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63-465: The Alano Español or Spanish Bulldog is a Spanish breed of medium to large sized dog of alaunt - bulldog type . It has at various times been used as a war dog , for bullfighting , for the management of cattle, for hunting and as a guard dog . In the later twentieth century it became an endangered breed ; a recovery project was launched, and numbers have since recovered. It was officially recognised under national law in 2004. The origins of

126-456: A breed standard which is a written description of the ideal specimen of the breed. Other uses of the term breed when referring to dogs include pure breeds , cross-breeds , mixed breeds and natural breeds . Prior to the standardization of dog breeds, there were many different types of dogs that were defined by their own, unique functions. Many different terms were used to describe dogs, such as breed, strain, type, kind, and variety. By

189-457: A lncRNA variant in the IGF1 region arisen in wolves from northern latitudes c. 53.000 years ago and fixed by natural and subsequently by human selection. Their behavioral traits include guarding, herding, and hunting, retrieving, and scent detection. Their personality traits include hypersocial behavior, boldness, and aggression, which demonstrates the functional and behavioral diversity of dogs. As

252-456: A "school" book about hawking , hunting, fishing, and heraldry , attributed to Juliana Berners (Barnes), lists dogs of the time mainly by function: "First there is a greyhound, a bastard, a mongrel , a mastiff, a limer, a spaniel, " raches " (small-to-medium-sized scenthounds), "kennets" (small hunting dogs), terriers , "butcher's hounds", dung-heap dogs, "trundel tails" ( lapdogs ?) and prick-eared curs , and small ladies puppies that bear away

315-450: A 0.46 kg (1.0 lb) teacup poodle to a 90 kg (200 lb) giant mastiff . The skull, body, and limb proportions vary significantly between breeds, with dogs displaying more phenotypic diversity than can be found within the entire order of carnivores. These breeds possess distinct traits related to morphology, which include body size, skull shape, tail phenotype, fur type and colour. The difference in size has been attributed to

378-570: A 2010 study ( Afghan Hound , Akita , Alaskan Malamute , American Eskimo , Basenji , Canaan dog , Chow Chow , Dingo , New Guinea singing dog , Saluki , Samoyed , Shar-Pei , Siberian Husky ), plus another three: the Eurasier , Finnish Spitz and Shiba Inu . The study referred to these 16 as basal breeds, as opposed to ancient breeds, as they exhibited genetic divergence but not all of them were historically considered to be "ancient breeds". The 2012 study found that modern breeds only emerged in

441-565: A base coat. Over top of the base color is either a lighter or darker color, giving the appearance of stripes. Brindle coloring exists in cattle . The Blue Wildebeest is a species of brindled gnu. For crested geckos , the term "brindle" is used to describe a morph with darker stripes of color. The word brindled is a variant of Middle English brended (early 15c.), from bren "brown color" (13c.), noun from past participle of brennen "burn" (from Proto-Germanic *brennan "to burn", from PIE root *gwher- "to heat, warm"). The etymological sense of

504-703: A known foundation stock may still not be recognized by some clubs as breeds. For instance, the feist is a hunting dog raised in the Southern United States for hunting small game. Feists have a consistent set of characteristics that reliably differentiate them from other dog types and breeds. However, the United Kennel Club recognizes one breed of feist, the Treeing Feist , while the American Kennel Club does not recognize any feist breed. A dog

567-464: A mix of two breeds and is not the result of intentional breeding . In the United States, the term mixed-breed is a favored synonym over mongrel among individuals who wish to avoid negative connotations associated with the latter term. The implication that such dogs must be a mix of defined breeds may stem from an inverted understanding of the origins of dog breeds. Purebred dogs have been, for

630-406: A particular environment and in isolation from other populations of the species. This environment included humans but with little or no selective breeding by humans . Brindle Brindle typically appears as black stripes on a red base. The stripes are eumelanin (black/brown pigment) and the base is phaeomelanin (red/yellow pigment), so the appearance of those pigments can be changed by any of

693-758: A recovery project was launched. There are two breed societies , the Asociación Nacional de Criadores de Alano Español (formed in 1995), and the Sociedad Española de Fomento y Cría del Alano Español. The Alano Español was officially recognised by the Ministerio de Agricultura, Pesca y Alimentación , the Spanish ministry of agriculture, in 2004; together with the Pastor Garafiano , the Ratonero Valenciano ,

