens , with a backdrop of high peaked mountains such as Kvasstinden . The coast is famous for its sheltered bays with white sandy shores. The area has a rich bird life and this is reflected in some good birding localities like Forfjorddalen Nature Reserve and Skogvoll,

10010-438: The pair's rough methods: for example, their diggers often used dynamite to unearth bones. Modern paleontologists would find such methods crude and unacceptable, since blasting easily destroys fossil and stratigraphic evidence. Despite their unrefined methods, the contributions of Cope and Marsh to paleontology were vast: Marsh unearthed 86 new species of dinosaur and Cope discovered 56, a total of 142 new species. Cope's collection

10120-408: The presence of proteins such as collagen, elastin , and laminin . Both specimens yielded collagen protein sequences that were viable for molecular phylogenetic analyses , which grouped them with birds as would be expected. The extraction of fragmentary DNA has also been reported for both of these fossils, along with a specimen of Hypacrosaurus . In 2015, Sergio Bertazzo and colleagues reported

10230-458: The preservation of collagen fibres and red blood cells in eight Cretaceous dinosaur specimens that did not show any signs of exceptional preservation, indicating that soft tissue may be preserved more commonly than previously thought. Suggestions that these structures represent bacterial biofilms have been rejected, but cross-contamination remains a possibility that is difficult to detect. Dinosaurs diverged from their archosaur ancestors during

10340-422: The rebbachisaurid Nigersaurus . There were three general dinosaur faunas in the Late Cretaceous. In the northern continents of North America and Asia, the major theropods were tyrannosaurids and various types of smaller maniraptoran theropods, with a predominantly ornithischian herbivore assemblage of hadrosaurids, ceratopsians, ankylosaurids, and pachycephalosaurians. In the southern continents that had made up

10450-1404: The rocks that produced the Jehol (Early Cretaceous) and Yanliao (Mid-to-Late Jurassic) biotas of northeastern China, from which hundreds of dinosaur specimens bearing impressions of feather-like structures (both closely related to birds and otherwise, see § Origin of birds ) have been described by Xing Xu and colleagues. In living reptiles and mammals, pigment-storing cellular structures known as melanosomes are partially responsible for producing colouration. Both chemical traces of melanin and characteristically shaped melanosomes have been reported from feathers and scales of Jehol and Yanliao dinosaurs, including both theropods and ornithischians. This has enabled multiple full-body reconstructions of dinosaur colouration , such as for Sinosauropteryx and Psittacosaurus by Jakob Vinther and colleagues, and similar techniques have also been extended to dinosaur fossils from other localities. (However, some researchers have also suggested that fossilized melanosomes represent bacterial remains. ) Stomach contents in some Jehol and Yanliao dinosaurs closely related to birds have also provided indirect indications of diet and digestive system anatomy (e.g., crops ). More concrete evidence of internal anatomy has been reported in Scipionyx from

10560-450: The sauropodomorphs Bagualosaurus , Buriolestes , Guaibasaurus , Macrocollum , Nhandumirim , Pampadromaeus , Saturnalia , and Unaysaurus . The Pebbly Arkose Formation, which is of uncertain age but was likely comparable to the other two, has produced the sauropodomorph Mbiresaurus , along with an unnamed herrerasaurid. Less well-preserved remains of the sauropodomorphs Jaklapallisaurus and Nambalia , along with

10670-416: The sauropodomorphs Chromogisaurus , Eodromaeus , and Panphagia . Eoraptor 's likely resemblance to the common ancestor of all dinosaurs suggests that the first dinosaurs would have been small, bipedal predators . The Santa Maria Formation (radiometrically dated to be older, at 233.23 million years old ) has produced the herrerasaurids Gnathovorax and Staurikosaurus , along with

10780-443: The scientist who identified it. Examples of the causes of reduced fertility in low population densities are: Population density is the number of people per unit of area, usually transcribed as "per square kilometer" or square mile, and which may include or exclude, for example, areas of water or glaciers. Commonly this is calculated for a county , city , country , another territory or the entire world . The world's population

10890-571: The skeleton now mounted and on display at the Museum für Naturkunde in Berlin ; this mount is 12 meters (39 ft) tall and 21.8 to 22.5 meters (72 to 74 ft) long, and would have belonged to an animal that weighed between 30 000 and 60 000  kilograms ( 70 000 and 130 000  lb). The longest complete dinosaur is the 27 meters (89 ft) long Diplodocus , which was discovered in Wyoming in