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756-561: A result, today dogs are the most abundant carnivore species and are dispersed around the world. The most striking example of this dispersal is that of the numerous modern breeds of European lineage during the Victorian era . A genetic study identified 51 regions of the dog genome which were associated with phenotype variation among breeds in the 57 traits studied, which included body, cranial, dental, and long bone shape and size. There were three quantitative trait loci that explained most of

819-488: A select set of ancestors who had the same characteristics. Dogs of a specific breed breed true , producing young that are very similar to their parents. An individual dog is identified as a member of a breed through proof of ancestry, using genetic analysis or written records of ancestry. Without such proof, identification of a specific breed is not reliable. Such records, called stud books , may be maintained by individuals, clubs, or other organizations. Kennel clubs provide

882-724: A sled dog is 20–25 kg (44–55 lb) based on thermo-regulation, and the ancient sled dogs were between 16–25 kg (35–55 lb). The same standard has been found in the remains of sled dogs from this region 2,000 years ago and in the modern Siberian Husky breed standard. Other dogs were larger at 30 kg (66 lb) and appear to be dogs that had been crossed with wolves and used for polar-bear hunting. Between 3,000 and 4,000 years ago greyhound -types of dogs were depicted on pottery and paintings in Egypt and Western Asia. Mastiff -type dogs were kept for guarding and hunting, and short-legged dogs were also bred. Most modern dog breeds are

945-462: A striped coat texture pattern. A Brindle1 test is available. Brindle coloring consists of irregular stripes extending vertically over the horse's body and horizontally around the legs. Brindle horses can also have a dorsal stripe when caused by dun or non-dun1 . It usually does not affect the head and legs as much as the body, with the heaviest concentrations of brindling being on the neck, shoulders and hindquarters. The coloring has been documented in

1008-407: A study indicated that some breed-temperaments, such as anxiety or fear, may be linked to gene mutations. Other temperaments may be due to the legacies of 'ancient' ancestry. Groups of owners that have dogs of the same breed and have an interest in dog breeding can form national Kennel clubs . Kennel Clubs maintain breed standards , record pedigrees in a breed registry (or studbook), and issue

1071-475: Is a global organisation with 98 members and contract partners (one member per country) that recognize 356 purebreds. A dog crossbreed is the result of mating two different breeds. "Designer Dog" became a fad in the late 20th century. Dog crossbreeds are combinations of lineages of the domestic dog ; they are distinguished from canid hybrids , which are interspecific crosses between Canis species (wolves, coyotes, jackals, etc.). Notable canid hybrids include

1134-731: Is a term formerly, but no longer, used for a particular group of dog breeds by the American Kennel Club. These breeds were referred to as "ancient", as opposed to modern, breeds because historically it was believed their origins dated back more than 500 years. In 2004, a study looked at the microsatellites of 414 purebred dogs representing 85 breeds. The study found that dog breeds were so genetically distinct that 99% of individual dogs could be correctly assigned to their breed based on their genotype , indicating that breeding barriers (purebred breeding) have led to distinct genetic units. The study identified 9 breeds that could be represented on

1197-439: Is discouraged. The head is large, strong, squarish and brachycephalic . The muzzle is short, ideally approximately 37% of the length of the head, with the lower jaw slightly concave; mild prognathism is tolerated. The ears are set high and are pendent if not cropped . The skin is very thick, with neck folds and some wrinkles on the face. Since the breed was used for hunting in packs, it is sociable with other dogs. The Alano

1260-565: Is neither a breed registry nor does it issue pedigrees or keep addresses for breeders. It is a global canine organization with member and contract partners (one member per country) that conduct international conformation shows, working/hunting/herding trials, and various other events. The results are submitted to FCI for processing, approval and international recognition. Each of the member and contract partners issue and maintain their own pedigrees and respective breed standards, and train their own judges. FCI ensures that each member mutually recognizes

1323-575: Is said to be purebred if their parents were purebred and if the dog meets the standards of the breed. The American Kennel Club allows mixed-breed dogs to be shown but under the condition the animals have been spayed or neutered, are not a wolf hybrid, and not eligible for the AKC Foundation Stock Service Program or an AKC Purebred Alternative Listing (PAL). In Canada, the Animal Pedigree Act lays out strict standards for

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1386-457: Is short and thick but never velvety, and is most often a brindle ; this may be of any colour, including grey or blue, with or without black. Other colours are fawn in any shade from sand-coloured to red, with or without black; and black-and-brindle – a black-and-tan in which the tan areas are brindled. The face may or may not have a black mask ; the nose is pigmented black. White markings to the neck, chest or paws are acceptable, but excessive white

1449-502: The Fédération Cynologique Internationale ." In 2017, a study showed that 9,000 years ago the domestic dog was present at what is now Zhokhov Island , arctic north-eastern Siberia, which was connected to the mainland at that time. The dogs were selectively bred as either sled dogs or as hunting dogs, which implies that a sled dog standard and a hunting dog standard existed at that time. The optimal maximum size for

1512-727: The Ratonero Mallorquín and the Ca Mè Mallorquí , it was added to the list of indigenous Spanish breeds. It is not recognised by the Fédération Cynologique Internationale . It has been suggested that the Cimarrón Uruguayo of Uruguay derives principally from the Alano Español. The Alano Español is a large dog of alaunt or bulldog type . Bitches stand some 55–60 cm at the withers , and weigh about 25–30 kg ; dogs are on average about 3 cm taller and 5 kg heavier. The coat

1575-484: The coydog , coywolf , dingo–dog hybrid , jackal–dog hybrid , and wolfdog .   Historically, crosses between dogs of different types were more well accepted at a time when modern purebred breeds (based on the eugenics principles) did not yet exist. Some crossbred dogs, created by breeding two purebred dogs of different breeds, may have the advantage of heterosis , or crossbreed vigor. This advantage can be progressively diluted when two crossbreeds are bred in

1638-418: The fleas and diverse small sorts." A study in 2012 examined 49,000 single nucleotide polymorphisms that gave a genome -wide coverage of 1,375 dogs representing 35 breeds, 19 wolves, and previous published genetic signatures of other breeds, giving a total of 121 breeds covered. The study found a deep genetic split between old-world and new-world wolves, and confirmed the genetic divergence of 13 breeds from

1701-500: The 19th century and that claims of their antiquity are based on little or no historical or empirical evidence. The study indicated that throughout history, global dog populations experienced numerous episodes of diversification and homogenization, with each round further reducing the power of genetic data derived from modern breeds to help infer their early history. Of the basal breeds, the American Eskimo Dog and Eurasier were

1764-744: The Alano Español are unknown. One hypothesis is that it derives from dogs brought to Spain in the Migration Period in the fifth century by the Alani , a nomadic pastoralist people from Central Europe . The first written reference to the breed in Spain is in a chapter of the fourteenth-century Libro de la Montería de Alfonso XI ("Book of the Hunt of Alfonso XI "), in which hunting dogs called Alani are described as having beautiful colours. Dogs of this type travelled with Spanish explorers and were used as war dogs in

1827-449: The English language to mention numbers of dog types are from the "Cynegetica" (hunting literature), namely, The Art of Venery (1327) by Twiti (Twici), a treatise that describes hunting with the limer (a leashed bloodhound type); the pack of running hounds , which included barcelets and brachetz (both scent hounds ); and the sighthound and greyhound . More significant in recording

1890-527: The adjective appears to be "marked as though by branding or burning". In William Shakespeare 's play Macbeth , the opening of act 4, scene 1, is often taken to refer to a brindled cat in the sense of coat pattern as described in this article, because it does contain the word "brinded": However, in this context – Elizabethan English – the word "brinded" was used to mean branded , as in marked by fire . The word in Elizabethans used to describe

1953-445: The attempt to create a breed, narrowing the gene pool. The best way to continue taking advantage of crossbreed vigor is from the breeding of dogs of purebred ancestry, as this vigor is typically seen only in the first generation cross of two purebred animals of separate breeds, thus taking advantage of genetic diversity . A mongrel, mixed-breed dog or mutt is a dog that does not belong to one officially recognized breed but can be