11000-406: The skull behind the eyes), and as members of the diapsid group Archosauria, had additional openings in the snout and lower jaw. Additionally, several characteristics once thought to be synapomorphies are now known to have appeared before dinosaurs, or were absent in the earliest dinosaurs and independently evolved by different dinosaur groups. These include an elongated scapula , or shoulder blade;

11110-434: The sole surviving dinosaurs. In traditional taxonomy , birds were considered a separate class that had evolved from dinosaurs, a distinct superorder . However, most contemporary paleontologists reject the traditional style of classification based on anatomical similarity, in favor of phylogenetic taxonomy based on deduced ancestry, in which each group is defined as all descendants of a given founding genus. Birds belong to

11220-515: The specimen's excavation. Owen and Hooley subsequently described skin impressions of Hypsilophodon and Iguanodon in 1885 and 1917. Since then, scale impressions have been most frequently found among hadrosaurids, where the impressions are known from nearly the entire body across multiple specimens. Starting from the 1990s, major discoveries of exceptionally preserved fossils in deposits known as conservation Lagerstätten contributed to research on dinosaur soft tissues. Chiefly among these were

11330-527: The spread of ankylosaurians, iguanodontians , and brachiosaurids through Europe, North America, and northern Africa . These were later supplemented or replaced in Africa by large spinosaurid and carcharodontosaurid theropods, and rebbachisaurid and titanosaurian sauropods, also found in South America . In Asia , maniraptoran coelurosaurians like dromaeosaurids, troodontids , and oviraptorosaurians became

11440-468: The study of dinosaur systematics and evolution. Cladistic analysis, among other techniques, helps to compensate for an often incomplete and fragmentary fossil record. Reference books summarizing the state of dinosaur research, such as David B. Weishampel and colleagues' The Dinosauria , made knowledge more accessible and spurred further interest in dinosaur research. The release of the first and second editions of The Dinosauria in 1990 and 2004, and of

11550-459: The taxonomic name has often been interpreted as a reference to dinosaurs' teeth, claws, and other fearsome characteristics, Owen intended it also to evoke their size and majesty. Owen recognized that the remains that had been found so far, Iguanodon , Megalosaurus and Hylaeosaurus , shared distinctive features, and so decided to present them as a distinct taxonomic group. As clarified by British geologist and historian Hugh Torrens, Owen had given

11660-849: The total number of dinosaur genera preserved in the fossil record at 1850, nearly 75% still undiscovered, and the number that ever existed (in or out of the fossil record) at 3,400. A 2016 estimate put the number of dinosaur species living in the Mesozoic at 1,543–2,468, compared to the number of modern-day birds (avian dinosaurs) at 10,806 species. Extinct dinosaurs, as well as modern birds, include genera that are herbivorous and others carnivorous, including seed-eaters, fish-eaters, insectivores, and omnivores. While dinosaurs were ancestrally bipedal (as are all modern birds), some evolved into quadrupeds, and others, such as Anchisaurus and Iguanodon , could walk as easily on two or four legs. Cranial modifications like horns and crests are common dinosaurian traits, and some extinct species had bony armor. Although

11770-452: Was an extremely important find: Hadrosaurus was one of the first nearly complete dinosaur skeletons found ( the first was in 1834, in Maidstone, England ), and it was clearly a bipedal creature. This was a revolutionary discovery as, until that point, most scientists had believed dinosaurs walked on four feet, like other lizards. Foulke's discoveries sparked a wave of interests in dinosaurs in

11880-437: Was an order of magnitude more massive than anything else that has since walked the Earth. Giant prehistoric mammals such as Paraceratherium (the largest land mammal ever) were dwarfed by the giant sauropods, and only modern whales approach or surpass them in size. There are several proposed advantages for the large size of sauropods, including protection from predation, reduction of energy use, and longevity, but it may be that

11990-481: Was established as a new municipality on 1 January 1964 due to the work of the Schei Committee . Andøy Municipality was created from the merger of Andenes Municipality (population: 3,812), Bjørnskinn Municipality (population: 1,835), and Dverberg Municipality (population: 1,719). Initially, the new Andøy Municipality had 7,366 residents. The municipal boundaries have not changed since that time. The municipality

12100-450: Was purportedly discovered in 1822 by Mary Ann Mantell , the wife of English geologist Gideon Mantell , though this is disputed and some historians say Gideon had acquired remains years earlier. Gideon Mantell recognized similarities between his fossils and the bones of modern iguanas and published his findings in 1825. The study of these "great fossil lizards" soon became of great interest to European and American scientists, and in 1842

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