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2016-540: The black and red hairs should intermingle evenly all over. Brindle guinea pigs' fur type is Abyssinian (rosetted). Brindle coloring in horses is extremely rare and most commonly caused by pigment concentrating along Blaschko's lines during fetal development. In two confirmed cases it has been linked to spontaneous chimerism , resulting in an animal with two sets of DNA , with the brindle pattern being an expression of two different sets of equine coat color genes in one horse. These forms are not heritable. In some horses

2079-548: The branches of a phylogenetic tree which grouped together with strong statistical support and could be separated from the other breeds with a modern European origin. These 9 breeds had been referred to as "ancient breeds". The study found that the Pharaoh Hound and Ibizan Hound were not as old as once believed; rather, they had been recreated from combinations of other breeds, and that the Norwegian Elkhound grouped with

2142-882: The breed club to be unacceptable (called faults ). In addition, most breed standards include a historical section, describing the place of origin and the original work done by the breed or its ancestor types . Dogs with a breed standard may be accepted into one or more of the major registries ( kennel clubs ) of dog breeds, including The Kennel Club (1873, UK), American Kennel Club (1884), New Zealand Kennel Club (1886), Canadian Kennel Club (1888), United Kennel Club (1898), United Kennel Clubs International (UCI, Germany 1976), Australian National Kennel Council (1958), and other national breed registries. Recognized dog breeds are classified by groups , such as Hound, Terrier, Working, Herding, Sporting, Non-Sporting, Toy and Miscellaneous; some groups may be further subdivided by some registries. Fédération Cynologique Internationale (FCI)

2205-498: The breeding, exhibiting and judging of pure-bred dogs. It now has 99 member countries. According to BigThink , over 40% of the world’s dog breeds come from the United Kingdom , France and Germany . It states: " Great Britain and France are the ground zero of dog fancying, with 57 registered breeds each. Germany is not far behind, with 47 breeds. These three countries alone represent more than 40% of all dog breeds recognized by

2268-399: The dilutions gray (called blue) or brown (sometimes called red). It is not uncommon for a brindled Cairn Terrier to become progressively more black or silver as it ages. Brindled domestic cats are also known as tortoiseshell cats , and in some cases, tabby cats ; they are almost exclusively female. Brindle is an old variety in guinea pigs . They are difficult to breed to perfection, as

2331-461: The documenting of what it calls "evolving breeds". The breed standard for each breed of dog is a detailed description of the appearance and behaviour of an idealized dog of that breed. Included in the breed standard description are externally observable aspects of appearance and behaviour that are considered by the breed club to be the most important for the breed, and externally observable details of appearance or temperament that are considered by

2394-556: The dog–wolf genetic divergence are not known. Dogs are the most variable mammal on earth with around 450 globally recognized dog breeds. In the Victorian era , directed human selection developed the modern dog breeds, which resulted in a vast range of phenotypes. Most breeds were derived from small numbers of founders within the last 200 years, and since then dogs have undergone rapid phenotypic change and were formed into today's modern breeds due to artificial selection imposed by humans. These breeds can vary in size and weight from

2457-504: The end of the Victorian era, society had changed and so did the role of dogs. The form was given a more prominent role than function. Different types or breeds of dog were being developed by breeders who wanted to define specific characteristics and desirable features in their dogs. Driven by dog shows and the groups that hosted them, the term dog breed took on an entirely new meaning. Dog show competitions included best-in-breed winners, and

2520-608: The genes which usually affect them. White markings and ticking can occur on any brindle dog. Brindle is caused by a complex gene process and is technically a form of mosaicism, where some cells express one allele (KB) and other cells express a different allele (ky), a little like tortoiseshell cats . This makes it very difficult to test for, and there are currently no commercially available tests that are able to detect brindle. Brindle dogs will usually test as KBky, and carriers (one dominant black allele, one brindle) cannot be identified without breeding. The brindle pattern may also take

2583-437: The last 200 years. As a result of their adaptability to many environments and breedability for human needs, today dogs are the most abundant carnivore species and are dispersed around the world. A dog breed will consistently produce the physical traits, movement and temperament that were developed over decades of selective breeding. For each breed they recognize, kennel clubs and breed registries usually maintain and publish

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2646-454: The modern breeds because the genetic heritage of the modern breeds has become blurred due to admixture, and the basal breeds have mostly avoided admixture with them due to geographic or cultural barriers. As dogs are a subspecies but their breeds are distinct genetic units, and because only certain breeds share the same type of cancers as humans, the differences in the genes of different breeds may be useful in human medical research. In 2014,

2709-416: The most part, artificially created from random-bred populations by human selective breeding with the purpose of enhancing desired physical, behavioral, or temperamental characteristics. Dogs that are not purebred are not necessarily a mix of such defined breeds. Therefore, among some experts and fans of such dogs, mongrel is still the preferred term. Natural breeds rose through time in response to

2772-458: The other European dogs despite reports of direct Scandinavian origins dating back 5,000 years. Dog types are broad categories of dogs based on form, function or style of work, lineage, or appearance. In contrast, modern dog breeds are particular breed standards, sharing a common set of heritable characteristics, determined by the kennel club that recognizes the breed. The spread of modern dog breeds has been difficult to resolve because many are

2835-583: The past. At the Zoological Museum of the Academy of Science in Saint Petersburg , a Russian cab horse of brindle coloring from the early 19th century was mounted and put on display due to its rarity. The brindled pattern found in horses could be described as vertical stripes that are found along the neck, back, hindquarters, and upper legs. The horse's head is usually a solid color and is not affected by

2898-504: The pattern seems to be inherited, indicating that one or more genes are responsible. One heritable brindle pattern in a family of American Quarter Horses was identified in 2016 and named Brindle1 (BR1). The Brindle1 phenotype has an X-linked , semidominant mode of inheritance. Female horses with this gene have a striped coat pattern; hairs from the stripes have a different texture as well as color, are less straight, and are more unruly. Male horses have sparse manes and tails but do not show

2961-441: The pedigrees and judges of all FCI members. Purebred dogs have more health problems than mongrel dogs, and require more veterinary visits, and tend to have lower longevity. Indeed, studies have reported lifespans that are shorter by between one and almost two years. Notably, dog breeds with flat faces and short noses have breathing difficulties, eye trouble and other health issues. The Fédération Cynologique Internationale

3024-412: The phenotype variation. Indicators of recent selection were shown by many of the 51 genomic regions that were associated with traits that define a breed, which include body size, coat characteristics, and ear floppiness. Wolf Shar Pei Shiba Inu Chow Chow Akita Inu Basenji Siberian Husky Alaskan Malamute Afghan Hound Saluki other breeds in the study "Ancient breed"

3087-505: The place of tan in tricolor coats of some dog breeds (such as Basenjis ). This coloration looks very similar to tricolor, and can be distinguished only at close range. Dogs of this color are often described as "trindle". It can also occur in combination with merle in the points, or as a brindle merle, in breeds such as the Cardigan Welsh Corgi , although the latter is not acceptable in the show ring. The "dark" markings are black or

3150-598: The product of the controlled breeding practices of the Victorian era (1830–1900). In 2010, a study looked at 48,000  single nucleotide polymorphisms that gave a genome -wide coverage of 912 dogs representing 85 breeds. The study found distinct genetic clusters within modern dogs that largely corresponded to phenotype or function. These included spitz-breeds, toy dogs, spaniels, Mastiff-like breeds, small terriers, retrievers, herding dogs, scent-hounds, and sight-hounds. There were 17 breeds that conflicted with phenotype or function and these were thought to be

3213-501: The products of the controlled breeding practices of the Victorian era (1830-1900), and the accurate documenting of pedigrees with the establishment of the English Kennel Club in 1873 in imitation of other stud book registries for cattle and horses. The domestic dog is the first species, and the only large carnivore , to have been domesticated. The first dogs were certainly wolflike, but the phenotypic changes that coincided with

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3276-470: The purebreds were winning. Breed standards are the reason the breed came to be, and with those standards are key features, including form, function and fitness for purpose. The Kennel Club in the UK was founded in 1873, and was the world's first national kennel club and breed registry. The International Canine Federation was founded in 1911 as a worldwide organization. Its objective is to bring global uniformity to

3339-534: The recognition of distinct dog breeds, but there are many independent clubs with differing, and sometimes inconsistent standards and they need not apply scientific standards. Four varieties of the Belgian Shepherd Dog are recognised as four distinct breeds by the New Zealand Kennel Club . Further, some groups of dogs which clearly share a persistent set of characteristics and documented descent from

3402-633: The result of crossing some of the other phenotypes. As in a 2004 study that found 9 'ancient breeds' to be genetically divergent, the study found 13 breeds that were genetically divergent from the modern breeds: the Basenji , Saluki , Afghan hound , Samoyed , Canaan dog , New Guinea singing dog , dingo , Chow Chow , Chinese Shar Pei , Akita , Alaskan malamute , Siberian husky and American Eskimo dog . The study found that there were three well-supported groups that were highly divergent and distinct from modern domestic dogs. The earliest books in

3465-455: The rules for conformation dog shows and trials and accreditation of judges. They often serve as registries , which are lists of adult purebred dogs and lists of litters of puppies born to purebred parents. A dog breed is represented by a sufficient number of individuals to stably transfer its specific characteristics over generations. Dogs of same breed have similar characteristics of appearance and behavior, primarily because they come from

3528-403: The striping. The brindled pattern has no effect on dark points on horses. Some brindle-colored horses are more eye-catching than others. With this coat pattern there is a base coat that covers the entire body of the horse. This base coat color can be any color. Recorded examples have been bay , chestnut , palomino , and dun . Earliest documented cases were said to have red dun or grulla as

3591-461: The subjugation of Native American peoples, as well as in the re-capture of slaves. Becerrillo , a ferocious war-dog owned by Juan Ponce de León , may have been of this type. Dogs of this type are shown in the bullfighting ring in an etching by Francisco de Goya in his series La Tauromaquia of 1816. From the late nineteenth century the Alano began to decline, for a variety of reasons: in 1880,

3654-542: The use and description of various dog types is The Master of Game (circa 1406) by Edward of York , a treatise that describes dogs and their work, such as the alaunt , greyhound, pack scent hounds, spaniel , and mastiff , used by the privileged and wealthy for hunting purposes. The Master of Game is a combination of the earlier Art of Venery and the French hunting treatise Livre de Chasse by Gaston Phoebus (circa 1387). The Book of Saint Albans , published in 1486,

3717-402: The use of dogs in the bull-ring was prohibited, and the new practice of bullfighting on foot became more widespread; the manner of hunting changed, and more use was made of imported hunting dogs of foreign breeds; extensive management of livestock became less common in parts of the country; and the handling of livestock in slaughterhouses was modernised and no longer made use of dogs. The Alano

3780-478: The very recent product of cross-breeding other basal breeds. Most basal breeds have hybridized with other lineages in the past. If those other lineages were other basal breeds then a basal genetic signature remains. The combination of introgression and past population bottlenecks suggested that basal breeds have little or no genetic connections to their ancestral populations and that their genetic distinctiveness does not signify ancient heritage. They are distinctive from

3843-635: Was no longer needed for its traditional tasks, and numbers fell rapidly, almost to the point of the disappearance or extinction of the breed. A surviving breeding population was identified in the mountains of Enkarterri /Las Encartaciones in the Basque Country in the 1980s. A breed standard was drawn up and a stud-book was started. In collaboration with the Real Sociedad Canina de España and municipal administrations including those of Alanís de la Sierra , Archidona , Cazalla and El Ronquillo ,

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3906-592: Was purposefully bred by humans to perform specific tasks, such as herding, hunting, and guarding. Dogs are the most variable mammal on Earth, with artificial selection producing upward of 360 globally recognized breeds. These breeds possess distinct traits related to morphology, which include body size and shape, tail phenotype , fur type, etc, but are only one species of dog. Their behavioral traits include guarding, herding, and hunting, and personality traits such as hyper-social behavior, boldness, and aggression. Most breeds were derived from small numbers of founders within

3969-417: Was used from Mediaeval times as a war dog . It has since been used for hunting of boar and deer , as a guard dog , and in the management of cattle, both at pasture and at slaughterhouses . Until about the end of the nineteenth century it was used in the bull-ring ; this use declined with the development of bullfighting on foot. Dog breed A dog breed is a particular type of dog that

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